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<document id="2E7B1135382A1259186C9BAF7D26B829" ID-CLB-Dataset="222448" ID-DOI="10.11646/zootaxa.5227.4.3" ID-GBIF-Dataset="ea087173-f3d9-491b-8157-366b194bb519" ID-ISSN="1175-5326" ID-Zenodo-Dep="7518924" ID-ZooBank="B28C27C8-7273-4294-8487-150075ABD889" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673311993872" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Khachikov, Eduard A." docDate="2023" docId="03BD8E09FFC4F6474F90FE9D142DA5D5" docLanguage="en" docName="zootaxa.5227.4.3.pdf" docOrigin="Zootaxa 5227 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omalium falsum Eppelsheim 1889" docType="treatment" docVersion="7" lastPageNumber="449" masterDocId="FF84F671FFC2F6414F07FFF31767A632" masterDocTitle="Faunistic and diagnostic notes on some species of the genus Omalium Gravenhorst, 1802 of Caucasus, Turkey and Iran (Coleoptera: Staphylinidae: Omaliinae, Omaliini)" masterLastPageNumber="458" masterPageNumber="443" pageNumber="449" updateTime="1699473653611" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C453E1EB429F798B43061687EB543593" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F25DFFE59795B7FE9EDD246E5FCC35E3">
<mods:title id="233B28E26A594CBAAEF074ED50B42EDF">Faunistic and diagnostic notes on some species of the genus Omalium Gravenhorst, 1802 of Caucasus, Turkey and Iran (Coleoptera: Staphylinidae: Omaliinae, Omaliini)</mods:title>
</mods:titleInfo>
<mods:name id="4260D45096601B56D0E4D71CC9FA6146" type="personal">
<mods:role id="7DB97266F5D68D835AB0727ABEB5F8BC">
<mods:roleTerm id="1CBFE0676C1F72F6357094B7D344E3EB">Author</mods:roleTerm>
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<mods:namePart id="5EEFE8F5FAF74313C5733EF3522194C1">Shavrin, Alexey V.</mods:namePart>
<mods:nameIdentifier id="D1E9E888815EA70111EDB2258999103D" type="ORCID">0000-0001-6487-4530</mods:nameIdentifier>
<mods:affiliation id="95B9D764D39A9E2248E8B7A7A4D99430">Daugavpils University, Institute of Life Sciences and Technologies, Coleopterological Research Centre, Vienibas 13, Daugavpils, LV- 5401, Latvia. ashavrin @ hotmail. com; https: // orcid. org / 0000 - 0001 - 6487 - 4530</mods:affiliation>
<mods:nameIdentifier id="940F0BA9B3DB21FEEE63481F2F00A0D9" type="email">ashavrin@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="C564FF54D35DEA293FAD8041D3336354" type="personal">
<mods:role id="3D5047526C11D3B19674882498D45528">
<mods:roleTerm id="DD92B1F633247B12E11B2F26CF5B648F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D8C34715CFACAD8983E838E19832025">Khachikov, Eduard A.</mods:namePart>
<mods:affiliation id="8A3AE8282B639C4FF3849A768BB23022">Southern Federal University, Bolshaya Sadovaya 105 / 42, Rostov-on-Don, 34006, Russia.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E0DAD9FB07240AEE822411AE9DD9DE5E">text</mods:typeOfResource>
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<mods:titleInfo id="5F9FB26A57F982D06076C205E6C4C825">
<mods:title id="2FA2BCFCB206A03FA13A973476FEFC24">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="69544102FE5949613C04078D08A714F8">
<mods:date id="C423E7405E7C900CE54054393F88DE20">2023</mods:date>
<mods:detail id="1ADED3FC4D7185229A8D0A1899268519" type="pubDate">
<mods:number id="5DA427DA6B7FE2CBBCD3F2D5414A9E02">2023-01-09</mods:number>
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<mods:detail id="919C5072C6D7A3B8421BCAC03480C65F" type="volume">
<mods:number id="F8FA3E11947ECD62FFCE8C72471F6078">5227</mods:number>
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<mods:number id="825DEB610050CD8390714C2835C31967">4</mods:number>
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<mods:start id="EA6AB2DADED32EF886540BB91351189F">443</mods:start>
