215 lines
36 KiB
XML
215 lines
36 KiB
XML
<document id="156A548695DCA360C8A461D0F1E04503" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF35FF45FDE6B83DA26EFD43" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Cheiloneurus danessa Noyes 2023, sp.nov." docType="treatment" docUuid="FF67506C-BC12-4674-94F6-07631357F03E" docUuidSource="ZooBank" docVersion="2" lastPageNumber="225" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="223" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="4A6670EB98ABB9E7C47CF3DC2A77F9CA">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="7E9D4754913F28EE93C5D6EA68CC395F">Noyes, John Stuart</mods:namePart>
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<mods:title id="B4C9E968AC4AC4A093A420FF83550A04">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="38CC280828DA0E32EAECC846CA0A4548">2023</mods:date>
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<taxonomicName id="4C134D32FF35FF47FDE6B83DA4FCFBF4" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[516,748,1049,1073]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="224" pageNumber="223" phylum="Arthropoda" rank="species" species="danessa" status="sp.nov.">
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<emphasis id="B967EAA3FF35FF47FDE6B83DA4FCFBF4" bold="true" box="[516,748,1049,1073]" italics="true" pageId="224" pageNumber="223">Cheiloneurus danessa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25457D8FF35FF47FD10B83DA52DFBF4" box="[754,829,1049,1073]" pageId="224" pageNumber="223" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFF35FF47FDFBB810A539FB89" box="[537,809,1076,1100]" pageId="224" pageNumber="223" type="description">
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<paragraph id="8BAC36B1FF35FF47FDFBB810A539FB89" blockId="224.[516,829,1049,1100]" box="[537,809,1076,1100]" pageId="224" pageNumber="223">
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(
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<figureCitation id="13282A34FF35FF47FDC2B810A4BBFB89" box="[544,683,1076,1100]" captionStart-0="Figs 538-552" captionStart-1="Figs 553-568" captionStartId-0="775.[181,226,1526,1550]" captionStartId-1="776.[181,226,1526,1550]" captionTargetBox-0="[181,1159,229,1508]" captionTargetBox-1="[185,1163,227,1508]" captionTargetId-0="figure-10@775.[181,1163,223,1508]" captionTargetId-1="figure-9@776.[181,1163,223,1508]" captionTargetPageId-0="775" captionTargetPageId-1="776" captionText-0="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." captionText-1="Figs 553-568. Cheiloneurus danessa, HT – (553) fore wing. Cheiloneurus idris – (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT – (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT – (567) antenna; (568) mesosoma, dorsal view." pageId="224" pageNumber="223">Figs 547-553</figureCitation>
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; Hab. E 85)
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</paragraph>
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</subSubSection>
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<subSubSection id="C309653AFF35FF45FF57B84DA771FEB9" lastPageId="226" lastPageNumber="225" pageId="224" pageNumber="223" type="diagnosis">
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<paragraph id="8BAC36B1FF35FF47FF57B84DA40CFA04" blockId="224.[181,1165,1129,1473]" pageId="224" pageNumber="223">
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DIAGNOSIS.
