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<document id="5F017CE0A38FC6ED456575426399D548" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF76FF03FDD8BF5DA29AFA4E" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Tico ryaspis Noyes 2023, sp.nov." docType="treatment" docUuid="065F8663-5E24-43C7-A708-D44266895DAE" docUuidSource="ZooBank" docVersion="2" lastPageNumber="163" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="162" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="415EA0589497840244B4A0C47F3D68FD">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="411F1B18B7B0B5FE0BF76CEF762FC661">Noyes, John Stuart</mods:namePart>
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<mods:date id="ADCA3C338F0E2564CBD6F63102302F67">2023</mods:date>
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<subSubSection id="C309653AFF76FF04FDD8BF5DA518FC54" box="[570,776,889,913]" pageId="163" pageNumber="162" type="nomenclature">
<paragraph id="8BAC36B1FF76FF04FDD8BF5DA518FC54" blockId="163.[537,808,889,940]" box="[570,776,889,913]" pageId="163" pageNumber="162">
<heading id="D0E481DDFF76FF04FDD8BF5DA518FC54" bold="true" box="[570,776,889,913]" centered="true" fontSize="10" level="2" pageId="163" pageNumber="162" reason="2">
<emphasis id="B967EAA3FF76FF04FDD8BF5DA518FC54" bold="true" box="[570,776,889,913]" pageId="163" pageNumber="162">
<taxonomicName id="4C134D32FF76FF04FDD8BF5DA4A7FC54" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[570,695,889,913]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="163" pageNumber="162" phylum="Arthropoda" rank="species" species="ryaspis" status="sp.nov.">
<emphasis id="B967EAA3FF76FF04FDD8BF5DA4A7FC54" bold="true" box="[570,695,889,913]" italics="true" pageId="163" pageNumber="162">Tico ryaspis</emphasis>
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<taxonomicNameLabel id="A25457D8FF76FF04FD5FBF5DA518FC54" box="[701,776,889,913]" pageId="163" pageNumber="162" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C309653AFF76FF04FDFBBFB0A538FC69" box="[537,808,916,940]" pageId="163" pageNumber="162" type="description">
<paragraph id="8BAC36B1FF76FF04FDFBBFB0A538FC69" blockId="163.[537,808,889,940]" box="[537,808,916,940]" pageId="163" pageNumber="162">
(
<figureCitation id="13282A34FF76FF04FDC2BFB0A4BBFC69" box="[544,683,916,940]" captionStart-0="Figs 396-411" captionStart-1="Figs 412-425" captionStartId-0="765.[181,226,1526,1550]" captionStartId-1="766.[181,226,1526,1550]" captionTargetBox-0="[185,1163,231,1508]" captionTargetBox-1="[182,1163,227,1508]" captionTargetId-0="figure-10@765.[181,1163,223,1508]" captionTargetId-1="figure-9@766.[181,1163,223,1508]" captionTargetPageId-0="765" captionTargetPageId-1="766" captionText-0="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." captionText-1="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="163" pageNumber="162">Figs 410-416</figureCitation>
; Hab. E 67)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF76FF03FF57BFEDA771FB59" lastPageId="164" lastPageNumber="163" pageId="163" pageNumber="162" type="diagnosis">
<paragraph id="8BAC36B1FF76FF04FF57BFEDA46DFB89" blockId="163.[181,1164,969,1100]" pageId="163" pageNumber="162">
DIAGNOSIS.
<emphasis id="B967EAA3FF76FF04FEABBFEDA785FC24" box="[329,405,969,993]" italics="true" pageId="163" pageNumber="162">Female</emphasis>
(length about
<quantity id="4CEB9B54FF76FF04FDC0BFEDA47EFC24" box="[546,622,969,993]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="163" pageNumber="162" unit="mm" value="0.8">0.8mm</quantity>
): hind tibia mostly dark brown; tangent to frontovertex forming an angle of about 90° with that of face; scape at least 1.2X as long as minimum width of frontovertex; funicle (
<figureCitation id="13282A34FF76FF04FE74BFDAA7FEFBD2" box="[406,494,1022,1047]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="163" pageNumber="162">Fig. 410</figureCitation>
) with F1-F3 at least slightly longer than broad and F4-F6 at least about 1.5X as long as broad; mesopleuron clearly extending past posterior margin of propodeum and touching base of gaster.