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<mods:classification id="BB447FE2FD5BBA6237BA792D2DCB6342">journal article</mods:classification>
<mods:identifier id="43569787DC343F7262D598E0844F1900" type="CLB-Dataset">222448</mods:identifier>
<mods:identifier id="7EE0B13E8AE0D26B824E11F9299D6ED7" type="DOI">10.11646/zootaxa.5227.4.3</mods:identifier>
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<mods:identifier id="876C457984F08472F43B80C701291319" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BD8E09FFC4F6474F90FE9D142DA5D5" ID-DOI="http://doi.org/10.5281/zenodo.7524761" ID-GBIF-Taxon="208788678" ID-Zenodo-Dep="7524761" LSID="urn:lsid:plazi:treatment:03BD8E09FFC4F6474F90FE9D142DA5D5" httpUri="http://treatment.plazi.org/id/03BD8E09FFC4F6474F90FE9D142DA5D5" lastPageNumber="449" pageId="6" pageNumber="449">
<subSubSection id="C30E6C94FFC4F6474F90FE9D1520A7BA" box="[151,583,366,393]" pageId="6" pageNumber="449" type="nomenclature">
<paragraph id="8BAB3F1FFFC4F6474F90FE9D1520A7BA" blockId="6.[151,583,366,429]" box="[151,583,366,393]" pageId="6" pageNumber="449">
<heading id="D0E38873FFC4F6474F90FE9D1520A7BA" bold="true" box="[151,583,366,393]" fontSize="11" level="1" pageId="6" pageNumber="449" reason="1">
<taxonomicName id="4C14449CFFC4F6474F90FE9D1520A7BA" ID-CoL="499TN" authority="Eppelsheim, 1889" authorityName="Eppelsheim" authorityYear="1889" box="[151,583,366,393]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="449" phylum="Arthropoda" rank="species" species="falsum">
<emphasis id="B960E30DFFC4F6474F90FE9D1520A7BA" bold="true" box="[151,583,366,393]" pageId="6" pageNumber="449">
<emphasis id="B960E30DFFC4F6474F90FE9D1603A7BB" bold="true" box="[151,356,366,393]" italics="true" pageId="6" pageNumber="449">Omalium falsum</emphasis>
<bibRefCitation id="EF8542EEFFC4F6474E6CFE9D1520A7BA" author="Eppelsheim, E." box="[363,583,366,392]" pageId="6" pageNumber="449" pagination="11 - 22" refId="ref10518" refString="Eppelsheim, E. (1889) Neue Staphylinen aus den Kaukasuslandern, besonders aus Circassien. Wiener Entomologische Zeitung, 8, 11 - 22. https: // doi. org / 10.5962 / bhl. part. 20020" type="journal article" year="1889">Eppelsheim, 1889</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E6C94FFC4F6474F90FE6016F9A79F" box="[151,414,403,429]" pageId="6" pageNumber="449" type="description">
<paragraph id="8BAB3F1FFFC4F6474F90FE6016F9A79F" blockId="6.[151,583,366,429]" box="[151,414,403,429]" pageId="6" pageNumber="449">
(
<figureCitation id="132F239AFFC4F6474F98FE601788A79F" box="[159,239,403,429]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1864,1888]" captionTargetBox="[234,1353,195,1823]" captionTargetId="figure-19@3.[220,1367,181,1840]" captionTargetPageId="3" captionText="FIGURES 1–10. Habitus of Omalium: 1—O. caesum (Turkey, Bursa: Uludağ), 2—O. cinnamomeum (Turkey, Antakya: Kişlak- Şenköy), 3—O. excavatum (Russia, Karachay-Cherkessia: Arkhyz), 4—O. falsum (Georgia, Tskhneti), 5—O. littorale (Russia, Adygea: Abago), 6—O. riparium impar (Russia, Krasnodar Territory: Khosta), 7—O. rivulare (Armenia, Alaverdi), 8—O. rugatum (Iran, Zagros Mts.), 9—O. turcicum (Russia, Adygea: Lagonaki), 10—O. wunderlei (Turkey, Gümüşhane: Erzincan- Kelkit). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7518926" httpUri="https://zenodo.org/record/7518926/files/figure.png" pageId="6" pageNumber="449">Figs. 4</figureCitation>
,
<figureCitation id="132F239AFFC4F6474FFCFE601624A79F" box="[251,323,403,429]" captionStart="FIGURES 19–26" captionStartId="9.[151,264,1887,1911]" captionTargetBox="[178,1428,438,1853]" captionTargetId="figure-103@9.[154,1433,417,1864]" captionTargetPageId="9" captionText="FIGURES 19–26. Aedeagus of Omalium falsum (Figs. 19–20: Georgia, Tskhneti), O. littorale (Figs. 21–22: Russia, North Ossetia-Alania: Kabardino-Sunzhenskiy Mts.), O. riparium impar (Figs. 23–24: Russia, Kransnodar Territory: Sochi National Park), O. rivulare (Figs. 25–26: Turkey, Artvin: Artvin): 19, 21, 23, 25—parameral view, 20, 22, 24, 26— lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7518930" httpUri="https://zenodo.org/record/7518930/files/figure.png" pageId="6" pageNumber="449">19–20</figureCitation>
