146 lines
19 KiB
XML
146 lines
19 KiB
XML
<document id="DF3CB89449A2D8B316F0E08EF664E656" ID-DOI="10.5281/zenodo.182518" ID-GBIF-Dataset="32ba7237-e044-423a-86ab-3cbfc911f978" ID-ISSN="1175-5326" ID-Zenodo-Dep="182518" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459868857621" checkinUser="plazi" docAuthor="Amorim, Dalton De Souza, Oliveira, Sarah Siqueira & Balbi, Maria Isabel P. A." docDate="2008" docId="03B187E8FFB9FFCDFF6FFCD795AF8475" docLanguage="en" docName="zt01789p065.pdf" docOrigin="Zootaxa 1789" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Azana atlantica Oliveira & Balbi, n.sp." docType="treatment" docVersion="9" lastPageNumber="60" masterDocId="FF88FF90FFBBFFCEFFF8FF8A9620877F" masterDocTitle="Azana atlantica, n. sp., with reduced mouthparts and two ocelli: first record of Azana for the Neotropical region (Diptera: Mycetophilidae: Sciophilinae)" masterLastPageNumber="65" masterPageNumber="57" pageNumber="59" updateTime="1698226407077" updateUser="plazi">
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<mods:mods id="57B82EB7263F8DD9C6D5A36876564C6B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="36626A0C2B1EFB24313E4AAF2A54C6B2">
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<mods:title id="0AE7898077AFC3ACE6812A28B956C81E">Azana atlantica, n. sp., with reduced mouthparts and two ocelli: first record of Azana for the Neotropical region (Diptera: Mycetophilidae: Sciophilinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="FD4A1D3EB075112A15EEE613FE79EF97" type="personal">
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<mods:role id="2156B065187AC39850E5C7A3C4F8DA5D">
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<mods:roleTerm id="1E1D3D869820B19A625C5F3AB7488BE2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="176E782290508F6ACC9FAB05ECDC34F4">Amorim, Dalton De Souza</mods:namePart>
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</mods:name>
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<mods:name id="81203F09D679116CF8C940F88FA4DF73" type="personal">
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<mods:role id="E54A739A9903A1CF58AE275269912A1F">
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<mods:roleTerm id="EFBE224B7722CACF59397B25299F6E38">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="426C60F9CA3149960862C084D73E5EF1">Oliveira, Sarah Siqueira</mods:namePart>
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</mods:name>
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<mods:name id="AF7B87DAFA0697BABF51BC613A187103" type="personal">
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<mods:role id="773C7225B8DB9BD9F6580431F37F1ED6">
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<mods:roleTerm id="114643BC2ECB1BE4A5BD03DFA6166C06">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D963C96E6FF27E033B9716B6CF77E147">Balbi, Maria Isabel P. A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B21BE9A313B95F283C33742EE6D997ED">text</mods:typeOfResource>
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<mods:titleInfo id="69FB0AE17D19D93E9D06B008C9920EA5">
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<mods:title id="7C8BE7A15ADCC4E9B71BF22DEEA170D6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="26351F2789FE2FE5CF4D4A27DFDCECBC">
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<mods:date id="65DD9726B09583004449BF65F7418E13">2008</mods:date>
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<mods:detail id="2C0750FD2B8E086AE2A1111B35E7D872" type="volume">
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<mods:number id="C1ABD539424ECF09314C7DF0AC04486C">1789</mods:number>
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</mods:detail>
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<mods:start id="3A15BFDDC04F40E8E62F6D838FE7C985">57</mods:start>
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<mods:end id="730F432020598B0016C6DE264BA6DBD5">65</mods:end>
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<mods:classification id="4BF6933408252ACE418BFA69461EDE08">journal article</mods:classification>
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<mods:identifier id="8692AFF157E73EBEA4FCD35D23D1D2BA" type="DOI">10.5281/zenodo.182518</mods:identifier>
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<mods:identifier id="9B0B501017DA667C9827B645BD98B64A" type="GBIF-Dataset">32ba7237-e044-423a-86ab-3cbfc911f978</mods:identifier>
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<mods:identifier id="37CF76B74C7C4284298E7F7790E5FB3E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D8E45EBE8226427020B24D5B78AC611B" type="Zenodo-Dep">182518</mods:identifier>