<emphasis id="B967EAA3FF76FF04FE34B810A41AFB89" box="[470,522,1076,1100]" italics="true" pageId="163" pageNumber="162">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FF76FF04FF57B84DA293FB44" blockId="163.[181,1165,1129,1687]" box="[181,1155,1129,1153]" pageId="163" pageNumber="162">
Female
<emphasis id="B967EAA3FF76FF04FEE4B84DA71EFB44" bold="true" box="[262,270,1129,1153]" pageId="163" pageNumber="162">(</emphasis>
<typeStatus id="54A88813FF76FF04FEEDB84DA778FB44" box="[271,360,1129,1153]" pageId="163" pageNumber="162">holotype</typeStatus>
<emphasis id="B967EAA3FF76FF04FE85B84DA77FFB44" bold="true" box="[359,367,1129,1153]" pageId="163" pageNumber="162">)</emphasis>
: length, including ovipositor,
<quantity id="4CEB9B54FF76FF04FD40B84DA4E8FB44" box="[674,760,1129,1153]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="163" pageNumber="162" unit="mm" value="0.84">0.84mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FF76FF04FC3DB84DA223FB44" box="[991,1075,1129,1153]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="163" pageNumber="162" unit="mm" value="0.82">0.82mm</quantity>
(CPD).
</paragraph>
<paragraph id="8BAC36B1FF76FF04FF39B8A0A6EDF9A4" blockId="163.[181,1165,1129,1687]" pageId="163" pageNumber="162">Head dark brown to black with a metallic sheen; occiput with a weak coppery purple sheen; frontovertex generally with a weak, mixed brassy, dark green and coppery purple sheen, mostly coppery purple above scrobes; scrobal area, temple and gena with a weak coppery purple and slightly brassy sheen; frontovertex, gena and interantennal prominence with a few scattered, inconspicuous, pale brown setae; maxillary palpus white; radicle dark brown; scape dark brown in proximal 0.7X, apex pale yellow; pedicel and flagellum mainly yellow, but pedicel brown at base and distal segments of clava slightly dusky; thorax dark brown to black with a weak metallic lustre; pronotum with a weak coppery purple sheen; mesoscutum and axilla with a weak brassy sheen and weak green and coppery purple reflections; tegula dark purple brown with a weak coppery purple sheen; scutellum dorsally moderately shiny with a weak coppery purple sheen with some dark green reflections, more dark green towards apex and side, extreme apex and side more strongly shining coppery and metallic green; mesopleuron with a moderate purple sheen; fore coxa white; fore femur white to very pale yellow, tibia and tarsus very pale yellow; mid coxa brown; mid leg otherwise pale yellow; hind coxa dark brown; hind femur dark brown, tibia brown, pale orange at apices, tarsus dusky very pale orange, apical tarsomere brown; wings hyaline; propodeum dark brown with a weak purple sheen, side with a weak metallic green and coppery sheen; gaster dark brown with coppery purple, brassy, green and blue reflections; gonostylus brown.</paragraph>
<paragraph id="8BAC36B1FF76FF04FF39BA40A29CF952" blockId="163.[181,1165,1129,1687]" pageId="163" pageNumber="162">
Head (
<figureCitation id="13282A34FF76FF04FEC3BA40A768F9B9" box="[289,376,1636,1660]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="163" pageNumber="162">Fig. 411</figureCitation>
) about 2.5X as wide as frontovertex, in profile about 1.5X as high as deep, tangent to frontovertex forming an angle of about 90° with that of face; occipital margin sharp,
</paragraph>
<paragraph id="8BAC36B1FF71FF03FBBCBCA6A29CFF5A" blockId="164.[1118,1164,130,159]" box="[1118,1164,130,159]" pageId="164" pageNumber="163">163</paragraph>
<paragraph id="8BAC36B1FF71FF03FF57BCFAA7D7FD43" blockId="164.[181,1165,222,1127]" pageId="164" pageNumber="163">
carinate; an indistinct, shallow, shiny-bottomed, elongate, oval depression present near occipital margin adjacent to eye; ocelli forming an angle of about 90°; frontovertex narrowest about level with anterior ocellus, with distinct, coarse, polygonally reticulate sculpture of mesh size generally a little larger than eye facet, not shallower above top of scrobes; temple and gena with much shallower, fine, elongate, polygonally reticulate sculpture; top of scrobes with shallow, regular, polygonally reticulate sculpture, scrobes smooth; interantennal prominence and mouth margin with slightly shallower, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, narrowly ∩-shaped; interantennal prominence dorsally acute; antenna as in
<figureCitation id="13282A34FF71FF03FC05BDBDA254FE74" box="[999,1092,409,433]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="164" pageNumber="163">Fig. 410</figureCitation>
; scape relatively short and stout, about 4.2X as long as broad; funicle with all segments clearly slightly longer than broad, distinctly increasing in size and width distad so that F6 is largest; clava about 3.5X as long as broad and about 1.2X as wide as F6, not curved downwards towards apex, slightly longer than F4-F6 combined, sutures parallel, sensory area small, at apex only giving it a rounded appearance; eye reaching occipital margin, clothed with sparse, inconspicuous setae, each clearly shorter than diameter of facet; clypeal margin clearly concave; mandible with 3 acute teeth. Relative measurements: HW 54,
<collectingRegion id="49D7F853FF71FF03FDE3BE70A436FDAE" box="[513,550,596,619]" country="Swaziland" name="Hhohho" pageId="164" pageNumber="163">HH</collectingRegion>
44.5, FV 22, POL 12, OOL 2, OCL 2.5, AOL 7.5, EL 31, EW 25, MS 18, SL 26.5, SW 7.
</paragraph>
<paragraph id="8BAC36B1FF71FF03FF39BEADA77EFC69" blockId="164.[181,1165,222,1127]" pageId="164" pageNumber="163">
Thorax (
<figureCitation id="13282A34FF71FF03FED6BEADA79DFD64" box="[308,397,649,673]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="164" pageNumber="163">Fig. 416</figureCitation>
) with pronotum hardly exposed behind head, its posterior margin broadly concave, not angular medially; pronotum and mesoscutum with sculpture generally similar to that in ocellar area but clearly shallower; axilla clearly raised, dorsally with shallow, polygonally reticulate sculpture, posterior face virtually completely smooth and shiny; scutellum mainly with fine, striate-reticulate sculpture that is clearly deeper than sculpture on mesoscutum and more or less arranged in whorls, extreme side and apex completely smooth and shiny; visible part of mesoscutum about 1.7X as broad as long; scutellum about 1.2X as broad as long, dorsally slightly convex, apex with a pair of slender, elongate setae; hind femur about 3.9X as long as broad; fore wing with venation and setation as in
<figureCitation id="13282A34FF71FF03FDD3BF7BA4ADFCB2" box="[561,701,863,887]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="164" pageNumber="163">Figs 412, 414</figureCitation>
; mesopleuron clearly extending past posterior margin of propodeum and touching base of gaster; propodeum (
<figureCitation id="13282A34FF71FF03FCDFBF5DA583FC54" box="[829,915,889,913]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="164" pageNumber="163">Fig. 416</figureCitation>
) medially about 0.1X as long as scutellum.
</paragraph>
<paragraph id="8BAC36B1FF71FF03FF39BF8BA51AFC39" blockId="164.[181,1165,222,1127]" pageId="164" pageNumber="163">Gaster with hypopygium reaching about 0.6X towards apex; last tergite about 0.7X as long as mid tibia, with apex broadly rounded; ovipositor hardly exserted, the exserted part about 0.1X as long as mid tibial spur or about 0.05X as long as mid tibia.</paragraph>
<paragraph id="8BAC36B1FF71FF03FF39BFDBA7B6FB89" blockId="164.[181,1165,222,1127]" pageId="164" pageNumber="163">
<typeStatus id="54A88813FF71FF03FF39BFDBA727FBD2" box="[219,311,1023,1047]" pageId="164" pageNumber="163">Paratype</typeStatus>
. Funicle with linear sensilla present only on F4-F6; hypopygium
<figureCitation id="13282A34FF71FF03FC27BFDBA20CFBD2" box="[965,1052,1023,1047]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="164" pageNumber="163">Fig. 413</figureCitation>
; ovipositor
<figureCitation id="13282A34FF71FF03FF57B83DA71EFBF4" box="[181,270,1049,1073]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="164" pageNumber="163">Fig. 415</figureCitation>
. Relative measurements (wings): FWL 67, FWW 29; HWL 44, HWW 10; (ovipositor): OL 43, GL 10 [MT 72].
</paragraph>
<paragraph id="8BAC36B1FF71FF03FF39B86BA4EEFBA2" blockId="164.[181,1165,222,1127]" box="[219,766,1103,1127]" pageId="164" pageNumber="163">Variation. Little variation noted in material examined.</paragraph>
<paragraph id="8BAC36B1FF71FF03FF57B8A0A771FB59" blockId="164.[181,353,1156,1180]" box="[181,353,1156,1180]" pageId="164" pageNumber="163">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF71FF03FF57B89DA7F9FB14" box="[181,489,1209,1233]" pageId="164" pageNumber="163" type="distribution">
<paragraph id="8BAC36B1FF71FF03FF57B89DA7F9FB14" blockId="164.[181,489,1209,1233]" box="[181,489,1209,1233]" pageId="164" pageNumber="163">
DISTRIBUTION.
<collectingCountry id="F3047621FF71FF03FE91B89DA7F5FB14" box="[371,485,1209,1233]" name="Costa Rica" pageId="164" pageNumber="163">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF71FF03FF57B8CBA76EFAC2" box="[181,382,1263,1287]" pageId="164" pageNumber="163" type="biology_ecology">
<paragraph id="8BAC36B1FF71FF03FF57B8CBA76EFAC2" blockId="164.[181,382,1263,1287]" box="[181,382,1263,1287]" pageId="164" pageNumber="163">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF71FF03FF57B902A29AFA4E" pageId="164" pageNumber="163" type="materials_examined">
<paragraph id="8BAC36B1FF71FF03FF57B902A7BDFAFE" blockId="164.[181,1165,1318,1419]" box="[181,429,1318,1339]" pageId="164" pageNumber="163">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FF71FF03FF39B965A29AFA4E" blockId="164.[181,1165,1318,1419]" pageId="164" pageNumber="163">
Type material.
<materialsCitation id="3B7B3CECFF71FF03FE89B964A77DFAB5" collectingDate="1996-01-26" collectingDateMax="1996-02-24" collectingDateMin="1996-01-26" collectionCode="MT, YPT" collectorName="J. S. Noyes" country="Costa Rica" elevation="1100" location="Guanacaste" pageId="164" pageNumber="163" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FF71FF03FE89B964A7AFFA93" box="[363,447,1344,1366]" pageId="164" pageNumber="163">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FF71FF03FE08B964A469FA93" bold="true" box="[490,633,1344,1366]" pageId="164" pageNumber="163">
<collectingCountry id="F3047621FF71FF03FE08B964A469FA93" box="[490,633,1344,1366]" name="Costa Rica" pageId="164" pageNumber="163">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF71FF03FD6AB964A4E9FA93" bold="true" box="[648,761,1344,1366]" pageId="164" pageNumber="163">
<collectingRegion id="49D7F853FF71FF03FD6AB964A4E9FA93" box="[648,761,1344,1366]" country="Costa Rica" name="Guanacaste" pageId="164" pageNumber="163">Guanacaste</collectingRegion>
</emphasis>
,
<taxonomicName id="4C134D32FF71FF03FCEAB964A59BFA93" authority="(ACG)" baseAuthorityName="ACG" box="[776,907,1344,1366]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="164" pageNumber="163" phylum="Tracheophyta" rank="species" species="cacao">Cacao (ACG)</taxonomicName>
,
<quantity id="4CEB9B54FF71FF03FC7AB964A5C9FA90" box="[920,985,1344,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="164" pageNumber="163" unit="m" value="1100.0">
<elevation id="003ED182FF71FF03FC7AB964A5C9FA90" box="[920,985,1344,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="164" pageNumber="163" unit="m" value="1100.0">1100m</elevation>
</quantity>
,
<collectionCode id="ED02AE74FF71FF03FC07B964A217FA90" box="[997,1031,1344,1365]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="164" pageNumber="163">MT</collectionCode>
/
<collectionCode id="ED02AE74FF71FF03FBECB964A229FA90" box="[1038,1081,1344,1365]" pageId="164" pageNumber="163">YPT</collectionCode>
,
<date id="FFAD1071FF71FF03FBA4B964A6E5FAB4" pageId="164" pageNumber="163" value="1996-01-26" valueMax="1996-02-24" valueMin="1996-01-26">
<collectingDate id="EFE9E999FF71FF03FBA4B964A6E5FAB4" pageId="164" pageNumber="163" value="1996-01-26" valueMax="1996-02-24" valueMin="1996-01-26">26.i-24. ii.1996</collectingDate>
</date>
(
<collectorName id="26E65367FF71FF03FEE1B97FA777FAB5" box="[259,359,1371,1392]" pageId="164" pageNumber="163">J.S. Noyes</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF71FF03FE94B97FA41EFA4F" collectingDate="1985-05-11" collectingDateMax="1985-06-01" collectingDateMin="1985-05-11" collectorName="D. Janzen &amp; I. D. Gauld" country="Costa Rica" location="Santa" pageId="164" pageNumber="163" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<typeStatus id="54A88813FF71FF03FE94B97FA7C2FAB5" box="[374,466,1371,1392]" pageId="164" pageNumber="163">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FF71FF03FE38B97FA475FAB4" bold="true" box="[474,613,1371,1393]" pageId="164" pageNumber="163">
<collectingCountry id="F3047621FF71FF03FE38B97FA475FAB4" box="[474,613,1371,1393]" name="Costa Rica" pageId="164" pageNumber="163">COSTA RICA</collectingCountry>
</emphasis>
, 1E,
<emphasis id="B967EAA3FF71FF03FD7BB97FA51AFAB4" bold="true" box="[665,778,1371,1393]" pageId="164" pageNumber="163">
<collectingRegion id="49D7F853FF71FF03FD7BB97FA51AFAB4" box="[665,778,1371,1393]" country="Costa Rica" name="Guanacaste" pageId="164" pageNumber="163">Guanacaste</collectingRegion>
</emphasis>
,
<location id="8ECC606AFF71FF03FCF4B97FA558FAB5" LSID="urn:lsid:plazi:treatment:03BA87A7FF76FF03FDD8BF5DA29AFA4E:8ECC606AFF71FF03FCF4B97FA558FAB5" box="[790,840,1371,1392]" country="Costa Rica" name="Santa" pageId="164" pageNumber="163" stateProvince="Guanacaste">Santa</location>
<taxonomicName id="4C134D32FF71FF03FCADB97FA20FFAB5" authority="NP, Bosque Hum" authorityName="NP, Bosque Hum" box="[847,1055,1371,1392]" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="164" pageNumber="163" phylum="Tracheophyta" rank="genus">Rosa NP, Bosque Hum</taxonomicName>
10-C,
<date id="FFAD1071FF71FF03FBBCB97FA71BFA4E" pageId="164" pageNumber="163" value="1985-05-11" valueMax="1985-06-01" valueMin="1985-05-11">
<collectingDate id="EFE9E999FF71FF03FBBCB97FA71BFA4E" pageId="164" pageNumber="163" value="1985-05-11" valueMax="1985-06-01" valueMin="1985-05-11">11.v- 1.vi.1985</collectingDate>
</date>
(
<collectorName id="26E65367FF71FF03FEFAB952A764FA4E" box="[280,372,1398,1419]" pageId="164" pageNumber="163">D. Janzen</collectorName>
,
<collectorName id="26E65367FF71FF03FE9FB952A7F1FA4E" box="[381,481,1397,1419]" pageId="164" pageNumber="163">I.D. Gauld</collectorName>
); 1E
</materialsCitation>
,
<materialsCitation id="3B7B3CECFF71FF03FDFBB952A4F3FA4E" box="[537,739,1397,1419]" collectingDate="1996-01-26" collectingDateMax="1996-02-24" collectingDateMin="1996-01-26" collectionCode="MT, YPT" collectorName="J. S. Noyes" country="Costa Rica" elevation="1100" location="Guanacaste" pageId="164" pageNumber="163" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">same data as holotype</materialsCitation>
.
<materialsCitation id="3B7B3CECFF71FF03FD0EB951A296FA4E" box="[748,1158,1397,1419]" collectionCode="MZUCR, NHMUK" country="Costa Rica" pageId="164" pageNumber="163" specimenCount="2" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FF71FF03FD0EB951A550FA4E" box="[748,832,1397,1419]" pageId="164" pageNumber="163">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FF71FF03FCBFB952A5A0FA4E" box="[861,944,1398,1419]" pageId="164" pageNumber="163">MZUCR</collectionCode>
,
<typeStatus id="54A88813FF71FF03FC5BB952A200FA4E" box="[953,1040,1398,1419]" pageId="164" pageNumber="163">paratypes</typeStatus>
in
<collectionCode id="ED02AE74FF71FF03FBCFB952A296FA4E" box="[1069,1158,1398,1419]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="164" pageNumber="163" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>