,
<figureCitation id="132F239AFFC4F6474E48FE60160AA79F" box="[335,365,403,429]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1503,1527]" captionTargetBox="[161,1431,318,1466]" captionTargetId="figure-33@11.[151,1435,304,1480]" captionTargetPageId="11" captionText="FIGURES 33–41. Female genital segment (ventral view) and spermatheca (shown by arrow in Fig. 40) of Omalium: 33—O. caesum (Russia, North Ossetia-Alania: Zincar), 34—O. cinnamomeum (Turkey, Antakya: Kişlak-Şenköy), 35—O. excavatum (Russia, Krasnodar Territory: Caucasus Nature Reserve), 36—O. falsum (Turkey, Rize: Ardeşen), 37—O. littorale (Russia, Volgograd Area: Elton Lake), 38—O. riparium impar (Georgia, Tskhneti), 39—O. rivulare (Russia, Krasnodar Territory: Derbiy), 40—O. rugatum (Russia, North Ossetia-Alania: Zincar), 41—O. turcicum (Russia, Krasnodar Territory, Caucasus Nature Reserve). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7518934" httpUri="https://zenodo.org/record/7518934/files/figure.png" pageId="6" pageNumber="449">36</figureCitation>
,
<figureCitation id="132F239AFFC4F6474E7EFE6016F1A79F" box="[377,406,403,429]" captionStart="FIGURE 42–44" captionStartId="12.[151,250,1982,2006]" captionTargetBox="[218,1359,381,1957]" captionTargetId="figure-124@12.[218,1369,381,1958]" captionTargetPageId="12" captionText="FIGURE 42–44. Distribution of Omalium in the Caucasus Region and Western Asia: 42—O. falsum, 43—O. turcicum, 44—O. wunderlei." figureDoi="http://doi.org/10.5281/zenodo.7518936" httpUri="https://zenodo.org/record/7518936/files/figure.png" pageId="6" pageNumber="449">42</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E6C94FFC4F6474F90FE241544A7DD" box="[151,547,471,495]" pageId="6" pageNumber="449" type="reference_group">
<paragraph id="8BAB3F1FFFC4F6474F90FE241544A7DD" blockId="6.[151,547,471,495]" box="[151,547,471,495]" pageId="6" pageNumber="449">
<treatmentCitationGroup id="AB041831FFC4F6474F90FE241544A7DD" box="[151,547,471,495]" pageId="6" pageNumber="449">
<treatmentCitation id="0AB5190EFFC4F6474F90FE241544A7DD" author="Eppelsheim, E." box="[151,547,471,495]" page="21" pageId="6" pageNumber="449" year="1889">
<taxonomicName id="4C14449CFFC4F6474F90FE241544A7DD" ID-CoL="499TN" authority="Eppelsheim, 1889: 21" authorityName="Eppelsheim" authorityPageNumber="21" authorityYear="1889" box="[151,547,471,495]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="449" phylum="Arthropoda" rank="species" species="falsum">
<emphasis id="B960E30DFFC4F6474F90FE241627A7DD" box="[151,320,471,495]" italics="true" pageId="6" pageNumber="449">Omalium falsum</emphasis>
<bibRefCitation id="EF8542EEFFC4F6474E41FE241544A7DD" author="Eppelsheim, E." box="[326,547,471,495]" pageId="6" pageNumber="449" pagination="11 - 22" refId="ref10518" refString="Eppelsheim, E. (1889) Neue Staphylinen aus den Kaukasuslandern, besonders aus Circassien. Wiener Entomologische Zeitung, 8, 11 - 22. https: // doi. org / 10.5962 / bhl. part. 20020" type="journal article" year="1889">Eppelsheim, 1889: 21</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30E6C94FFC4F6474F90FDEE134DA56A" pageId="6" pageNumber="449" type="materials_examined">
<paragraph id="8BAB3F1FFFC4F6474F90FDEE1799A53D" blockId="6.[151,1437,541,1000]" pageId="6" pageNumber="449">
<emphasis id="B960E30DFFC4F6474F90FDEE151BA405" bold="true" box="[151,636,541,567]" pageId="6" pageNumber="449">
Material examined.