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<treatment id="03B187E8FFB9FFCDFF6FFCD795AF8475" ID-DOI="http://doi.org/10.5281/zenodo.6229075" ID-GBIF-Taxon="119369075" ID-Zenodo-Dep="6229075" LSID="urn:lsid:plazi:treatment:03B187E8FFB9FFCDFF6FFCD795AF8475" httpUri="http://treatment.plazi.org/id/03B187E8FFB9FFCDFF6FFCD795AF8475" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59">
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<subSubSection id="C3026575FFB9FFCCFF6FFCD7973784E5" pageId="2" pageNumber="59" type="nomenclature">
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<paragraph id="8BA736FEFFB9FFCCFF6FFCD794568408" blockId="2.[151,630,861,922]" box="[151,630,861,887]" pageId="2" pageNumber="59">
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<heading id="D0EF8192FFB9FFCCFF6FFCD794568408" bold="true" box="[151,630,861,887]" fontSize="11" level="1" pageId="2" pageNumber="59" reason="1">
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<emphasis id="B96CEAECFFB9FFCCFF6FFCD794568408" bold="true" box="[151,630,861,887]" pageId="2" pageNumber="59">
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<taxonomicName id="4C184D7DFFB9FFCCFF6FFCD7940D8408" ID-CoL="K5GT" authority="Oliveira & Balbi" authorityName="Oliveira & Balbi" box="[151,557,861,887]" class="Insecta" family="Mycetophilidae" genus="Azana" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="atlantica" status="sp. nov.">
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<emphasis id="B96CEAECFFB9FFCCFF6FFCD797758408" bold="true" box="[151,341,861,887]" italics="true" pageId="2" pageNumber="59">Azana atlantica</emphasis>
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Oliveira & Balbi
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25F5797FFB9FFCCFDC3FCD794568408" box="[571,630,861,887]" pageId="2" pageNumber="59" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA736FEFFB9FFCCFF6FFC0A973784E5" blockId="2.[151,630,861,922]" box="[151,279,896,922]" pageId="2" pageNumber="59">
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(
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<figureCitation id="13232A7BFFB9FFCCFF67FC0A972E84E5" box="[159,270,896,922]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURES 5 – 6" captionStartId-0="3.[151,255,1310,1334]" captionStartId-1="4.[151,255,1166,1190]" captionStartId-2="5.[151,255,1746,1770]" captionStartId-3="6.[151,255,1166,1190]" captionStartId-4="7.[151,269,1944,1968]" captionTargetBox-1="[243,1344,194,1136]" captionTargetBox-2="[151,1436,194,1721]" captionTargetBox-3="[151,1421,477,1142]" captionTargetBox-4="[293,1216,197,1880]" captionTargetId-1="figure@4.[243,1345,194,1138]" captionTargetId-2="figure@5.[151,1436,194,1722]" captionTargetId-3="figure@6.[151,1436,477,1142]" captionTargetId-4="figure@7.[288,1303,197,1924]" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Azana atlantica, sp. n. Holotype. Head, lateral view." captionText-1="FIGURE 2. Azana atlantica, sp. n. Holotype. Detail of mouthparts, lateral view" captionText-2="FIGURE 3. Azana atlantica, sp. n. Holotype. Thorax, lateral view." captionText-3="FIGURE 4. Azana atlantica, sp. n. Holotype. Wing." captionText-4="FIGURES 5 – 6. Azana atlantica, sp. n. Holotype. Male terminalia. 5. Ventral view. 6. Dorsal view. Abbreviations: Ce, cercus; Gc, gonocoxite; GcAp, gonocoxal apodeme; Gs, gonostyle; T 9, tergite 9; Te, tegmen." httpUri-1="https://zenodo.org/record/182519/files/figure.png" httpUri-2="https://zenodo.org/record/182520/files/figure.png" httpUri-3="https://zenodo.org/record/182521/files/figure.png" httpUri-4="https://zenodo.org/record/182522/files/figure.png" pageId="2" pageNumber="59">Figs. 1–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026575FFB9FFCCFF6FFC5A96C98317" pageId="2" pageNumber="59" type="diagnosis">
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<paragraph id="8BA736FEFFB9FFCCFF6FFC5A96C98317" blockId="2.[151,1437,976,2015]" pageId="2" pageNumber="59">
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<emphasis id="B96CEAECFFB9FFCCFF6FFC5A97378495" bold="true" box="[151,279,976,1002]" pageId="2" pageNumber="59">Diagnosis.</emphasis>
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Two ocelli, mouthparts reduced, maxillary palpus 1-segmented. Wing fumose brown, slightly darker across the mid of the wing and close to the apex; first section of Rs not sclerotized; M4 entirely absent; A1 scarcely visible. Gonostyle triangular, with a row of spines along the entire inner margin; parameres absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026575FFB9FFCCFF3EFBFC937B8277" pageId="2" pageNumber="59" type="materials_examined">
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<paragraph id="8BA736FEFFB9FFCCFF3EFBFC937B8277" blockId="2.[151,1437,976,2015]" pageId="2" pageNumber="59">
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<emphasis id="B96CEAECFFB9FFCCFF3EFBFC946083EF" bold="true" box="[198,576,1142,1168]" pageId="2" pageNumber="59">
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Material examined.
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<typeStatus id="54A3885CFFB9FFCCFE36FBFC946083EF" box="[462,576,1142,1168]" pageId="2" pageNumber="59" type="holotype">Holotype</typeStatus>
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</emphasis>
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, ɗ,
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<collectingCountry id="F30F766EFFB9FFCCFD86FBFC94CB83EF" box="[638,747,1142,1168]" name="Brazil" pageId="2" pageNumber="59">BRAZIL</collectingCountry>
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, State of São Paulo, Salesópolis, Estação Biológica de Boracéia, Trilha dos Pilões,
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<geoCoordinate id="EE2C5039FFB9FFCCFE01FB1494E183C7" box="[505,705,1182,1208]" direction="south" orientation="latitude" pageId="2" pageNumber="59" precision="1" value="-23.651611">23º 39’ 05.8” S</geoCoordinate>
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,
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<geoCoordinate id="EE2C5039FFB9FFCCFD2BFB14958583C7" box="[723,933,1182,1208]" direction="west" orientation="longitude" pageId="2" pageNumber="59" precision="1" value="-45.895725">45º 53’ 44.6” W</geoCoordinate>
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, Malaise Trap (“trilha Ponto 4”),
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<date id="FFA6103EFFB9FFCCFAA8FB149735839F" pageId="2" pageNumber="59" value="2001-03-30" valueMax="2001-04-02" valueMin="2001-03-30">30.iii– 02.iv.2001</date>
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, STP Amarante & eq. cols.
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<typeStatus id="54A3885CFFB9FFCCFD95FB4C94C3839F" box="[621,739,1222,1248]" pageId="2" pageNumber="59" type="paratype">Paratypes</typeStatus>
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: 1 ɗ, same data as
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<typeStatus id="54A3885CFFB9FFCCFC2BFB4C921D839F" box="[979,1085,1222,1248]" pageId="2" pageNumber="59" type="holotype">holotype</typeStatus>
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; 1 ɗ, same data as
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<typeStatus id="54A3885CFFB9FFCCFAD6FB4C93B7839F" box="[1326,1431,1222,1248]" pageId="2" pageNumber="59" type="holotype">holotype</typeStatus>
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, except
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<geoCoordinate id="EE2C5039FFB9FFCCFF12FB6497808277" box="[234,416,1262,1288]" direction="south" orientation="latitude" pageId="2" pageNumber="59" precision="1" value="-23.65175">23º 39’ 06.3” S</geoCoordinate>
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,
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<geoCoordinate id="EE2C5039FFB9FFCCFE55FB64944F8277" box="[429,623,1262,1288]" direction="west" orientation="longitude" pageId="2" pageNumber="59" precision="1" value="-45.89692">45º 53’ 48.9” W</geoCoordinate>
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; 1 ɗ,
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<emphasis id="B96CEAECFFB9FFCCFD4BFB6494CC8278" box="[691,748,1262,1287]" italics="true" pageId="2" pageNumber="59">idem</emphasis>
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, except
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<geoCoordinate id="EE2C5039FFB9FFCCFCB6FB6492248277" box="[846,1028,1262,1288]" direction="south" orientation="latitude" pageId="2" pageNumber="59" precision="1" value="-23.651417">23º 39’ 05.1” S</geoCoordinate>
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,
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<geoCoordinate id="EE2C5039FFB9FFCCFBE9FB6492F18277" box="[1041,1233,1262,1288]" direction="west" orientation="longitude" pageId="2" pageNumber="59" precision="1" value="-45.897724">45º 53’ 51.8” W</geoCoordinate>
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,
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<date id="FFA6103EFFB9FFCCFB24FB6493768277" box="[1244,1366,1262,1288]" pageId="2" pageNumber="59" value="2005-02-26">26.ii.2005</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026575FFB9FFCDFF3EFA9C978B85C5" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59" type="description">
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<paragraph id="8BA736FEFFB9FFCDFF3EFA9C92E485ED" blockId="2.[151,1437,976,2015]" lastBlockId="3.[151,1437,151,778]" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59">
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<emphasis id="B96CEAECFFB9FFCCFF3EFA9C97D9824F" bold="true" box="[198,505,1302,1328]" pageId="2" pageNumber="59">Description. Male. Head</emphasis>
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(
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<figureCitation id="13232A7BFFB9FFCCFDF2FA9C945A824F" box="[522,634,1302,1328]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,255,1310,1334]" captionStartId-1="4.[151,255,1166,1190]" captionTargetBox-1="[243,1344,194,1136]" captionTargetId-1="figure@4.[243,1345,194,1138]" captionTargetPageId-1="4" captionText-0="FIGURE 1. Azana atlantica, sp. n. Holotype. Head, lateral view." captionText-1="FIGURE 2. Azana atlantica, sp. n. Holotype. Detail of mouthparts, lateral view" httpUri-1="https://zenodo.org/record/182519/files/figure.png" pageId="2" pageNumber="59">Figs. 1–2</figureCitation>
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). Vertex light brown, two ocelli, apart from the eye margin, ocellar area darker. Frons light yellowish brown, setose. Clypeus brown, setose. Mouthparts reduced, a single light brown palpomere, labella and palpi setose. Scape and pedicel light brown, scape trapezoidal, enlarging to the apex, elongated, pedicel more rounded, shorter; flagellum whitish yellow, with 14 flagellomeres; flagellomeres cylindrical, short, wider than long, with scattered setae. Lateral and ventral occiput setae separated from the larger, dorsal setae.
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<emphasis id="B96CEAECFFB9FFCCFE57FA54942C8287" bold="true" box="[431,524,1502,1528]" pageId="2" pageNumber="59">Thorax</emphasis>
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(
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<figureCitation id="13232A7BFFB9FFCCFDE5FA5494448287" box="[541,612,1502,1528]" captionStart="FIGURE 3" captionStartId="5.[151,255,1746,1770]" captionTargetBox="[151,1436,194,1721]" captionTargetId="figure@5.[151,1436,194,1722]" captionTargetPageId="5" captionText="FIGURE 3. Azana atlantica, sp. n. Holotype. Thorax, lateral view." httpUri="https://zenodo.org/record/182520/files/figure.png" pageId="2" pageNumber="59">Fig. 3</figureCitation>
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). Mesonotum strongly arched, setose, light brown, with a pair of yellowish brown longitudinal bands connected anteriorly in each half of the scutum. Scutellum brown, setose. Pleural sclerites light brown, mesepimeron lighter dorsally. Pronotum with about 7 large setae. Primary anapleural suture complete, no secondary anapleural suture. Anepisternum with about 25 setae on dorsal half, divided dorsoventrally by an unsclerotized slender band. Laterotergite not specially protruded, with about 20 setae concentrated on posterior half; mediotergite with more than 20 setae distributed longitudinally along the entire midline. Mesepimeron produced on the ventral half, slender. Pedicel of halter whitish, knob brown, small setae on the entire pedicel.
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<emphasis id="B96CEAECFFB9FFCCFDDBF97C9446806F" bold="true" box="[547,614,1782,1808]" pageId="2" pageNumber="59">Wing</emphasis>
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(
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<figureCitation id="13232A7BFFB9FFCCFD8FF97C949F806F" box="[631,703,1782,1808]" captionStart="FIGURE 4" captionStartId="6.[151,255,1166,1190]" captionTargetBox="[151,1421,477,1142]" captionTargetId="figure@6.[151,1436,477,1142]" captionTargetPageId="6" captionText="FIGURE 4. Azana atlantica, sp. n. Holotype. Wing." httpUri="https://zenodo.org/record/182521/files/figure.png" pageId="2" pageNumber="59">Fig. 4</figureCitation>
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). Length, 4.0 mm. Membrane densely covered with macrotrichia and microtricha. Membrane slightly fumose, with a brown, rounded macula from C to slightly posterior of M at the mid of the wing, and darker at the wing apex; cubital fold conspicuously brown. C ending almost halfway between R5 and M. Sc incomplete. R1 short, reaching C slightly beyond middle of the wing. R5 reaching C well before wing apex, base of Rs indistinct, r-m perfectly longitudinal, indistinct from Rs. Medial fork absent, insertion of M on r-m very basal in the wing, close to origin of CuA. CuA unforked. A1 weak but present, ending half way to wing margin.
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<emphasis id="B96CEAECFFB8FFCDFD84FF1D949E87CE" bold="true" box="[636,702,151,177]" pageId="3" pageNumber="60">Legs.</emphasis>
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Anterior coxa yellowish, basal and distal ends brown, mid and hind coxae brown, setae on distal half; trochanters brown; anterior femur yellowish, mid and hind femora brown; all tibiae and tarsi yellowish, first tarsomere about three times the sum of the length of the remaining tarsomeres. Tibial spurs 1:2:2, brown, mid and hind legs with longer spur about twice the length of the smaller one.
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<emphasis id="B96CEAECFFB8FFCDFF37FEBD9771862E" bold="true" box="[207,337,311,337]" pageId="3" pageNumber="60">Abdomen.</emphasis>
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Tergites 1–2 yellowish, brownish laterally, tergites 3–7 light brown; tergite 7 arched on posterior margin. Sternites 1–3 yellowish, sternites 4–7 light brown, with a pair of longitudinal, weakly sclerotized bands. T8 and S8 brown, short.
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<emphasis id="B96CEAECFFB8FFCDFDE2FE0D948186DE" bold="true" box="[538,673,391,417]" pageId="3" pageNumber="60">Terminalia</emphasis>
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(
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<figureCitation id="13232A7BFFB8FFCDFD4EFE02950886DD" box="[694,808,392,418]" captionStart="FIGURES 5 – 6" captionStartId="7.[151,269,1944,1968]" captionTargetBox="[293,1216,197,1880]" captionTargetId="figure@7.[288,1303,197,1924]" captionTargetPageId="7" captionText="FIGURES 5 – 6. Azana atlantica, sp. n. Holotype. Male terminalia. 5. Ventral view. 6. Dorsal view. Abbreviations: Ce, cercus; Gc, gonocoxite; GcAp, gonocoxal apodeme; Gs, gonostyle; T 9, tergite 9; Te, tegmen." httpUri="https://zenodo.org/record/182522/files/figure.png" pageId="3" pageNumber="60">Figs. 5–6</figureCitation>
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). Terminalia brown. Gonocoxites not fused mesally, well separated from each other, with no projections on distal margin; gonostyle subtriangular, with a digitiform basal projection and a row of 21–24 elongated spines along inner margin. Tegmen relatively unmodified, rounded distally, with a pair of apodemes directed anteriorly; gonocoxal apodemes very long, projected anteriorly beyond the margin of the terminalia. No evidence of parameres (referred to as “inner styles” by
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<bibRefCitation id="EF894B0FFFB8FFCDFAACFDA296F68515" author="Coher" pageId="3" pageNumber="65" refString="Coher, E. I. (1995) A contribution to a revision of the genus Azana Walker, 1856 (Insecta: Diptera: Mycetophilidae: Sciophilinae). Reichenbachia, 31 (17), 83 - 91." type="journal article" year="1995">Coher 1995</bibRefCitation>
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). T9 well developed, almost rectangular. T10 not produced, cerci produced as a pair of lobes more or less fused to each other distally to T9, rather membranous, covered with dense microtricha.
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</paragraph>
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<paragraph id="8BA736FEFFB8FFCDFF3EFD15978B85C5" blockId="3.[151,1437,151,778]" box="[198,427,671,698]" pageId="3" pageNumber="60">
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<emphasis id="B96CEAECFFB8FFCDFF3EFD15970985C6" bold="true" box="[198,297,671,697]" pageId="3" pageNumber="60">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026575FFB8FFCDFF3EFD4D95AF8475" pageId="3" pageNumber="60" type="etymology">
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<paragraph id="8BA736FEFFB8FFCDFF3EFD4D95AF8475" blockId="3.[151,1437,151,778]" pageId="3" pageNumber="60">
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||
<emphasis id="B96CEAECFFB8FFCDFF3EFD4D9773859E" bold="true" box="[198,339,711,737]" pageId="3" pageNumber="60">Etymology.</emphasis>
|
||
The species name is feminine and comes from “Atlantic”, as a reference to the fact that the species is known from the Atlantic Forest of southeastern
|
||
<collectingCountry id="F30F766EFFB8FFCDFCB8FD7A95AB8475" box="[832,907,752,778]" name="Brazil" pageId="3" pageNumber="60">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |