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<mods:title id="8C8B53F89EFE98F6B0850BFAB6B2C512">New species of Carpophilus Stephens from Eastern Arunachal Pradesh, India (Coleoptera: Nitidulidae: Carpophilinae)</mods:title>
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Key to the subgenera and species of
<taxonomicName id="4C184D4F1F41FFB3FDDEFBF3FD66FBFE" ID-CoL="62KJB" box="[603,753,1106,1133]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEADE1F41FFB3FDDEFBF3FD66FBFE" bold="true" box="[603,753,1106,1133]" italics="true" pageId="1" pageNumber="295">Carpophilus</emphasis>
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of Northeast
<collectingCountry id="F30F765C1F41FFB3FC19FBF3FC48FBFF" box="[924,991,1106,1132]" name="India" pageId="1" pageNumber="295">India</collectingCountry>
</emphasis>
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<paragraph id="8BA736CC1F41FFB3FF12FB39FA0AFB3E" blockId="1.[151,1437,1176,2019]" box="[151,1437,1176,1198]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12FB39FA0AFB3E" box="[151,1437,1176,1198]" pageId="1" pageNumber="295">
1. Mesoventrite with raised median longitudinal carina (see
<figureCitation id="13232A491F41FFB3FC8FFB39FCD5FB3E" box="[778,834,1176,1198]" captionStart="FIGURES 89" captionStartId="4.[151,264,1496,1520]" captionTargetBox="[208,1380,1065,1464]" captionTargetId="figure-337@4.[208,1380,1065,1473]" captionTargetPageId="4" captionText="FIGURES 89. Carpophilus spp.: 8, Mesoventrite with median longitudinal carina (C. jelineki); 9, Mesoventrite without median longitudinal carina (C. venkataramani)." figureDoi="http://doi.org/10.5281/zenodo.13849877" httpUri="https://zenodo.org/record/13849877/files/figure.png" pageId="1" pageNumber="295">Fig. 8</figureCitation>
)................................................2.
</keyLead>
</paragraph>
<paragraph id="8BA736CC1F41FFB3FF12FB14FA0AFB59" blockId="1.[151,1437,1176,2019]" box="[151,1437,1205,1227]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12FB14FA0AFB59" box="[151,1437,1205,1227]" pageId="1" pageNumber="295">
- Mesoventrite devoid of median longitudinal carina (see
<figureCitation id="13232A491F41FFB3FD79FB14FCA2FB59" box="[764,821,1205,1227]" captionStart="FIGURES 89" captionStartId="4.[151,264,1496,1520]" captionTargetBox="[208,1380,1065,1464]" captionTargetId="figure-337@4.[208,1380,1065,1473]" captionTargetPageId="4" captionText="FIGURES 89. Carpophilus spp.: 8, Mesoventrite with median longitudinal carina (C. jelineki); 9, Mesoventrite without median longitudinal carina (C. venkataramani)." figureDoi="http://doi.org/10.5281/zenodo.13849877" httpUri="https://zenodo.org/record/13849877/files/figure.png" pageId="1" pageNumber="295">Fig. 9</figureCitation>
)................................................10.