<collectingCountry id="F3037F8FFFC4F6474E82FDEE1697A405" box="[389,496,541,567]" name="Russia" pageId="6" pageNumber="449">RUSSIA</collectingCountry>
:
<collectingRegion id="49D0F1FDFFC4F6474EFBFDEE151FA405" box="[508,632,541,567]" country="Russia" name="Dagestan" pageId="6" pageNumber="449">DAGeSTAN</collectingRegion>
:
</emphasis>
1 ♁: Kurush.
<quantity id="4CEC92FAFFC4F6474C1FFDEE140BA40A" box="[792,876,541,568]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="6" pageNumber="449" unit="m" value="1850.0">1850 m</quantity>
a.s.l.
<date id="FFAA19DFFFC4F6474CABFDEE134AA40A" box="[940,1069,541,568]" pageId="6" pageNumber="449" value="1991-05-07">05.07.1991</date>
. J. Martens &amp; W. Schawaller leg. (
<collectionCode id="ED05A7DAFFC4F6474F98FDB21796A469" box="[159,241,577,603]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="6" pageNumber="449">SMNS</collectionCode>
);
<emphasis id="B960E30DFFC4F6474E01FDB2169EA469" bold="true" box="[262,505,577,603]" pageId="6" pageNumber="449">
<collectingCountry id="F3037F8FFFC4F6474E01FDB216F5A469" box="[262,402,577,603]" name="Georgia" pageId="6" pageNumber="449">GEORGIA</collectingCountry>
:
<collectingRegion id="49D0F1FDFFC4F6474E9AFDB21697A469" box="[413,496,577,603]" country="Georgia" name="Tbilisi" pageId="6" pageNumber="449">TBILISI</collectingRegion>
:
</emphasis>
1 ♁,
<specimenCount id="9D12F496FFC4F6474D3DFDB21500A46E" box="[570,615,577,604]" pageId="6" pageNumber="449" type="female">1 ♀</specimenCount>
: Tskhneti.
<quantity id="4CEC92FAFFC4F6474DE4FDB2141AA46E" box="[739,893,577,604]" metricMagnitude="3" metricUnit="m" metricValue="1.225" metricValueMax="1.3" metricValueMin="1.15" pageId="6" pageNumber="449" unit="m" value="1225.0" valueMax="1300.0" valueMin="1150.0">1150–1300 m</quantity>
a.s.l., forest with
<taxonomicName id="4C14449CFFC4F6474B4FFDB113E9A469" box="[1096,1166,578,603]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="449" phylum="Tracheophyta" rank="genus">
<emphasis id="B960E30DFFC4F6474B4FFDB113E9A469" box="[1096,1166,578,603]" italics="true" pageId="6" pageNumber="449">Fagus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C14449CFFC4F6474B9AFDB11262A46E" box="[1181,1285,578,604]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="449" phylum="Tracheophyta" rank="genus">
<emphasis id="B960E30DFFC4F6474B9AFDB11262A46E" box="[1181,1285,578,604]" italics="true" pageId="6" pageNumber="449">Carpinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C14449CFFC4F6474A13FDB1122EA46E" box="[1300,1353,578,604]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="449" phylum="Tracheophyta" rank="genus">
<emphasis id="B960E30DFFC4F6474A13FDB1122EA46E" box="[1300,1353,578,604]" italics="true" pageId="6" pageNumber="449">Acer</emphasis>
</taxonomicName>
etc., in litter. 16,
<date id="FFAA19DFFFC4F6474E04FD9616E3A4B2" box="[259,388,613,640]" pageId="6" pageNumber="449" value="1987-05-18">18.05.1987</date>
. S.I. Golovatch &amp; K.Yu. Eskov leg. (cSh);
<emphasis id="B960E30DFFC4F6474C60FD96131FA44D" bold="true" box="[871,1144,613,639]" pageId="6" pageNumber="449">
<collectingCountry id="F3037F8FFFC4F6474C60FD961346A44D" box="[871,1057,613,639]" name="Azerbaijan" pageId="6" pageNumber="449">AZERBAIJAN</collectingCountry>
:
<collectingRegion id="49D0F1FDFFC4F6474B2AFD961313A44D" box="[1069,1140,613,639]" country="Azerbaijan" name="Baku" pageId="6" pageNumber="449">BAKU</collectingRegion>
:
</emphasis>
1 ♁: NW Baku. Altiagak.
<quantity id="4CEC92FAFFC4F6474F90FD7A178CA496" box="[151,235,649,676]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="449" unit="m" value="1200.0">1200 m</quantity>
a.s.l. 21–
<date id="FFAA19DFFFC4F6474E53FD7A16B0A491" box="[340,471,649,675]" pageId="6" pageNumber="449" value="1996-06-23">23.06.1996</date>
. W. Schawaller leg. (
<collectionCode id="ED05A7DAFFC4F6474DB9FD7A1477A491" box="[702,784,649,675]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="6" pageNumber="449">SMNS</collectionCode>
);
<emphasis id="B960E30DFFC4F6474C24FD7A148EA491" bold="true" box="[803,1001,649,675]" pageId="6" pageNumber="449">
<collectingCountry id="F3037F8FFFC4F6474C24FD7A14FAA491" box="[803,925,649,675]" name="Turkey" pageId="6" pageNumber="449">TURKEY</collectingCountry>
:
<collectingRegion id="49D0F1FDFFC4F6474CAFFD7A1482A491" box="[936,997,649,675]" country="Turkey" name="Rize" pageId="6" pageNumber="449">RIZe</collectingRegion>
:
</emphasis>
<specimenCount id="9D12F496FFC4F6474CE8FD7A137DA496" box="[1007,1050,649,676]" pageId="6" pageNumber="449" type="female">1 ♀</specimenCount>
: ca.
<quantity id="4CEC92FAFFC4F6474B4BFD7A13F5A491" box="[1100,1170,649,676]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="449" unit="km" value="25.0">25 km</quantity>
S Ardeşen,
<geoCoordinate id="EE2059D8FFC4F6474A11FD7812FBA494" box="[1302,1436,651,678]" degrees="40" direction="north" minutes="59" orientation="latitude" pageId="6" pageNumber="449" precision="15" seconds="30" value="40.99167">40°59′30′′N</geoCoordinate>
<geoCoordinate id="EE2059D8FFC4F6474F90FD5C1679A4F8" box="[151,286,687,714]" degrees="40" direction="east" minutes="58" orientation="longitude" pageId="6" pageNumber="449" precision="15" seconds="42" value="40.978333">40°58′42′′E</geoCoordinate>
.
<date id="FFAA19DFFFC4F6474E2EFD5E16CCA4F5" box="[297,427,685,711]" pageId="6" pageNumber="449" value="2006-03-08">
03.08.
<quantity id="4CEC92FAFFC4F6474E68FD5E16CCA4F5" box="[367,427,685,711]" metricMagnitude="3" metricUnit="m" metricValue="2.00683" pageId="6" pageNumber="449" unit="m" value="2006.83">2006</quantity>
</date>
. 830 m a.s.l., mixed forest with
<taxonomicName id="4C14449CFFC4F6474C1FFD5E14DAA4F5" box="[792,957,685,711]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="6" pageNumber="449" phylum="Tracheophyta" rank="genus">
<emphasis id="B960E30DFFC4F6474C1FFD5E14DAA4F5" box="[792,957,685,711]" italics="true" pageId="6" pageNumber="449">Rhododendron</emphasis>
</taxonomicName>
. M. Schülke leg. (cSh);
<emphasis id="B960E30DFFC4F6474BD3FD5E1626A4D9" bold="true" pageId="6" pageNumber="449">
<collectingCountry id="F3037F8FFFC4F6474BD3FD5E1247A4F5" box="[1236,1312,685,711]" name="Iran" pageId="6" pageNumber="449">IRAN</collectingCountry>
: ÂZARBẲIẲN-e SHARQI:
</emphasis>
1 ♁: Arasbaran Biosphere Reserve.
<quantity id="4CEC92FAFFC4F6474DDEFD221413A4DE" box="[729,884,721,748]" metricMagnitude="3" metricUnit="m" metricValue="1.725" metricValueMax="1.8" metricValueMin="1.65" pageId="6" pageNumber="449" unit="m" value="1725.0" valueMax="1800.0" valueMin="1650.0">1650–1800 m</quantity>
a.s.l. 04–
<date id="FFAA19DFFFC4F6474CE6FD221305A4DE" box="[993,1122,721,748]" pageId="6" pageNumber="449" value="1978-06-15">15.06.1978</date>
. J. Martens &amp; R. Pieper leg. (
<collectionCode id="ED05A7DAFFC4F6474F98FD061796A53D" box="[159,241,757,783]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="6" pageNumber="449">SMNS</collectionCode>
).
</paragraph>
<paragraph id="8BAB3F1FFFC4F6474FC0FCEA134DA56A" blockId="6.[151,1437,541,1000]" pageId="6" pageNumber="449">
<emphasis id="B960E30DFFC4F6474FC0FCEA150BA501" bold="true" box="[199,620,793,819]" pageId="6" pageNumber="449">
Published records.
<collectingCountry id="F3037F8FFFC4F6474EABFCEA1501A501" box="[428,614,793,819]" name="Azerbaijan" pageId="6" pageNumber="449">AZERBAIJAN</collectingCountry>
:
</emphasis>
<bibRefCitation id="EF8542EEFFC4F6474D74FCEA1402A501" author="Eppelsheim, E." box="[627,869,793,819]" pageId="6" pageNumber="449" pagination="11 - 22" refId="ref10518" refString="Eppelsheim, E. (1889) Neue Staphylinen aus den Kaukasuslandern, besonders aus Circassien. Wiener Entomologische Zeitung, 8, 11 - 22. https: // doi. org / 10.5962 / bhl. part. 20020" type="journal article" year="1889">Eppelsheim 1889: 21</bibRefCitation>
,
<bibRefCitation id="EF8542EEFFC4F6474C75FCEA1307A501" author="Bernhauer, M." box="[882,1120,793,819]" pageId="6" pageNumber="449" pagination="186 - 192" refId="ref10405" refString="Bernhauer, M. (1903) Neue Staphyliniden der palaarktischen Fauna. Munchener Koleopterologische Zeitschrift, 1, 186 - 192." type="journal article" year="1903">Bernhauer 1903: 190</bibRefCitation>
(as
<taxonomicName id="4C14449CFFC4F6474B97FCE917B4A565" authority="Bernhauer, 1903" authorityName="Bernhauer" authorityYear="1903" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="449" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="B960E30DFFC4F6474B97FCE913C3A506" box="[1168,1188,794,820]" italics="true" pageId="6" pageNumber="449">O</emphasis>
.
<emphasis id="B960E30DFFC4F6474BB4FCEA127DA501" box="[1203,1306,793,819]" italics="true" pageId="6" pageNumber="449">excellens</emphasis>
<bibRefCitation id="EF8542EEFFC4F6474A25FCEA17B4A565" author="Bernhauer, M." pageId="6" pageNumber="449" pagination="186 - 192" refId="ref10405" refString="Bernhauer, M. (1903) Neue Staphyliniden der palaarktischen Fauna. Munchener Koleopterologische Zeitschrift, 1, 186 - 192." type="journal article" year="1903">Bernhauer, 1903</bibRefCitation>
</taxonomicName>
);
<emphasis id="B960E30DFFC4F6474FE0FCCE161FA565" bold="true" box="[231,376,829,855]" pageId="6" pageNumber="449">
<collectingCountry id="F3037F8FFFC4F6474FE0FCCE1614A565" box="[231,371,829,855]" name="Georgia" pageId="6" pageNumber="449">GEORGIA</collectingCountry>
:
</emphasis>
<bibRefCitation id="EF8542EEFFC4F6474E78FCCE15BFA565" author="Schulke, M. &amp; Smetana, A." box="[383,728,829,856]" pageId="6" pageNumber="449" refId="ref11577" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana 2015: 346</bibRefCitation>
;
<emphasis id="B960E30DFFC4F6474DE3FCCE1404A565" bold="true" box="[740,867,829,855]" pageId="6" pageNumber="449">
<collectingCountry id="F3037F8FFFC4F6474DE3FCCE1439A565" box="[740,862,829,855]" name="Turkey" pageId="6" pageNumber="449">TURKEY</collectingCountry>
:
</emphasis>
<bibRefCitation id="EF8542EEFFC4F6474C6EFCCE1341A56A" author="Zanetti, A." box="[873,1062,829,856]" pageId="6" pageNumber="449" pagination="45 - 63" refId="ref11791" refString="Zanetti, A. (2002) Studies on Omalium Gravenhorst, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). Bollettino del Museo Civivo di Storia Naturale di Verona. Botanica Zoologia, 26, 45 - 63." type="journal article" year="2002">Zanetti 2002: 55</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E6C94FFC4F6474FC0FC92147FA5AD" pageId="6" pageNumber="449" type="distribution">
<paragraph id="8BAB3F1FFFC4F6474FC0FC92147FA5AD" blockId="6.[151,1437,541,1000]" pageId="6" pageNumber="449">
<emphasis id="B960E30DFFC4F6474FC0FC921606A549" bold="true" box="[199,353,865,891]" pageId="6" pageNumber="449">Distribution.</emphasis>
<taxonomicName id="4C14449CFFC4F6474E6EFC921543A549" authorityName="Eppelsheim" authorityYear="1889" box="[361,548,865,891]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="449" phylum="Arthropoda" rank="species" species="falsum">
<emphasis id="B960E30DFFC4F6474E6EFC921543A549" box="[361,548,865,891]" italics="true" pageId="6" pageNumber="449">Omalium falsum</emphasis>
</taxonomicName>
is known from the Caucasus (
<collectingCountry id="F3037F8FFFC4F6474C79FC9214BBA549" box="[894,988,865,891]" name="Georgia" pageId="6" pageNumber="449">Georgia</collectingCountry>
,
<collectingCountry id="F3037F8FFFC4F6474CEFFC921302A549" box="[1000,1125,865,891]" name="Azerbaijan" pageId="6" pageNumber="449">Azerbaijan</collectingCountry>
) and
<collectingCountry id="F3037F8FFFC4F6474BA3FC921393A549" box="[1188,1268,865,891]" name="Turkey" pageId="6" pageNumber="449">Turkey</collectingCountry>
(
<bibRefCitation id="EF8542EEFFC4F6474A03FC9212FFA549" author="Zanetti, A." box="[1284,1432,865,891]" pageId="6" pageNumber="449" pagination="45 - 63" refId="ref11791" refString="Zanetti, A. (2002) Studies on Omalium Gravenhorst, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). Bollettino del Museo Civivo di Storia Naturale di Verona. Botanica Zoologia, 26, 45 - 63." type="journal article" year="2002">Zanetti 2002</bibRefCitation>
,
<bibRefCitation id="EF8542EEFFC4F6474F90FC7616DDA592" author="Schulke, M. &amp; Smetana, A." box="[151,442,901,928]" pageId="6" pageNumber="449" refId="ref11577" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana 2015</bibRefCitation>
). The distribution as in
<figureCitation id="132F239AFFC4F6474DC6FC761473A5AD" box="[705,788,901,927]" captionStart="FIGURE 42–44" captionStartId="12.[151,250,1982,2006]" captionTargetBox="[218,1359,381,1957]" captionTargetId="figure-124@12.[218,1369,381,1958]" captionTargetPageId="12" captionText="FIGURE 42–44. Distribution of Omalium in the Caucasus Region and Western Asia: 42—O. falsum, 43—O. turcicum, 44—O. wunderlei." figureDoi="http://doi.org/10.5281/zenodo.7518936" httpUri="https://zenodo.org/record/7518936/files/figure.png" pageId="6" pageNumber="449">Fig. 42</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E6C94FFC4F6474FC0FC5A142DA5D5" pageId="6" pageNumber="449" type="discussion">
<paragraph id="8BAB3F1FFFC4F6474FC0FC5A1269A5F1" blockId="6.[151,1437,541,1000]" box="[199,1294,937,964]" pageId="6" pageNumber="449">
<emphasis id="B960E30DFFC4F6474FC0FC5A165CA5F1" bold="true" box="[199,315,937,963]" pageId="6" pageNumber="449">Remarks.</emphasis>
Habitus as in
<figureCitation id="132F239AFFC4F6474EDCFC5A1547A5F1" box="[475,544,937,963]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1864,1888]" captionTargetBox="[234,1353,195,1823]" captionTargetId="figure-19@3.[220,1367,181,1840]" captionTargetPageId="3" captionText="FIGURES 1–10. Habitus of Omalium: 1—O. caesum (Turkey, Bursa: Uludağ), 2—O. cinnamomeum (Turkey, Antakya: Kişlak- Şenköy), 3—O. excavatum (Russia, Karachay-Cherkessia: Arkhyz), 4—O. falsum (Georgia, Tskhneti), 5—O. littorale (Russia, Adygea: Abago), 6—O. riparium impar (Russia, Krasnodar Territory: Khosta), 7—O. rivulare (Armenia, Alaverdi), 8—O. rugatum (Iran, Zagros Mts.), 9—O. turcicum (Russia, Adygea: Lagonaki), 10—O. wunderlei (Turkey, Gümüşhane: Erzincan- Kelkit). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7518926" httpUri="https://zenodo.org/record/7518926/files/figure.png" pageId="6" pageNumber="449">Fig. 4</figureCitation>
. Aedeagus as in
<figureCitation id="132F239AFFC4F6474DDDFC5A1406A5F1" box="[730,865,937,963]" captionStart="FIGURES 19–26" captionStartId="9.[151,264,1887,1911]" captionTargetBox="[178,1428,438,1853]" captionTargetId="figure-103@9.[154,1433,417,1864]" captionTargetPageId="9" captionText="FIGURES 19–26. Aedeagus of Omalium falsum (Figs. 19–20: Georgia, Tskhneti), O. littorale (Figs. 21–22: Russia, North Ossetia-Alania: Kabardino-Sunzhenskiy Mts.), O. riparium impar (Figs. 23–24: Russia, Kransnodar Territory: Sochi National Park), O. rivulare (Figs. 25–26: Turkey, Artvin: Artvin): 19, 21, 23, 25—parameral view, 20, 22, 24, 26— lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7518930" httpUri="https://zenodo.org/record/7518930/files/figure.png" pageId="6" pageNumber="449">Figs. 19–20</figureCitation>
. Female genital segment as in
<figureCitation id="132F239AFFC4F6474BBFFC5A126DA5F1" box="[1208,1290,937,963]" captionStart="FIGURES 33–41" captionStartId="11.[151,264,1503,1527]" captionTargetBox="[161,1431,318,1466]" captionTargetId="figure-33@11.[151,1435,304,1480]" captionTargetPageId="11" captionText="FIGURES 33–41. Female genital segment (ventral view) and spermatheca (shown by arrow in Fig. 40) of Omalium: 33—O. caesum (Russia, North Ossetia-Alania: Zincar), 34—O. cinnamomeum (Turkey, Antakya: Kişlak-Şenköy), 35—O. excavatum (Russia, Krasnodar Territory: Caucasus Nature Reserve), 36—O. falsum (Turkey, Rize: Ardeşen), 37—O. littorale (Russia, Volgograd Area: Elton Lake), 38—O. riparium impar (Georgia, Tskhneti), 39—O. rivulare (Russia, Krasnodar Territory: Derbiy), 40—O. rugatum (Russia, North Ossetia-Alania: Zincar), 41—O. turcicum (Russia, Krasnodar Territory, Caucasus Nature Reserve). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7518934" httpUri="https://zenodo.org/record/7518934/files/figure.png" pageId="6" pageNumber="449">Fig. 36</figureCitation>
.
</paragraph>
<paragraph id="8BAB3F1FFFC4F6474FC0FC3D142DA5D5" blockId="6.[151,1437,541,1000]" box="[199,842,973,1000]" pageId="6" pageNumber="449">
It is here recorded from
<collectingCountry id="F3037F8FFFC4F6474ED0FC3E1545A5D5" box="[471,546,973,999]" name="Russia" pageId="6" pageNumber="449">Russia</collectingCountry>
and
<collectingCountry id="F3037F8FFFC4F6474D5FFC3D15E2A5DA" box="[600,645,974,1000]" name="Iran" pageId="6" pageNumber="449">Iran</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>
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