treatments-xml/data/03/BA/87/03BA87E7905BC8117BC6C458FBC606C6.xml

<document id="40A9C4CAAA6DCD9C119C4E7FBEE3B594" ID-DOI="10.5281/zenodo.183067" ID-GBIF-Dataset="6b6ecc7d-3573-4900-b8a6-0b5039143c20" ID-ISSN="1175-5326" ID-Zenodo-Dep="183067" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459890306794" checkinUser="plazi" docAuthor="Mceachran, John D. &amp; Dewitt, Thomas J." docDate="2008" docId="03BA87E7905BC8117BC6C458FBC606C6" docLanguage="en" docName="zt01824p054.pdf" docOrigin="Zootaxa 1824" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Heterandria tuxtlaensis Mceachran &amp; Dewitt, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="FF83FF9F905FC8187B51C334FFC8061C" masterDocTitle="A new livebearing fish, Heterandria tuxtlaensis, from Lake Catemaco, Veracruz, Mexico (Cyprinodontiformes: Poeciliidae)" masterLastPageNumber="54" masterPageNumber="45" pageNumber="49" updateTime="1698227986178" updateUser="plazi">
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<mods:title id="7E8579D6C86C8912C44C9A91908D0371">A new livebearing fish, Heterandria tuxtlaensis, from Lake Catemaco, Veracruz, Mexico (Cyprinodontiformes: Poeciliidae)</mods:title>
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<mods:namePart id="A48BF50CC345658905FBCD98153DD12A">Mceachran, John D.</mods:namePart>
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<mods:namePart id="6990AFFB86F7433F1618787BDBEA9BFE">Dewitt, Thomas J.</mods:namePart>
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<mods:date id="A5D791660675FAFC32C46247D3BBC965">2008</mods:date>
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<mods:number id="214BC93C71F199B1EF6CD64FC7C65146">1824</mods:number>
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<treatment id="03BA87E7905BC8117BC6C458FBC606C6" ID-DOI="http://doi.org/10.5281/zenodo.5678545" ID-GBIF-Taxon="119371871" ID-Zenodo-Dep="5678545" LSID="urn:lsid:plazi:treatment:03BA87E7905BC8117BC6C458FBC606C6" httpUri="http://treatment.plazi.org/id/03BA87E7905BC8117BC6C458FBC606C6" lastPageId="9" lastPageNumber="54" pageId="4" pageNumber="49">
<subSubSection id="C309657A905BC81C7BC6C458FE6E01B5" pageId="4" pageNumber="49" type="nomenclature">
<paragraph id="8BAC36F1905BC81C7BC6C458FDD1019A" blockId="4.[151,537,1900,1961]" box="[151,537,1900,1926]" pageId="4" pageNumber="49">
<heading id="D0E4819D905BC81C7BC6C458FDD1019A" bold="true" box="[151,537,1900,1926]" fontSize="11" level="1" pageId="4" pageNumber="49" reason="1">
<emphasis id="B967EAE3905BC81C7BC6C458FDD1019A" bold="true" box="[151,537,1900,1926]" pageId="4" pageNumber="49">
<taxonomicName id="4C134D72905BC81C7BC6C458FE7C019A" ID-CoL="6M2Y7" box="[151,436,1900,1926]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="4" pageNumber="49" phylum="Chordata" rank="species" species="tuxtlaensis" status="sp. nov.">
<emphasis id="B967EAE3905BC81C7BC6C458FE7C019A" bold="true" box="[151,436,1900,1926]" italics="true" pageId="4" pageNumber="49">Heterandria tuxtlaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2545798905BC81C7AEDC458FDD1019A" box="[444,537,1900,1926]" pageId="4" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC36F1905BC81C7BC6C4BBFE6E01B5" blockId="4.[151,537,1900,1961]" box="[151,422,1935,1961]" pageId="4" pageNumber="49">
(
<figureCitation id="13282A74905BC81C7BCEC4BBFEC701B5" box="[159,271,1935,1961]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,255,1695,1719]" captionStartId-1="3.[151,255,1745,1769]" captionStartId-2="5.[151,255,663,687]" captionStartId-3="7.[151,255,750,774]" captionTargetBox-0="[273,1305,1273,1656]" captionTargetBox-1="[191,1393,570,1713]" captionTargetBox-2="[291,1296,371,638]" captionTargetBox-3="[265,1323,263,733]" captionTargetId-0="figure@1.[254,1313,1271,1668]" captionTargetId-1="figure@3.[191,1401,557,1713]" captionTargetId-2="figure@5.[291,1296,371,639]" captionTargetId-3="figure@7.[265,1324,263,733]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="7" captionText-0="FIGURE 1. Radiograph negative of a male H. tuxtlaensis showing landmarks used for the body shape analysis. 1 — anterio-dorsal-most position on the snout; 2 — top of head where skull breaks away from the body outline; 3, 4 — origin and insertion of the dorsal fin, respectively; 5 — dorsal origin of the caudal fin (anterio-dorsal-most procurrent ray); 6 — middle of caudal fin base (between hypural plates); 7 — ventral origin of caudal fin (anterio-ventral-most procurrent ray; 8, 9 — insertion and origin of anal fin, respectively; 10 — anterior margin of pelvic fins; 11 — anterio-ventral corner of interoperculum; 12 — first branchiostegal ray at the body outline; 13 — center of orbit (eye position); 14 — junction between cranium and first vertebral centrum; 15 — vertebral centrum bearing third gonapophysis; 16 — reticular point of lower jaw." captionText-1="FIGURE 2. Shape difference between H. tuxtlaensis from Lake Catemaco and H. bimaculata from Sontecomapan. Transformation grids show the linear regression of shape on habitat controlling for allometry. A) Males. The shape gradient for males explained 29.6 % of total shape variance. B) Females. The gradient for females explained 30.5 % of total shape variance. Multivariate least squares means (i. e. group centroids) are plotted and their 95 % confidence limits are given as ellipses." captionText-2="FIGURE 3. Most extreme individuals on the species axis, for males, from our sample of 30 fish. Left: the new species, H. tuxtlaensis. Right: the presumed sister taxon H. bimaculata. Differences in size are incidental and are not responsible for the shape differences observed. The correlation between the species axis and the axis of allometry was r = 0.08." captionText-3="FIGURE 4. Heterandria tuxtlaensis. A. Mature male (Holotype, UMMZ 248620, 44.0 mm SL). B. Mature female (Paratype, TCWC 762.03, 41.3 mm SL). C. Gonopodium tip of Holotype." httpUri-0="https://zenodo.org/record/183068/files/figure.png" httpUri-1="https://zenodo.org/record/183069/files/figure.png" httpUri-2="https://zenodo.org/record/183070/files/figure.png" httpUri-3="https://zenodo.org/record/183071/files/figure.png" pageId="4" pageNumber="49">Figs. 1–4</figureCitation>
, 
<tableCitation id="C691034A905BC81C7A4DC4BBFE5401B5" box="[284,412,1935,1961]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="2.[151,239,1309,1333]" captionStartId-1="4.[151,239,151,175]" captionStartId-2="6.[151,239,271,295]" captionTargetBox-2="[159,1411,405,599]" captionTargetPageId-2="6" captionText-0="TABLE 1. MANCOVA results for morphometric variation associated with habitat and body size." captionText-1="TABLE 2. Species axis loadings for traditional and geometric morphometrics. A. Partial regression coefficients of traditional morphometric characters regressed on species. B. Correlations of superimposed landmark coordinates with the shape gradient separating species." captionText-2="TABLE 3. Traditional morphometrics and meristics of Heterandria tuxtlaensis and H. bimaculata. Except for SL and meristics, data are given in percentage of standard length." httpUri-2="http://table.plazi.org/id/DF6C66799059C81E7BC6C23BFD360755" pageId="4" pageNumber="49" tableUuid-2="DF6C66799059C81E7BC6C23BFD360755">Tables 1–3</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C309657A905BC81D7BC6C4E9FD4606D3" lastPageId="5" lastPageNumber="50" pageId="4" pageNumber="49" type="reference_group">
<paragraph id="8BAC36F1905BC81D7BC6C4E9FD4606D3" blockId="4.[151,1436,2012,2036]" lastBlockId="5.[151,654,152,207]" lastPageId="5" lastPageNumber="50" pageId="4" pageNumber="49">
<taxonomicName id="4C134D72905BC81C7BC6C4E9FE5101E8" ID-CoL="6M2WB" box="[151,409,2013,2036]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="4" pageNumber="49" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE3905BC81C7BC6C4E9FE5101E8" box="[151,409,2013,2036]" italics="true" pageId="4" pageNumber="49">Heterandria bimaculata</emphasis>
</taxonomicName>
[
<emphasis id="B967EAE3905BC81C7AF9C4E9FD2201E8" box="[424,746,2013,2036]" italics="true" pageId="4" pageNumber="49">
non 
<taxonomicName id="4C134D72905BC81C7A86C4E9FD2201E8" ID-CoL="7FWX6" authority="Heckel" authorityName="Heckel" box="[471,746,2013,2036]" class="Actinopterygii" family="Poeciliidae" genus="Xiphophorus" kingdom="Animalia" order="Cyprinodontiformes" pageId="4" pageNumber="49" phylum="Chordata" rank="species" species="bimaculatus">Xiphophorus bimaculatus</taxonomicName>
</emphasis>
Heckel] 
<bibRefCitation id="EF824B00905BC81C781AC4E8FC0701EF" author="Rosen" box="[843,975,2012,2035]" pageId="4" pageNumber="54" refString="Rosen, D. E. (1979) Fishes of the uplands and intermontane basins of Guatemala: revisionary studies and comparative geography. Bulletin of the American Museum of Natural History, 162, 267 - 376." type="journal article" year="1979">Rosen, 1979</bibRefCitation>
(in part, specimens listed as 
<taxonomicName id="4C134D72905BC81C7E55C4E9FA5401E8" ID-CoL="6M2WB" box="[1284,1436,2013,2036]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="4" pageNumber="49" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE3905BC81C7E55C4E9FA5401E8" box="[1284,1436,2013,2036]" italics="true" pageId="4" pageNumber="49">H. bimaculata</emphasis>
</taxonomicName>
from Arroyo Esciunopan, USNM 214151) 
<taxonomicName id="4C134D72905AC81D7BC6C38CFD4606D3" ID-CoL="6LQXZ" authority="Miller and Conner, 1997" authorityName="Miller and Conner" authorityYear="1997" box="[151,654,184,207]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="50" phylum="Chordata" rank="species" species="jonesi">
<emphasis id="B967EAE3905AC81D7BC6C38CFE4A06D3" box="[151,386,184,207]" italics="true" pageId="5" pageNumber="50">Heterandria cf. jonesi</emphasis>
<bibRefCitation id="EF824B00905AC81D7AD9C38CFD4606D3" author="Miller" box="[392,654,184,207]" pageId="5" pageNumber="54" refString="Miller, R. R. &amp; Van Conner, J. (1997) Peces de Catemaco. In: Gonzalez-Soriano, E., Dirzo, R. &amp; Vogt, R. C. (Eds.), Historia natural de Los Tuxtlas. Universidad Nacional Autonoma de Mexico, pp. 451 - 456." type="book chapter" year="1997">Miller and Conner, 1997</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C309657A905AC81D7BC6C3CBFBF10095" pageId="5" pageNumber="50" type="materials_examined">
<paragraph id="8BAC36F1905AC81D7BC6C3CBFD15075E" blockId="5.[151,1436,255,322]" pageId="5" pageNumber="50">
<emphasis id="B967EAE3905AC81D7BC6C3CBFEDA0705" bold="true" box="[151,274,255,281]" pageId="5" pageNumber="50">
<typeStatus id="54A88853905AC81D7BC6C3CBFEC60705" box="[151,270,255,281]" pageId="5" pageNumber="50" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED02AE34905AC81D7A4DC234FEBF0706" box="[284,375,256,282]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="5" pageNumber="50">UMMZ</collectionCode>
248620, 44.0 mm SL, mature male; Veracruz: Laguna Catemaco, south shore near Miniagua; J.V. Conner and J.R. Mayer, 
<date id="FFAD1031905AC81D7967C21CFD11075E" box="[566,729,296,322]" pageId="5" pageNumber="50" value="1964-08-11">11 Aug. 1964</date>
.
</paragraph>
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<paragraph id="8BAC36F1905AC81D7BC6C1A3FAAB04EF" blockId="5.[151,1436,663,755]" pageId="5" pageNumber="50">
<emphasis id="B967EAE3905AC81D7BC6C1A3FED104B3" bold="true" box="[151,281,663,687]" pageId="5" pageNumber="50">FIGURE 3.</emphasis>
Most extreme individuals on the species axis, for males, from our sample of 30 fish. Left: the new species, 
<taxonomicName id="4C134D72905AC81D7BC6C18EFEE204CD" box="[151,298,698,721]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="50" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE3905AC81D7BC6C18EFEE204CD" box="[151,298,698,721]" italics="true" pageId="5" pageNumber="50">H. tuxtlaensis</emphasis>
</taxonomicName>
. Right: the presumed sister taxon 
<taxonomicName id="4C134D72905AC81D79C9C18EFCF904CD" box="[664,817,698,721]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="50" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE3905AC81D79C9C18EFCF904CD" box="[664,817,698,721]" italics="true" pageId="5" pageNumber="50">H. bimaculata</emphasis>
</taxonomicName>
. Differences in size are incidental and are not responsible for the shape differences observed. The correlation between the species axis and the axis of allometry was r = 0.08.
</paragraph>
</caption>
<paragraph id="8BAC36F1905AC81D7B97C013FB13058D" blockId="5.[151,1437,807,2033]" pageId="5" pageNumber="50">
<emphasis id="B967EAE3905AC81D7B97C013FE85055D" bold="true" box="[198,333,807,833]" pageId="5" pageNumber="50">
<typeStatus id="54A88853905AC81D7B97C013FE80055D" box="[198,328,807,833]" pageId="5" pageNumber="50" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="ED02AE34905AC81D7A04C013FE67055D" LSID="urn:lsid:biocol.org:col:34925" box="[341,431,807,833]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="50">AMNH</collectionCode>
242324 (
<specimenCount id="9D15FD78905AC81D7970C013FD0C055D" box="[545,708,807,833]" pageId="5" pageNumber="50" type="generic">15 specimens</specimenCount>
, 
<quantity id="4CEB9B14905AC81D7980C013FC5E055D" box="[721,918,807,833]" metricMagnitude="-2" metricUnit="m" metricValue="5.529999999999999" metricValueMax="7.22" metricValueMin="3.84" pageId="5" pageNumber="50" unit="mm" value="55.3" valueMax="72.2" valueMin="38.4">38.4 to 72.2 mm</quantity>
SL), 
<collectionCode id="ED02AE34905AC81D788BC013FBE7055D" box="[986,1071,807,833]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
762.03 (
<specimenCount id="9D15FD78905AC81D7FC9C013FAF4055D" box="[1176,1340,807,833]" pageId="5" pageNumber="50" type="generic">77 specimens</specimenCount>
, 
<quantity id="4CEB9B14905AC81D7E18C013FF370575" metricMagnitude="-2" metricUnit="m" metricValue="4.255" metricValueMax="6.959999999999999" metricValueMin="1.55" pageId="5" pageNumber="50" unit="mm" value="42.55" valueMax="69.6" valueMin="15.5">15.5 to 69.6 mm</quantity>
SL). 
<collectionCode id="ED02AE34905AC81D7A10C07BFE540575" box="[321,412,847,873]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="5" pageNumber="50">UMMZ</collectionCode>
248621 (
<specimenCount id="9D15FD78905AC81D795DC07BFD660575" box="[524,686,847,873]" pageId="5" pageNumber="50" type="generic">15 specimens</specimenCount>
, 
<quantity id="4CEB9B14905AC81D79EBC07BFCB30575" box="[698,891,847,873]" metricMagnitude="-2" metricUnit="m" metricValue="4.95" metricValueMax="6.15" metricValueMin="3.75" pageId="5" pageNumber="50" unit="mm" value="49.5" valueMax="61.5" valueMin="37.5">37.5 to 61.5 mm</quantity>
SL), 
<collectionCode id="ED02AE34905AC81D78ECC07BFBFF0575" box="[957,1079,847,873]" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="5" pageNumber="50">IBUNAM</collectionCode>
, 14632 (
<specimenCount id="9D15FD78905AC81D7FCDC07BFAF70575" box="[1180,1343,847,873]" pageId="5" pageNumber="50" type="generic">15 specimens</specimenCount>
, 
<quantity id="4CEB9B14905AC81D7E1BC07BFF37058D" metricMagnitude="-2" metricUnit="m" metricValue="4.735" metricValueMax="6.05" metricValueMin="3.42" pageId="5" pageNumber="50" unit="mm" value="47.35" valueMax="60.5" valueMin="34.2">34.2 to 60.5 mm</quantity>
SL), and 
<collectionCode id="ED02AE34905AC81D7A22C043FE00058D" LSID="urn:lsid:biocol.org:col:34871" box="[371,456,887,913]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="50">USNM</collectionCode>
14632 (
<specimenCount id="9D15FD78905AC81D7979C043FD0F058D" box="[552,711,887,913]" pageId="5" pageNumber="50" type="generic">15 specimens</specimenCount>
<quantity id="4CEB9B14905AC81D799EC043FC58058D" box="[719,912,887,913]" metricMagnitude="-2" metricUnit="m" metricValue="4.459999999999999" metricValueMax="5.93" metricValueMin="2.9899999999999998" pageId="5" pageNumber="50" unit="mm" value="44.599999999999994" valueMax="59.3" valueMin="29.9">29.9 to 59.3 mm</quantity>
SL), same data as 
<typeStatus id="54A88853905AC81D7F3CC043FB1F058D" box="[1133,1239,887,913]" pageId="5" pageNumber="50" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BAC36F1905AC81D7B97C0ABFBF10095" blockId="5.[151,1437,807,2033]" pageId="5" pageNumber="50">
<emphasis id="B967EAE3905AC81D7B97C0ABFE4605A5" bold="true" box="[198,398,927,953]" pageId="5" pageNumber="50">Other material:</emphasis>
<collectionCode id="ED02AE34905AC81D7AC8C0ABFE3B05A5" LSID="urn:lsid:biocol.org:col:34925" box="[409,499,927,953]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="50">AMNH</collectionCode>
78581 (
<specimenCount id="9D15FD78905AC81D7909C0ABFD3505A5" box="[600,765,927,953]" pageId="5" pageNumber="50" type="generic">26 specimens</specimenCount>
) Veracruz, small stream through a pasture 
<quantity id="4CEB9B14905AC81D7E54C0ABFA9105A5" box="[1285,1369,927,953]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="50" unit="km" value="0.5">0.5 km</quantity>
from Lake Catemaco on road beyond Coyame; 
<collectionCode id="ED02AE34905AC81D79D3C0F3FD1405FD" LSID="urn:lsid:biocol.org:col:34925" box="[642,732,967,993]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="50">AMNH</collectionCode>
78593 (
<specimenCount id="9D15FD78905AC81D786DC0F3FC2305FD" box="[828,1003,967,993]" pageId="5" pageNumber="50" type="generic">113 specimens</specimenCount>
) Veracruz, small stream running into Lake Catemaco, just S of outflow; 
<collectionCode id="ED02AE34905AC81D796BC0DBFD5C0215" LSID="urn:lsid:biocol.org:col:34925" box="[570,660,1007,1033]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="50">AMNH</collectionCode>
78597 (
<specimenCount id="9D15FD78905AC81D79A6C0DBFC520215" box="[759,922,1007,1033]" pageId="5" pageNumber="50" type="generic">61 specimens</specimenCount>
) Veracruz, stream below Sihuapan flowing into outflow (Río Grande) of Lake Catemaco above falls; 
<collectionCode id="ED02AE34905AC81D786CC723FC5F022D" LSID="urn:lsid:biocol.org:col:34925" box="[829,919,1047,1073]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="5" pageNumber="50">AMNH</collectionCode>
78604 (
<specimenCount id="9D15FD78905AC81D78A7C723FB50022D" box="[1014,1176,1047,1073]" pageId="5" pageNumber="50" type="generic">44 specimens</specimenCount>
) Veracruz, small river near Quetzalan flowing into Lake Catemaco; 
<collectionCode id="ED02AE34905AC81D79E9C70BFCCE0245" box="[696,774,1087,1113]" httpUri="http://grbio.org/cool/v821-4x4f" name="Gulf Coast Research Laboratory" pageId="5" pageNumber="50">GCRL</collectionCode>
<date id="FFAD1031905AC81D785EC70BFC8E0245" box="[783,838,1087,1113]" pageId="5" pageNumber="50" value="1971-05">V71</date>
: 6828 (
<specimenCount id="9D15FD78905AC81D78F1C70BFB9A0245" box="[928,1106,1087,1113]" pageId="5" pageNumber="50" type="generic">107 specimens</specimenCount>
) Veracruz, Lake Catemaco, road from San Andres Tuxtla to Tehuantepec, 
<quantity id="4CEB9B14905AC81D79EEC753FCC2029D" box="[703,778,1127,1153]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="50" unit="km" value="10.0">10 km</quantity>
South of San Andres Tuxtla; 
<collectionCode id="ED02AE34905AC81D7F3EC753FB0C029D" box="[1135,1220,1127,1153]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
752.01 (
<specimenCount id="9D15FD78905AC81D7E7EC753FF1F02B5" pageId="5" pageNumber="50" type="generic">22 specimens</specimenCount>
) Veracruz, Río Quezelapam, 
<quantity id="4CEB9B14905AC81D7962C7BBFDB802B5" box="[563,624,1167,1193]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="50" unit="km" value="3.0">3 km</quantity>
east of Lake Catemaco; 
<collectionCode id="ED02AE34905AC81D78C6C7BBFC2402B5" box="[919,1004,1167,1193]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
756.02 (
<specimenCount id="9D15FD78905AC81D7F06C7BBFB2202B5" box="[1111,1258,1167,1193]" pageId="5" pageNumber="50" type="generic">8 specimens</specimenCount>
) Veracruz, Río Grande at hydroelectric plant, about 
<quantity id="4CEB9B14905AC81D7903C783FD5802CD" box="[594,656,1207,1233]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="50" unit="km" value="9.0">9 km</quantity>
southwest of San Andrés Tuxtla; 
<collectionCode id="ED02AE34905AC81D7F7CC783FB4A02CD" box="[1069,1154,1207,1233]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
757.02 (
<specimenCount id="9D15FD78905AC81D7FBEC783FA5C02CD" box="[1263,1428,1207,1233]" pageId="5" pageNumber="50" type="generic">30 specimens</specimenCount>
) Veracruz, Lake Catemaco at dam spillway; 
<collectionCode id="ED02AE34905AC81D79C9C7EBFD2502E5" box="[664,749,1247,1273]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
760.01 (
<specimenCount id="9D15FD78905AC81D7804C7EBFC3002E5" box="[853,1016,1247,1273]" pageId="5" pageNumber="50" type="generic">94 specimens</specimenCount>
) Veracruz, Lake Encantada, 3.0 km northeast of San Andrés Tuxtla; 
<collectionCode id="ED02AE34905AC81D7949C633FDA5033D" box="[536,621,1287,1313]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
762 (
<specimenCount id="9D15FD78905AC81D79E3C633FCAC033D" box="[690,868,1287,1313]" pageId="5" pageNumber="50" type="generic">139 specimens</specimenCount>
) Veracruz, Lake Catemaco, south shore of lake; 
<collectionCode id="ED02AE34905AC81D7BC6C61BFF240355" box="[151,236,1327,1353]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
775.01 (
<specimenCount id="9D15FD78905AC81D7A02C61BFE2D0355" box="[339,485,1327,1353]" pageId="5" pageNumber="50" type="generic">9 specimens</specimenCount>
) Veracruz, Río Grande, at spillway below dam, about 
<quantity id="4CEB9B14905AC81D7F0FC61BFB520355" box="[1118,1178,1327,1353]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="50" unit="km" value="1.0">1 km</quantity>
west of highway 180; 
<collectionCode id="ED02AE34905AC81D7BC6C663FF24036D" box="[151,236,1367,1393]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
776.01 (
<specimenCount id="9D15FD78905AC81D7A0BC663FDC8036D" box="[346,512,1367,1393]" pageId="5" pageNumber="50" type="generic">52 specimens</specimenCount>
) Veracruz, Río Grande, reservoir of hydroelectric dam, about 
<quantity id="4CEB9B14905AC81D7FA8C663FAF0036D" box="[1273,1336,1367,1393]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="50" unit="km" value="1.0">1 km</quantity>
west of highway 180; 
<collectionCode id="ED02AE34905AC81D7A11C64BFE5D0385" box="[320,405,1407,1433]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
809.03 (
<specimenCount id="9D15FD78905AC81D7AAFC64BFD690385" box="[510,673,1407,1433]" pageId="5" pageNumber="50" type="generic">12 specimens</specimenCount>
) Veracruz, Lake Catemaco, south shore of lake; 
<collectionCode id="ED02AE34905AC81D7FB1C64BFAFD0385" box="[1248,1333,1407,1433]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
1847.03 (
<specimenCount id="9D15FD78905AC81D7BF3C693FEE603DD" box="[162,302,1447,1473]" pageId="5" pageNumber="50" type="generic">1 specimen</specimenCount>
) Veracruz, Lake Catemaco, at Cemolapan; 
<collectionCode id="ED02AE34905AC81D7809C693FC6503DD" box="[856,941,1447,1473]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
1849.02 (
<specimenCount id="9D15FD78905AC81D7F63C693FB0503DD" box="[1074,1229,1447,1473]" pageId="5" pageNumber="50" type="generic">2 specimens</specimenCount>
) Veracruz, Lake Catemaco, at Coyame; 
<collectionCode id="ED02AE34905AC81D7AFEC6FBFDCC03F5" box="[431,516,1487,1513]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
1850.01 (
<specimenCount id="9D15FD78905AC81D792EC6FBFCEB03F5" box="[639,803,1487,1513]" pageId="5" pageNumber="50" type="generic">14 specimens</specimenCount>
) Veracruz, Lake Chalchoapan, just west of mouth of Río Grande west of Lake Catemaco; 
<collectionCode id="ED02AE34905AC81D7916C6C3FD54000D" box="[583,668,1527,1553]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="5" pageNumber="50">TCWC</collectionCode>
1854.01 (
<specimenCount id="9D15FD78905AC81D7840C6C3FC6C000D" box="[785,932,1527,1553]" pageId="5" pageNumber="50" type="generic">6 specimens</specimenCount>
) Veracruz, Lake Catemaco; TU uncat. Veracruz, Lake Catemaco, at outlet of Río San Andres, about 
<quantity id="4CEB9B14905AC81D7815C52BFC6D0025" box="[836,933,1567,1593]" metricMagnitude="4" metricUnit="m" metricValue="2.18" pageId="5" pageNumber="50" unit="km" value="21.8">21.8 km</quantity>
east of San Andres, highway 180; 
<collectionCode id="ED02AE34905AC81D7E11C52BFA530025" box="[1344,1435,1567,1593]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="5" pageNumber="50">UMMZ</collectionCode>
178556 (
<specimenCount id="9D15FD78905AC81D7A50C573FE6E007D" box="[257,422,1607,1633]" pageId="5" pageNumber="50" type="generic">30 specimens</specimenCount>
) Veracruz, Lake Catemaco at Playa Azul, ca 
<quantity id="4CEB9B14905AC81D789EC573FBC6007D" box="[975,1038,1607,1633]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="50" unit="km" value="3.0">3 km</quantity>
E of Catemaco; 
<collectionCode id="ED02AE34905AC81D7F8EC573FAF2007D" box="[1247,1338,1607,1633]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="5" pageNumber="50">UMMZ</collectionCode>
178557 (
<specimenCount id="9D15FD78905AC81D7BCEC55BFE980095" box="[159,336,1647,1673]" pageId="5" pageNumber="50" type="generic">164 specimens</specimenCount>
) Veracruz, tributary of Lake Catemaco on hwy 
<quantity id="4CEB9B14905AC81D782FC55BFC180095" box="[894,976,1647,1673]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="50" unit="km" value="3.2">3.2 km</quantity>
SE lake.
</paragraph>
</subSubSection>
<subSubSection id="C309657A905AC81D7B97C5A3FB560165" pageId="5" pageNumber="50" type="diagnosis">
<paragraph id="8BAC36F1905AC81D7B97C5A3FB560165" blockId="5.[151,1437,807,2033]" pageId="5" pageNumber="50">
<emphasis id="B967EAE3905AC81D7B97C5A3FE8E00AD" bold="true" box="[198,326,1687,1713]" pageId="5" pageNumber="50">Diagnosis.</emphasis>
The following characters distinguished 
<taxonomicName id="4C134D72905AC81D7874C5ACFC0F00AD" box="[805,967,1688,1713]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="50" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE3905AC81D7874C5ACFC0F00AD" box="[805,967,1688,1713]" italics="true" pageId="5" pageNumber="50">H. tuxtlaensis</emphasis>
</taxonomicName>
from the other species of 
<taxonomicName id="4C134D72905AC81D7E52C5ACFA5C00AD" box="[1283,1428,1688,1713]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B967EAE3905AC81D7E52C5ACFA5C00AD" box="[1283,1428,1688,1713]" italics="true" pageId="5" pageNumber="50">Heterandria</emphasis>
</taxonomicName>
: body depth in males 23.2 to 27.1% (=25.3%) of SL; dorsal fin base length 26.5 to 31.3% (=28.9%) of SL in males and 21.9 to 28.2% (=25.6%) SL in females; dorsal fin rays 12 or 13 (=12.7); basicaudal spot relatively small, subcircular to bar-shaped, and limited to area above midlateral line and not extending to dorsal midline; serrae on gonopodial ray 4p arched away from ray 4a; subdistal small segments on gonopodial ray 4a number 5 to 10; and terminal segment on gonopodial ray 4a long and strongly recurved.
</paragraph>
</subSubSection>
<subSubSection id="C309657A905AC81E7B97C4B3FDB0018E" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50" type="description">
<paragraph id="8BAC36F1905AC81E7B97C4B3FE1906C6" blockId="5.[151,1437,807,2033]" lastBlockId="6.[151,1436,152,218]" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
<emphasis id="B967EAE3905AC81D7B97C4B3FE9601BD" bold="true" box="[198,350,1927,1953]" pageId="5" pageNumber="50">Description.</emphasis>
<taxonomicName id="4C134D72905AC81D7A37C4BCFDB301BD" box="[358,635,1928,1953]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="50" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE3905AC81D7A37C4BCFDB301BD" box="[358,635,1928,1953]" italics="true" pageId="5" pageNumber="50">Heterandria tuxtlaensis</emphasis>
</taxonomicName>
is a moderate-sized 
<taxonomicName id="4C134D72905AC81D783FC4BCFC3701BD" box="[878,1023,1928,1953]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="5" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="B967EAE3905AC81D783FC4BCFC3701BD" box="[878,1023,1928,1953]" italics="true" pageId="5" pageNumber="50">Heterandria</emphasis>
</taxonomicName>
(up to 
<quantity id="4CEB9B14905AC81D7F02C4B3FB6E01BD" box="[1107,1190,1927,1953]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="50" unit="mm" value="49.0">49 mm</quantity>
SL for males and 
<quantity id="4CEB9B14905AC81D7E2EC4B3FF0D01D5" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="50" unit="mm" value="75.0">75 mm</quantity>
SL for females). Meristic and morphometric variables are summarized in 
<tableCitation id="C691034A905AC81D7F15C49BFB5701D5" box="[1092,1183,1967,1993]" captionStart="TABLE 3" captionStartId="6.[151,239,271,295]" captionTargetBox="[159,1411,405,599]" captionTargetPageId="6" captionText="TABLE 3. Traditional morphometrics and meristics of Heterandria tuxtlaensis and H. bimaculata. Except for SL and meristics, data are given in percentage of standard length." httpUri="http://table.plazi.org/id/DF6C66799059C81E7BC6C23BFD360755" pageId="5" pageNumber="50" tableUuid="DF6C66799059C81E7BC6C23BFD360755">Table 3</tableCitation>
. This species is relatively slender, with a short dorsal fin, shallow caudal peduncle, and a short caudal fin. In females head is relatively narrow and anal fin is relatively short. Pectoral fin rays 15 or 16, lateral line scales 28 to 32, and vertebrae 29 to 33 (
<figureCitation id="13282A749059C81E7A2AC3F4FE0C06C6" box="[379,452,192,218]" captionStart="FIGURE 4" captionStartId="7.[151,255,750,774]" captionTargetBox="[265,1323,263,733]" captionTargetId="figure@7.[265,1324,263,733]" captionTargetPageId="7" captionText="FIGURE 4. Heterandria tuxtlaensis. A. Mature male (Holotype, UMMZ 248620, 44.0 mm SL). B. Mature female (Paratype, TCWC 762.03, 41.3 mm SL). C. Gonopodium tip of Holotype." httpUri="https://zenodo.org/record/183071/files/figure.png" pageId="6" pageNumber="51">Fig. 4</figureCitation>
).
</paragraph>
<caption id="DF6C66799059C81E7BC6C23BFD360755" ID-Table-UUID="DF6C66799059C81E7BC6C23BFD360755" httpUri="http://table.plazi.org/id/DF6C66799059C81E7BC6C23BFD360755" pageId="6" pageNumber="51" targetBox="[159,1411,405,599]" targetIsTable="true" targetPageId="6">
<paragraph id="8BAC36F19059C81E7BC6C23BFD360755" blockId="6.[151,1435,271,329]" pageId="6" pageNumber="51">
<emphasis id="B967EAE39059C81E7BC6C23BFEC2073B" bold="true" box="[151,266,271,295]" pageId="6" pageNumber="51">TABLE 3.</emphasis>
Traditional morphometrics and meristics of 
<taxonomicName id="4C134D729059C81E79A2C224FC39073B" box="[755,1009,272,295]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="51" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39059C81E79A2C224FC39073B" box="[755,1009,272,295]" italics="true" pageId="6" pageNumber="51">Heterandria tuxtlaensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C134D729059C81E7F7BC224FB0C073B" box="[1066,1220,272,295]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="51" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39059C81E7F7BC224FB0C073B" box="[1066,1220,272,295]" italics="true" pageId="6" pageNumber="51">H. bimaculata</emphasis>
</taxonomicName>
. Except for SL and meristics, data are given in percentage of standard length.
</paragraph>
</caption>
<paragraph id="8BAC36F19059C81E7BCEC25EFE2D004E" blockId="6.[159,1285,362,385]" lastBlockId="6.[151,1437,1312,2018]" pageId="6" pageNumber="51">
Measurements 
<typeStatus id="54A888539059C81E7AFAC25EFDC6079D" box="[427,526,362,385]" pageId="6" pageNumber="51" type="holotype">Holotype</typeStatus>
<typeStatus id="54A888539059C81E7972C25EFD43079D" box="[547,651,362,385]" pageId="6" pageNumber="51" type="paratype">Paratypes</typeStatus>
(
<taxonomicName id="4C134D729059C81E79CAC25EFCE5079D" box="[667,813,362,385]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="51" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39059C81E79CAC25EFCE5079D" box="[667,813,362,385]" italics="true" pageId="6" pageNumber="51">H. tuxtlaensis</emphasis>
</taxonomicName>
) Non-types (
<taxonomicName id="4C134D729059C81E7F35C25EFB35079D" box="[1124,1277,362,385]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="6" pageNumber="51" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39059C81E7F35C25EFB35079D" box="[1124,1277,362,385]" italics="true" pageId="6" pageNumber="51">H. bimaculata</emphasis>
</taxonomicName>
) Ray 3 of gonopodium terminating at level of second to last subterminal segment of ray 4a in 
<typeStatus id="54A888539059C81E7FADC614FAAB0326" box="[1276,1379,1312,1338]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
(terminating at level of last subterminal or terminal segment in male 
<typeStatus id="54A888539059C81E78C3C67CFBCF037E" box="[914,1031,1352,1378]" pageId="6" pageNumber="51" type="paratype">paratypes</typeStatus>
), distal segments of ray 3 enlarged ventrally to form elbow and running contiguous with ray 4a, segments distal to elbow number 7 to 9 (
<figureCitation id="13282A749059C81E7E14C644FA460396" box="[1349,1422,1392,1418]" captionStart="FIGURE 4" captionStartId="7.[151,255,750,774]" captionTargetBox="[265,1323,263,733]" captionTargetId="figure@7.[265,1324,263,733]" captionTargetPageId="7" captionText="FIGURE 4. Heterandria tuxtlaensis. A. Mature male (Holotype, UMMZ 248620, 44.0 mm SL). B. Mature female (Paratype, TCWC 762.03, 41.3 mm SL). C. Gonopodium tip of Holotype." httpUri="https://zenodo.org/record/183071/files/figure.png" pageId="6" pageNumber="51">Fig. 4</figureCitation>
). Subdistal 7 to 10 segments of ray 4a abruptly shortened, anterior 4 to 6 forming peg-like processes and posterior 3 to 5 squarish. Distal segment of ray 4a greatly elongated, about 5 to 7 times length of proximal squarish segment, and decurved about 110º to 120º. Subdistal 10 to 14 segments of ray 4p bear serrae, distal segments of ray 4p number 5 to 10 and lack serrae. Ray 5a arches downward toward upwardly arched ray 4p and terminates medial to tip of ray 4a.
</paragraph>
<paragraph id="8BAC36F19059C81E7AFAC2A1FABD044B" pageId="6" pageNumber="51">
<table id="F913C451905937E77BCEC2A1FA4B044B" box="[159,1411,405,599]" gridcols="6" gridrows="4" pageId="6" pageNumber="51">
<tr id="352334B3905937E77BCEC2A1FA4B07B0" box="[159,1411,405,428]" gridrow="0" pageId="6" pageNumber="51" rowspan-0="1">
<th id="76F25DCF905937E77AFAC2A1FE2A07B0" box="[427,482,405,428]" gridcol="1" gridrow="0" pageId="6" pageNumber="51">Male</th>
<th id="76F25DCF905937E77972C2A1FD2807B0" box="[547,736,405,428]" gridcol="2" gridrow="0" pageId="6" pageNumber="51">Males</th>
<th id="76F25DCF905937E77855C2A1FC0907B0" box="[772,961,405,428]" gridcol="3" gridrow="0" pageId="6" pageNumber="51">Females</th>
<th id="76F25DCF905937E778B4C2A1FB6A07B0" box="[997,1186,405,428]" gridcol="4" gridrow="0" pageId="6" pageNumber="51">Males</th>
<th id="76F25DCF905937E77F97C2A1FA4B07B0" box="[1222,1411,405,428]" gridcol="5" gridrow="0" pageId="6" pageNumber="51">Females</th>
</tr>
<tr id="352334B3905937E77BCEC2F4FA4B041D" box="[159,1411,448,513]" gridrow="1" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC2F4FE85041D" box="[159,333,448,513]" gridcol="0" gridrow="1" pageId="6" pageNumber="51">Standard length</th>
<td id="76F25DCF905937E77AFAC2F4FE2A041D" box="[427,482,448,513]" gridcol="1" gridrow="1" pageId="6" pageNumber="51">44</td>
<td id="76F25DCF905937E77972C2F4FD28041D" box="[547,736,448,513]" gridcol="2" gridrow="1" pageId="6" pageNumber="51">Range Mean 31.8–49.4 43</td>
<td id="76F25DCF905937E77855C2F4FC09041D" box="[772,961,448,513]" gridcol="3" gridrow="1" pageId="6" pageNumber="51">Range Mean 33.7–74.5 55.8</td>
<td id="76F25DCF905937E778B4C2F4FB6A041D" box="[997,1186,448,513]" gridcol="4" gridrow="1" pageId="6" pageNumber="51">Range Mean 30.1–40.0 36.3</td>
<td id="76F25DCF905937E77F97C2F4FA4B041D" box="[1222,1411,448,513]" gridcol="5" gridrow="1" pageId="6" pageNumber="51">Range Mean 39.6–64.6 52.2</td>
</tr>
<tr id="352334B3905937E77BCEC121FA4B0430" box="[159,1411,533,556]" gridrow="2" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC121FE850430" box="[159,333,533,556]" gridcol="0" gridrow="2" pageId="6" pageNumber="51">Depth</th>
<td id="76F25DCF905937E77AFAC121FE2A0430" box="[427,482,533,556]" gridcol="1" gridrow="2" pageId="6" pageNumber="51">26.4</td>
<td id="76F25DCF905937E77972C121FD280430" box="[547,736,533,556]" gridcol="2" gridrow="2" pageId="6" pageNumber="51">23.2–27.1 25.3</td>
<td id="76F25DCF905937E77855C121FC090430" box="[772,961,533,556]" gridcol="3" gridrow="2" pageId="6" pageNumber="51">23.9–27.6 24.3</td>
<td id="76F25DCF905937E778B4C121FB6A0430" box="[997,1186,533,556]" gridcol="4" gridrow="2" pageId="6" pageNumber="51">26.2–29.9 28.1</td>
<td id="76F25DCF905937E77F97C121FA4B0430" box="[1222,1411,533,556]" gridcol="5" gridrow="2" pageId="6" pageNumber="51">25.7–29.5 27.5</td>
</tr>
<tr id="352334B3905937E77BCEC174FA4B044B" box="[159,1411,576,599]" gridrow="3" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC174FE85044B" box="[159,333,576,599]" gridcol="0" gridrow="3" pageId="6" pageNumber="51">Predorsal length</th>
<td id="76F25DCF905937E77AFAC174FE2A044B" box="[427,482,576,599]" gridcol="1" gridrow="3" pageId="6" pageNumber="51">50.9</td>
<td id="76F25DCF905937E77972C174FD28044B" box="[547,736,576,599]" gridcol="2" gridrow="3" pageId="6" pageNumber="51">48.3–53.3 51.1</td>
<td id="76F25DCF905937E77855C174FC09044B" box="[772,961,576,599]" gridcol="3" gridrow="3" pageId="6" pageNumber="51">54.7–58.1 56</td>
<td id="76F25DCF905937E778B4C174FB6A044B" box="[997,1186,576,599]" gridcol="4" gridrow="3" pageId="6" pageNumber="51">49.3–54.2 51.9</td>
<td id="76F25DCF905937E77F97C174FA4B044B" box="[1222,1411,576,599]" gridcol="5" gridrow="3" pageId="6" pageNumber="51">53.1–59.6 55.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAC36F19059C81E7BCEC15EFABD02D3" pageId="6" pageNumber="51">
<table id="F913C451905937E77BCEC15EFA4B02D3" box="[159,1411,618,1231]" gridcols="5" gridrows="10" pageId="6" pageNumber="51">
<tr id="352334B3905937E77BCEC15EFA4B049D" box="[159,1411,618,641]" gridrow="0" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC15EFE12049D" box="[159,474,618,641]" gridcol="0" gridrow="0" pageId="6" pageNumber="51">Dorsal-caudal peduncle 49.6</th>
<th id="76F25DCF905937E77972C15EFD1A049D" box="[547,722,618,641]" gridcol="1" gridrow="0" pageId="6" pageNumber="51">45.4–53.6 50.2</th>
<th id="76F25DCF905937E77855C15EFC7B049D" box="[772,947,618,641]" gridcol="2" gridrow="0" pageId="6" pageNumber="51">44.2–48.5 46</th>
<th id="76F25DCF905937E778B4C15EFB5C049D" box="[997,1172,618,641]" gridcol="3" gridrow="0" pageId="6" pageNumber="51">50.0–55.4 52.7</th>
<th id="76F25DCF905937E77F97C15EFA4B049D" box="[1222,1411,618,641]" gridcol="4" gridrow="0" pageId="6" pageNumber="51">43.5–50.0 47.5</th>
</tr>
<tr id="352334B3905937E77BCEC1A1FA4B04B0" box="[159,1411,661,684]" gridrow="1" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC1A1FE1204B0" box="[159,474,661,684]" gridcol="0" gridrow="1" pageId="6" pageNumber="51">Anal-caudal peduncle -</th>
<td id="76F25DCF905937E77972C1A1FD1A04B0" box="[547,722,661,684]" gridcol="1" gridrow="1" pageId="6" pageNumber="51">- -</td>
<td id="76F25DCF905937E77855C1A1FC7B04B0" box="[772,947,661,684]" gridcol="2" gridrow="1" pageId="6" pageNumber="51">44.5–47.9 46</td>
<td id="76F25DCF905937E778B4C1A1FB5C04B0" box="[997,1172,661,684]" gridcol="3" gridrow="1" pageId="6" pageNumber="51">- -</td>
<td id="76F25DCF905937E77F97C1A1FA4B04B0" box="[1222,1411,661,684]" gridcol="4" gridrow="1" pageId="6" pageNumber="51">43.2–47.6 44.9</td>
</tr>
<tr id="352334B3905937E77BCEC1F4FA4B051D" box="[159,1411,704,769]" gridrow="2" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC1F4FE12051D" box="[159,474,704,769]" gridcol="0" gridrow="2" pageId="6" pageNumber="51">Head length 27.1 Head width 13.5</th>
<td id="76F25DCF905937E77972C1F4FD1A051D" box="[547,722,704,769]" gridcol="1" gridrow="2" pageId="6" pageNumber="51">24.0–27.0 25.5 12.0–14.1 13.3</td>
<td id="76F25DCF905937E77855C1F4FC7B051D" box="[772,947,704,769]" gridcol="2" gridrow="2" pageId="6" pageNumber="51">25.0–27.7 26 15.2–16.7 15.8</td>
<td id="76F25DCF905937E778B4C1F4FB5C051D" box="[997,1172,704,769]" gridcol="3" gridrow="2" pageId="6" pageNumber="51">25.3–27.9 26.5 12.9–15.3 14</td>
<td id="76F25DCF905937E77F97C1F4FA4B051D" box="[1222,1411,704,769]" gridcol="4" gridrow="2" pageId="6" pageNumber="51">26.6–29.9 27.6 16.1–18.2 17.4</td>
</tr>
<tr id="352334B3905937E77BCEC021FA4B0530" box="[159,1411,789,812]" gridrow="3" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC021FE120530" box="[159,474,789,812]" gridcol="0" gridrow="3" pageId="6" pageNumber="51">Snout length 7.9</th>
<td id="76F25DCF905937E77972C021FD1A0530" box="[547,722,789,812]" gridcol="1" gridrow="3" pageId="6" pageNumber="51">5.9–8.4 7.4</td>
<td id="76F25DCF905937E77855C021FC7B0530" box="[772,947,789,812]" gridcol="2" gridrow="3" pageId="6" pageNumber="51">6.9–9.7 8.3</td>
<td id="76F25DCF905937E778B4C021FB5C0530" box="[997,1172,789,812]" gridcol="3" gridrow="3" pageId="6" pageNumber="51">7.5–9.2 8.4</td>
<td id="76F25DCF905937E77F97C021FA4B0530" box="[1222,1411,789,812]" gridcol="4" gridrow="3" pageId="6" pageNumber="51">8.1–10.4 9.3</td>
</tr>
<tr id="352334B3905937E77BCEC074FA4B059D" box="[159,1411,832,897]" gridrow="4" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC074FE12059D" box="[159,474,832,897]" gridcol="0" gridrow="4" pageId="6" pageNumber="51">Peduncle depth 17.8 Dorsal base length 28.5</th>
<td id="76F25DCF905937E77972C074FD1A059D" box="[547,722,832,897]" gridcol="1" gridrow="4" pageId="6" pageNumber="51">15.4–18.1 16.8 26.5–31.3 28.9</td>
<td id="76F25DCF905937E77855C074FC7B059D" box="[772,947,832,897]" gridcol="2" gridrow="4" pageId="6" pageNumber="51">14.5–16.4 15.5 21.9–28.2 25.6</td>
<td id="76F25DCF905937E778B4C074FB5C059D" box="[997,1172,832,897]" gridcol="3" gridrow="4" pageId="6" pageNumber="51">16.6–19.1 17.7 29.8–33.4 31.7</td>
<td id="76F25DCF905937E77F97C074FA4B059D" box="[1222,1411,832,897]" gridcol="4" gridrow="4" pageId="6" pageNumber="51">15.1–16.7 16 27.1–31.7 29.1</td>
</tr>
<tr id="352334B3905937E77BCEC0A1FA4B05E5" box="[159,1411,917,1017]" gridrow="5" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC0A1FE1205E5" box="[159,474,917,1017]" gridcol="0" gridrow="5" pageId="6" pageNumber="51">Depressed dorsal 40.4 length Depressed anal length -</th>
<td id="76F25DCF905937E77972C0A1FD1A05E5" box="[547,722,917,1017]" gridcol="1" gridrow="5" pageId="6" pageNumber="51">38.7–48.8 41.6 - -</td>
<td id="76F25DCF905937E77855C0A1FC7B05E5" box="[772,947,917,1017]" gridcol="2" gridrow="5" pageId="6" pageNumber="51">33.2–38.2 36 21.0–24.6 22.7</td>
<td id="76F25DCF905937E778B4C0A1FB5C05E5" box="[997,1172,917,1017]" gridcol="3" gridrow="5" pageId="6" pageNumber="51">44.7–47.9 46.7 - -</td>
<td id="76F25DCF905937E77F97C0A1FA4B05E5" box="[1222,1411,917,1017]" gridcol="4" gridrow="5" pageId="6" pageNumber="51">39.3–45.1 42 23.1–27.1 24.7</td>
</tr>
<tr id="352334B3905937E77BCEC739FA4B0238" box="[159,1411,1037,1060]" gridrow="6" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC739FE120238" box="[159,474,1037,1060]" gridcol="0" gridrow="6" pageId="6" pageNumber="51">Caudal length 23.7</th>
<td id="76F25DCF905937E77972C739FD1A0238" box="[547,722,1037,1060]" gridcol="1" gridrow="6" pageId="6" pageNumber="51">21.2–24.9 23.1</td>
<td id="76F25DCF905937E77855C739FC7B0238" box="[772,947,1037,1060]" gridcol="2" gridrow="6" pageId="6" pageNumber="51">19.1–22.8 20.4</td>
<td id="76F25DCF905937E778B4C739FB5C0238" box="[997,1172,1037,1060]" gridcol="3" gridrow="6" pageId="6" pageNumber="51">25.5–30.8 27.9</td>
<td id="76F25DCF905937E77F97C739FA4B0238" box="[1222,1411,1037,1060]" gridcol="4" gridrow="6" pageId="6" pageNumber="51">23.7–30.2 26.4</td>
</tr>
<tr id="352334B3905937E77BCEC70CFA4B0265" box="[159,1411,1080,1145]" gridrow="7" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC70CFE120265" box="[159,474,1080,1145]" gridcol="0" gridrow="7" pageId="6" pageNumber="51">Dorsal rays 13 Pectoral rays 16</th>
<td id="76F25DCF905937E77972C70CFD1A0265" box="[547,722,1080,1145]" gridcol="1" gridrow="7" pageId="6" pageNumber="51">12–13 12.6 15–16 15.5</td>
<td id="76F25DCF905937E77855C70CFC7B0265" box="[772,947,1080,1145]" gridcol="2" gridrow="7" pageId="6" pageNumber="51">12–13 12.9 15–16 15.3</td>
<td id="76F25DCF905937E778B4C70CFB5C0265" box="[997,1172,1080,1145]" gridcol="3" gridrow="7" pageId="6" pageNumber="51">14–15 14.5 14–16 14.5</td>
<td id="76F25DCF905937E77F97C70CFA4B0265" box="[1222,1411,1080,1145]" gridcol="4" gridrow="7" pageId="6" pageNumber="51">14–15 14.3 14–16 14.6</td>
</tr>
<tr id="352334B3905937E77BCEC7B9FA4B02B8" box="[159,1411,1165,1188]" gridrow="8" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC7B9FE1202B8" box="[159,474,1165,1188]" gridcol="0" gridrow="8" pageId="6" pageNumber="51">Lateral line scales 29</th>
<td id="76F25DCF905937E77972C7B9FD1A02B8" box="[547,722,1165,1188]" gridcol="1" gridrow="8" pageId="6" pageNumber="51">28–32 29.6</td>
<td id="76F25DCF905937E77855C7B9FC7B02B8" box="[772,947,1165,1188]" gridcol="2" gridrow="8" pageId="6" pageNumber="51">28–30 29.2</td>
<td id="76F25DCF905937E778B4C7B9FB5C02B8" box="[997,1172,1165,1188]" gridcol="3" gridrow="8" pageId="6" pageNumber="51">27–29 27.8</td>
<td id="76F25DCF905937E77F97C7B9FA4B02B8" box="[1222,1411,1165,1188]" gridcol="4" gridrow="8" pageId="6" pageNumber="51">27–29 28</td>
</tr>
<tr id="352334B3905937E77BCEC78CFA4B02D3" box="[159,1411,1208,1231]" gridrow="9" pageId="6" pageNumber="51">
<th id="76F25DCF905937E77BCEC78CFE1202D3" box="[159,474,1208,1231]" gridcol="0" gridrow="9" pageId="6" pageNumber="51">Vertebrae number 33</th>
<td id="76F25DCF905937E77972C78CFD1A02D3" box="[547,722,1208,1231]" gridcol="1" gridrow="9" pageId="6" pageNumber="51">29–33 30.6</td>
<td id="76F25DCF905937E77855C78CFC7B02D3" box="[772,947,1208,1231]" gridcol="2" gridrow="9" pageId="6" pageNumber="51">29–31 30.4</td>
<td id="76F25DCF905937E778B4C78CFB5C02D3" box="[997,1172,1208,1231]" gridcol="3" gridrow="9" pageId="6" pageNumber="51">30–32 31.2</td>
<td id="76F25DCF905937E77F97C78CFA4B02D3" box="[1222,1411,1208,1231]" gridcol="4" gridrow="9" pageId="6" pageNumber="51">29–31 30.1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAC36F19059C81E7B97C56BFDB0018E" blockId="6.[151,1437,1312,2018]" pageId="6" pageNumber="51">
<emphasis id="B967EAE39059C81E7B97C56BFE1C0065" bold="true" box="[198,468,1631,1657]" pageId="6" pageNumber="51">Coloration in alcohol.</emphasis>
Body is tan, with edges of scale pockets dark brown producing cross hatching pattern on dorsal and lateral surfaces, dark pigment thickest along center of scale pockets. Scale pockets, except for ventral most, are about equally pigmented and mid lateral stripe is absent. Single irregular dark spots are located on upper anterior aspect of operculum and on flank above base of operculum. Dark pigment spots along basal section and mid section of dorsal fin membranes produce two irregular dark stripes on fin. Dark spots located on anal fin membranes of females produce streak-like patterns. Subcircular to bar-shaped dark spot (basicaudal spot) is located on caudal peduncle and base of caudal fin, it is restricted to area above lateral midline and fails to reach dorsal midline.
</paragraph>
</subSubSection>
<subSubSection id="C309657A9059C81E7B97C4ABFF3001FE" pageId="6" pageNumber="51" type="etymology">
<paragraph id="8BAC36F19059C81E7B97C4ABFF3001FE" blockId="6.[151,1437,1312,2018]" pageId="6" pageNumber="51">
<emphasis id="B967EAE39059C81E7B97C4ABFE9B01A5" bold="true" box="[198,339,1951,1977]" pageId="6" pageNumber="51">Etymology.</emphasis>
Named for the 
<typeStatus id="54A888539059C81E795BC494FDF501A6" box="[522,573,1952,1978]" pageId="6" pageNumber="51">type</typeStatus>
and only known locality of the species the Tuxtla Mountains of Veracruz, 
<collectingCountry id="F30476619059C81E7BC6C4FCFF3D01FE" box="[151,245,1992,2018]" name="Mexico" pageId="6" pageNumber="51">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309657A9058C8107B97C3A3FD1707D6" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52" type="distribution">
<paragraph id="8BAC36F19058C81F7B97C3A3FD2A06C6" blockId="7.[151,1437,151,218]" pageId="7" pageNumber="52">
<emphasis id="B967EAE39058C81F7B97C3A3FEAF06AD" bold="true" box="[198,359,151,177]" pageId="7" pageNumber="52">Distribution.</emphasis>
This species is endemic to Lake Catemaco and tributaries of the lake and the Río Grande de Catemaco above the falls at El Salto de Eyipantla.
</paragraph>
<caption id="DF6C66799058C81F7BC6C1DAFC64053A" httpUri="https://zenodo.org/record/183071/files/figure.png" pageId="7" pageNumber="52" targetBox="[265,1323,263,733]" targetPageId="7">
<paragraph id="8BAC36F19058C81F7BC6C1DAFC64053A" blockId="7.[151,1436,750,806]" pageId="7" pageNumber="52">
<emphasis id="B967EAE39058C81F7BC6C1DAFED5051A" bold="true" box="[151,285,750,774]" pageId="7" pageNumber="52">FIGURE 4.</emphasis>
<taxonomicName id="4C134D729058C81F7A79C1DBFDE0051A" box="[296,552,751,774]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F7A79C1DBFDE0051A" box="[296,552,751,774]" italics="true" pageId="7" pageNumber="52">Heterandria tuxtlaensis</emphasis>
</taxonomicName>
. A. Mature male (Holotype, UMMZ 248620, 44.0 mm SL). B. Mature female (Paratype, TCWC 762.03, 41.3 mm SL). C. Gonopodium tip of Holotype.
</paragraph>
</caption>
<paragraph id="8BAC36F19058C81F7B97C065FF3B01E8" blockId="7.[151,1437,849,2036]" pageId="7" pageNumber="52">
<emphasis id="B967EAE39058C81F7B97C065FD8F0577" bold="true" box="[198,583,849,875]" pageId="7" pageNumber="52">Comparison and relationships.</emphasis>
Within 
<taxonomicName id="4C134D729058C81F79F7C066FCFF0577" box="[678,823,850,875]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B967EAE39058C81F79F7C066FCFF0577" box="[678,823,850,875]" italics="true" pageId="7" pageNumber="52">Heterandria</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C134D729058C81F7817C066FC2F0577" box="[838,999,850,875]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F7817C066FC2F0577" box="[838,999,850,875]" italics="true" pageId="7" pageNumber="52">H. tuxtlaensis</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C134D729058C81F7F93C066FAA20577" box="[1218,1386,850,875]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7F93C066FAA20577" box="[1218,1386,850,875]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
and can be identified as this species in Rosen’s (1979) key to species of 
<taxonomicName id="4C134D729058C81F7893C04EFB9B058F" box="[962,1107,890,915]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B967EAE39058C81F7893C04EFB9B058F" box="[962,1107,890,915]" italics="true" pageId="7" pageNumber="52">Heterandria</emphasis>
</taxonomicName>
. Interestingly, however, the state of the basicaudal spot does not agree with the states of this character in any of the other species of 
<taxonomicName id="4C134D729058C81F7E1EC096FF2D05FF" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B967EAE39058C81F7E1EC096FF2D05FF" italics="true" pageId="7" pageNumber="52">Heterandria</emphasis>
</taxonomicName>
. The basal caudal spot of 
<taxonomicName id="4C134D729058C81F7945C0FEFD7C05FF" box="[532,692,970,995]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F7945C0FEFD7C05FF" box="[532,692,970,995]" italics="true" pageId="7" pageNumber="52">H. tuxtlaensis</emphasis>
</taxonomicName>
is not difuse, pale, or obsolescent as in the case of 
<taxonomicName id="4C134D729058C81F7E52C0FEFA5D05FF" box="[1283,1429,970,995]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="attenuata">
<emphasis id="B967EAE39058C81F7E52C0FEFA5D05FF" box="[1283,1429,970,995]" italics="true" pageId="7" pageNumber="52">H. attenuata</emphasis>
</taxonomicName>
, is not bordered by a clear area and broken dark pigment as in the case of 
<taxonomicName id="4C134D729058C81F78B5C0C6FBA70217" box="[996,1135,1010,1035]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="litoperas">
<emphasis id="B967EAE39058C81F78B5C0C6FBA70217" box="[996,1135,1010,1035]" italics="true" pageId="7" pageNumber="52">H. litoperas</emphasis>
</taxonomicName>
, is not elongated as in the case of 
<taxonomicName id="4C134D729058C81F7BA0C72EFEA7022F" box="[241,367,1050,1075]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="obliqua">
<emphasis id="B967EAE39058C81F7BA0C72EFEA7022F" box="[241,367,1050,1075]" italics="true" pageId="7" pageNumber="52">H. obliqua</emphasis>
</taxonomicName>
, does not have ventral and posterior comet-like extensions as in the case of 
<taxonomicName id="4C134D729058C81F7FA3C72EFAB1022F" box="[1266,1401,1050,1075]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="anzuetoi">
<emphasis id="B967EAE39058C81F7FA3C72EFAB1022F" box="[1266,1401,1050,1075]" italics="true" pageId="7" pageNumber="52">H. anzuetoi</emphasis>
</taxonomicName>
, is not irregular and horizontally elongated as in the case of 
<taxonomicName id="4C134D729058C81F7813C776FC2D0247" box="[834,997,1090,1115]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="cataractae">
<emphasis id="B967EAE39058C81F7813C776FC2D0247" box="[834,997,1090,1115]" italics="true" pageId="7" pageNumber="52">H. cataractae</emphasis>
</taxonomicName>
, and is not large and round as in the case of 
<taxonomicName id="4C134D729058C81F7BA3C75EFE48029F" box="[242,384,1130,1155]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="dirempta">
<emphasis id="B967EAE39058C81F7BA3C75EFE48029F" box="[242,384,1130,1155]" italics="true" pageId="7" pageNumber="52">H. dirempta</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C134D729058C81F7AEAC75EFDAC029F" box="[443,612,1130,1155]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7AEAC75EFDAC029F" box="[443,612,1130,1155]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
. However, the other character states in the key, e.g. body depth, head width, caudal peduncle depth, number of lateral line scales, number of pectoral fin rays, vertebral number, and gonopodial traits are more similar to the states in 
<taxonomicName id="4C134D729058C81F7987C78EFCB502CF" box="[726,893,1210,1235]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7987C78EFCB502CF" box="[726,893,1210,1235]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
than to the states in any of the other species of 
<taxonomicName id="4C134D729058C81F7BC6C7D6FEE002E7" box="[151,296,1250,1275]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B967EAE39058C81F7BC6C7D6FEE002E7" box="[151,296,1250,1275]" italics="true" pageId="7" pageNumber="52">Heterandria</emphasis>
</taxonomicName>
. The gonopodium of 
<taxonomicName id="4C134D729058C81F797EC7D6FD1B02E7" box="[559,723,1250,1275]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F797EC7D6FD1B02E7" box="[559,723,1250,1275]" italics="true" pageId="7" pageNumber="52">H. tuxtlaensis</emphasis>
</taxonomicName>
cannot be distinguished from that of 
<taxonomicName id="4C134D729058C81F7FC9C7D6FA8A02E7" box="[1176,1346,1250,1275]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7FC9C7D6FA8A02E7" box="[1176,1346,1250,1275]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EF824B009058C81F7E02C7D6FF2C0338" author="Rosen" pageId="7" pageNumber="54" refString="Rosen, D. E. (1979) Fishes of the uplands and intermontane basins of Guatemala: revisionary studies and comparative geography. Bulletin of the American Museum of Natural History, 162, 267 - 376." type="journal article" year="1979">Rosen (1979)</bibRefCitation>
also provides 12 tables comparing meristic and morphometric values of the species of 
<taxonomicName id="4C134D729058C81F7F8BC63EFAA3033F" box="[1242,1387,1290,1315]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B967EAE39058C81F7F8BC63EFAA3033F" box="[1242,1387,1290,1315]" italics="true" pageId="7" pageNumber="52">Heterandria</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C134D729058C81F7BC6C606FEFF0357" box="[151,311,1330,1355]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F7BC6C606FEFF0357" box="[151,311,1330,1355]" italics="true" pageId="7" pageNumber="52">H. tuxtlaensis</emphasis>
</taxonomicName>
falls within the ranges of 
<taxonomicName id="4C134D729058C81F7939C606FCC70357" box="[616,783,1330,1355]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7939C606FCC70357" box="[616,783,1330,1355]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
, with exception of values related to size and position of the dorsal fin, the number of pectoral fin rays, and width of the head in females. 
<taxonomicName id="4C134D729058C81F7F07C66EFAA4036F" box="[1110,1388,1370,1395]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F7F07C66EFAA4036F" box="[1110,1388,1370,1395]" italics="true" pageId="7" pageNumber="52">Heterandria tuxtlaensis</emphasis>
</taxonomicName>
has fewer than the modal number of dorsal fin rays of 
<taxonomicName id="4C134D729058C81F79B1C6B6FC4F0387" box="[736,903,1410,1435]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F79B1C6B6FC4F0387" box="[736,903,1410,1435]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
, although the dorsal fin ray counts for 
<taxonomicName id="4C134D729058C81F7E1AC6B6FF3903DF" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F7E1AC6B6FF3903DF" italics="true" pageId="7" pageNumber="52">H. tuxtlaensis</emphasis>
</taxonomicName>
are within the lower part of the range of values for 
<taxonomicName id="4C134D729058C81F7816C69EFC2603DF" box="[839,1006,1450,1475]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7816C69EFC2603DF" box="[839,1006,1450,1475]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
from 
<collectingCountry id="F30476619058C81F7F65C69EFB5903D8" box="[1076,1169,1450,1476]" name="Mexico" pageId="7" pageNumber="52">Mexico</collectingCountry>
. Dorsal fin rays counts for Mexican 
<taxonomicName id="4C134D729058C81F7A7CC6E6FE1C03F7" box="[301,468,1490,1515]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7A7CC6E6FE1C03F7" box="[301,468,1490,1515]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
given by Rosen are from 
<bibRefCitation id="EF824B009058C81F7851C6E6FC6A03F0" author="Miller" box="[768,930,1490,1516]" pageId="7" pageNumber="54" refString="Miller, R. R. (1974) Mexican species of the genus Heterandria, subgenus Pseudoxiphophorus (Pisces: Poeciliidae). Transactions of the San Diego Society of Natural History, 17, 235 - 250." type="journal article" year="1974">Miller (1974)</bibRefCitation>
. Specimens of 
<taxonomicName id="4C134D729058C81F7F00C6E6FB3003F7" box="[1105,1272,1490,1515]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7F00C6E6FB3003F7" box="[1105,1272,1490,1515]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
from Miller’s study are from Veracruz and Oaxaca, and include specimens from the Río Papaloapan but not from Lake Catemaco. 
<taxonomicName id="4C134D729058C81F7A4AC516FDF80027" box="[283,560,1570,1595]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F7A4AC516FDF80027" box="[283,560,1570,1595]" italics="true" pageId="7" pageNumber="52">Heterandria tuxtlaensis</emphasis>
</taxonomicName>
has a smaller range and mean number of dorsal fin rays than any of the 50 collections of 
<taxonomicName id="4C134D729058C81F7A6DC57EFE2B007F" box="[316,483,1610,1635]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7A6DC57EFE2B007F" box="[316,483,1610,1635]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
examined in this study. 
<taxonomicName id="4C134D729058C81F79ADC57EFBD9007F" box="[764,1041,1610,1635]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F79ADC57EFBD9007F" box="[764,1041,1610,1635]" italics="true" pageId="7" pageNumber="52">Heterandria tuxtlaensis</emphasis>
</taxonomicName>
can be further distinguished from 
<taxonomicName id="4C134D729058C81F7BC6C546FE880097" box="[151,320,1650,1675]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7BC6C546FE880097" box="[151,320,1650,1675]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
by its shorter and more forwardly placed dorsal fin base, greater number of pectoral fin rays, narrower head width in female specimens, and smaller size of the basicaudal spot. Furthermore the specimens of 
<taxonomicName id="4C134D729058C81F7BE7C5F6FE9600C7" box="[182,350,1730,1755]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7BE7C5F6FE9600C7" box="[182,350,1730,1755]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
from the Río San Juan of the Papaloapan Drainage below the falls of the Río Grande at Eyipantla, the remainder of the Río Papaloapan drainage, and the coastal drainages of the Tuxtlas are deeper bodied, have a longer dorsal fin base, longer depressed dorsal fin, longer caudal fin, higher dorsal fin ray count and a basicaudal spot typical of 
<taxonomicName id="4C134D729058C81F795FC40EFD7E014F" box="[526,694,1850,1875]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F795FC40EFD7E014F" box="[526,694,1850,1875]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
(
<tableCitation id="C691034A9058C81F7995C40EFCE20148" box="[708,810,1850,1876]" captionStart="TABLE 2" captionStartId="4.[151,239,151,175]" captionText="TABLE 2. Species axis loadings for traditional and geometric morphometrics. A. Partial regression coefficients of traditional morphometric characters regressed on species. B. Correlations of superimposed landmark coordinates with the shape gradient separating species." pageId="7" pageNumber="52">Tables 2</tableCitation>
, 
<tableCitation id="C691034A9058C81F7867C40EFC8F0148" box="[822,839,1850,1876]" captionStart="TABLE 3" captionStartId="6.[151,239,271,295]" captionTargetBox="[159,1411,405,599]" captionTargetPageId="6" captionText="TABLE 3. Traditional morphometrics and meristics of Heterandria tuxtlaensis and H. bimaculata. Except for SL and meristics, data are given in percentage of standard length." httpUri="http://table.plazi.org/id/DF6C66799059C81E7BC6C23BFD360755" pageId="7" pageNumber="52" tableUuid="DF6C66799059C81E7BC6C23BFD360755">3</tableCitation>
; 
<figureCitation id="13282A749058C81F7805C40EFC6E0148" box="[852,934,1850,1876]" captionStart="FIGURE 2" captionStartId="3.[151,255,1745,1769]" captionTargetBox="[191,1393,570,1713]" captionTargetId="figure@3.[191,1401,557,1713]" captionTargetPageId="3" captionText="FIGURE 2. Shape difference between H. tuxtlaensis from Lake Catemaco and H. bimaculata from Sontecomapan. Transformation grids show the linear regression of shape on habitat controlling for allometry. A) Males. The shape gradient for males explained 29.6 % of total shape variance. B) Females. The gradient for females explained 30.5 % of total shape variance. Multivariate least squares means (i. e. group centroids) are plotted and their 95 % confidence limits are given as ellipses." httpUri="https://zenodo.org/record/183069/files/figure.png" pageId="7" pageNumber="52">Figs. 2</figureCitation>
, 
<figureCitation id="13282A749058C81F78E3C40EFC0A0148" box="[946,962,1850,1876]" captionStart="FIGURE 3" captionStartId="5.[151,255,663,687]" captionTargetBox="[291,1296,371,638]" captionTargetId="figure@5.[291,1296,371,639]" captionTargetPageId="5" captionText="FIGURE 3. Most extreme individuals on the species axis, for males, from our sample of 30 fish. Left: the new species, H. tuxtlaensis. Right: the presumed sister taxon H. bimaculata. Differences in size are incidental and are not responsible for the shape differences observed. The correlation between the species axis and the axis of allometry was r = 0.08." httpUri="https://zenodo.org/record/183070/files/figure.png" pageId="7" pageNumber="52">3</figureCitation>
). These differences may have led 
<bibRefCitation id="EF824B009058C81F7E03C40EFEBE0160" author="Miller" pageId="7" pageNumber="54" refString="Miller, R. R. &amp; Van Conner, J. (1997) Peces de Catemaco. In: Gonzalez-Soriano, E., Dirzo, R. &amp; Vogt, R. C. (Eds.), Historia natural de Los Tuxtlas. Universidad Nacional Autonoma de Mexico, pp. 451 - 456." type="book chapter" year="1997">Miller and Conner (1997)</bibRefCitation>
to consider the Lake Catemaco population to be close to or identical with 
<taxonomicName id="4C134D729058C81F7FB4C456FA870167" box="[1253,1359,1890,1915]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="jonesi">
<emphasis id="B967EAE39058C81F7FB4C456FA870167" box="[1253,1359,1890,1915]" italics="true" pageId="7" pageNumber="52">H. jonesi</emphasis>
</taxonomicName>
rather than a species most similar to 
<taxonomicName id="4C134D729058C81F7951C4BEFD6101BF" box="[512,681,1930,1955]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39058C81F7951C4BEFD6101BF" box="[512,681,1930,1955]" italics="true" pageId="7" pageNumber="52">H. bimaculata</emphasis>
</taxonomicName>
. Apparently they based this decision on a collection of 
<taxonomicName id="4C134D729058C81F7E10C4BEFF2101D7" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39058C81F7E10C4BEFF2101D7" italics="true" pageId="7" pageNumber="52">H. tuxtlaensis</emphasis>
</taxonomicName>
(UMMZ 178556) that lacks mature males. Thus they were unaware of the gonopodial characters of the species.
</paragraph>
<paragraph id="8BAC36F19057C8107B97C3ACFD1707D6" blockId="8.[151,1437,152,2018]" pageId="8" pageNumber="53">
It is likely that the new species shares an ancestor with 
<taxonomicName id="4C134D729057C810781EC3ACFC3F06AD" box="[847,1015,152,177]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39057C810781EC3ACFC3F06AD" box="[847,1015,152,177]" italics="true" pageId="8" pageNumber="53">H. bimaculata</emphasis>
</taxonomicName>
because of the above listed similarities between the two species and because 
<taxonomicName id="4C134D729057C81079D6C3F4FCE706C5" box="[647,815,192,217]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39057C81079D6C3F4FCE706C5" box="[647,815,192,217]" italics="true" pageId="8" pageNumber="53">H. bimaculata</emphasis>
</taxonomicName>
is the only other species of 
<taxonomicName id="4C134D729057C8107F27C3F4FACF06C5" box="[1142,1287,192,217]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B967EAE39057C8107F27C3F4FACF06C5" box="[1142,1287,192,217]" italics="true" pageId="8" pageNumber="53">Heterandria</emphasis>
</taxonomicName>
found in the Río Papaloapan drainage. Only one other species, 
<taxonomicName id="4C134D729057C81079A9C3DCFCAD071D" box="[760,869,232,257]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="jonesi">
<emphasis id="B967EAE39057C81079A9C3DCFCAD071D" box="[760,869,232,257]" italics="true" pageId="8" pageNumber="53">H. jonesi</emphasis>
</taxonomicName>
occurs in 
<collectingCountry id="F30476619057C81078B7C3DCFBF7071E" box="[998,1087,232,258]" name="Mexico" pageId="8" pageNumber="53">Mexico</collectingCountry>
and it has not been reported south of the Río Nautla in central Veracruz (
<bibRefCitation id="EF824B009057C81079CDC224FCFB0736" author="Miller" box="[668,819,272,298]" pageId="8" pageNumber="54" refString="Miller, R. R. (1974) Mexican species of the genus Heterandria, subgenus Pseudoxiphophorus (Pisces: Poeciliidae). Transactions of the San Diego Society of Natural History, 17, 235 - 250." type="journal article" year="1974">Miller, 1974</bibRefCitation>
). The other species of 
<taxonomicName id="4C134D729057C8107F6DC224FB050735" box="[1084,1229,272,297]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B967EAE39057C8107F6DC224FB050735" box="[1084,1229,272,297]" italics="true" pageId="8" pageNumber="53">Heterandria</emphasis>
</taxonomicName>
are either limited to the southern 
<collectingCountry id="F30476619057C8107A1EC20CFE23074E" box="[335,491,312,338]" name="United States of America" pageId="8" pageNumber="53">United States</collectingCountry>
(
<taxonomicName id="4C134D729057C8107AAFC20CFD4A074D" box="[510,642,312,337]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="formosa">
<emphasis id="B967EAE39057C8107AAFC20CFD4A074D" box="[510,642,312,337]" italics="true" pageId="8" pageNumber="53">H. formosa</emphasis>
</taxonomicName>
) or are endemic to 
<collectingCountry id="F30476619057C810783AC20CFC25074E" box="[875,1005,312,338]" name="Guatemala" pageId="8" pageNumber="53">Guatemala</collectingCountry>
, and of all of these, the most similar congeners are limited to the Río Senizo in the head waters of the Río Usumacinta drainage, in Alta Verapaz, 
<collectingCountry id="F30476619057C8107BC6C2BCFEDE07BE" box="[151,278,392,418]" name="Guatemala" pageId="8" pageNumber="53">Guatemala</collectingCountry>
(
<taxonomicName id="4C134D729057C8107A79C2BCFE0207BD" box="[296,458,392,417]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cataractae">
<emphasis id="B967EAE39057C8107A79C2BCFE0207BD" box="[296,458,392,417]" italics="true" pageId="8" pageNumber="53">H. cataractae</emphasis>
</taxonomicName>
) and the Río Chajmaic, in the headwaters of the Río Usumacinta drainage in Alta Verapaz, 
<collectingCountry id="F30476619057C8107A55C284FE4B07D6" box="[260,387,432,458]" name="Guatemala" pageId="8" pageNumber="53">Guatemala</collectingCountry>
(
<taxonomicName id="4C134D729057C8107AC5C284FDE907D5" box="[404,545,432,457]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="dirempta">
<emphasis id="B967EAE39057C8107AC5C284FDE907D5" box="[404,545,432,457]" italics="true" pageId="8" pageNumber="53">H. dirempta</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EF824B009057C8107968C284FD1A07D6" author="Rosen" box="[569,722,432,458]" pageId="8" pageNumber="54" refString="Rosen, D. E. (1979) Fishes of the uplands and intermontane basins of Guatemala: revisionary studies and comparative geography. Bulletin of the American Museum of Natural History, 162, 267 - 376." type="journal article" year="1979">Rosen, 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C309657A9057C8107B97C2E3FE4103C6" pageId="8" pageNumber="53" type="discussion">
<paragraph id="8BAC36F19057C8107B97C2E3FBEF05B6" blockId="8.[151,1437,152,2018]" pageId="8" pageNumber="53">
<emphasis id="B967EAE39057C8107B97C2E3FEF707ED" bold="true" box="[198,319,471,497]" pageId="8" pageNumber="53">Remarks.</emphasis>
<taxonomicName id="4C134D729057C8107A16C2ECFD9407ED" box="[327,604,472,497]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39057C8107A16C2ECFD9407ED" box="[327,604,472,497]" italics="true" pageId="8" pageNumber="53">Heterandria tuxtlaensis</emphasis>
</taxonomicName>
constitutes the sixth endemic species of the 12 indigenous fish species of Lake Catemaco. According to 
<bibRefCitation id="EF824B009057C8107973C134FC9B0406" author="Miller" box="[546,851,512,538]" pageId="8" pageNumber="54" refString="Miller, R. R. &amp; Van Conner, J. (1997) Peces de Catemaco. In: Gonzalez-Soriano, E., Dirzo, R. &amp; Vogt, R. C. (Eds.), Historia natural de Los Tuxtlas. Universidad Nacional Autonoma de Mexico, pp. 451 - 456." type="book chapter" year="1997">Miller and Conner (1997)</bibRefCitation>
four of the 12 indigenous fish species in the lake were endemic and that future research might reveal that six of the remaining species were undescribed, and thus also endemic to the lake. One of these six putative new species (
<taxonomicName id="4C134D729057C81078ECC164FB0A0475" box="[957,1218,592,617]" class="Actinopterygii" family="Poeciliidae" genus="Xiphophorus" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="kallmani">
<emphasis id="B967EAE39057C81078ECC164FB0A0475" box="[957,1218,592,617]" italics="true" pageId="8" pageNumber="53">Xiphophorus kallmani</emphasis>
</taxonomicName>
) was described by 
<bibRefCitation id="EF824B009057C8107BC6C14CFE05048E" author="Meyer" box="[151,461,632,658]" pageId="8" pageNumber="54" refString="Meyer, M. K. &amp; Schartl, M. (2003) Xiphophorus kallmani sp. n. - a new species of swordtail from Mexico (Teleostei, Cyprinodontiformes, Poeciliidae). Zoologische Abhandlungen, Staatliches Museum fur Tierkunde Dresden, 53, 57 - 64." type="journal article" year="2003">Meyer and Schartl (2003)</bibRefCitation>
and herein we describe the sixth endemic species. Thus endemicity within Lake Catemaco lies between 50% and 83%, depending on the status of the possibly new species. This range of endemicity is high considering the lake is a maximum of 2 million years old (
<bibRefCitation id="EF824B009057C8107F06C1FCFB2C04FE" author="West" box="[1111,1252,712,738]" pageId="8" pageNumber="54" refString="West, R. C. (1964) Surface configuration and associated geology of Middle America .. In: Wauchope, R. &amp; West, R. C., Handbook of Middle American Indians. University of Texas Press, Austin, pp. 33 - 83." type="book chapter" year="1964">West, 1964</bibRefCitation>
). 
<bibRefCitation id="EF824B009057C8107FADC1FCFF2D0516" pageId="8" pageNumber="54" refString="Mateos, M., Sanjur, O. I. &amp; Vrijenhoek, R. C. (2002) Historical biogeography of the livebearing fish genus Poeciliopsis (Poeciliidae: Cyprinodontiformes). Evolution, 56, 972 - 984." type="journal article">
Mateos 
<emphasis id="B967EAE39057C8107E0CC1FCFA5404FD" box="[1373,1436,712,737]" italics="true" pageId="8" pageNumber="53">et al.</emphasis>
(2002)
</bibRefCitation>
investigated the historical biogeography of the poeciliid genus 
<taxonomicName id="4C134D729057C8107882C1C4FBAA0515" box="[979,1122,752,777]" class="Actinopterygii" family="Poeciliidae" genus="Poeciliopsis" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B967EAE39057C8107882C1C4FBAA0515" box="[979,1122,752,777]" italics="true" pageId="8" pageNumber="53">Poeciliopsis</emphasis>
</taxonomicName>
, which includes a species, 
<emphasis id="B967EAE39057C8107BC6C02CFEE8052D" box="[151,288,792,817]" italics="true" pageId="8" pageNumber="53">P. c a t e m a c o</emphasis>
, endemic to the lake. Based on a molecular clock for 
<emphasis id="B967EAE39057C81078CAC02CFC1B052D" box="[923,979,792,817]" italics="true" pageId="8" pageNumber="53">cyt b</emphasis>
divergences, 
<bibRefCitation id="EF824B009057C8107F25C02CFAAA052E" box="[1140,1378,792,818]" pageId="8" pageNumber="54" refString="Mateos, M., Sanjur, O. I. &amp; Vrijenhoek, R. C. (2002) Historical biogeography of the livebearing fish genus Poeciliopsis (Poeciliidae: Cyprinodontiformes). Evolution, 56, 972 - 984." type="journal article">
Mateos 
<emphasis id="B967EAE39057C8107F83C02CFAC5052D" box="[1234,1293,792,817]" italics="true" pageId="8" pageNumber="53">et al.</emphasis>
(2002)
</bibRefCitation>
estimated that 
<emphasis id="B967EAE39057C8107A48C074FE6A0545" box="[281,418,832,857]" italics="true" pageId="8" pageNumber="53">P. c a t e m a c o</emphasis>
has been isolated from other members of its 
<emphasis id="B967EAE39057C81078E4C074FBC70545" box="[949,1039,832,857]" italics="true" pageId="8" pageNumber="53">gracilis</emphasis>
species group for 0.75 to 1.5 million years. This estimate is within the age given by 
<bibRefCitation id="EF824B009057C81079BCC05CFCB4059E" author="West" box="[749,892,872,898]" pageId="8" pageNumber="54" refString="West, R. C. (1964) Surface configuration and associated geology of Middle America .. In: Wauchope, R. &amp; West, R. C., Handbook of Middle American Indians. University of Texas Press, Austin, pp. 33 - 83." type="book chapter" year="1964">West (1964)</bibRefCitation>
for the lake and it is reasonable to assume that all of the endemic species to Lake Catemaco are 0.75 to 1.5 million years old.
</paragraph>
<paragraph id="8BAC36F19057C8107B97C08CFB8F026E" blockId="8.[151,1437,152,2018]" pageId="8" pageNumber="53">
The endemic fishes in Lake Catemaco are predominantly poeciliids (five species in four genera). The other endemic species is a characid, 
<taxonomicName id="4C134D729057C8107919C0D4FCA105E5" box="[584,873,992,1017]" class="Actinopterygii" family="Characidae" genus="Bramocharax" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="caballeroi">
<emphasis id="B967EAE39057C8107919C0D4FCA105E5" box="[584,873,992,1017]" italics="true" pageId="8" pageNumber="53">Bramocharax caballeroi</emphasis>
</taxonomicName>
. The putative new species in the lake include a clupeid of the genus 
<emphasis id="B967EAE39057C8107ADDC73CFDC6023D" box="[396,526,1032,1057]" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="4C134D729057C8107ADDC73CFDC2023D" box="[396,522,1032,1057]" class="Actinopterygii" family="Clupeidae" genus="Dorosoma" kingdom="Animalia" order="Clupeiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="genus">Dorosoma</taxonomicName>
,
</emphasis>
similar to 
<emphasis id="B967EAE39057C81079DCC73CFCE1023D" box="[653,809,1032,1057]" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="4C134D729057C81079DCC73CFCEC023D" box="[653,804,1032,1057]" class="Actinopterygii" family="Clupeidae" genus="Dorosoma" kingdom="Animalia" order="Clupeiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="mexicana">D. mexicana</taxonomicName>
,
</emphasis>
two pimelodids of the genus 
<taxonomicName id="4C134D729057C8107FD4C73CFB27023D" box="[1157,1263,1032,1057]" class="Actinopterygii" family="Heptapteridae" genus="Rhamdia" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B967EAE39057C8107FD4C73CFB27023D" box="[1157,1263,1032,1057]" italics="true" pageId="8" pageNumber="53">Rhamdia</emphasis>
</taxonomicName>
, and a cichlid, similar to 
<emphasis id="B967EAE39057C8107A43C704FE1A0255" box="[274,466,1072,1097]" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="4C134D729057C8107A43C704FE070255" box="[274,463,1072,1097]" family="Cichlidae" genus="Vieja" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="fenestrate">Vieja fenestrate</taxonomicName>
.
</emphasis>
This high level of endemicity complements genetic and geological findings suggesting the lake is ancient and well separated from surrounding fish populations.
</paragraph>
<paragraph id="8BAC36F19057C8107B97C7B4FE4103C6" blockId="8.[151,1437,152,2018]" pageId="8" pageNumber="53">
Finally we hope to be indulged to make a methodological point, be it removed from our major goal of describing 
<taxonomicName id="4C134D729057C8107A4AC79CFE7702DD" box="[283,447,1192,1217]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="tuxtlaensis">
<emphasis id="B967EAE39057C8107A4AC79CFE7702DD" box="[283,447,1192,1217]" italics="true" pageId="8" pageNumber="53">H. tuxtlaensis</emphasis>
</taxonomicName>
. We wish to advocate general use of geometric morphometrics in describing new taxa. Such information is a useful addition to traditional measures if reported in a manner to facilitate comparisons among taxa. To this end we report the direction of individual landmark deviations across the species gradient. Correlations of partial warps or principle components with species gradients would not be general, because these variables change in each new analysis. Landmarks retain homology across studies. We also report effect sizes in a manner conducive to comparison with other taxa, and to meta-analysis (
<bibRefCitation id="EF824B009057C8107FA2C644FEFD03AE" author="Gurevitch" pageId="8" pageNumber="54" refString="Gurevitch, J. &amp; Hedges, L. V. (2001) Meta-analysis: Combining the results of independent experiments. Pp. 347 - 370 In Scheiner, SM &amp; Gurevitch, J (Eds.) Design and Analysis of Ecological Experiments. Oxford: New York. 415 pp." type="book chapter" year="2001">Gurevitch and Hedges 2001</bibRefCitation>
). Thus we hope this approach can be used or expanded in future work in the valuable service of describing new taxa.
</paragraph>
</subSubSection>
<subSubSection id="C309657A9057C8107B97C6D3FC710176" pageId="8" pageNumber="53" type="materials_examined">
<paragraph id="8BAC36F19057C8107B97C6D3FE27015E" blockId="8.[151,1437,152,2018]" pageId="8" pageNumber="53">
<emphasis id="B967EAE39057C8107B97C6D3FDD9001D" bold="true" box="[198,529,1511,1537]" pageId="8" pageNumber="53">Other material examined.</emphasis>
<taxonomicName id="4C134D729057C810794DC6DCFCF4001D" box="[540,828,1512,1537]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="B967EAE39057C810794DC6DCFCF4001D" box="[540,828,1512,1537]" italics="true" pageId="8" pageNumber="53">Heterandria bimaculata</emphasis>
</taxonomicName>
(
<emphasis id="B967EAE39057C8107801C6D3FBF8001D" bold="true" box="[848,1072,1511,1537]" pageId="8" pageNumber="53">
Veracruz, 
<collectingCountry id="F30476619057C8107884C6D3FBF8001D" box="[981,1072,1511,1537]" name="Mexico" pageId="8" pageNumber="53">Mexico</collectingCountry>
</emphasis>
): 
<collectionCode id="ED02AE349057C8107F1CC6DCFB6A001E" LSID="urn:lsid:biocol.org:col:34795" box="[1101,1186,1512,1538]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="53">FMNH</collectionCode>
4601, 3726, 
<collectionCode id="ED02AE349057C8107E16C6DCFA54001E" box="[1351,1436,1512,1538]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="53">TCWC</collectionCode>
755.02, 759.01, 779.02, 1852.01, 
<collectionCode id="ED02AE349057C810796FC524FD510036" box="[574,665,1552,1578]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="53">UMMZ</collectionCode>
108614, 184547; (
<emphasis id="B967EAE39057C81078DFC53BFBAA0035" bold="true" box="[910,1122,1551,1577]" pageId="8" pageNumber="53">
Tabasco, 
<collectingCountry id="F30476619057C8107F58C53BFBAA0035" box="[1033,1122,1551,1577]" name="Mexico" pageId="8" pageNumber="53">Mexico</collectingCountry>
</emphasis>
): 
<collectionCode id="ED02AE349057C8107FD5C524FB160036" LSID="urn:lsid:biocol.org:col:34925" box="[1156,1246,1552,1578]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="53">AMNH</collectionCode>
20401, 27489, 
<collectionCode id="ED02AE349057C8107BC6C50CFF3A004E" box="[151,242,1592,1618]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="53">UMMZ</collectionCode>
184718, 209339, 
<date id="FFAD10319057C8107A9BC50CFDEE004E" box="[458,550,1592,1618]" pageId="8" pageNumber="53" value="42-08-21">210842</date>
; (
<emphasis id="B967EAE39057C810796DC503FCCE004D" bold="true" box="[572,774,1591,1617]" pageId="8" pageNumber="53">
Oaxaca, 
<collectingCountry id="F30476619057C81079FAC503FCCE004D" box="[683,774,1591,1617]" name="Mexico" pageId="8" pageNumber="53">Mexico</collectingCountry>
</emphasis>
): 
<collectionCode id="ED02AE349057C8107871C50CFCB3004E" box="[800,891,1592,1618]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="53">UMMZ</collectionCode>
178533, 178546, 183902, 234799; (
<emphasis id="B967EAE39057C8107E7FC503FF3A0065" bold="true" pageId="8" pageNumber="53">
Chiapas, 
<collectingCountry id="F30476619057C8107BC6C56BFF3A0065" box="[151,242,1631,1657]" name="Mexico" pageId="8" pageNumber="53">Mexico</collectingCountry>
</emphasis>
): NMW-59796 (
<typeStatus id="54A888539057C8107AE6C554FDEC0066" box="[439,548,1632,1658]" pageId="8" pageNumber="53" type="syntype">Syntypes</typeStatus>
of 
<taxonomicName id="4C134D729057C810791AC554FCBA0065" box="[587,882,1632,1657]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="B967EAE39057C810791AC554FCBA0065" box="[587,882,1632,1657]" italics="true" pageId="8" pageNumber="53">Heterandria bimaculatus</emphasis>
</taxonomicName>
), 
<collectionCode id="ED02AE349057C81078DAC554FC2D0066" LSID="urn:lsid:biocol.org:col:34925" box="[907,997,1632,1658]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="53">AMNH</collectionCode>
24649, 59805, 59806, 69807, 59808, 
<collectionCode id="ED02AE349057C8107BC6C5BCFF3A00BE" box="[151,242,1672,1698]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="53">UMMZ</collectionCode>
163778, 186370, 191730, 196644, 209324, 209353, 209361, 
<date id="FFAD10319057C8107883C5BCFBE500BE" box="[978,1069,1672,1698]" pageId="8" pageNumber="53" value="14-08-21">210814</date>
, 
<collectionCode id="ED02AE349057C8107F6BC5BCFB4700BE" box="[1082,1167,1672,1698]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="53">TCWC</collectionCode>
3207.01; (
<emphasis id="B967EAE39057C8107E40C5B3FF3A00D5" bold="true" pageId="8" pageNumber="53">
Campeche, 
<collectingCountry id="F30476619057C8107BC6C59BFF3A00D5" box="[151,242,1711,1737]" name="Mexico" pageId="8" pageNumber="53">Mexico</collectingCountry>
</emphasis>
): 
<collectionCode id="ED02AE349057C8107A5DC584FEAF00D6" box="[268,359,1712,1738]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="53">UMMZ</collectionCode>
<date id="FFAD10319057C8107A3EC584FE0200D6" box="[367,458,1712,1738]" pageId="8" pageNumber="53" value="56-08-19">190856</date>
, 196612; (
<emphasis id="B967EAE39057C810791EC59BFCE800D5" bold="true" box="[591,800,1711,1737]" pageId="8" pageNumber="53">
Yucatan, 
<collectingCountry id="F30476619057C8107994C59BFCE800D5" box="[709,800,1711,1737]" name="Mexico" pageId="8" pageNumber="53">Mexico</collectingCountry>
</emphasis>
): 
<collectionCode id="ED02AE349057C810786BC584FC5D00D6" box="[826,917,1712,1738]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="53">UMMZ</collectionCode>
<date id="FFAD10319057C81078CCC584FC3D00D6" box="[925,1013,1712,1738]" pageId="8" pageNumber="53" value="78-10-20">102078</date>
(
<typeStatus id="54A888539057C8107F55C584FBBB00D6" box="[1028,1139,1712,1738]" pageId="8" pageNumber="53" type="holotype">Holotype</typeStatus>
of 
<emphasis id="B967EAE39057C8107FCAC584FF1D00ED" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="4C134D729057C8107FCAC584FAE400D5" box="[1179,1324,1712,1737]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="genus">Heterandria</taxonomicName>
b. peninsulae
</emphasis>
), 
<date id="FFAD10319057C8107BBFC5ECFE8E00EE" box="[238,326,1752,1778]" pageId="8" pageNumber="53" value="79-10-20">102079</date>
(
<typeStatus id="54A888539057C8107A06C5ECFE0800EE" box="[343,448,1752,1778]" pageId="8" pageNumber="53" type="paratype">Paratype</typeStatus>
of 
<emphasis id="B967EAE39057C8107AB8C5ECFD6300ED" box="[489,683,1752,1777]" italics="true" pageId="8" pageNumber="53">H. b. peninsulae</emphasis>
); (
<emphasis id="B967EAE39057C810799EC5E3FC2400ED" bold="true" box="[719,1004,1751,1777]" pageId="8" pageNumber="53">
<taxonomicName id="4C134D729057C810799EC5E3FC4B00ED" authority="Roo" authorityName="Roo" box="[719,899,1751,1777]" class="Actinopterygii" family="Poeciliidae" genus="Quintana" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="genus">Quintana Roo</taxonomicName>
, 
<collectingCountry id="F30476619057C81078C0C5E3FC2400ED" box="[913,1004,1751,1777]" name="Mexico" pageId="8" pageNumber="53">Mexico</collectingCountry>
</emphasis>
): 
<collectionCode id="ED02AE349057C8107F56C5ECFBAA00EE" box="[1031,1122,1752,1778]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="53">UMMZ</collectionCode>
<date id="FFAD10319057C8107F3AC5ECFB0E00EE" box="[1131,1222,1752,1778]" pageId="8" pageNumber="53" value="81-08-21">210881</date>
; (
<collectingCountry id="F30476619057C8107F8CC5E3FAE000ED" box="[1245,1320,1751,1777]" name="Belize" pageId="8" pageNumber="53">
<emphasis id="B967EAE39057C8107F8CC5E3FAE000ED" bold="true" box="[1245,1320,1751,1777]" pageId="8" pageNumber="53">Belize</emphasis>
</collectingCountry>
): 
<collectionCode id="ED02AE349057C8107E13C5ECFA5300EE" LSID="urn:lsid:biocol.org:col:34925" box="[1346,1435,1752,1778]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="53">AMNH</collectionCode>
78703, 
<collectionCode id="ED02AE349057C8107BA1C434FE8D0106" LSID="urn:lsid:biocol.org:col:34795" box="[240,325,1792,1818]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="53">FMNH</collectionCode>
82180, 
<collectionCode id="ED02AE349057C8107AF4C434FDC80106" box="[421,512,1792,1818]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="53">UMMZ</collectionCode>
<date id="FFAD10319057C8107958C434FDAC0106" box="[521,612,1792,1818]" pageId="8" pageNumber="53" value="24-04-19">190424</date>
, 202737; (
<collectingCountry id="F30476619057C81079B3C5CBFCA40105" box="[738,876,1791,1817]" name="Guatemala" pageId="8" pageNumber="53">
<emphasis id="B967EAE39057C81079B3C5CBFCA40105" bold="true" box="[738,876,1791,1817]" pageId="8" pageNumber="53">Guatemala</emphasis>
</collectingCountry>
): 
<collectionCode id="ED02AE349057C81078D7C434FC280106" LSID="urn:lsid:biocol.org:col:34925" box="[902,992,1792,1818]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="53">AMNH</collectionCode>
24584, 24663, 24486, 36339, 36353, 36357, 36361, 36362, 36363.
</paragraph>
<paragraph id="8BAC36F19057C8107B94C464FC710176" blockId="8.[151,1437,152,2018]" box="[197,953,1872,1898]" pageId="8" pageNumber="53">
<taxonomicName id="4C134D729057C8107B94C464FD970176" authority="(Guatemala)" baseAuthorityName="Guatemala" box="[197,607,1872,1898]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="anzuetoi">
<emphasis id="B967EAE39057C8107B94C464FE080175" box="[197,448,1872,1897]" italics="true" pageId="8" pageNumber="53">Heterandria anzuetoi</emphasis>
(
<collectingCountry id="F30476619057C8107A81C464FD9F0176" box="[464,599,1872,1898]" name="Guatemala" pageId="8" pageNumber="53">Guatemala</collectingCountry>
)
</taxonomicName>
: 
<collectionCode id="ED02AE349057C810793BC464FD0D0176" box="[618,709,1872,1898]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="53">UMMZ</collectionCode>
197389 (
<typeStatus id="54A888539057C8107865C464FC650176" box="[820,941,1872,1898]" pageId="8" pageNumber="53" type="paratype">Paratypes</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C309657A9057C8117B94C44CFBC606C6" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53" type="description">
<paragraph id="8BAC36F19057C8107B94C44CFC75018E" blockId="8.[151,1437,152,2018]" box="[197,957,1912,1938]" pageId="8" pageNumber="53">
<taxonomicName id="4C134D729057C8107B94C44CFDAB018E" authority="(Guatemala)" baseAuthorityName="Guatemala" box="[197,611,1912,1938]" class="Actinopterygii" family="Poeciliidae" genus="Heternadria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="attenuata">
<emphasis id="B967EAE39057C8107B94C44CFE04018D" box="[197,460,1912,1937]" italics="true" pageId="8" pageNumber="53">Heternadria attenuata</emphasis>
(
<collectingCountry id="F30476619057C8107A84C44CFD94018E" box="[469,604,1912,1938]" name="Guatemala" pageId="8" pageNumber="53">Guatemala</collectingCountry>
)
</taxonomicName>
: UMMZ 197091 (
<typeStatus id="54A888539057C8107869C44CFC79018E" box="[824,945,1912,1938]" pageId="8" pageNumber="53" type="paratype">Paratypes</typeStatus>
).
</paragraph>
<paragraph id="8BAC36F19057C8107B94C494FC0301A6" blockId="8.[151,1437,152,2018]" box="[197,971,1952,1978]" pageId="8" pageNumber="53">
<taxonomicName id="4C134D729057C8107B94C494FDB101A6" authority="(Guatemala)" baseAuthorityName="Guatemala" box="[197,633,1952,1978]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="cataractae">
<emphasis id="B967EAE39057C8107B94C494FE1201A5" box="[197,474,1952,1977]" italics="true" pageId="8" pageNumber="53">Heterandria cataractae</emphasis>
(
<collectingCountry id="F30476619057C8107ABBC494FDB901A6" box="[490,625,1952,1978]" name="Guatemala" pageId="8" pageNumber="53">Guatemala</collectingCountry>
)
</taxonomicName>
: UMMZ 193888 (
<typeStatus id="54A888539057C810781FC494FC0B01A6" box="[846,963,1952,1978]" pageId="8" pageNumber="53" type="paratype">Paratypes</typeStatus>
)
</paragraph>
<paragraph id="8BAC36F19057C8107B94C4FCFC7F01FE" blockId="8.[151,1437,152,2018]" box="[197,951,1992,2018]" pageId="8" pageNumber="53">
<taxonomicName id="4C134D729057C8107B94C4FCFDAC01FE" authority="(Guatemala)" baseAuthorityName="Guatemala" box="[197,612,1992,2018]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="dirempta">
<emphasis id="B967EAE39057C8107B94C4FCFE0E01FD" box="[197,454,1992,2017]" italics="true" pageId="8" pageNumber="53">Heterandria dirempta</emphasis>
(
<collectingCountry id="F30476619057C8107A87C4FCFD9501FE" box="[470,605,1992,2018]" name="Guatemala" pageId="8" pageNumber="53">Guatemala</collectingCountry>
)
</taxonomicName>
: UMMZ 187950 (
<typeStatus id="54A888539057C8107868C4FCFC6701FE" box="[825,943,1992,2018]" pageId="8" pageNumber="53" type="paratype">Paratypes</typeStatus>
)
</paragraph>
<paragraph id="8BAC36F19056C8117B94C3ACFBC606C6" blockId="9.[197,1053,152,218]" pageId="9" pageNumber="54">
<taxonomicName id="4C134D729056C8117B94C3ACFDAC06AE" authority="(Guatemala)" baseAuthorityName="Guatemala" box="[197,612,152,178]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="litoperas">
<emphasis id="B967EAE39056C8117B94C3ACFE0D06AD" box="[197,453,152,177]" italics="true" pageId="9" pageNumber="54">Heterandria litoperas</emphasis>
(
<collectingCountry id="F30476619056C8117A84C3ACFD9406AE" box="[469,604,152,178]" name="Guatemala" pageId="9" pageNumber="54">Guatemala</collectingCountry>
)
</taxonomicName>
: UMMZ 197134, 146107 (
<typeStatus id="54A888539056C81178CEC3ACFBDD06AE" box="[927,1045,152,178]" pageId="9" pageNumber="54" type="paratype">Paratypes</typeStatus>
) 
<taxonomicName id="4C134D729056C8117B94C3F4FD9E06C6" authority="(Guatemala)" baseAuthorityName="Guatemala" box="[197,598,192,218]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="obliqua">
<emphasis id="B967EAE39056C8117B94C3F4FE7F06C5" box="[197,439,192,217]" italics="true" pageId="9" pageNumber="54">Heterandria obliqua</emphasis>
(
<collectingCountry id="F30476619056C8117A96C3F4FD8606C6" box="[455,590,192,218]" name="Guatemala" pageId="9" pageNumber="54">Guatemala</collectingCountry>
)
</taxonomicName>
: UMMZ 197079, 193893 (
<typeStatus id="54A888539056C81178C0C3F4FBCF06C6" box="[913,1031,192,218]" pageId="9" pageNumber="54" type="paratype">Paratypes</typeStatus>
)
</paragraph>
</subSubSection>
</treatment>
</document>