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<keyStep id="30EC2E691F41FFB3FF12FB73FA0AFA04" pageId="1" pageNumber="295">
<paragraph id="8BA736CC1F41FFB3FF12FB73FA0BFAAC" blockId="1.[151,1437,1176,2019]" pageId="1" pageNumber="295">
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2. Antennal club segments (911) somewhat loosely arranged; punctures on mesoventrite shallow, less distinct; distal plate of mesoventrite not divided by median ridge; mesocoxal border regular, uniform, mesocoxal line on metaventrite rectilinear, reaching the metepisternum at 3/4
<superScript id="7C6D9B841F41FFB3FCC8FAADFCCFFA8B" attach="left" box="[845,856,1292,1304]" fontSize="5" pageId="1" pageNumber="295">th</superScript>
of its length, forming a large and conspicuous axillary space……………………
<taxonomicName id="4C184D4F1F41FFB3FE59FA8BFC0EFAAC" authority="Kirejtshuk, 2008" authorityName="Kirejtshuk" authorityYear="2008" box="[476,921,1322,1344]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="subGenus" subGenus="Semocarpolus">
<emphasis id="B96CEADE1F41FFB3FE59FA8BFD65FAAC" box="[476,754,1322,1343]" italics="true" pageId="1" pageNumber="295">Carpophilus (Semocarpolus)</emphasis>
<bibRefCitation id="EF894B3D1F41FFB3FD7DFA8BFC0EFAAC" author="Kirejtshuk, A. G." box="[760,921,1322,1344]" pageId="1" pageNumber="295" pagination="107 - 122" refId="ref5504" refString="Kirejtshuk, A. G. (2008) A current generic classification of sap beetles (Coleoptera, Nitidulidae). Zoosystematica Rossica, 17 (1), 107 - 122. https: // doi. org / 10.31610 / zsr / 2008.17.1.107" type="journal article" year="2008">Kirejtshuk, 2008</bibRefCitation>
</taxonomicName>
.........................................3.
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- Antennal club segments (911) rather compact or closely arranged; strong and distinct punctation on mesoventrite; mesoventrite with longitudinal carina diverging anteriorly and continued through sides; mesocoxal line on metaventrite indistinct; mesocoxal border irregular…………………….
<emphasis id="B96CEADE1F41FFB3FDC0FA22FD7EFA04" box="[581,745,1410,1432]" italics="true" pageId="1" pageNumber="295">
C. (
<taxonomicName id="4C184D4F1F41FFB3FDEDFA23FD75FA04" ID-CoL="62KJB" box="[616,738,1410,1431]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">Carpophilus</taxonomicName>
)
</emphasis>
<bibRefCitation id="EF894B3D1F41FFB3FD6AFA23FC15FA04" author="Stephens, J. F." box="[751,898,1410,1432]" pageId="1" pageNumber="295" refId="ref5709" refString="Stephens, J. F. (1829) A systematic catalogue of British Insects: Being an attempt to arrange all the hitherto discovered indigenous insects in accordance with their natural affinities. Containing also the references to every English writer on entomology, and to the principal foreign authors. With all the published British genera to the present time. Part 1. Baldwin and Cradock, London, xxxiv + 416 + 388 pp. https: // doi. org / 10.5962 / t. 174460" type="book" year="1829">Stephens, 1829</bibRefCitation>
...........................................4.
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<paragraph id="8BA736CC1F41FFB3FF12FA01FA0BFA7C" blockId="1.[151,1437,1176,2019]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12FA01FA0BFA7C" pageId="1" pageNumber="295">
3. Body subquadrate; dorsum rather dull, lack luster; pronotum 1.5× as wide as long; mesocoxal line diverging from mesocoxae at the middle of its outer margin; lateral lobes of male genitalia in ventral view gradually converging and narrowing towards apices, apices slightly upturned, not bending towards each other............
<taxonomicName id="4C184D4F1F41FFB3FC5BFA7AFA0BFA7C" authority="Kirejtshuk, 2018" authorityName="Kirejtshuk" authorityYear="2018" box="[990,1436,1498,1520]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="adjunctus" subGenus="Semocarpolus">
<emphasis id="B96CEADE1F41FFB3FC5BFA7AFB62FA7C" box="[990,1269,1498,1520]" italics="true" pageId="1" pageNumber="295">C. (Semocarpolus) adjunctus</emphasis>
<bibRefCitation id="EF894B3D1F41FFB3FB7EFA7BFA0BFA7C" author="Kirejtshuk, A. G." box="[1275,1436,1498,1520]" pageId="1" pageNumber="295" pagination="1186 - 1216" refId="ref5550" refString="Kirejtshuk, A. G. (2018) New taxa of Carpophilinae (Coleoptera, Nitidulidae) from the Himalaya and Northern Indochina. Part 1. Entomological Review, 98 (9), 1186 - 1216. https: // doi. org / 10.1134 / S 0013873818090063" type="journal article" year="2018">Kirejtshuk, 2018</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8BA736CC1F41FFB3FF12FA59FA0BF9DB" blockId="1.[151,1437,1176,2019]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12FA59FA0BF9DB" pageId="1" pageNumber="295">
- Body moderately elongate; dorsum shiny; pronotum 1.25× as wide as long; mesocoxal line on metaventrite diverging from mesocoxae at one-third of its outer margin; lateral lobes of male genitalia in ventral view subparallel upto apex, apices bending and facing each other like closing ends of a gripping clip..............
<taxonomicName id="4C184D4F1F41FFB3FC31F992FA0BF9DB" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[948,1436,1586,1608]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="marginellus" subGenus="Semocarpolus">
<emphasis id="B96CEADE1F41FFB3FC31F992FB77F9D4" box="[948,1248,1586,1608]" italics="true" pageId="1" pageNumber="295">C. (Semocarpolus) marginellus</emphasis>
Motschulsky, 1858
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<paragraph id="8BA736CC1F41FFB3FF12F9F1FA0BF9F6" blockId="1.[151,1437,1176,2019]" box="[151,1436,1615,1638]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12F9F1FA0BF9F6" box="[151,1436,1615,1638]" pageId="1" pageNumber="295">4. Dorsum dark brownish with macula or pale patch on elytra; lateral margin of pronotum devoid of any sinuation..........5.</keyLead>
</paragraph>
<paragraph id="8BA736CC1F41FFB3FF12F9CCFA0BF90C" blockId="1.[151,1437,1176,2019]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12F9CCFA0BF90C" pageId="1" pageNumber="295">- Dorsum unicolorous, chestnut brown to black; lateral margin of pronotum arcuate with feeble sinuation near apical one-third....................................................................................................8.</keyLead>
</paragraph>
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<paragraph id="8BA736CC1F41FFB3FF12F906FA0AF92D" blockId="1.[151,1437,1176,2019]" box="[151,1437,1703,1726]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12F906FA0AF92D" box="[151,1437,1703,1726]" pageId="1" pageNumber="295">5. Humeral angles of elytra devoid of yellowish spots..........................................................6.</keyLead>
</paragraph>
<paragraph id="8BA736CC1F41FFB3FF12F964FA0AF949" blockId="1.[151,1437,1176,2019]" box="[151,1437,1733,1755]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12F964FA0AF949" box="[151,1437,1733,1755]" pageId="1" pageNumber="295">- Humeral angles of elytra with yellowish spots...............................................................7.</keyLead>
</paragraph>
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<keyStep id="30EC2E691F41FFB3FF12F943FA0BF8FD" pageId="1" pageNumber="295">
<paragraph id="8BA736CC1F41FFB3FF12F943FA0BF8A0" blockId="1.[151,1437,1176,2019]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12F943FA0BF8A0" pageId="1" pageNumber="295">
6. Elytra bear yellowish spots near middle of elytra not reaching apex; lateral lobes of tegmen in male genitalia in ventral view with upturned apices; in lateral view broad base abruptly narrowed behind basal one-fourth, moderately bent with blunt apices, with tuft of moderately long setae on inner margins near apices..............
<taxonomicName id="4C184D4F1F41FFB3FC6BF8BCFA0BF8A0" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" box="[1006,1436,1821,1843]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="bifenestratus" subGenus="Carpophilus">
<emphasis id="B96CEADE1F41FFB3FC6BF8BCFA83F8A1" box="[1006,1300,1821,1842]" italics="true" pageId="1" pageNumber="295">C. (Carpophilus) bifenestratus</emphasis>
<bibRefCitation id="EF894B3D1F41FFB3FA9FF8BCFA0BF8A0" author="Murray, A." box="[1306,1436,1821,1843]" pageId="1" pageNumber="295" pagination="211 - 414" refId="ref5601" refString="Murray, A. (1864) Monograph of the Family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414. https: // doi. org / 10.1111 / j. 1096 - 3642.1863. tb 00163. x" type="journal article" year="1864">Murray, 1864</bibRefCitation>
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</paragraph>
<paragraph id="8BA736CC1F41FFB3FF12F89AFA0BF8FD" blockId="1.[151,1437,1176,2019]" pageId="1" pageNumber="295">
<keyLead id="30E995F91F41FFB3FF12F89AFA0BF8FD" pageId="1" pageNumber="295">
- Elytra bear rectangular yellowish patch extending from base to apex along the suture, sides brown; dorsum nearly glabrous.......................................................................
<taxonomicName id="4C184D4F1F41FFB3FBADF8F9FA0BF8FD" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" box="[1064,1436,1880,1902]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="ligatus" subGenus="Carpophilus">
<emphasis id="B96CEADE1F41FFB3FBADF8F9FA84F8FE" box="[1064,1299,1880,1901]" italics="true" pageId="1" pageNumber="295">C. (Carpophilus) ligatus</emphasis>
<bibRefCitation id="EF894B3D1F41FFB3FA9FF8F9FA0BF8FD" author="Murray, A." box="[1306,1436,1880,1902]" pageId="1" pageNumber="295" pagination="211 - 414" refId="ref5601" refString="Murray, A. (1864) Monograph of the Family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414. https: // doi. org / 10.1111 / j. 1096 - 3642.1863. tb 00163. x" type="journal article" year="1864">Murray, 1864</bibRefCitation>
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7. A distinct, dark M-shaped patch separates the humeral spots and apical spots on elytra; pygidium of female truncate at apex; lateral lobes of male genitalia in ventral view feebly elongate, with slightly broader apices; in lateral view, distinctly bent beyond basal third, narrower towards apex but terminally slightly broader and truncate..............................................................................................
<taxonomicName id="4C184D4F1F41FFB3FC5BF86CFA0BF870" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[990,1436,1997,2019]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="hemipterus" subGenus="Carpophilus">
<emphasis id="B96CEADE1F41FFB3FC5BF86CFB64F871" box="[990,1267,1997,2018]" italics="true" pageId="1" pageNumber="295">C. (Carpophilus) hemipterus</emphasis>
(Linnaeus, 1758)
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<paragraph id="8BA736CC1F42FFB0FF12FF39FA0BFF74" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FF39FA0BFF74" pageId="2" pageNumber="296">
- No distinct M-shaped patch separating humeral spots and apical spots on elytra; pygidium of female rounded at apex; lateral lobes of male genitalia in ventral view broadly elongate, with distinctly broader apices; in lateral view, moderately bent, gradually narrowed beyond basal third with apices rather acute.......
<taxonomicName id="4C184D4F1F42FFB0FC1FFF73FA0BFF74" authority="Audisio &amp; Kirejtshuk, 1989" authorityName="Audisio &amp; Kirejtshuk" authorityYear="1989" box="[922,1436,210,232]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="jelineki" subGenus="Carpophilus">
<emphasis id="B96CEADE1F42FFB0FC1FFF73FB1CFF74" box="[922,1163,210,231]" italics="true" pageId="2" pageNumber="296">C. (Carpophilus) jelineki</emphasis>
Audisio &amp; Kirejtshuk, 1989
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E691F42FFB0FF12FF51FA0AFEB1" pageId="2" pageNumber="296">
<paragraph id="8BA736CC1F42FFB0FF12FF51FA0BFE96" blockId="2.[151,1437,151,1376]" box="[151,1436,239,261]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FF51FA0BFE96" box="[151,1436,239,261]" pageId="2" pageNumber="296">
8. Legs distinctly paler than rest of the body; scutellar shield distinctly tranverse......
<taxonomicName id="4C184D4F1F42FFB0FB9AFF51FA0BFE96" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" box="[1055,1436,239,261]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="flavipes" subGenus="Carpophilus">
<emphasis id="B96CEADE1F42FFB0FB9AFF51FA83FE97" box="[1055,1300,239,261]" italics="true" pageId="2" pageNumber="296">C. (Carpophilus) flavipes</emphasis>
<bibRefCitation id="EF894B3D1F42FFB0FA9FFF51FA0BFE96" author="Murray, A." box="[1306,1436,239,261]" pageId="2" pageNumber="296" pagination="211 - 414" refId="ref5601" refString="Murray, A. (1864) Monograph of the Family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414. https: // doi. org / 10.1111 / j. 1096 - 3642.1863. tb 00163. x" type="journal article" year="1864">Murray, 1864</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA736CC1F42FFB0FF12FEACFA0AFEB1" blockId="2.[151,1437,151,1376]" box="[151,1437,269,291]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FEACFA0AFEB1" box="[151,1437,269,291]" pageId="2" pageNumber="296">- Legs not paler than the rest of the body; scutellar shield marginally transverse.....................................9.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E691F42FFB0FF12FE8BFA0BFE0B" pageId="2" pageNumber="296">
<paragraph id="8BA736CC1F42FFB0FF12FE8BFA0BFECE" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FE8BFA0BFECE" pageId="2" pageNumber="296">
9. Posterior angles of pronotum obtusely pointed; lateral lobes of tegmen of male genitalia in ventral view, narrowly elongate, lobes converging apicad; in lateral view, tegmen moderately bent..............
<taxonomicName id="4C184D4F1F42FFB0FB85FEE9FA0BFECE" authority="Erichson, 1843" authorityName="Erichson" authorityYear="1843" box="[1024,1436,327,349]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="obsoletus" subGenus="Carpophilus">
<emphasis id="B96CEADE1F42FFB0FB85FEE9FA92FECF" box="[1024,1285,327,349]" italics="true" pageId="2" pageNumber="296">C. (Carpophilus) obsoletus</emphasis>
Erichson, 1843
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA736CC1F42FFB0FF12FEC4FA0BFE0B" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FEC4FA0BFE0B" pageId="2" pageNumber="296">
- Posterior angles of pronotum acute; lateral lobes of tegmen of male genitalia in ventral view, broadly elongate, lobes diverging apicad; in lateral view tegmen strongly bent.................................
<taxonomicName id="4C184D4F1F42FFB0FB9DFE23FA0BFE0B" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" box="[1048,1436,386,408]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="cribratus" subGenus="Carpophilus">
<emphasis id="B96CEADE1F42FFB0FB9DFE23FA83FE04" box="[1048,1300,386,407]" italics="true" pageId="2" pageNumber="296">C.(Carpophilus) cribratus</emphasis>
<bibRefCitation id="EF894B3D1F42FFB0FA9FFE22FA0BFE0B" author="Murray, A." box="[1306,1436,386,408]" pageId="2" pageNumber="296" pagination="211 - 414" refId="ref5601" refString="Murray, A. (1864) Monograph of the Family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414. https: // doi. org / 10.1111 / j. 1096 - 3642.1863. tb 00163. x" type="journal article" year="1864">Murray, 1864</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E691F42FFB0FF12FE01FA0AFDDB" pageId="2" pageNumber="296">
<paragraph id="8BA736CC1F42FFB0FF12FE01FA0AFE7C" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FE01FA0AFE7C" pageId="2" pageNumber="296">
10. Body ovoid, dorsum subdepressed; last two segments of antennal club somewhat closely fit, basal antennomere of club loosely articulated; pronotal disc flattened, anterior margin of pronotum narrower than posterior margin; elytra wider than base of prothorax………
<taxonomicName id="4C184D4F1F42FFB0FE02FE7BFCC0FE63" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[391,855,474,496]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="subGenus" subGenus="Ecnomorphus">
<emphasis id="B96CEADE1F42FFB0FE02FE7BFD0CFE7C" box="[391,667,474,495]" italics="true" pageId="2" pageNumber="296">Carpophilus (Ecnomorphus)</emphasis>
Motschulsky, 1858
</taxonomicName>
..............................................11.
</keyLead>
</paragraph>
<paragraph id="8BA736CC1F42FFB0FF12FE59FA0AFDDB" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FE59FA0AFDDB" pageId="2" pageNumber="296">
- Body oblong, subparallel, dorsum semiconvex; segments of antennal club closely arranged, compact; pronotal disc moderately convex, pronotum subquadrate, anterior margin as broad as posterior margin; elytra about as wide as base of prothorax…………
<taxonomicName id="4C184D4F1F42FFB0FF67FD93FDF5FDDB" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" box="[226,610,562,584]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="subGenus" subGenus="Myothorax">
<emphasis id="B96CEADE1F42FFB0FF67FD93FE4EFDD4" box="[226,473,562,583]" italics="true" pageId="2" pageNumber="296">Carpophilus (Myothorax)</emphasis>
<bibRefCitation id="EF894B3D1F42FFB0FE5AFD92FDF5FDDB" author="Murray, A." box="[479,610,562,584]" pageId="2" pageNumber="296" pagination="211 - 414" refId="ref5601" refString="Murray, A. (1864) Monograph of the Family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414. https: // doi. org / 10.1111 / j. 1096 - 3642.1863. tb 00163. x" type="journal article" year="1864">Murray, 1864</bibRefCitation>
</taxonomicName>
..................................................................15.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E691F42FFB0FF12FDF1FA0AFD2E" pageId="2" pageNumber="296">
<paragraph id="8BA736CC1F42FFB0FF12FDF1FA0AFD11" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FDF1FA0AFD11" pageId="2" pageNumber="296">11. Anterior and posterior angles of pronotum broadly rounded; anterior margin almost straight; posterior margin almost straight with a median notch above scutellum....................................................................12.</keyLead>
</paragraph>
<paragraph id="8BA736CC1F42FFB0FF12FD2AFA0AFD2E" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FD2AFA0AFD2E" pageId="2" pageNumber="296">- Anterior and posterior angles of pronotum angulate; anterior margin emarginate; posterior margin feebly emarginate and sinuate on either side of near extremity.........................................................................13.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E691F42FFB0FF12FD64FA0BFC34" pageId="2" pageNumber="296">
<paragraph id="8BA736CC1F42FFB0FF12FD64FA0BFCA1" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FD64FA0BFCA1" pageId="2" pageNumber="296">
12. Body moderately convex; dorsum reddish-brown, with dark rectangular patch on elytra in the middle; prosternal projection more than half as wide as front coxae, apex slightly wider; lateral lobes of tegmen of male genitalia in ventral view broadly elongate, having irregular outer margins, and converge apically like folded hands, apex not upturned; in lateral view, broad base abruptly narrowed behind basal third with acute apices.....................
<taxonomicName id="4C184D4F1F42FFB0FC76FCBCFA0BFCA1" authority="Grouvelle, 1908" authorityName="Grouvelle" authorityYear="1908" box="[1011,1436,797,819]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="signatus" subGenus="Ecnomorphus">
<emphasis id="B96CEADE1F42FFB0FC76FCBCFB6DFCA1" box="[1011,1274,797,818]" italics="true" pageId="2" pageNumber="296">C. (Ecnomorphus) signatus</emphasis>
<bibRefCitation id="EF894B3D1F42FFB0FA85FCBCFA0BFCA1" author="Grouvelle, A." box="[1280,1436,797,819]" pageId="2" pageNumber="296" pagination="325 - 397" refId="ref5121" refString="Grouvelle, A. (1908) Coleopteres de la region indienne. Rhysodidae, Trogositidae, Nitidulidae, Colydiidae, Cucujidae. (1 er memoire). Nitidulidae. Annales de la Societe Entomologique de France, 77, 325 - 397. [https: // biostor. org / reference / 97297]" type="journal article" year="1908">Grouvelle, 1908</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA736CC1F42FFB0FF12FC9AFA0BFC34" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FC9AFA0BFC34" pageId="2" pageNumber="296">
- Body subdepressed; dorsum yellowish-brown, with faint irregular dark patch in middle of elytra; prosternal projection less than one-third as wide as front coxae, apex widened, bulbous; lateral lobes of tegmen of male genitalia in ventral view elongate, having uniform outer margins, gradually narrowed to apex, apex little upturned; in lateral view, lateral lobes gradually narrowed to apex and bent, apex bent upward....................
<taxonomicName id="4C184D4F1F42FFB0FCA9FC33FA0BFC34" authority="Dasgupta, Pal &amp; Powell, 2021" authorityName="Dasgupta, Pal &amp; Powell" authorityYear="2021" box="[812,1436,914,936]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="venkataramani" subGenus="Ecnomorphus">
<emphasis id="B96CEADE1F42FFB0FCA9FC33FBE3FC34" box="[812,1140,914,935]" italics="true" pageId="2" pageNumber="296">C. (Ecnomorphus) venkataramani</emphasis>
<bibRefCitation id="EF894B3D1F42FFB0FBFFFC32FA0BFC34" author="Dasgupta, J. &amp; Pal, T. K. &amp; Powell, G. S." box="[1146,1436,914,936]" pageId="2" pageNumber="296" pagination="627 - 649" refId="ref5014" refString="Dasgupta, J., Pal, T. K. &amp; Powell, G. S. (2021) Taxonomy of Carpophilinae (Coleoptera: Nitidulidae) from Tripura, India with a new species. Annales Zoologici, 71 (3), 627 - 649. https: // doi. org / 10.3161 / 00034541 ANZ 2021.71.3.003" type="journal article" year="2021">Dasgupta, Pal &amp; Powell, 2021</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E691F42FFB0FF12FC11FA0AFBA9" pageId="2" pageNumber="296">
<paragraph id="8BA736CC1F42FFB0FF12FC11FA0BFB93" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FC11FA0BFB93" pageId="2" pageNumber="296">
13. Body slightly elongate; lateral sides of pronotum uniformly arcuate, mild sinuation on posterior margin near corners; male genitalia in lateral view shows partly recurved, somewhat hook-shaped parameres, with rounded apex, in ventral view, parameres conjointly almost as long as broad, apices slightly upturned........
<taxonomicName id="4C184D4F1F42FFB0FC74FC4BFADFFC6C" authorityName="Dasgupta" authorityYear="2024" box="[1009,1352,1002,1023]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="dhritibanerjeeae" status="sp. nov." subGenus="Ecnomorphus">
<emphasis id="B96CEADE1F42FFB0FC74FC4BFADFFC6C" box="[1009,1352,1002,1023]" italics="true" pageId="2" pageNumber="296">C. (Ecnomorphus) dhritibanerjeeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57A51F42FFB0FACBFC4BFA0BFB93" box="[1358,1436,1002,1024]" pageId="2" pageNumber="296" rank="species">
<emphasis id="B96CEADE1F42FFB0FACBFC4BFA01FC6C" bold="true" box="[1358,1430,1002,1023]" pageId="2" pageNumber="296">sp. nov</emphasis>
.
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA736CC1F42FFB0FF12FBA9FA0AFBA9" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FBA9FA0AFBA9" pageId="2" pageNumber="296">- Body somewhat ovate; lateral sides of pronotum slightly arcuate in anterior half and subparallel in posterior half, distinct sinuation on posterior margin near corners................................................................14.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E691F42FFB0FF12FBE3FA0BFB79" pageId="2" pageNumber="296">
<paragraph id="8BA736CC1F42FFB0FF12FBE3FA0BFB00" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FBE3FA0BFB00" pageId="2" pageNumber="296">
14. Paired elliptical patches on elytra with well-defined edges, borders delineated; male genitalia in lateral view somewhat sickle-shaped, with apices acuminate; in ventral view, lateral lobes of tegmen somewhat diverging apicad, apices widely set.................................................................
<taxonomicName id="4C184D4F1F42FFB0FC1AFBDCFA0BFB00" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[927,1436,1149,1171]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="plagiatipennis" subGenus="Ecnomorphus">
<emphasis id="B96CEADE1F42FFB0FC1AFBDCFB48FB01" box="[927,1247,1149,1170]" italics="true" pageId="2" pageNumber="296">C. (Ecnomorphus) plagiatipennis</emphasis>
Motschulsky, 1858
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA736CC1F42FFB0FF12FB3AFA0BFB79" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FB3AFA0BFB79" pageId="2" pageNumber="296">
- Paired elliptical patches on elytra with diffused edges, borders of which not delineated; male genitalia in lateral view somewhat L shaped, with corners curved, apex rounded; in ventral view, lateral lobes of tegmen somewhat parallel lengthwise, apices closely set......................................................
<taxonomicName id="4C184D4F1F42FFB0FC55FB74FA0BFB79" authority="Dasgupta &amp; Pal, 2019" authorityName="Dasgupta &amp; Pal" authorityYear="2019" box="[976,1436,1237,1259]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="jahari" subGenus="Ecnomorphus">
<emphasis id="B96CEADE1F42FFB0FC55FB74FB54FB79" box="[976,1219,1237,1258]" italics="true" pageId="2" pageNumber="296">C. (Ecnomorphus) jahari</emphasis>
<bibRefCitation id="EF894B3D1F42FFB0FB4DFB74FA0BFB79" author="Dasgupta, J. &amp; Pal, T. K." box="[1224,1436,1237,1259]" pageId="2" pageNumber="296" pagination="23 - 28" refId="ref4874" refString="Dasgupta, J. &amp; Pal, T. K. (2019) New species of the genus Carpophilus Stephens, 1829 (Coleoptera: Nitidulidae: Carpophilinae) from India. Far Eastern Entomologist, 378, 23 - 28. [https: // www. biosoil. ru / FEE / Publication / 1810] https: // doi. org / 10.25221 / fee. 378.4" type="book chapter" year="2019">Dasgupta &amp; Pal, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EC2E691F42FFB0FF12FB53FA0BFAF3" pageId="2" pageNumber="296">
<paragraph id="8BA736CC1F42FFB0FF12FB53FA0BFAB6" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FB53FA0BFAB6" pageId="2" pageNumber="296">
15. Tibiae distinctly and conspicuously dilated from basal one-third of hind leg towards apices in males; in male, finger-like lateral lobes of tegmen, gradually narrowed towards apices...........................
<taxonomicName id="4C184D4F1F42FFB0FB9BFAB1FA0BFAB6" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" box="[1054,1436,1295,1317]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="truncatus" subGenus="Myothorax">
<emphasis id="B96CEADE1F42FFB0FB9BFAB1FA83FAB6" box="[1054,1300,1295,1317]" italics="true" pageId="2" pageNumber="296">C. (Myothorax) truncatus</emphasis>
<bibRefCitation id="EF894B3D1F42FFB0FA9FFAB1FA0BFAB6" author="Murray, A." box="[1306,1436,1295,1317]" pageId="2" pageNumber="296" pagination="211 - 414" refId="ref5601" refString="Murray, A. (1864) Monograph of the Family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414. https: // doi. org / 10.1111 / j. 1096 - 3642.1863. tb 00163. x" type="journal article" year="1864">Murray, 1864</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA736CC1F42FFB0FF12FA8CFA0BFAF3" blockId="2.[151,1437,151,1376]" pageId="2" pageNumber="296">
<keyLead id="30E995F91F42FFB0FF12FA8CFA0BFAF3" pageId="2" pageNumber="296">
- Tibiae gradually dilated from base to apex in both sexes; in male, lateral lobes of tegmen abruptly narrowed behind basal fourth, slender apicad and distinctly bent, few marginal setae near apices...................
<taxonomicName id="4C184D4F1F42FFB0FBC4FAEBFA0BFAF3" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" box="[1089,1436,1354,1376]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="nepos" subGenus="Myothorax">
<emphasis id="B96CEADE1F42FFB0FBC4FAEBFA83FAF3" box="[1089,1300,1354,1376]" italics="true" pageId="2" pageNumber="296">C. (Myothorax) nepos</emphasis>
<bibRefCitation id="EF894B3D1F42FFB0FA9FFAEAFA0BFAF3" author="Murray, A." box="[1306,1436,1354,1376]" pageId="2" pageNumber="296" pagination="211 - 414" refId="ref5601" refString="Murray, A. (1864) Monograph of the Family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414. https: // doi. org / 10.1111 / j. 1096 - 3642.1863. tb 00163. x" type="journal article" year="1864">Murray, 1864</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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