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<emphasis id="B967EAA3FF35FF47FEABB84DA785FB44" box="[329,405,1129,1153]" italics="true" pageId="224" pageNumber="223">Female</emphasis>
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(length about
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<quantity id="4CEB9B54FF35FF47FDC7B84DA465FB44" box="[549,629,1129,1153]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="224" pageNumber="223" unit="mm" value="0.9">0.9 mm</quantity>
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): body generally dark brown with a weak to moderate metallic sheen; mesoscutum with uniform brown setae; fore wing (
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<figureCitation id="13282A34FF35FF47FC88B8A0A5D3FB59" box="[874,963,1156,1180]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT – (553) fore wing. Cheiloneurus idris – (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT – (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT – (567) antenna; (568) mesosoma, dorsal view." pageId="224" pageNumber="223">Fig. 553</figureCitation>
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) hyaline; fore coxa white; head about 2.7X as wide as frontovertex; antenna (
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<figureCitation id="13282A34FF35FF47FCE3B8BAA547FB73" box="[769,855,1182,1206]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="224" pageNumber="223">Fig. 547</figureCitation>
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) with pedicel about as long as F1-F2 combined; all funicle segments very slightly longer than broad and with linear sensilla; clava slightly longer than F4-F6 combined, with apex rounded; mandible with 1 acute tooth and a broad, upper truncation; mesoscutum (
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<figureCitation id="13282A34FF35FF47FDDFB8CAA483FAC3" box="[573,659,1262,1286]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="224" pageNumber="223">Fig. 552</figureCitation>
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) with longitudinally elongate, lineolate sculpture, some elongate, polygonally reticulate sculpture medially and rounded cells near posterior margin; scutellum with longitudinally elongate, lineolate sculpture, a few polygonally reticulate cells in middle; scutellum without tuft of setae medially in apical half; fore wing (
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<figureCitation id="13282A34FF35FF47FC56B91AA21BFA93" box="[948,1035,1342,1366]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT – (553) fore wing. Cheiloneurus idris – (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT – (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT – (567) antenna; (568) mesosoma, dorsal view." pageId="224" pageNumber="223">Fig. 553</figureCitation>
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) about 2.3X as long as broad; parastigma slightly swollen, not downcurved; stigmal vein (
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<figureCitation id="13282A34FF35FF47FC03B97DA229FAB4" box="[993,1081,1369,1393]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="224" pageNumber="223">Fig. 550</figureCitation>
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) with 3 campaniform sensilla at apex; gaster (
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<figureCitation id="13282A34FF35FF47FDA5B950A4B1FA49" box="[583,673,1396,1420]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="224" pageNumber="223">Fig. 551</figureCitation>
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) with “glands” on Gt1 and Gt5, those on Gt1 continuous, not separated, those on Gt5 separated by less than 0.2X their own lengths; syntergum broader than long.
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<emphasis id="B967EAA3FF35FF47FE97B98DA7B9FA04" box="[373,425,1449,1473]" italics="true" pageId="224" pageNumber="223">Male</emphasis>
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: unknown.
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</paragraph>
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Female
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<emphasis id="B967EAA3FF35FF47FEE4B9FAA71EFA33" bold="true" box="[262,270,1502,1526]" pageId="224" pageNumber="223">(</emphasis>
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<typeStatus id="54A88813FF35FF47FEEDB9FBA778FA32" box="[271,360,1503,1527]" pageId="224" pageNumber="223">holotype</typeStatus>
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<emphasis id="B967EAA3FF35FF47FE85B9FAA77FFA33" bold="true" box="[359,367,1502,1526]" pageId="224" pageNumber="223">)</emphasis>
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: length, including ovipositor,
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<quantity id="4CEB9B54FF35FF47FD40B9FBA4E8FA32" box="[674,760,1503,1527]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="224" pageNumber="223" unit="mm" value="0.9">0.90mm</quantity>
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; excluding ovipositor,
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<quantity id="4CEB9B54FF35FF47FC3DB9FAA223FA33" box="[991,1075,1502,1526]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="224" pageNumber="223" unit="mm" value="0.86">0.86mm</quantity>
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(
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<collectionCode id="ED02AE74FF35FF47FBA3B9FBA268FA32" box="[1089,1144,1503,1527]" pageId="224" pageNumber="223">CPD</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BAC36B1FF35FF47FF39B9DDA29DF952" blockId="224.[181,1165,1502,1687]" pageId="224" pageNumber="223">Head dark brown to black with a metallic sheen; occiput with a weak coppery sheen, some metallic blue-green reflections below occipital margin; frontovertex generally slightly shiny with a weak coppery purple sheen along occipital margin and above scrobes, some dull coppery purple elsewhere, with some hardly discernible metallic dark blue-green; scrobal area with a moderate coppery purple sheen, area between eye and scrobes dull coppery purple with a hint of metallic dark blue-green; temple and gena with a dull coppery purple sheen; frontovertex, gena and</paragraph>
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<paragraph id="8BAC36B1FF34FF46FF57BCA6A6F3FF5A" blockId="225.[181,227,130,159]" box="[181,227,130,159]" pageId="225" pageNumber="224">
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</pageNumber>
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</paragraph>
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<pageTitle id="CB8CEED6FF34FF46FEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="225" pageNumber="224">
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Taxonomic Monographs on Neotropical
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<taxonomicName id="4C134D32FF34FF46FD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="225" pageNumber="224" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
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2
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||
</pageTitle>
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||
</paragraph>
|
||
<paragraph id="8BAC36B1FF34FF46FF57BCFAA7F8FCE2" blockId="225.[181,1165,222,1687]" pageId="225" pageNumber="224">interantennal prominence with a few scattered, inconspicuous, pale brown setae; palpi white; radicle orange-brown; scape mostly dark brown on external surface with apical 0.4X ventrally pale orange, internal surface similar but with most of apical half pale orange; pedicel mainly dark brown, apex very pale orange; flagellum slightly dusky, yellow, clava dusky pale orange; thorax dark brown to black with a weak to moderate metallic lustre; pronotum with a moderate coppery sheen; mesoscutum quite dull with a slight brassy and weak purple sheen, posteriorly a little more shiny and coppery along posterior margin; tegula dark brown with a slight coppery sheen; axilla with a weak to moderate coppery purple sheen; scutellum dorsally about as shiny as mesoscutum, medially with a weak coppery purple sheen, laterally mixed with weak to moderate metallic dark blue-green, extreme side and apex with a moderate metallic dark blue-green sheen mixed coppery; mesopleuron fairly shiny with a slight brassy and coppery purple sheen with a hint of metallic dark blue-green; fore coxa white, mid and hind coxae dark brown; fore femur white to very pale yellow with a very thin dark brown line along dorsal margin in middle half and a very faint, shorter one opposite, tibia white at extreme base, otherwise pale orange with a very weak dusky spot subbasally on outer surface, tarsus pale orange with apical tarsomere slightly dusky; mid femur white with a narrow, incomplete, dark brown subapical band, tibia white with knee very narrowly dark brown and with a narrow, dark brown subbasal band, spur very pale yellow, tarsus white to very pale yellow; hind femur dark brown with extreme base and apex white, tibia dark brown with extreme base white and apical two-fifths white, tarsus white to pale yellow with apical tarsomere dusky; wings hyaline; propodeum dark brown with a weak purple sheen, side with a moderate coppery sheen; gaster dark brown with coppery purple, brassy, green and blue reflections; gonostylus brown.</paragraph>
|
||
<paragraph id="8BAC36B1FF34FF46FF39BF0DA7CAFAC2" blockId="225.[181,1165,222,1687]" pageId="225" pageNumber="224">
|
||
Head about 2.7X as wide as frontovertex, in profile about 1.4X as high as deep, tangent to frontovertex forming an angle of about 95° with that of face; occipital margin sharp, not carinate; no distinct oval depression adjacent to eye near occipital margin; ocelli forming an angle of about 80°; inner eye margins subparallel; frontovertex with moderately deep, polygonally reticulate sculpture of mesh size generally subequal to eye facet, not shallower above top of scrobes; temple and gena with shallower, fine, elongate, polygonally reticulate sculpture; top of scrobes with shallow, regular, polygonally reticulate sculpture, scrobes smooth; interantennal prominence and mouth margin with slightly shallower, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, broadly ∩-shaped; interantennal prominence dorsally narrowly rounded; antenna as in
|
||
<figureCitation id="13282A34FF34FF46FED8B83DA780FBF4" box="[314,400,1049,1073]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="225" pageNumber="224">Fig. 547</figureCitation>
|
||
; scape relatively short and stout, slightly broadened and flattened, about 3.4X as long as broad; funicle with segments all slightly longer than broad and becoming progressively larger distad so that F6 is largest; linear sensilla present on all funicle segments; clava about 3.2X as long as broad and about 1.1X as wide as F6, not curved downwards towards apex, slightly longer than F4-F6 combined, sutures parallel, sensory area small, at apex only giving it a rounded appearance; eye slightly overreaching occipital margin, appearing naked; malar sulcus absent; clypeal margin weakly concave; mandible with one tooth and a broad, slightly oblique, truncation. Relative measurements: HW 79,
|
||
<collectingRegion id="49D7F853FF34FF46FDEBB8F1A43EFB29" box="[521,558,1237,1260]" country="Swaziland" name="Hhohho" pageId="225" pageNumber="224">HH</collectingRegion>
|
||
61, FV 29, POL 14.5, OOL 2, OCL 4, AOL 9, EL 42, EW 38.5, MS 24, SL 27.5, SW 8.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FF34FF46FF39B92DA784F9A7" blockId="225.[181,1165,222,1687]" pageId="225" pageNumber="224">
|
||
Thorax (
|
||
<figureCitation id="13282A34FF34FF46FED6B92DA79DFAE4" box="[308,397,1289,1313]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="225" pageNumber="224">Fig. 552</figureCitation>
|
||
) with pronotum hardly exposed behind head, its posterior margin broadly concave, with a shallow median invagination; pronotum with shallower, finer polygonally reticulate sculpture than ocellar area; mesoscutum with conspicuously longitudinally lineolate striate-reticulate to striate sculpture that is finer than that in ocellar area and about as deep, a few rounded cells near posterior margin; axilla with slightly transversely elongate, polygonally reticulate sculpture that is slightly shallower than that on mesoscutum; scutellum dorsally with striate to striate-reticulate sculpture that is slightly deeper and coarser than that on mesoscutum, cellular sculpture along mid-line more rounded, extreme side and apex quite smooth and shiny; visible part of mesoscutum about 1.8X as broad as long; scutellum about 1.2X as broad as long, apex with a pair of slender, elongate setae; hind femur about 3.4X as long as broad; fore wing with venation and setation as in
|
||
<figureCitation id="13282A34FF34FF46FE19BA30A444F9E9" box="[507,596,1556,1580]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="225" pageNumber="224">Figs 550</figureCitation>
|
||
,
|
||
<figureCitation id="13282A34FF34FF46FDBDBA30A498F9E9" box="[607,648,1556,1580]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT – (553) fore wing. Cheiloneurus idris – (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT – (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT – (567) antenna; (568) mesosoma, dorsal view." pageId="225" pageNumber="224">553</figureCitation>
|
||
; linea calva entire and open; propodeum (
|
||
<figureCitation id="13282A34FF34FF46FBCDBA30A295F9E9" box="[1071,1157,1556,1580]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="225" pageNumber="224">Fig. 552</figureCitation>
|
||
) medially less than 0.1X as long as scutellum. Relative measurements: FWL 67.5, FWW 29.5; HWL 47.5, HWW 11.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FF34FF46FF39BA40A29CF953" blockId="225.[181,1165,222,1687]" pageId="225" pageNumber="224">
|
||
Gaster with hypopygium reaching about 0.5X to apex; last tergite about 0.9X as long as mid tibia, with apex acute; “glands” (
|
||
<figureCitation id="13282A34FF34FF46FDE7BA5BA44CF952" box="[517,604,1663,1687]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="225" pageNumber="224">Fig. 551</figureCitation>
|
||
) present on Gt1 and Gt5, those on Gt1 continuous, not
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FF37FF45FBBCBCA6A29CFF5A" blockId="226.[1118,1164,130,159]" box="[1118,1164,130,159]" pageId="226" pageNumber="225">225</paragraph>
|
||
<paragraph id="8BAC36B1FF37FF45FF57BCFAA5E5FEE9" blockId="226.[181,1164,222,326]" pageId="226" pageNumber="225">
|
||
separated, those on Gt5 separated by less than 0.2X their own lengths; hypopygium
|
||
<figureCitation id="13282A34FF37FF45FBCCBCFAA298FF33" box="[1070,1160,222,246]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="226" pageNumber="225">Fig. 551</figureCitation>
|
||
; ovipositor (
|
||
<figureCitation id="13282A34FF37FF45FEC9BCDDA792FED4" box="[299,386,249,273]" captionStart="Figs 538-552" captionStartId="775.[181,226,1526,1550]" captionTargetBox="[181,1159,229,1508]" captionTargetId="figure-10@775.[181,1163,223,1508]" captionTargetPageId="775" captionText="Figs 538-552. Cheiloneurus carita, HT – (538) ovipositor; (539) mesosoma, dorsal view. Cheiloneurus epytus – (540) antenna; (541) gastral tergites, showing “glands” on Gt1 and Gt5; (542) fore wing; (543) apex of fore wing venation; (544) mesosoma, dorsal view; (545) hypopygium; (546) head, facial view. Cheiloneurus danessa, HT – (547) antenna; (548) ovipositor; (549) hypopygium; (550) apex of fore wing venation; (551) gastral tergites showing “glands” on Gt1 and Gt5; (552) mesosoma, dorsal view." pageId="226" pageNumber="225">Fig. 548</figureCitation>
|
||
) slightly exserted, the exserted part about 0.5X as long as mid tibial spur or about 0.15X as long as mid tibia. Relative measurements: OL 75, GL 20 [MT 48].
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FF37FF45FF39BD0AA457FE83" blockId="226.[181,1164,222,326]" box="[219,583,302,326]" pageId="226" pageNumber="225">
|
||
Variation. Only
|
||
<typeStatus id="54A88813FF37FF45FE9CBD0AA7C7FE83" box="[382,471,302,326]" pageId="226" pageNumber="225">holotype</typeStatus>
|
||
examined.
|
||
</paragraph>
|
||
<paragraph id="8BAC36B1FF37FF45FF57BD40A771FEB9" blockId="226.[181,353,356,380]" box="[181,353,356,380]" pageId="226" pageNumber="225">Male. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFF37FF45FF57BDBDA7F9FE74" box="[181,489,409,433]" pageId="226" pageNumber="225" type="distribution">
|
||
<paragraph id="8BAC36B1FF37FF45FF57BDBDA7F9FE74" blockId="226.[181,489,409,433]" box="[181,489,409,433]" pageId="226" pageNumber="225">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="F3047621FF37FF45FE91BDBDA7F5FE74" box="[371,485,409,433]" name="Costa Rica" pageId="226" pageNumber="225">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFF37FF45FF57BDEBA76EFE23" box="[181,382,462,487]" pageId="226" pageNumber="225" type="biology_ecology">
|
||
<paragraph id="8BAC36B1FF37FF45FF57BDEBA76EFE23" blockId="226.[181,382,462,487]" box="[181,382,462,487]" pageId="226" pageNumber="225">HOSTS. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFF37FF45FF57BE22A50AFD95" pageId="226" pageNumber="225" type="materials_examined">
|
||
<paragraph id="8BAC36B1FF37FF45FF57BE22A7BDFDDE" blockId="226.[181,1164,518,593]" box="[181,429,518,539]" pageId="226" pageNumber="225">MATERIAL EXAMINED.</paragraph>
|
||
<paragraph id="8BAC36B1FF37FF45FF39BE05A50AFD95" blockId="226.[181,1164,518,593]" pageId="226" pageNumber="225">
|
||
Type material.
|
||
<materialsCitation id="3B7B3CECFF37FF45FE80BE04A453FD95" collectingDate="2016-02-15" collectingDateMax="2016-02-17" collectingDateMin="2016-02-15" collectorName="J. S. Noyes" country="Costa Rica" elevation="770" latitude="10.8" location="Guanacaste" longLatPrecision="1297" longitude="-85.4" pageId="226" pageNumber="225" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
|
||
<typeStatus id="54A88813FF37FF45FE80BE04A7A6FDF3" box="[354,438,544,566]" pageId="226" pageNumber="225">Holotype</typeStatus>
|
||
E:
|
||
<emphasis id="B967EAA3FF37FF45FE3ABE04A472FDF3" bold="true" box="[472,610,544,566]" pageId="226" pageNumber="225">
|
||
<collectingCountry id="F3047621FF37FF45FE3ABE04A472FDF3" box="[472,610,544,566]" name="Costa Rica" pageId="226" pageNumber="225">COSTA RICA</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B967EAA3FF37FF45FD8EBE04A4CDFDF3" bold="true" box="[620,733,544,566]" pageId="226" pageNumber="225">
|
||
<collectingRegion id="49D7F853FF37FF45FD8EBE04A4CDFDF3" box="[620,733,544,566]" country="Costa Rica" name="Guanacaste" pageId="226" pageNumber="225">Guanacaste</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<quantity id="4CEB9B54FF37FF45FD0ABE04A50CFDF3" box="[744,796,544,566]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="226" pageNumber="225" unit="km" value="18.0">18km</quantity>
|
||
<taxonomicName id="4C134D32FF37FF45FCC3BE05A225FDF3" authority=", Buena Vista Lodge" authorityName="Buena Vista Lodge" box="[801,1077,544,566]" class="Insecta" family="Encyrtidae" genus="Neococcidencyrtus" kingdom="Animalia" order="Hymenoptera" pageId="226" pageNumber="225" phylum="Arthropoda" rank="species" species="liberia">N. Liberia, Buena Vista Lodge</taxonomicName>
|
||
,
|
||
<geoCoordinate id="EE275076FF37FF45FBDCBE04A29CFDF0" box="[1086,1164,544,565]" degrees="10" direction="north" minutes="48" orientation="latitude" pageId="226" pageNumber="225" precision="925" value="10.8">10°48’N</geoCoordinate>
|
||
<geoCoordinate id="EE275076FF37FF45FF57BE1FA718FD94" box="[181,264,571,593]" degrees="85" direction="west" minutes="24" orientation="longitude" pageId="226" pageNumber="225" precision="925" value="-85.4">85°24’W</geoCoordinate>
|
||
,
|
||
<quantity id="4CEB9B54FF37FF45FEF0BE1FA757FD95" box="[274,327,571,592]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="226" pageNumber="225" unit="m" value="770.0">
|
||
<elevation id="003ED182FF37FF45FEF0BE1FA757FD95" box="[274,327,571,592]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="226" pageNumber="225" unit="m" value="770.0">770m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFAD1071FF37FF45FEB3BE1FA7DCFD94" box="[337,460,571,593]" pageId="226" pageNumber="225" value="2016-02-15" valueMax="2016-02-17" valueMin="2016-02-15">
|
||
<collectingDate id="EFE9E999FF37FF45FEB3BE1FA7DCFD94" box="[337,460,571,593]" pageId="226" pageNumber="225" value="2016-02-15" valueMax="2016-02-17" valueMin="2016-02-15">15-17.ii.2016</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="26E65367FF37FF45FE3BBE1FA42DFD95" box="[473,573,571,592]" pageId="226" pageNumber="225">J.S. Noyes</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7B3CECFF37FF45FDAEBE1FA506FD95" box="[588,790,571,593]" collectionCode="NHMUK" country="Costa Rica" pageId="226" pageNumber="225" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
|
||
<typeStatus id="54A88813FF37FF45FDAEBE1FA4B0FD94" box="[588,672,571,593]" pageId="226" pageNumber="225">Holotype</typeStatus>
|
||
in
|
||
<collectionCode id="ED02AE74FF37FF45FD5FBE1FA506FD95" box="[701,790,571,592]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="226" pageNumber="225" type="Museum">NHMUK</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309653AFF37FF45FF57BE4BA26EFD43" box="[181,1150,622,647]" pageId="226" pageNumber="225" type="discussion">
|
||
<paragraph id="8BAC36B1FF37FF45FF57BE4BA26EFD43" blockId="226.[181,1150,622,647]" box="[181,1150,622,647]" pageId="226" pageNumber="225">
|
||
COMMENTS. The
|
||
<typeStatus id="54A88813FF37FF45FE9DBE4AA7C8FD43" box="[383,472,622,646]" pageId="226" pageNumber="225">holotype</typeStatus>
|
||
has the right flagellum missing but is otherwise in good condition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |