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<document id="CBCDFA6AA9BA1843DA05DE590AE1D221" ID-CLB-Dataset="7762" ID-DOI="10.5852/ejt.2021.739.1263" ID-GBIF-Dataset="79e0ebb9-6412-4821-879a-9ec760579681" ID-ISSN="2118-9773" ID-Zenodo-Dep="4600344" ID-ZooBank="C333D3B5-7B4B-4A53-8548-E74DFEE9EBC2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615533921823" checkinUser="plazi" docAuthor="Linský, Marek, Čiamporová-Zaťovičová, Zuzana &amp; Čiampor, Fedor" docDate="2021" docId="03B187D7E832FFDAFE48FA2FFD39FD40" docLanguage="en" docName="ejt-739_1-35.pdf" docOrigin="European Journal of Taxonomy 739 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.739.1263" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onychelmis whiteheadi Spangler &amp; Santiago 1991" docType="treatment" docVersion="8" lastPageNumber="11" masterDocId="FF88FFAFE83AFFD0FFA2FFFFF916FF92" masterDocTitle="A revision of Onychelmis Hinton, 1941 (Coleoptera: Elmidae), with description of new species, DNA barcoding and notes on the geography of the genus" masterLastPageNumber="35" masterPageNumber="1" pageNumber="9" updateTime="1698933319914" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="C70137AC5051B5A6840FA8F83B943C5B">
<mods:title id="F46FEE82C211FF9AD6D84D818890EC6A">A revision of Onychelmis Hinton, 1941 (Coleoptera: Elmidae), with description of new species, DNA barcoding and notes on the geography of the genus</mods:title>
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<mods:name id="5ECEEDC0B9D1C0AB419CF7227EDE772E" type="personal">
<mods:role id="16DE2661B865027DC2AA69A02A9FCC4E">
<mods:roleTerm id="7AD7440753331C91DB85AF1122AF1FEA">Author</mods:roleTerm>
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<mods:namePart id="2811DF4CC2F2FA12B8584AEB3677A1C1">Linský, Marek</mods:namePart>
<mods:nameIdentifier id="160308F5CB19313D0D5BB2C342E4A507" type="LSID">E2AC74B9-FA5F-4D43-9168-4513D3E80A04</mods:nameIdentifier>
<mods:affiliation id="009E5C36F8267983E6DD9D1FB43744FE">Zoology Lab, Institute of Botany, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, SK- 84523, Bratislava, Slovakia. &amp; Department of Zoology, Faculty of Natural Science, Comenius University, Mlynská dolina B- 1, SK- 84215 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="3F5B8C961C1ADDA8D3D2A0F777AD26E9" type="email">marek.linsky@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C501274503C241F94B25E122B2F8B0F5">Čiamporová-Zaťovičová, Zuzana</mods:namePart>
<mods:nameIdentifier id="EB413C41793C2F7B5E2E4A65B50CC6CC" type="LSID">8D6A1FDB-06AB-44A7-B1F9-3FC86B7C9804</mods:nameIdentifier>
<mods:affiliation id="84B37926C8C741D48B9994874B685EC3">Zoology Lab, Institute of Botany, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, SK- 84523, Bratislava, Slovakia. &amp; Department of Ecology, Faculty of Natural Science, Comenius University, Mlynská dolina B- 1, SK- 84215 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="F2AC8941C8DC3961FC669F5BE46DBD66" type="email">zuzana.zatovicova@savba.sk</mods:nameIdentifier>
</mods:name>
<mods:name id="6582EC6D168B1C90A1F3B2F5D02BEE4A" type="personal">
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<mods:namePart id="A429B00B6023B2EAA4D8A06A4D25D344">Čiampor, Fedor</mods:namePart>
<mods:nameIdentifier id="3669579913108E042C19259A5C4C0A75" type="LSID">977CE415-B2E7-45DA-9DBE-B6D7CCCC58E4</mods:nameIdentifier>
<mods:affiliation id="2B1B998505683BA8F2D62430D0760AFE">Zoology Lab, Institute of Botany, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, SK- 84523, Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="15484812B98B851323D7775463F2D94B" type="email">f.ciampor@savba.sk</mods:nameIdentifier>
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<mods:title id="533596B842F9A65EBAE02F0427006D06">European Journal of Taxonomy</mods:title>
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<treatment id="03B187D7E832FFDAFE48FA2FFD39FD40" ID-DOI="http://doi.org/10.5281/zenodo.4602497" ID-GBIF-Taxon="180331473" ID-Zenodo-Dep="4602497" LSID="urn:lsid:plazi:treatment:03B187D7E832FFDAFE48FA2FFD39FD40" httpUri="http://treatment.plazi.org/id/03B187D7E832FFDAFE48FA2FFD39FD40" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="C302654AE832FFD8FE48FA2FFD5CFA79" box="[490,1098,1488,1515]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BA736C1E832FFD8FE48FA2FFD5CFA79" blockId="8.[490,1098,1488,1551]" box="[490,1098,1488,1515]" pageId="8" pageNumber="9">
<heading id="D0EF81ADE832FFD8FE48FA2FFD5CFA79" box="[490,1098,1488,1515]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="4C184D42E832FFD8FE48FA2FFD5CFA79" ID-CoL="49QNY" authority="Spangler &amp; Santiago, 1991" authorityName="Spangler &amp; Santiago" authorityYear="1991" box="[490,1098,1488,1515]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B96CEAD3E832FFD8FE48FA2FFA13FA79" bold="true" box="[490,773,1488,1515]" italics="true" pageId="8" pageNumber="9">Onychelmis whiteheadi</emphasis>
<bibRefCitation id="EF894B30E832FFD8FCAEFA2EFD5CFA79" author="Spangler P. &amp; Santiago S." box="[780,1098,1489,1515]" pageId="8" pageNumber="9" pagination="495 - 498" refId="ref17990" refString="Spangler P. &amp; Santiago S. 1991. A new species and new records from Colombia of the water beetle genus Onychelmis Hinton (Coleoptera: Elmidae: Elminae). Proceedings of the Entomological Society of Washington 93 (2): 495 - 498." type="journal article" year="1991">Spangler &amp; Santiago, 1991</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302654AE832FFD8FDD4FA0AFAA8F99D" box="[630,958,1525,1551]" pageId="8" pageNumber="9" type="description">
<paragraph id="8BA736C1E832FFD8FDD4FA0AFAA8F99D" blockId="8.[490,1098,1488,1551]" box="[630,958,1525,1551]" pageId="8" pageNumber="9">
<figureCitation id="13232A44E832FFD8FDD4FA0AFBA8F99D" box="[630,702,1525,1551]" captionStart="Fig" captionStartId="7.[188,231,1928,1954]" captionTargetBox="[248,1355,704,1872]" captionTargetId="figure-147@7.[189,1398,684,1890]" captionTargetPageId="7" captionText="Fig. 2. Habitus of Onychelmis spp. A–B. O. longicollis (Sharp, 1882), holotype, ♂ (NHM). A. Dorsal view. B. Ventral view. C–D. O. whiteheadi Spangler &amp; Santiago, 1991, holotype, ♂ (NMNH). C. Dorsal view. D. Ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4600350" httpUri="https://zenodo.org/record/4600350/files/figure.png" pageId="8" pageNumber="9">Figs 2</figureCitation>
C–D, 6H, 7A–B, 12B
</paragraph>
</subSubSection>
<subSubSection id="C302654AE832FFD8FF1FF9C8FAD2F9C3" box="[189,964,1591,1618]" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="8BA736C1E832FFD8FF1FF9C8FAD2F9C3" blockId="8.[189,964,1591,1618]" box="[189,964,1591,1618]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="AB0811EFE832FFD8FF1FF9C8FAD2F9C3" box="[189,964,1591,1618]" pageId="8" pageNumber="9">
<treatmentCitation id="0AB910D0E832FFD8FF1FF9C8FA45F9C0" author="Spangler P. &amp; Santiago S." box="[189,851,1591,1618]" page="495" pageId="8" pageNumber="9" year="1991">
<taxonomicName id="4C184D42E832FFD8FF1FF9C8FA45F9C0" ID-CoL="49QNY" authority="Spangler &amp; Santiago, 1991: 495" authorityName="Spangler &amp; Santiago" authorityYear="1991" box="[189,851,1591,1618]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B96CEAD3E832FFD8FF1FF9C8F8C6F9C3" box="[189,464,1591,1617]" italics="true" pageId="8" pageNumber="9">Onychelmis whiteheadi</emphasis>
<bibRefCitation id="EF894B30E832FFD8FE75F9C8FA45F9C0" author="Spangler P. &amp; Santiago S." box="[471,851,1591,1618]" pageId="8" pageNumber="9" pagination="495 - 498" refId="ref17990" refString="Spangler P. &amp; Santiago S. 1991. A new species and new records from Colombia of the water beetle genus Onychelmis Hinton (Coleoptera: Elmidae: Elminae). Proceedings of the Entomological Society of Washington 93 (2): 495 - 498." type="journal article" year="1991">Spangler &amp; Santiago, 1991: 495</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1–2.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C302654AE832FFD8FF1FF986FAB9F8BD" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BA736C1E832FFD8FF1FF986F8D0F901" blockId="8.[189,454,1657,1683]" box="[189,454,1657,1683]" pageId="8" pageNumber="9">
<heading id="D0EF81ADE832FFD8FF1FF986F8D0F901" bold="true" box="[189,454,1657,1683]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B96CEAD3E832FFD8FF1FF986F8D0F901" bold="true" box="[189,454,1657,1683]" pageId="8" pageNumber="9">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736C1E832FFD8FF1FF956FAB9F8BD" blockId="8.[189,1399,1705,1840]" pageId="8" pageNumber="9">
<taxonomicName id="4C184D42E832FFD8FF1FF956F8C6F951" authorityName="Spangler &amp; Santiago" authorityYear="1991" box="[189,464,1705,1731]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B96CEAD3E832FFD8FF1FF956F8C6F951" box="[189,464,1705,1731]" italics="true" pageId="8" pageNumber="9">Onychelmis whiteheadi</emphasis>
</taxonomicName>
can be distinguished from other species of the genus by the combination of the following characters: 1) larger size (CL:
<quantity id="4CE09B24E832FFD8FD0EF933FA49F975" box="[684,863,1740,1767]" metricMagnitude="-3" metricUnit="m" metricValue="1.6549999999999998" metricValueMax="1.66" metricValueMin="1.65" pageId="8" pageNumber="9" unit="mm" value="1.6549999999999998" valueMax="1.66" valueMin="1.65">1.65–1.66 mm</quantity>
); 2) pro- and mesofemora with tomentum at base (
<figureCitation id="13232A44E832FFD8FEA0F90EF877F89E" box="[258,353,1777,1804]" captionStart="Fig" captionStartId="18.[189,232,1893,1919]" captionTargetBox="[199,1389,1223,1846]" captionTargetId="figure-257@18.[189,1398,1213,1855]" captionTargetPageId="18" captionText="Fig. 6. Tomentum on pro- and mesofemora of Onychelmis spp. A. O. leleupi Delève, 1968. B. O. lenkae sp. nov. C. O. lobata sp. nov. D. O. longicollis (Sharp, 1882). E. O. minor sp. nov. F. O. onorei sp. nov. G. O. splendida sp. nov. H. O. whiteheadi Spangler &amp; Santiago, 1991. Drawings are schematic, bodies are divided to identical halves: left = male, right = female (note: females unknown for D and H)." figureDoi="http://doi.org/10.5281/zenodo.4600362" httpUri="https://zenodo.org/record/4600362/files/figure.png" pageId="8" pageNumber="9">Fig. 6H</figureCitation>
); 3) humeri produced; 4) prominent carina on sixth interval present; 5) elytral punctures shallowly impressed; 6) apeX of aedeagus trilobate (
<figureCitation id="13232A44E832FFD8FC83F8EAFA73F8BD" box="[801,869,1813,1839]" captionStart="Fig" captionStartId="20.[189,232,1926,1952]" captionTargetBox="[252,1337,285,1878]" captionTargetId="figure-16@20.[234,1353,265,1886]" captionTargetPageId="20" captionText="Fig. 7. Aedeagi of Onychelmis spp. A. O. whiteheadi Spangler &amp; Santiago, 1991, ventral view. B. O. whiteheadi, lateral view. C. O. longicollis (Sharp, 1882), ventral view. D. O. longicollis, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4600366" httpUri="https://zenodo.org/record/4600366/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
A–B).
</paragraph>
</subSubSection>
<subSubSection id="C302654AE832FFD9FF1FF8A8FD44FEFC" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BA736C1E832FFD8FF1FF8A8F8BCF8E3" blockId="8.[189,426,1879,1905]" box="[189,426,1879,1905]" pageId="8" pageNumber="9">
<heading id="D0EF81ADE832FFD8FF1FF8A8F8BCF8E3" bold="true" box="[189,426,1879,1905]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B96CEAD3E832FFD8FF1FF8A8F8BCF8E3" bold="true" box="[189,426,1879,1905]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736C1E832FFD8FF48F879FD20F87B" blockId="8.[189,1398,1926,2025]" pageId="8" pageNumber="9">
<materialsCitation id="3B703C9CE832FFD8FF48F879FD20F87B" ID-GBIF-Occurrence="3051860317" collectingDate="1984-02-26" collectionCode="NMNH" collectorName="Maria T. Szauer" country="Colombia" location="COLOMBIA" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Bogota" typeStatus="holotype">
<emphasis id="B96CEAD3E832FFD8FF48F879F84AF832" bold="true" box="[234,348,1926,1952]" pageId="8" pageNumber="9">
<typeStatus id="54A38863E832FFD8FF48F879F84AF832" box="[234,348,1926,1952]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F30F7651E832FFD8FF1FF855F84EF856" box="[189,344,1962,1988]" name="Colombia" pageId="8" pageNumber="9">COLOMBIA</collectingCountry>
•
<specimenCount id="9D1EFD48E832FFD8FED3F854F89FF857" box="[369,393,1963,1989]" pageId="8" pageNumber="9" type="male">♂</specimenCount>
; “
<collectingCountry id="F30F7651E832FFD8FE02F855FB52F856" box="[416,580,1962,1988]" name="Colombia" pageId="8" pageNumber="9">COLOMBIA</collectingCountry>
: [CUND.]:
<locationDeviation id="9A152FE3E832FFD8FD6FF855FAE1F857" box="[717,1015,1962,1989]" pageId="8" pageNumber="9">Río Cachéta near Bogota</locationDeviation>
,
<date id="FFA61001E832FFD8FBA0F855FD81F856" box="[1026,1175,1962,1988]" pageId="8" pageNumber="9" value="1984-02-26">
<collectingDate id="EFE2E9E9E832FFD8FBA0F855FD81F856" box="[1026,1175,1962,1988]" pageId="8" pageNumber="9" value="1984-02-26">26 Feb 1984</collectingDate>
</date>
,
<collectorName id="26ED5317E832FFD8FB06F855FC49F856" box="[1188,1375,1962,1989]" pageId="8" pageNumber="9">Maria T. Szauer</collectorName>
// [red] HOLOTYPE
<taxonomicName id="4C184D42E832FFD8FE3DF831FA33F87A" authority="Spangler" authorityName="Spangler &amp; Santiago" authorityYear="1991" box="[415,805,1998,2024]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="whiteheadi">Onychelmis whiteheadi Spangler</taxonomicName>
&amp; Santiago.”;
<collectionCode id="ED09AE04E832FFD8FC77F830FD25F87B" box="[981,1075,1999,2025]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9" type="Museum">NMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA736C1E833FFD9FF48FEF6FD44FEFC" blockId="9.[189,1399,265,366]" pageId="9" pageNumber="10">
<materialsCitation id="3B703C9CE833FFD9FF48FEF6FD58FEFC" ID-GBIF-Occurrence="3051860304" collectingDate="1984-02-26" collectionCode="NMNH" collectorName="Maria T. Szauer" country="Colombia" location="COLOMBIA" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Bogota" typeStatus="paratype">
<emphasis id="B96CEAD3E833FFD9FF48FEF6F84AFEB1" bold="true" box="[234,348,265,291]" pageId="9" pageNumber="10">
<typeStatus id="54A38863E833FFD9FF48FEF6F84AFEB1" box="[234,348,265,291]" pageId="9" pageNumber="10" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F30F7651E833FFD9FF1FFED1F84EFEDA" box="[189,344,302,328]" name="Colombia" pageId="9" pageNumber="10">COLOMBIA</collectingCountry>
•
<specimenCount id="9D1EFD48E833FFD9FED3FED1F889FEDA" box="[369,415,302,328]" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
; “
<collectingCountry id="F30F7651E833FFD9FE15FED1FB4DFEDA" box="[439,603,302,328]" name="Colombia" pageId="9" pageNumber="10">COLOMBIA</collectingCountry>
: [CUND.]:
<locationDeviation id="9A152FE3E833FFD9FD46FED1FD18FEDA" box="[740,1038,302,328]" pageId="9" pageNumber="10">Río Cachéta near Bogota</locationDeviation>
,
<date id="FFA61001E833FFD9FBBBFED1FDB8FEDA" box="[1049,1198,302,328]" pageId="9" pageNumber="10" value="1984-02-26">
<collectingDate id="EFE2E9E9E833FFD9FBBBFED1FDB8FEDA" box="[1049,1198,302,328]" pageId="9" pageNumber="10" value="1984-02-26">26 Feb 1984</collectingDate>
</date>
,
<collectorName id="26ED5317E833FFD9FB19FED1FC61FEDA" box="[1211,1399,302,328]" pageId="9" pageNumber="10">Maria T. Szauer</collectorName>
// [blue] PARATYPE
<taxonomicName id="4C184D42E833FFD9FE19FEACFA56FEFF" authority="Spangler" authorityName="Spangler &amp; Santiago" authorityYear="1991" box="[443,832,339,365]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="whiteheadi">Onychelmis whiteheadi Spangler</taxonomicName>
&amp; Santiago.”;
<collectionCode id="ED09AE04E833FFD9FC52FEABFD58FEFC" box="[1008,1102,340,366]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">NMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C302654AE833FFDAFF1FFE69F82FFDF2" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="8BA736C1E833FFD9FF1FFE69F87DFE22" blockId="9.[189,363,406,432]" box="[189,363,406,432]" pageId="9" pageNumber="10">
<heading id="D0EF81ADE833FFD9FF1FFE69F87DFE22" bold="true" box="[189,363,406,432]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B96CEAD3E833FFD9FF1FFE69F87DFE22" bold="true" box="[189,363,406,432]" pageId="9" pageNumber="10">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736C1E833FFD9FF48FE38F83CFE73" blockId="9.[189,1398,455,631]" box="[234,298,455,481]" pageId="9" pageNumber="10">
<emphasis id="B96CEAD3E833FFD9FF48FE38F83CFE73" bold="true" box="[234,298,455,481]" pageId="9" pageNumber="10">Male</emphasis>
</paragraph>
<paragraph id="8BA736C1E833FFD9FF1FFE13FB4AFDE4" blockId="9.[189,1398,455,631]" pageId="9" pageNumber="10">
BODY. Obovate, black (
<figureCitation id="13232A44E833FFD9FE42FE13FB3AFD94" box="[480,556,492,518]" captionStart="Fig" captionStartId="7.[188,231,1928,1954]" captionTargetBox="[248,1355,704,1872]" captionTargetId="figure-147@7.[189,1398,684,1890]" captionTargetPageId="7" captionText="Fig. 2. Habitus of Onychelmis spp. A–B. O. longicollis (Sharp, 1882), holotype, ♂ (NHM). A. Dorsal view. B. Ventral view. C–D. O. whiteheadi Spangler &amp; Santiago, 1991, holotype, ♂ (NMNH). C. Dorsal view. D. Ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4600350" httpUri="https://zenodo.org/record/4600350/files/figure.png" pageId="9" pageNumber="10">Fig. 2</figureCitation>
C–D); length
<quantity id="4CE09B24E833FFD9FD7BFE13FA9BFD95" box="[729,909,492,519]" metricMagnitude="-3" metricUnit="m" metricValue="1.6549999999999998" metricValueMax="1.66" metricValueMin="1.65" pageId="9" pageNumber="10" unit="mm" value="1.6549999999999998" valueMax="1.66" valueMin="1.65">1.65–1.66 mm</quantity>
; width
<quantity id="4CE09B24E833FFD9FC4DFE13FDB5FD95" box="[1007,1187,492,519]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="8.1" metricValueMin="7.9" pageId="9" pageNumber="10" unit="mm" value="0.8" valueMax="0.81" valueMin="0.79">0.79–0.81 mm</quantity>
; dorsum conveX, glabrous, with sparse, light yellowish setae. Hairy or scale-like tomentum distributed on following areas: genae, sides of prosternum, mesoventrite, metaventrite and abdomen, epipleura, and medial and lateral portions on bases of femora.
</paragraph>
<paragraph id="8BA736C1E833FFD9FF1FFD5FF85EFC96" blockId="9.[189,1401,671,772]" pageId="9" pageNumber="10">COLOUR. Head, pronotum and elytra black; venter dark brown to black; coxae, femora and tarsi dark brown with reddish tinge; trochanters and tibiae brown; basal segments of antennae and tarsal claws pale brown.</paragraph>
<paragraph id="8BA736C1E833FFD9FF1FFCD1F9ECFBE0" blockId="9.[189,1399,813,1138]" pageId="9" pageNumber="10">
HEAD. Partly retractable into prothoraX, dorsally shiny. Antennae filiform, 11-segmented; pedicel about twice as long as scape; remaining segments about 4 times as long as scape and pedicel combined, segments 3–10 subrectangular, subequal in length; terminal segment longest, suboval, with pointed apex. Labrum with anterior margin very slightly emarginate medially; anterolateral angles broadly arcuate with numerous golden, recumbent hair-like setae; clypeus shorter and wider than labrum, about 3.5 times as wide as long, anterior margin slightly concave, anterolateral angles rounded; frontoclypeal suture almost straight. Eyes well developed, HW:
<quantity id="4CE09B24E833FFD9FCA9FBF2FAAEFBB5" box="[779,952,1037,1063]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="9" pageNumber="10" unit="mm" value="0.315" valueMax="0.32" valueMin="0.31">0.31–0.32 mm</quantity>
, ID:
<quantity id="4CE09B24E833FFD9FC51FBF2FDB7FBB5" box="[1011,1185,1037,1064]" metricMagnitude="-4" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="9" pageNumber="10" unit="mm" value="0.155" valueMax="0.16" valueMin="0.15">0.15–0.16 mm</quantity>
, suboval in lateral view, protruding from head outline in dorsal view, circumocular surface raised. Frons convex between eyes.
</paragraph>
<paragraph id="8BA736C1E833FFD9FF1FFB64FD06F87A" blockId="9.[188,1401,1179,2025]" pageId="9" pageNumber="10">
THORAX. Pronotum widest behind middle, PW:
<quantity id="4CE09B24E833FFD9FD53FB64FA89FB27" box="[753,927,1179,1205]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.699999999999999" pageId="9" pageNumber="10" unit="mm" value="0.475" valueMax="0.48" valueMin="0.47">0.47–0.48 mm</quantity>
, PL:
<quantity id="4CE09B24E833FFD9FC7CFB64FD9AFB27" box="[990,1164,1179,1206]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.4" metricValueMin="5.1" pageId="9" pageNumber="10" unit="mm" value="0.525" valueMax="0.54" valueMin="0.51">0.51–0.54 mm</quantity>
; surface shiny, with narrow reticulation along basal margin and posterolateral angles, with dense tiny punctures; sublateral carinae never well-developed, but a short, very fine raised line in basal ¼ present; disc conveX, divided by a broad, deep transverse impression before middle; two prescutellar foveae separated by a raised line extending from base to apical discal half, connecting both halves and merging into them; anterior margin arcuate; posterior margin bisinuate; sides of pronotum convex before and after transverse constriction; lateral margins narrowly rimmed; posterolateral angles orthogonal; anterolateral angles slightly produced. Hypomeron finely microreticulate, widest in middle. Prosternum moderately long in front of procoxae, with anterior margin concave; sides raised around procoxae, forming carinae, not reaching anterior margin, prosternal process long, moderately broad and with posterior margin broadly rounded. Mesoventrite coarse, short and wide, with deep triangular groove for reception of prosternal process; posterior margin around mesocoxae raised. Metaventrite slightly wider than long, wrinkled, more or less shiny with setigerous punctures; disc convex with deep, medial, triangular depression in posterior half; discrimen in basal ¾; in males with one prebasal fovea on each side. Elytra (EL:
<quantity id="4CE09B24E833FFD9FB67F980FC65F908" box="[1221,1395,1663,1690]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="9" pageNumber="10" unit="mm" value="0.135" valueMax="0.15" valueMin="0.12">0.12–0.15 mm</quantity>
, EW:
<quantity id="4CE09B24E833FFD9FF58F95BF8BFF92D" box="[250,425,1700,1727]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="8.1" metricValueMin="7.9" pageId="9" pageNumber="10" unit="mm" value="0.8" valueMax="0.81" valueMin="0.79">0.79–0.81 mm</quantity>
) conveX, widest in anterior ¾; sides strongly declivous; surface shiny, with dense tiny punctures; elytral margin narrowly rimmed; humeri protruding from outline; epipleuron tapering posteriorly. Prominent carina on siXth interval in ⅘ of elytron. Elytra with ten rows of small, shallowly impressed punctures, separated by 2–4 times puncture diameter, diminishing toward lateral margins and elytral apeX. Scutellum subovate, flat. Legs moderately long; femora clavate; tibiae longest. Protibiae with anterior cleaning fringe on apical ⅓; mesotibiae with two cleaning fringes – anterior on apical ⅕ and posterior on apical ⅓; metatibiae with posterior cleaning fringe on apical ⅓. Tarsi 5-segmented, first four segments each with one fine pale, recumbent seta, fifth segment slightly shorter than remaining segments combined; claws with a large subbasal and smaller basal teeth.
</paragraph>
<paragraph id="8BA736C1E830FFDAFF1FFEF6FA4BFE68" blockId="10.[189,1400,265,506]" pageId="10" pageNumber="11">
ABDOMEN. With 5 ventrites. First ventrite with basal margin broadly rounded, fifth ventrite longest, apically setose, with posterior margin broadly arcuate. Aedeagus (
<figureCitation id="13232A44E830FFDAFC72FED2FD00FED5" box="[976,1046,301,327]" captionStart="Fig" captionStartId="20.[189,232,1926,1952]" captionTargetBox="[252,1337,285,1878]" captionTargetId="figure-16@20.[234,1353,265,1886]" captionTargetPageId="20" captionText="Fig. 7. Aedeagi of Onychelmis spp. A. O. whiteheadi Spangler &amp; Santiago, 1991, ventral view. B. O. whiteheadi, lateral view. C. O. longicollis (Sharp, 1882), ventral view. D. O. longicollis, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4600366" httpUri="https://zenodo.org/record/4600366/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
A–B) elongate. Penis without fibula; corona below apeX, well developed; in ventral view concave, from apical half widening toward laterally projected, apically nearly acute sides, apical margin medially with wide arcuate projection; in lateral view thick, evenly narrowed from base to curved apex, with apical sides pointing backwards. Parameres absent. Phallobase about 1.5 times as long as penis, subparallel, distal portion markedly narrowed, in ventral view straight, curved in lateral view.
</paragraph>
<paragraph id="8BA736C1E830FFDAFF48FDDDF853FDAE" blockId="10.[189,325,546,608]" box="[234,325,546,572]" pageId="10" pageNumber="11">
<emphasis id="B96CEAD3E830FFDAFF48FDDDF853FDAE" bold="true" box="[234,325,546,572]" pageId="10" pageNumber="11">Female</emphasis>
</paragraph>
<paragraph id="8BA736C1E830FFDAFF1FFDB9F82FFDF2" blockId="10.[189,325,546,608]" box="[189,313,582,608]" pageId="10" pageNumber="11">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C302654AE830FFDAFF1FFD77FD39FD40" box="[189,1071,648,722]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BA736C1E830FFDAFF1FFD77F840FD30" blockId="10.[189,342,648,674]" box="[189,342,648,674]" pageId="10" pageNumber="11">
<heading id="D0EF81ADE830FFDAFF1FFD77F840FD30" bold="true" box="[189,342,648,674]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B96CEAD3E830FFDAFF1FFD77F840FD30" bold="true" box="[189,342,648,674]" pageId="10" pageNumber="11">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736C1E830FFDAFF1FFD47FD39FD40" blockId="10.[189,1071,695,722]" box="[189,1071,695,722]" pageId="10" pageNumber="11">
Known only from the
<typeStatus id="54A38863E830FFDAFE60FD47F8E3FD40" box="[450,501,696,722]" pageId="10" pageNumber="11">type</typeStatus>
locality in
<collectingRegion id="49DCF823E830FFDAFDDBFD48FA31FD43" box="[633,807,695,721]" country="Colombia" name="Cundinamarca" pageId="10" pageNumber="11">Cundinamarca</collectingRegion>
,
<collectingCountry id="F30F7651E830FFDAFC91FD48FAB1FD43" box="[819,935,695,721]" name="Colombia" pageId="10" pageNumber="11">Colombia</collectingCountry>
(
<figureCitation id="13232A44E830FFDAFC15FD48FD34FD40" box="[951,1058,695,722]" captionStart="Fig" captionStartId="28.[189,232,1916,1942]" captionTargetBox="[189,1398,1158,1876]" captionTargetId="figure-274@28.[189,1398,1158,1876]" captionTargetPageId="28" captionText="Fig. 12. Distribution map of Onychelmis Hinton, 1941. A. New species described within this study. B. Published records of Onychelmis and potential distribution of the genus." figureDoi="http://doi.org/10.5281/zenodo.4600376" httpUri="https://zenodo.org/record/4600376/files/figure.png" pageId="10" pageNumber="11">Fig. 12B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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