168 lines
19 KiB
XML
168 lines
19 KiB
XML
<document id="92355AD6A09C90E1E8DEA0BDBA2C815C" ID-DOI="10.12651/JSR.2013.2.1.015" ID-ISSN="2713-8615" ID-Zenodo-Dep="13136911" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722360455222" checkinUser="felipe" docAuthor="Blakemore, Robert J." docDate="2013" docId="03BA87B0F648D616A9DBFFB5FC28B52A" docLanguage="en" docName="JSpeciesRes.2.1.15-54.pdf" docOrigin="Journal of Species Research 2 (1)" docSource="http://dx.doi.org/10.12651/jsr.2013.2.1.015" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Amynthas phaselus subsp. maculosus" docType="treatment" docVersion="2" lastPageNumber="31" masterDocId="FF83FFC8F658D606A946F91BFF97B017" masterDocTitle="Jeju-do earthworms (Oligochaeta: Megadrilacea) - Quelpart Island revisited" masterLastPageNumber="54" masterPageNumber="15" pageNumber="31" updateTime="1722361037191" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="E2CFAEFEF47B1B2CDA86A2FFEA83FDEF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A2D99590FC6DCE33A991FC306F10D805">
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<mods:title id="5963B10ED930ED47BC9A5840EAE00ED5">Jeju-do earthworms (Oligochaeta: Megadrilacea) - Quelpart Island revisited</mods:title>
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</mods:titleInfo>
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<mods:name id="EBF66E6831EB239A00A4A4C6E159CA0B" type="personal">
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</mods:role>
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<mods:namePart id="2FB24522180837B3BF6955015AE2FD75">Blakemore, Robert J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B983121351E5A5930D3C3C9C8EBEDF8B">text</mods:typeOfResource>
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<mods:title id="87AEF8B03018829518D2826C8C0C5EB3">Journal of Species Research</mods:title>
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</mods:titleInfo>
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<mods:part id="0E461CA703C44E1E148D8991DF43A843">
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<mods:date id="676191ED92CB480517B4B65FB21699E6">2013</mods:date>
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<mods:detail id="B57353F28971CDF88DE71EB14DA92DD6" type="pubDate">
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<mods:number id="4F312C813582EF83E3EE47995BB31B45">2013-02-28</mods:number>
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</mods:detail>
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<mods:detail id="A7644597C1097190D77805C5D24AF833" type="volume">
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<mods:number id="15D30A273DFF906351AD5A08DAD1E340">2</mods:number>
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</mods:detail>
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<mods:detail id="0FC435F4B0D7154F7238C2C99E0C243E" type="issue">
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<mods:number id="DEF50FF662100294B4CD0A605D9F73B1">1</mods:number>
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<mods:extent id="2E120DE6E11E2D33A749D1AF4973C28B" unit="page">
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<mods:start id="D2038AE54D5E999A7AF3F5820DD06CC2">15</mods:start>
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<mods:end id="0FA2CE65ABF05576D776747D2D1DBA97">54</mods:end>
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<mods:location id="3DA75BBC4B94D61DBAA6AA9DD69175AE">
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<mods:url id="0AE3661379F39846DC288EA611602049">http://dx.doi.org/10.12651/jsr.2013.2.1.015</mods:url>
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</mods:location>
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<mods:classification id="E1114F8717F4DB1100731CF8C2110E4C">journal article</mods:classification>
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<mods:identifier id="D4E24DC5ECFED594E2F04297461338D0" type="DOI">10.12651/JSR.2013.2.1.015</mods:identifier>
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<mods:identifier id="53FDA96FF34BB369E30C6A7C1FC22679" type="ISSN">2713-8615</mods:identifier>
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<mods:identifier id="4A1D90FD39A6F9F56C89466E7371ABD1" type="Zenodo-Dep">13136911</mods:identifier>
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</mods:mods>
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<treatment id="03BA87B0F648D616A9DBFFB5FC28B52A" LSID="urn:lsid:plazi:treatment:03BA87B0F648D616A9DBFFB5FC28B52A" httpUri="http://treatment.plazi.org/id/03BA87B0F648D616A9DBFFB5FC28B52A" lastPageNumber="31" pageId="16" pageNumber="31">
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<subSubSection id="C309652DF648D616A9DBFFB5FEE9B6F0" pageId="16" pageNumber="31" type="nomenclature">
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<paragraph id="8BAC36A6F648D616A9DBFFB5FD28B6D1" blockId="16.[157,703,1710,1767]" box="[157,703,1710,1735]" pageId="16" pageNumber="31">
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<heading id="D0E481CAF648D616A9DBFFB5FD28B6D1" bold="true" box="[157,703,1710,1735]" fontSize="10" level="2" pageId="16" pageNumber="31" reason="4">
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<taxonomicName id="4C134D25F648D616A9DBFFB5FD28B6D1" authority="(Hatai, 1930: 661)" baseAuthorityName="Hatai" baseAuthorityPageNumber="661" baseAuthorityYear="1930" box="[157,703,1710,1735]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="subSpecies" species="phaselus" subSpecies="maculosus">
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<emphasis id="B967EAB4F648D616A9DBFFB5FD28B6D1" bold="true" box="[157,703,1710,1735]" pageId="16" pageNumber="31">
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<emphasis id="B967EAB4F648D616A9DBFFB5FE78B6D1" bold="true" box="[157,495,1710,1734]" italics="true" pageId="16" pageNumber="31">Amynthas phaselus maculosus</emphasis>
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(Hatai, 1930: 661)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BAC36A6F648D616A9DBFFD4FEE9B6F0" blockId="16.[157,703,1710,1767]" box="[157,382,1743,1767]" pageId="16" pageNumber="31">
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<emphasis id="B967EAB4F648D616A9DBFFD4FEE9B6F0" bold="true" box="[157,382,1743,1767]" pageId="16" pageNumber="31">
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comb. nov. [
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<figureCitation id="13282A23F648D616A860FFD4FEE1B6F0" box="[294,374,1743,1767]" captionStart="Fig" captionStartId="17.[165,198,1779,1800]" captionTargetBox="[279,1161,256,1758]" captionTargetId="figure-105@17.[253,1203,243,1826]" captionTargetPageId="17" captionText="Fig. 10. Amynthas phaselus maculosus (Hatai, 1930: 661). Ventral view with prostomium, spermathecae and prostates in situ; caeca 27rhs." figureDoi="http://doi.org/10.5281/zenodo.13136929" httpUri="https://zenodo.org/record/13136929/files/figure.png" pageId="16" pageNumber="31">Fig. 10</figureCitation>
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]
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C309652DF648D616A9DBFE1FFDFEB78A" pageId="16" pageNumber="31" type="materials_examined">
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<paragraph id="8BAC36A6F648D616A9DBFE1FFDFEB78A" blockId="16.[157,762,1796,1981]" pageId="16" pageNumber="31">
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<emphasis id="B967EAB4F648D616A9DBFE1FFE13B70B" bold="true" box="[157,388,1796,1820]" pageId="16" pageNumber="31">Material examined.</emphasis>
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IV0000261231 mature specimen collected from besides a stream near Sanbansan SW
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<collectingRegion id="49D7F844F648D616AB8FFE3FFD61B72B" box="[713,758,1828,1852]" country="South Korea" name="Jeju" pageId="16" pageNumber="31">Jeju</collectingRegion>
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, 12
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<superScript id="7C669BEEF648D616A9F1FE59FF53B745" attach="left" box="[183,196,1858,1874]" fontSize="7" pageId="16" pageNumber="31">th</superScript>
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June, 2012 by
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<collectionCode id="ED02AE63F648D616A836FE5EFE09B74B" box="[368,414,1861,1884]" pageId="16" pageNumber="31">RJB</collectionCode>
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- a mature that was figured and dissected, providing DNA samples (
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<collectionCode id="ED02AE63F648D616AB7EFE7EFDFEB76A" box="[568,617,1893,1917]" pageId="16" pageNumber="31">WO</collectionCode>
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51 redone as w27). Included in same jar is a tail portion.
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</paragraph>
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</subSubSection>
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<subSubSection id="C309652DF648D616A9DBFEBEFC22B26C" pageId="16" pageNumber="31" type="description">
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<paragraph id="8BAC36A6F648D616A9DBFEBEFD6DB7AA" blockId="16.[157,762,1796,1981]" box="[157,762,1957,1981]" pageId="16" pageNumber="31">
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<emphasis id="B967EAB4F648D616A9DBFEBEFEB1B7AA" bold="true" box="[157,294,1957,1981]" pageId="16" pageNumber="31">Description.</emphasis>
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(
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<collectingRegion id="49D7F844F648D616A877FEBEFECAB7AA" box="[305,349,1957,1981]" country="South Korea" name="Jeju" pageId="16" pageNumber="31">Jeju</collectingRegion>
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specimen). Length
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<quantity id="4CEB9B43F648D616AB76FEBEFD1EB7AB" box="[560,649,1957,1981]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="31" unit="mm" value="100.0">100 mm</quantity>
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. Segments
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</paragraph>
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<paragraph id="8BAC36A6F648D616AA70F81AFC22B26C" blockId="16.[822,1428,257,1341]" pageId="16" pageNumber="31">99. Colour brownish, clitellum buff. First dorsal pore 11/ 12. Genital markings absent. Setae ca. 36 on segment 10. Spermathecae ca. 0.3 C apart in 5/6/7/8. Male pores superficial on 18 with 12 setae intervening. Septa 8/9/10 are aborted around gizzard. Spermathecae in 6-8 with elongate and corrugated ampulla on shorter duct with clavate diverticulum. Hearts are in 13. Holandric: testis in 10 & 11, seminal vesicles in 11 & 12 the latter pair with finger-like dorsal projections. Ovaries in 12, no ovisacs. Prostates large in 18. Oesophagus dilated in 11-13. Intestinal origin in 15, no typhlosole to about 35. Gut contains mainly soil.</paragraph>
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</subSubSection>
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<subSubSection id="C309652DF648D616AA70FB98FC28B52A" pageId="16" pageNumber="31" type="discussion">
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<paragraph id="8BAC36A6F648D616AA70FB98FC28B52A" blockId="16.[822,1428,257,1341]" pageId="16" pageNumber="31">
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<emphasis id="B967EAB4F648D616AA70FB98FC32B28C" bold="true" box="[822,933,643,667]" pageId="16" pageNumber="31">Remarks.</emphasis>
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The current specimen agrees with earlier descriptions of
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<taxonomicName id="4C134D25F648D616AA8EFBB8FAC1B2AC" authority="(Kobayashi, 1934)" baseAuthorityName="Kobayashi" baseAuthorityYear="1934" box="[968,1366,675,699]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="kamitai">
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<emphasis id="B967EAB4F648D616AA8EFBB8FB1DB2AC" box="[968,1162,675,699]" italics="true" pageId="16" pageNumber="31">Amynthas kamitai</emphasis>
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(
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<bibRefCitation id="EF824B57F648D616ADD3FBB8FAD8B2AC" author="Kobayashi, S." box="[1173,1359,675,699]" pageId="16" pageNumber="31" pagination="1 - 11" refId="ref17097" refString="Kobayashi, S. 1934. Three new Korean earthworms belonging to the genus Pheretima, together with the wider range of the distribution of Pheretima hilgendorfi (Michaelsen). Journal of Chosen Natural History Society 19: 1 - 11." type="journal article" year="1934">Kobayashi, 1934</bibRefCitation>
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)
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</taxonomicName>
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from Jeju that appears now to correspond with
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<taxonomicName id="4C134D25F648D616AC44FBD8FC51B2EC" authority="(Hatai, 1930)" baseAuthorityName="Hatai" baseAuthorityYear="1930" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="maculosus">
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<emphasis id="B967EAB4F648D616AC44FBD8FA04B2CC" box="[1282,1427,707,731]" italics="true" pageId="16" pageNumber="31">A. maculosus</emphasis>
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(Hatai, 1930)
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</taxonomicName>
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from northern
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<collectingCountry id="F3047636F648D616AD2CFBFFFB3EB2EC" box="[1130,1193,740,763]" name="Japan" pageId="16" pageNumber="31">Japan</collectingCountry>
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, that itself was previously included in synonymy with
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<taxonomicName id="4C134D25F648D616ADFBFA1FFCE0B32C" authority="(Hatai, 1930)" baseAuthorityName="Hatai" baseAuthorityYear="1930" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="phaselus">
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<emphasis id="B967EAB4F648D616ADFBFA1FFAA8B30C" box="[1213,1343,771,795]" italics="true" pageId="16" pageNumber="31">A. phaselus</emphasis>
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(Hatai, 1930)
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</taxonomicName>
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, e.g. by Blakemore (2008).
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<taxonomicName id="4C134D25F648D616ADEFFA3FFAADB32C" baseAuthorityName="Hatai" baseAuthorityYear="1930" box="[1193,1338,803,827]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="maculosus">
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<emphasis id="B967EAB4F648D616ADEFFA3FFAADB32C" box="[1193,1338,803,827]" italics="true" pageId="16" pageNumber="31">A. maculosus</emphasis>
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</taxonomicName>
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was revived based on a
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<collectingRegion id="49D7F844F648D616AAB8FA58FBD3B34C" box="[1022,1092,835,859]" country="Japan" name="Tokyo" pageId="16" pageNumber="31">Tokyo</collectingRegion>
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<typeStatus id="54A88804F648D616AD08FA5FFB35B34B" box="[1102,1186,836,860]" pageId="16" pageNumber="31" type="syntype">syntype</typeStatus>
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by Blakemore (2012: 113, fig. 12). Its new synonyms are
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<taxonomicName id="4C134D25F648D616AD8CFA7FFB93B38B" authority="Kobayashi (1934: 5" authorityName="Kobayashi" authorityYear="1934" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="kamitai">
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<emphasis id="B967EAB4F648D616AD8CFA7FFA04B36B" box="[1226,1427,868,892]" italics="true" pageId="16" pageNumber="31">Pheretima kamitai</emphasis>
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<bibRefCitation id="EF824B57F648D616AA70FA9FFB93B38B" author="Kobayashi, S." box="[822,1028,900,924]" pageId="16" pageNumber="31" pagination="1 - 11" refId="ref17097" refString="Kobayashi, S. 1934. Three new Korean earthworms belonging to the genus Pheretima, together with the wider range of the distribution of Pheretima hilgendorfi (Michaelsen). Journal of Chosen Natural History Society 19: 1 - 11." type="journal article" year="1934">Kobayashi (1934: 5</bibRefCitation>
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</taxonomicName>
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, figs. 4-6; 1938: 146, fig. 11) that was from Seoul and latterly included
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<taxonomicName id="4C134D25F648D616ADE6FABFFC3BB3CB" authority="Kobayashi (1936: 165" authorityName="Kobayashi" authorityYear="1936" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="serrata">
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<emphasis id="B967EAB4F648D616ADE6FABFFA8DB3AB" box="[1184,1306,932,956]" italics="true" pageId="16" pageNumber="31">Ph. serrata</emphasis>
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<bibRefCitation id="EF824B57F648D616AC64FABFFC3BB3CB" author="Kobayashi, S." pageId="16" pageNumber="31" pagination="115 - 138" refId="ref17140" refString="Kobayashi, S. 1936. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi (1936: 165</bibRefCitation>
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</taxonomicName>
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, text fig. 10) in synonymy. Referred to as
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<taxonomicName id="4C134D25F648D616AC29FADFFB8AB3EB" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="variety" species="phaselus" variety="kamitai">
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<emphasis id="B967EAB4F648D616AC29FADFFC04B3EB" italics="true" pageId="16" pageNumber="31">Ph. phaselus</emphasis>
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var.
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<emphasis id="B967EAB4F648D616AA8BFAFFFB8AB3EB" box="[973,1053,996,1020]" italics="true" pageId="16" pageNumber="31">kamitai</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="EF824B57F648D616AD0FFAFFFAD6B3EB" author="Kobayashi, S." box="[1097,1345,996,1020]" pageId="16" pageNumber="31" pagination="89 - 170" refId="ref17216" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">Kobayashi (1938: 411)</bibRefCitation>
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and by
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<bibRefCitation id="EF824B57F648D616AA70FD1FFBACB40B" author="Song, M. & K. Y. Paik" box="[822,1083,1028,1052]" pageId="16" pageNumber="31" pagination="135 - 144" refId="ref17447" refString="Song, M. J and K. Y. Paik. 1969. Supplemental note of Pheretima kanrazana and Ph. soulensis. Theses Colloquium Communication on 60 th Birthday of Dr In Suck Yang, pp. 135 - 144 [Not currently accessible]." type="book chapter" year="1969">Song & Paik (1969: 16</bibRefCitation>
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,
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<bibRefCitation id="EF824B57F648D616AD0FFD1FFB3BB40B" author="Song, M. - J. & K. - Y. Paik" box="[1097,1196,1028,1052]" pageId="16" pageNumber="31" pagination="9 - 14" refId="ref17495" refString="Song, M. - J. and K. - Y. Paik. 1970. Earthworms from Chejoo-do Island, Korea. Korean Journal of Zoology 13 (1): 9 - 14." type="journal article" year="1970">1970: 11</bibRefCitation>
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) on specimens from
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<collectingRegion id="49D7F844F648D616AA70FD3FFC08B42B" box="[822,927,1060,1084]" country="Japan" name="Hokkaido" pageId="16" pageNumber="31">Hokkaido</collectingRegion>
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and Jeju, respectively. Since nothing of substance separates
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<taxonomicName id="4C134D25F648D616AAA8FD5EFB3CB44B" authorityName="Hong" authorityYear="2001" box="[1006,1195,1093,1117]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="minjae">
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<emphasis id="B967EAB4F648D616AAA8FD5EFB3CB44B" box="[1006,1195,1093,1117]" italics="true" pageId="16" pageNumber="31">Amynthas minjae</emphasis>
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</taxonomicName>
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Hong, in Hong, Lee & Kim (2001) from these prior taxa, it too is included in synonymy. Reduced to a sub-species of the nominal taxon, the question of its relationship to
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<taxonomicName id="4C134D25F648D616ADBFFDBEFAE0B4AA" baseAuthorityName="Hatai" baseAuthorityYear="1930" box="[1273,1399,1189,1213]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="phaselus">
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<emphasis id="B967EAB4F648D616ADBFFDBEFAE0B4AA" box="[1273,1399,1189,1213]" italics="true" pageId="16" pageNumber="31">A. phaselus</emphasis>
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</taxonomicName>
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is not fully resolved. This Jeju specimen (initially thought similar to
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<taxonomicName id="4C134D25F648D616AAEDFDFDFB88B4EA" baseAuthorityName="Kinberg" baseAuthorityYear="1867" box="[939,1055,1253,1277]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="gracilis">
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<emphasis id="B967EAB4F648D616AAEDFDFDFB88B4EA" box="[939,1055,1253,1277]" italics="true" pageId="16" pageNumber="31">A. gracilis</emphasis>
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</taxonomicName>
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) complies with
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<taxonomicName id="4C134D25F648D616AD9EFDFDFAFCB4EA" baseAuthorityName="Hatai" baseAuthorityYear="1930" box="[1240,1387,1253,1277]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="maculosus">
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<emphasis id="B967EAB4F648D616AD9EFDFDFAFCB4EA" box="[1240,1387,1253,1277]" italics="true" pageId="16" pageNumber="31">A. maculosus</emphasis>
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</taxonomicName>
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, or rather its synonym
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<taxonomicName id="4C134D25F648D616AD45FC1DFBF8B50A" authorityName="Kobayashi" authorityYear="1934" box="[1027,1135,1285,1309]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="kamitai">
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<emphasis id="B967EAB4F648D616AD45FC1DFBF8B50A" box="[1027,1135,1285,1309]" italics="true" pageId="16" pageNumber="31">A. kamitai</emphasis>
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</taxonomicName>
|
|
, more so than with
|
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<taxonomicName id="4C134D25F648D616AC03FC1DFCFCB52A" baseAuthorityName="Hatai" baseAuthorityYear="1930" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="31" phylum="Annelida" rank="species" species="phaselus">
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|
<emphasis id="B967EAB4F648D616AC03FC1DFCFCB52A" italics="true" pageId="16" pageNumber="31">A. phaselus</emphasis>
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</taxonomicName>
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proper.
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |