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<figureCitation id="13232A2DFFD3FD555D9CF9FAFEB3440E" box="[151,262,1616,1642]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="3.[152,255,1145,1169]" captionStartId-1="4.[152,255,1974,1998]" captionStartId-10="15.[152,255,1908,1932]" captionStartId-11="16.[152,255,1908,1932]" captionStartId-2="5.[152,255,1943,1967]" captionStartId-3="6.[152,255,1943,1967]" captionStartId-4="7.[152,255,1075,1099]" captionStartId-5="8.[152,255,1867,1891]" captionStartId-6="9.[152,255,1908,1932]" captionStartId-7="10.[152,255,1943,1967]" captionStartId-8="11.[152,255,1908,1932]" captionStartId-9="12.[152,255,1484,1508]" captionTargetBox-0="[181,1410,310,1115]" captionTargetBox-1="[177,1411,698,1926]" captionTargetBox-10="[166,1419,184,1878]" captionTargetBox-11="[164,1423,181,1884]" captionTargetBox-2="[154,1435,195,1910]" captionTargetBox-3="[160,1431,185,1919]" captionTargetBox-4="[153,1428,189,1051]" captionTargetBox-5="[151,1427,287,1838]" captionTargetBox-6="[166,1422,186,1884]" captionTargetBox-7="[153,1435,185,1919]" captionTargetBox-8="[164,1423,181,1884]" captionTargetBox-9="[310,1278,181,1460]" captionTargetId-0="figure-70@3.[176,1410,306,1119]" captionTargetId-1="figure-263@4.[164,1423,667,1950]" captionTargetId-10="figure-18@15.[164,1423,181,1884]" captionTargetId-11="figure-27@16.[164,1423,181,1884]" captionTargetId-2="figure-18@5.[151,1435,181,1919]" captionTargetId-3="figure-27@6.[151,1435,181,1919]" captionTargetId-4="figure-18@7.[151,1435,181,1051]" captionTargetId-5="figure-62@8.[151,1435,272,1844]" captionTargetId-6="figure-18@9.[164,1423,181,1884]" captionTargetId-7="figure-27@10.[151,1435,181,1919]" captionTargetId-8="figure-18@11.[164,1423,181,1884]" captionTargetId-9="figure-27@12.[310,1278,181,1460]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="15" captionTargetPageId-11="16" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="FIGURE 1. Map showing the known distribution of the Pselaphacus procerus Kuhnt, 1910. (A–G) Brazil: (A) Benjamin Constant, Amazonas state: 4° 23′ 0″ S, 70° 1′ 53″W. (B) Uberlândia, Minas Gerais state: 18° 54′41″ S, 48° 15′ 44″W. (C) Três Lagoas, Mato Grosso do Sul state: 20° 45′ 35″ S, 51° 41′ 42″W. (D) Castilho, São Paulo state: 20° 52′ 12″ S, 51° 29′ 13″W. (E) Marília, São Paulo state: 22° 13′ 15″ S, 49° 56′ 55″W. (F) Salto Grande, São Paulo state: 22° 52′ 53″ S, 49° 57′ 44″W. (G) Piracicaba, São Paulo state: 22° 43’ 30” S, 47° 38’ 56” W. (H) Misiones, Argentina: 26° 55′ S, 54° 31′ W, [type locality]." captionText-1="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." captionText-10="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." captionText-11="FIGURE 12. Pselaphacus procerus Kuhnt, 1910. Early larvae under SEM. (A–B) head, ventral and lateral views; (C) mouthparts, ventral; (D) antenna; (E) mesothoracic tubercle near coxa; (F) mesonotum; (G–H) thorax, lateral and ventral views; (I) thoracic spiracle; (J) abdominal spiracle; (K) segments VIII–X, lateral view. Scale bar: A, B, F, G, H, K = 0.1 mm; C, D, E = 0.03 mm; I, J = 0.01 mm." captionText-2="FIGURE 3. Pselaphacus procerus Kuhnt, 1910. Male. Mouthparts. (A) front and clypeus; (B) epipharynx; (C–D) mandible, dorsal and ventral views; (E) maxilla and labium; (F–G) lacinia and galea, dorsal and ventral views. Scale bar: A, C, D, E = 0.5 mm, B, F, G = 0.1 mm." captionText-3="FIGURE 4. Pselaphacus procerus Kuhnt, 1910. Male. (A) sternite IX; (B) tergite and sternite IX (C–G) aedeagus: (C–D) tegmen and parameres, ventral and lateral views; (E–F) penis, lateral and ventral views; (G) apex of accessory muscle. Scale bar: A, B, C, F = 0.5 mm, D, E = 0.1 mm." captionText-4="FIGURE 5. Pselaphacus procerus Kuhnt, 1910. Female. (A) sternite VIII; (B) tergite VIII; (C) spermatheca; (D) ovipositor, dorsal view. Scale bar: A, B, D = 0.5 mm, C = 0.1 mm." captionText-5="FIGURE 6. Pselaphacus procerus Kuhnt, 1910. Mature larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." captionText-6="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." captionText-7="FIGURE 8. Pselaphacus procerus Kuhnt, 1910. Mature larva: (A) pro and mesonotum; (C) prothoracic leg, internal view; (D) pretarsus. Early larva: (B) pro and mesonotum; (E) prothoracic leg, internal view; (F) middle leg, external view; (G) thoracic spiracle; (H) segment IX, dorsal view; (I) urogomphi, apex. Scale bar: A, B = 1.0 mm, C, G, H = 0.1 mm, E, F = 0.5 mm." captionText-8="FIGURE 9. Pselaphacus procerus Kuhnt, 1910. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm." captionText-9="FIGURE 10. Pselaphacus procerus Kuhnt, 1910. Early larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14022033" figureDoi-1="http://doi.org/10.5281/zenodo.14022035" figureDoi-10="http://doi.org/10.5281/zenodo.14022059" figureDoi-11="http://doi.org/10.5281/zenodo.14022063" figureDoi-2="http://doi.org/10.5281/zenodo.14022037" figureDoi-3="http://doi.org/10.5281/zenodo.14022041" figureDoi-4="http://doi.org/10.5281/zenodo.14022043" figureDoi-5="http://doi.org/10.5281/zenodo.14022045" figureDoi-6="http://doi.org/10.5281/zenodo.14022049" figureDoi-7="http://doi.org/10.5281/zenodo.14022051" figureDoi-8="http://doi.org/10.5281/zenodo.14022053" figureDoi-9="http://doi.org/10.5281/zenodo.14022057" httpUri-0="https://zenodo.org/record/14022033/files/figure.png" httpUri-1="https://zenodo.org/record/14022035/files/figure.png" httpUri-10="https://zenodo.org/record/14022059/files/figure.png" httpUri-11="https://zenodo.org/record/14022063/files/figure.png" httpUri-2="https://zenodo.org/record/14022037/files/figure.png" httpUri-3="https://zenodo.org/record/14022041/files/figure.png" httpUri-4="https://zenodo.org/record/14022043/files/figure.png" httpUri-5="https://zenodo.org/record/14022045/files/figure.png" httpUri-6="https://zenodo.org/record/14022049/files/figure.png" httpUri-7="https://zenodo.org/record/14022051/files/figure.png" httpUri-8="https://zenodo.org/record/14022053/files/figure.png" httpUri-9="https://zenodo.org/record/14022057/files/figure.png" pageId="2" pageNumber="75">Figs 1–12</figureCitation>
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; 
<treatmentCitation id="0AB910B9FFD3FD555E6BF930FBE044D6" author="Blackwelder, R. E." box="[864,1109,1690,1714]" page="464" pageId="2" pageNumber="75" year="1945">
<bibRefCitation id="EF894B59FFD3FD555E6BF930FBE044D6" author="Blackwelder, R. E." box="[864,1109,1690,1714]" pageId="2" pageNumber="75" pagination="456 - 469" refId="ref8827" refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 3. Erotylidae. Bulletin of the United States National Museum, 184, 456 - 469. https: // doi. org / 10.5479 / si. 03629236.185.3" type="journal article" year="1945">Blackwelder, 1945: 464</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0AB910B9FFD3FD555955F930FA9644D5" author="Alvarenga, M." box="[1118,1315,1690,1714]" page="7" pageId="2" pageNumber="75" year="1994">
<bibRefCitation id="EF894B59FFD3FD555955F930FA9644D5" author="Alvarenga, M." box="[1118,1315,1690,1714]" pageId="2" pageNumber="75" pagination="1 - 175" refId="ref8785" refString="Alvarenga, M. (1994) Catalogo dos Erotylidae (Coleoptera) Neotropicais. Revista Brasileira de Zoologia, 11 (1), 1 - 175. https: // doi. org / 10.1590 / S 0101 - 81751994000100001" type="journal article" year="1994">Alvarenga, 1994: 7</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3026523FFD3FD545D9CF975FDE04379" lastPageId="3" lastPageNumber="76" pageId="2" pageNumber="75" type="diagnosis">
<paragraph id="8BA736A8FFD3FD545D9CF975FDE04379" blockId="2.[151,1436,1759,2039]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="76" pageId="2" pageNumber="75">
<emphasis id="B96CEABAFFD3FD555D9CF975FEE7449E" bold="true" box="[151,338,1759,1786]" italics="true" pageId="2" pageNumber="75">Adult diagnosis.</emphasis>
Pronotum marginate laterally by a narrow black band; anterior margin with irregular black band, projected at middle forming a triangular narrow band almost reaching mid-length; two rounded black patches on anterior third; posterior margin with irregular black band, projected at middle forming a wide short band with one shorter triangular projection on each side. Scutellar shield black. Elytron with black marks as follow: base marginated by a narrow black band; a band on scutellar region, reaching 5th stria, length about ⅕ of elytron length, with irregular posterior margin; an irregular band on middle, reaching 4th stria, length approximately the same as scutellar band; a longitudinal stripe between 5th and 6th striae, at the same level as middle band; a longitudinal stripe on the elytral suture, reaching the 1st stria between scutellar and middle bands and reaching the 2nd stria after middle band, slightly narrowing toward apex; an oblong stripe near apex, between 3th and 7th striae, slightly oblique, length approximately the same as previous marks; external margin entirely bordered, thicker on the first half. Epipleura black. Ventral coloration almost totally black, abdominal ventrites II-IV laterally brownish-orange, and V almost entirely brownish-orange.
</paragraph>
</subSubSection>
<caption id="DF676620FFD2FD545D93FBD3FAE94745" ID-DOI="http://doi.org/10.5281/zenodo.14022033" ID-Zenodo-Dep="14022033" httpUri="https://zenodo.org/record/14022033/files/figure.png" pageId="3" pageNumber="76" startId="3.[152,255,1145,1169]" targetBox="[181,1410,310,1115]" targetPageId="3" targetType="figure">
<paragraph id="8BA736A8FFD2FD545D93FBD3FAE94745" blockId="3.[151,1437,1145,1316]" pageId="3" pageNumber="76">
<emphasis id="B96CEABAFFD2FD545D93FBD3FEAE46F5" bold="true" box="[152,283,1145,1169]" pageId="3" pageNumber="76">FIGURE 1.</emphasis>
Map showing the known distribution of the 
<taxonomicName id="4C184D2BFFD2FD545FFCFBD3FBD246F6" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[759,1127,1145,1170]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFD2FD545FFCFBD3FC6C46F5" box="[759,985,1145,1169]" italics="true" pageId="3" pageNumber="76">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFD2FD545EE9FBD3FBD246F6" author="Kuhnt, P." box="[994,1127,1145,1170]" pageId="3" pageNumber="76" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. (A–G) Brazil: (A) Benjamin Constant, Amazonas state: 4° 23′ 0″ S, 70° 1′ 53″W. (B) Uberlândia, Minas Gerais state: 18° 54′41″ S, 48° 15′ 44″W. (C) Três Lagoas, Mato Grosso do Sul state: 20° 45′ 35″ S, 51° 41′ 42″W. (D) Castilho, São Paulo state: 20° 52′ 12″ S, 51° 29′ 13″W. (E) Marília, São Paulo state: 22° 13′ 15″ S, 49° 56′ 55″W. (F) Salto Grande, São Paulo state: 22° 52′ 53″ S, 49° 57′ 44″W. (G) Piracicaba, São Paulo state: 22° 43’ 30” S, 47° 38’ 56” W. (H) Misiones, Argentina: 26° 55′ S, 54° 31′ W, [type locality].
</paragraph>
</caption>
<subSubSection id="C3026523FFD2FD5B5DCCFAD9FB664560" lastPageId="12" lastPageNumber="85" pageId="3" pageNumber="76" type="description">
<paragraph id="8BA736A8FFD2FD545DCCFAD9FB264426" blockId="3.[151,1437,1395,2034]" pageId="3" pageNumber="76">
<emphasis id="B96CEABAFFD2FD545DCCFAD9FE1247EA" bold="true" box="[199,423,1395,1422]" italics="true" pageId="3" pageNumber="76">Adult redescription</emphasis>
(
<figureCitation id="13232A2DFFD2FD545CBCFADEFDA947EB" box="[439,540,1396,1423]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="4.[152,255,1974,1998]" captionStartId-1="5.[152,255,1943,1967]" captionStartId-2="6.[152,255,1943,1967]" captionStartId-3="7.[152,255,1075,1099]" captionTargetBox-0="[177,1411,698,1926]" captionTargetBox-1="[154,1435,195,1910]" captionTargetBox-2="[160,1431,185,1919]" captionTargetBox-3="[153,1428,189,1051]" captionTargetId-0="figure-263@4.[164,1423,667,1950]" captionTargetId-1="figure-18@5.[151,1435,181,1919]" captionTargetId-2="figure-27@6.[151,1435,181,1919]" captionTargetId-3="figure-18@7.[151,1435,181,1051]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." captionText-1="FIGURE 3. Pselaphacus procerus Kuhnt, 1910. Male. Mouthparts. (A) front and clypeus; (B) epipharynx; (C–D) mandible, dorsal and ventral views; (E) maxilla and labium; (F–G) lacinia and galea, dorsal and ventral views. Scale bar: A, C, D, E = 0.5 mm, B, F, G = 0.1 mm." captionText-2="FIGURE 4. Pselaphacus procerus Kuhnt, 1910. Male. (A) sternite IX; (B) tergite and sternite IX (C–G) aedeagus: (C–D) tegmen and parameres, ventral and lateral views; (E–F) penis, lateral and ventral views; (G) apex of accessory muscle. Scale bar: A, B, C, F = 0.5 mm, D, E = 0.1 mm." captionText-3="FIGURE 5. Pselaphacus procerus Kuhnt, 1910. Female. (A) sternite VIII; (B) tergite VIII; (C) spermatheca; (D) ovipositor, dorsal view. Scale bar: A, B, D = 0.5 mm, C = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14022035" figureDoi-1="http://doi.org/10.5281/zenodo.14022037" figureDoi-2="http://doi.org/10.5281/zenodo.14022041" figureDoi-3="http://doi.org/10.5281/zenodo.14022043" httpUri-0="https://zenodo.org/record/14022035/files/figure.png" httpUri-1="https://zenodo.org/record/14022037/files/figure.png" httpUri-2="https://zenodo.org/record/14022041/files/figure.png" httpUri-3="https://zenodo.org/record/14022043/files/figure.png" pageId="3" pageNumber="76">Figs 2–5</figureCitation>
) Length: 
<quantity id="4CE09B4DFFD2FD545F86FADEFC8447EA" box="[653,817,1396,1423]" metricMagnitude="-2" metricUnit="m" metricValue="1.1949999999999998" metricValueMax="1.34" metricValueMin="1.05" pageId="3" pageNumber="76" unit="mm" value="11.95" valueMax="13.4" valueMin="10.5">10.5–13.4 mm</quantity>
(n = 11). Body broad, elongated, wide, parallel-sided. Glabrous and shiny, except the anterior edge of prosternum, ventrally with a row of yellow setae, and tibiae and tarsi with yellow setae. General coloration black, with pronotum and elytra brownish-orange with black marks; labrum, reddish-brown and apices of palpi and labium, brownish. Punctuation fine, weak, and dense; frons with punctuation fine and heterogeneous, thicker at base; gena ventrally with punctuation irregular and thicker; pronotum with punctuation heterogeneous, denser laterally on anterior third, and thicker near middle.
</paragraph>
<paragraph id="8BA736A8FFD2FD535DCCF9E6FACF40E7" blockId="3.[151,1437,1395,2034]" lastBlockId="4.[151,1436,151,643]" lastPageId="4" lastPageNumber="77" pageId="3" pageNumber="76">
<emphasis id="B96CEABAFFD2FD545DCCF9E6FEB14402" box="[199,260,1612,1638]" italics="true" pageId="3" pageNumber="76">Head</emphasis>
(
<figureCitation id="13232A2DFFD2FD545C18F9E6FE384402" box="[275,397,1612,1638]" captionStart="FIGURE 2" captionStartId="4.[152,255,1974,1998]" captionTargetBox="[177,1411,698,1926]" captionTargetId="figure-263@4.[164,1423,667,1950]" captionTargetPageId="4" captionText="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="3" pageNumber="76">Fig. 2A–D</figureCitation>
) glabrous; frontoclypeal suture weak with lateral margins convergent posteriorly, interrupted at middle. Frons (
<figureCitation id="13232A2DFFD2FD545C54F9DAFE0D44EE" box="[351,440,1648,1674]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[154,1435,195,1910]" captionTargetId="figure-18@5.[151,1435,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Pselaphacus procerus Kuhnt, 1910. Male. Mouthparts. (A) front and clypeus; (B) epipharynx; (C–D) mandible, dorsal and ventral views; (E) maxilla and labium; (F–G) lacinia and galea, dorsal and ventral views. Scale bar: A, C, D, E = 0.5 mm, B, F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022037" httpUri="https://zenodo.org/record/14022037/files/figure.png" pageId="3" pageNumber="76">Fig. 3A</figureCitation>
) narrow and marginate laterally on basal half; prominent behind antennal insertions and on eyes, forming canthus, and then gradually narrowed apically, anterior margin not carinate; clypeus anteriorly strongly emarginate at middle forming two well-developed rounded lobes; ocular striae and subocular pores present. Subgenal braces well developed with pores present. Antennae (
<figureCitation id="13232A2DFFD2FD545E65F976FC5C4492" box="[878,1001,1756,1782]" captionStart="FIGURE 2" captionStartId="4.[152,255,1974,1998]" captionTargetBox="[177,1411,698,1926]" captionTargetId="figure-263@4.[164,1423,667,1950]" captionTargetPageId="4" captionText="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="3" pageNumber="76">Fig. 2A–C</figureCitation>
) not reaching base of pronotum, with 11 antennomeres: 1 
<superScript id="7C6D9BE0FFD2FD545C79F8AAFEC8456A" attach="left" box="[370,381,1792,1806]" fontSize="6" pageId="3" pageNumber="76">st</superScript>
slightly elongate, wide, almost cylindrical; 2 
<superScript id="7C6D9BE0FFD2FD545E72F955FC3C4569" attach="left" box="[889,905,1791,1805]" fontSize="6" pageId="3" pageNumber="76">nd</superScript>
short, globular; 3 
<superScript id="7C6D9BE0FFD2FD545959F955FBD54569" attach="left" box="[1106,1120,1791,1805]" fontSize="6" pageId="3" pageNumber="76">rd</superScript>
longer, almost as long as 4 
<superScript id="7C6D9BE0FFD2FD54589BF955FA284569" attach="right" box="[1424,1437,1791,1805]" fontSize="6" pageId="3" pageNumber="76">th</superScript>
and 5 
<superScript id="7C6D9BE0FFD2FD545DDDF889FF564555" attach="left" box="[214,227,1827,1841]" fontSize="6" pageId="3" pageNumber="76">th</superScript>
together, narrowed at base; 4 
<superScript id="7C6D9BE0FFD2FD545F3FF889FDF44555" attach="none" box="[564,577,1827,1841]" fontSize="6" pageId="3" pageNumber="76">th</superScript>
–8 
<superScript id="7C6D9BE0FFD2FD545F56F889FDDF4555" attach="left" box="[605,618,1827,1841]" fontSize="6" pageId="3" pageNumber="76">th</superScript>
slightly increasing in width and decreasing in length; three last forming a club. Eyes glabrous, coarsely granulate. Labrum (
<figureCitation id="13232A2DFFD2FD545FF8F8E2FCF84506" box="[755,845,1864,1890]" captionStart="FIGURE 2" captionStartId="4.[152,255,1974,1998]" captionTargetBox="[177,1411,698,1926]" captionTargetId="figure-263@4.[164,1423,667,1950]" captionTargetPageId="4" captionText="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="3" pageNumber="76">Fig. 2C</figureCitation>
) transverse, trapezoidal, wider at middle; swelling transversely near base; distal margin arched; setae of varied sizes, more concentrated near angles and shorter near middle. Epipharynx (
<figureCitation id="13232A2DFFD2FD545C82F83AFE5445CE" box="[393,481,1936,1962]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[154,1435,195,1910]" captionTargetId="figure-18@5.[151,1435,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Pselaphacus procerus Kuhnt, 1910. Male. Mouthparts. (A) front and clypeus; (B) epipharynx; (C–D) mandible, dorsal and ventral views; (E) maxilla and labium; (F–G) lacinia and galea, dorsal and ventral views. Scale bar: A, C, D, E = 0.5 mm, B, F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022037" httpUri="https://zenodo.org/record/14022037/files/figure.png" pageId="3" pageNumber="76">Fig. 3B</figureCitation>
) transverse with one row of wide and short setae near each fore angle, and several campaniform sensilla below; a stout long seta at middle, marginate anteriorly by campaniform sensilla; fine and dense setae inwardly directed surrounded laterally and posteriorly median seta. Mandibles (
<figureCitation id="13232A2DFFD2FD54598DF872FB484596" box="[1158,1277,2008,2034]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[154,1435,195,1910]" captionTargetId="figure-18@5.[151,1435,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Pselaphacus procerus Kuhnt, 1910. Male. Mouthparts. (A) front and clypeus; (B) epipharynx; (C–D) mandible, dorsal and ventral views; (E) maxilla and labium; (F–G) lacinia and galea, dorsal and ventral views. Scale bar: A, C, D, E = 0.5 mm, B, F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022037" httpUri="https://zenodo.org/record/14022037/files/figure.png" pageId="3" pageNumber="76">Fig. 3C–D</figureCitation>
) symmetrical, elongated, strongly curved with two apical and one wide rounded subapical dorsal teeth; with a brush of setae at base; prostheca elongate, fleshy, formed by short setae; long setae between prostheca and mola; mola with lateral short setae, more concentrated dorsally; mesal area microspined internally dorsal and ventrally. Maxilla (
<figureCitation id="13232A2DFFD5FD53583CFF74FA3A429C" box="[1335,1423,222,248]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[154,1435,195,1910]" captionTargetId="figure-18@5.[151,1435,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Pselaphacus procerus Kuhnt, 1910. Male. Mouthparts. (A) front and clypeus; (B) epipharynx; (C–D) mandible, dorsal and ventral views; (E) maxilla and labium; (F–G) lacinia and galea, dorsal and ventral views. Scale bar: A, C, D, E = 0.5 mm, B, F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022037" httpUri="https://zenodo.org/record/14022037/files/figure.png" pageId="4" pageNumber="77">Fig. 3E</figureCitation>
): cardo transverse with an elongate basal condylar projection and with one pore opening at middle; basistipes and mediostipes elongate and triangular; basistipes with one long seta; galea and lacinia (
<figureCitation id="13232A2DFFD5FD53596BFE8CFB624324" box="[1120,1239,294,320]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[154,1435,195,1910]" captionTargetId="figure-18@5.[151,1435,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Pselaphacus procerus Kuhnt, 1910. Male. Mouthparts. (A) front and clypeus; (B) epipharynx; (C–D) mandible, dorsal and ventral views; (E) maxilla and labium; (F–G) lacinia and galea, dorsal and ventral views. Scale bar: A, C, D, E = 0.5 mm, B, F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022037" httpUri="https://zenodo.org/record/14022037/files/figure.png" pageId="4" pageNumber="77">Fig. 3F–G</figureCitation>
) separated; galea narrowed at base, rounded externally; lacinia elongate with narrowed apex; galea and lacinia covered by stout long setae, more concentrate near apex. Maxillary palpi with four palpomeres: basal one elongate and curved; second and third short, wider than long; distal wide, securiform, 1.8x wider than long. Labium (
<figureCitation id="13232A2DFFD5FD53594FFE38FB2C43C8" box="[1092,1177,402,428]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[154,1435,195,1910]" captionTargetId="figure-18@5.[151,1435,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Pselaphacus procerus Kuhnt, 1910. Male. Mouthparts. (A) front and clypeus; (B) epipharynx; (C–D) mandible, dorsal and ventral views; (E) maxilla and labium; (F–G) lacinia and galea, dorsal and ventral views. Scale bar: A, C, D, E = 0.5 mm, B, F, G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022037" httpUri="https://zenodo.org/record/14022037/files/figure.png" pageId="4" pageNumber="77">Fig. 3E</figureCitation>
) short and wide; ligula subtrapezoidal, emarginate anteriorly, marginate by short setae; with four pair of setae and many campaniform sensilla near middle (distal and basal shorter); prementum band-like; mentum almost as wide as long with lateral margins sinuous; with a pentagonal elevated plate at middle of anterior third area; basal ⅔ widely concave; concavity rounded anteriorly, including four long setae and about four pairs of short setae. Labial papi with three palpomeres; palpiger band-like with crescent sclerotized area; basal elongate, gradually widened to apex; median transverse and wide, with lateral margin curved and four setae; apical elongate, slightly securiform (slightly longer than wide).
</paragraph>
<caption id="DF676620FFD5FD535D93F81CFC644595" ID-DOI="http://doi.org/10.5281/zenodo.14022035" ID-Zenodo-Dep="14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="4" pageNumber="77" startId="4.[152,255,1974,1998]" targetBox="[177,1411,698,1926]" targetPageId="4" targetType="figure">
<paragraph id="8BA736A8FFD5FD535D93F81CFC644595" blockId="4.[151,1436,1974,2034]" pageId="4" pageNumber="77">
<emphasis id="B96CEABAFFD5FD535D93F81CFEAC45AA" bold="true" box="[152,281,1974,1998]" pageId="4" pageNumber="77">FIGURE 2.</emphasis>
<taxonomicName id="4C184D2BFFD5FD535C14F81CFD3D45AA" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[287,648,1974,1998]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFD5FD535C14F81CFE4A45AA" box="[287,511,1974,1998]" italics="true" pageId="4" pageNumber="77">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFD5FD535F0DF81CFD3D45AA" author="Kuhnt, P." box="[518,648,1974,1998]" pageId="4" pageNumber="77" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Male. 
<emphasis id="B96CEABAFFD5FD535FD8F81CFC9645AA" box="[723,803,1974,1998]" italics="true" pageId="4" pageNumber="77">Habitus</emphasis>
. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF676620FFD4FD525D93F83DFE364592" ID-DOI="http://doi.org/10.5281/zenodo.14022037" ID-Zenodo-Dep="14022037" httpUri="https://zenodo.org/record/14022037/files/figure.png" pageId="5" pageNumber="78" startId="5.[152,255,1943,1967]" targetBox="[154,1435,195,1910]" targetPageId="5" targetType="figure">
<paragraph id="8BA736A8FFD4FD525D93F83DFE364592" blockId="5.[151,1437,1943,2039]" pageId="5" pageNumber="78">
<emphasis id="B96CEABAFFD4FD525D93F83DFEAC45CB" bold="true" box="[152,281,1943,1967]" pageId="5" pageNumber="78">FIGURE 3.</emphasis>
<taxonomicName id="4C184D2BFFD4FD525C2AF83DFD3945CB" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[289,652,1943,1967]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="78" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFD4FD525C2AF83DFDB445CB" box="[289,513,1943,1967]" italics="true" pageId="5" pageNumber="78">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFD4FD525F02F83DFD3945CB" author="Kuhnt, P." box="[521,652,1943,1967]" pageId="5" pageNumber="78" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Male. Mouthparts. (A) front and clypeus; (B) epipharynx; (C–D) mandible, dorsal and ventral views; (E) maxilla and labium; (F–G) lacinia and galea, dorsal and ventral views. Scale bar: A, C, D, E = 0.5 mm, B, F, G = 0.1 mm.
</paragraph>
</caption>
<caption id="DF676620FFD7FD515D93F83DFD8A4592" ID-DOI="http://doi.org/10.5281/zenodo.14022041" ID-Zenodo-Dep="14022041" httpUri="https://zenodo.org/record/14022041/files/figure.png" pageId="6" pageNumber="79" startId="6.[152,255,1943,1967]" targetBox="[160,1431,185,1919]" targetPageId="6" targetType="figure">
<paragraph id="8BA736A8FFD7FD515D93F83DFD8A4592" blockId="6.[151,1436,1943,2039]" pageId="6" pageNumber="79">
<emphasis id="B96CEABAFFD7FD515D93F83DFEAE45CB" bold="true" box="[152,283,1943,1967]" pageId="6" pageNumber="79">FIGURE 4.</emphasis>
<taxonomicName id="4C184D2BFFD7FD515C2FF83DFD2645CB" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[292,659,1943,1967]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="79" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFD7FD515C2FF83DFDB345CB" box="[292,518,1943,1967]" italics="true" pageId="6" pageNumber="79">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFD7FD515F04F83DFD2645CB" author="Kuhnt, P." box="[527,659,1943,1967]" pageId="6" pageNumber="79" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Male. (A) sternite IX; (B) tergite and sternite IX (C–G) aedeagus: (C–D) tegmen and parameres, ventral and lateral views; (E–F) penis, lateral and ventral views; (G) apex of accessory muscle. Scale bar: A, B, C, F = 0.5 mm, D, E = 0.1 mm.
</paragraph>
</caption>
<caption id="DF676620FFD6FD505D93FB99FD73460B" ID-DOI="http://doi.org/10.5281/zenodo.14022043" ID-Zenodo-Dep="14022043" httpUri="https://zenodo.org/record/14022043/files/figure.png" pageId="7" pageNumber="80" startId="7.[152,255,1075,1099]" targetBox="[153,1428,189,1051]" targetPageId="7" targetType="figure">
<paragraph id="8BA736A8FFD6FD505D93FB99FD73460B" blockId="7.[151,1437,1075,1136]" pageId="7" pageNumber="80">
<emphasis id="B96CEABAFFD6FD505D93FB99FEAC4628" bold="true" box="[152,281,1075,1100]" pageId="7" pageNumber="80">FIGURE 5.</emphasis>
<taxonomicName id="4C184D2BFFD6FD505C2AFB99FD394628" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[289,652,1075,1100]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="80" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFD6FD505C2AFB99FDB4462F" box="[289,513,1075,1099]" italics="true" pageId="7" pageNumber="80">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFD6FD505F02FB99FD394628" author="Kuhnt, P." box="[521,652,1075,1100]" pageId="7" pageNumber="80" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Female. (A) sternite VIII; (B) tergite VIII; (C) spermatheca; (D) ovipositor, dorsal view. Scale bar: A, B, D = 0.5 mm, C = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA736A8FFD6FD505DCCFB6BFD914491" blockId="7.[151,1437,1217,2033]" pageId="7" pageNumber="80">
<emphasis id="B96CEABAFFD6FD505DCCFB6BFEAD46BF" box="[199,280,1217,1243]" italics="true" pageId="7" pageNumber="80">Thorax</emphasis>
. Pronotum (
<figureCitation id="13232A2DFFD6FD505CA8FB6BFD8F46BF" box="[419,570,1217,1243]" captionStart="FIGURE 2" captionStartId="4.[152,255,1974,1998]" captionTargetBox="[177,1411,698,1926]" captionTargetId="figure-263@4.[164,1423,667,1950]" captionTargetPageId="4" captionText="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="7" pageNumber="80">Fig. 2A, C–E</figureCitation>
) wider than long (1.5x wider than long), slightly convex; slightly narrowed at fore angles; lateral margins almost straight, with narrow flap; stronger convex at middle inside lateral weak grooves; fore angles prominent; posterior margin prominent on median third, forming three rounded weak lobes. Marginal pores opening on posterior third and at posterior and anterior angles. Scutellar shield short (
<figureCitation id="13232A2DFFD6FD505992FA87FAA04723" box="[1177,1301,1325,1351]" captionStart="FIGURE 2" captionStartId="4.[152,255,1974,1998]" captionTargetBox="[177,1411,698,1926]" captionTargetId="figure-263@4.[164,1423,667,1950]" captionTargetPageId="4" captionText="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="7" pageNumber="80">Fig. 2A, D</figureCitation>
), triangular, wider than long; fore angles rounded; glabrous with a few punctures. Elytra (
<figureCitation id="13232A2DFFD6FD505EE1FAFBFBEA470F" box="[1002,1119,1361,1387]" captionStart="FIGURE 2" captionStartId="4.[152,255,1974,1998]" captionTargetBox="[177,1411,698,1926]" captionTargetId="figure-263@4.[164,1423,667,1950]" captionTargetPageId="4" captionText="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="7" pageNumber="80">Fig. 2A, E</figureCitation>
) elongate, 1.55x longer than wide, slightly wider than the basal margin of pronotum; lateral margins parallel on anterior half and slightly and roundly narrowed apically; apices conjointly and widely rounded; moderately convex, lateral margins marginated; nine striae marked by row of coarse punctures; interstices flat and almost same width. Prosternum (
<figureCitation id="13232A2DFFD6FD505812FA16FA2147B2" box="[1305,1428,1468,1494]" captionStart="FIGURE 2" captionStartId="4.[152,255,1974,1998]" captionTargetBox="[177,1411,698,1926]" captionTargetId="figure-263@4.[164,1423,667,1950]" captionTargetPageId="4" captionText="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="7" pageNumber="80">Fig. 2B, E</figureCitation>
) wide; prosternal process gradually expanded apically, with lateral carinae and bilobed apex; two pore openings, one on each side near margin of the prosternal process; procoxal lines straight, not surpassing coxae. Mesoventrite small, slightly convex, anterior edge sinuate, mesocoxal lines curved. Metaventrite convex, glabrous, punctures fine and sparse; metacoxal lines conspicuous; discrimen 0.71x as long as metaventrite. Legs (
<figureCitation id="13232A2DFFD6FD5059A1F9E6FA974402" box="[1194,1314,1612,1638]" captionStart="FIGURE 2" captionStartId="4.[152,255,1974,1998]" captionTargetBox="[177,1411,698,1926]" captionTargetId="figure-263@4.[164,1423,667,1950]" captionTargetPageId="4" captionText="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="7" pageNumber="80">Fig. 2B, E</figureCitation>
): procoxae oval; mesocoxae almost globular; metacoxae transverse. Femora wide, smooth, without spines or other outgrowths, slightly constricted subapically, making ventral angles rounded. Tibiae flat, widened apically, internal margin and apex somewhat setose, setae more concentrated on protibiae; two apical spurs. Tarsomeres 1–3 widened and densely setose beneath, tarsal claws simple.
</paragraph>
<paragraph id="8BA736A8FFD6FD505DCCF955FAD34559" blockId="7.[151,1437,1217,2033]" pageId="7" pageNumber="80">
<emphasis id="B96CEABAFFD6FD505DCCF955FE84457D" box="[199,305,1791,1817]" italics="true" pageId="7" pageNumber="80">Abdomen</emphasis>
(
<figureCitation id="13232A2DFFD6FD505C49F955FE0C457D" box="[322,441,1791,1817]" captionStart="FIGURE 2" captionStartId="4.[152,255,1974,1998]" captionTargetBox="[177,1411,698,1926]" captionTargetId="figure-263@4.[164,1423,667,1950]" captionTargetPageId="4" captionText="FIGURE 2. Pselaphacus procerus Kuhnt, 1910. Male. Habitus. (A) dorsal view; (B) ventral view; (C) frontal view; (D) head and pronotum, and anterior region of elytra; (E) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022035" httpUri="https://zenodo.org/record/14022035/files/figure.png" pageId="7" pageNumber="80">Fig. 2B, E</figureCitation>
). Ventrites I–V transverse; I–II longer; III–V almost same length; apical narrower, with distal margin widely curved; ventrites with punctuation fine and irregular, coarser laterally, bearing tiny setae.
</paragraph>
<paragraph id="8BA736A8FFD6FD5F5DCCF8EDFF46429C" blockId="7.[151,1437,1217,2033]" lastBlockId="8.[151,1436,151,248]" lastPageId="8" lastPageNumber="81" pageId="7" pageNumber="80">
Male (
<figureCitation id="13232A2DFFD6FD505C06F8EDFEE54505" box="[269,336,1863,1889]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[160,1431,185,1919]" captionTargetId="figure-27@6.[151,1435,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Pselaphacus procerus Kuhnt, 1910. Male. (A) sternite IX; (B) tergite and sternite IX (C–G) aedeagus: (C–D) tegmen and parameres, ventral and lateral views; (E–F) penis, lateral and ventral views; (G) apex of accessory muscle. Scale bar: A, B, C, F = 0.5 mm, D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022041" httpUri="https://zenodo.org/record/14022041/files/figure.png" pageId="7" pageNumber="80">Fig. 4</figureCitation>
): Tergite IX (
<figureCitation id="13232A2DFFD6FD505CD4F8EDFD864505" box="[479,563,1863,1889]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[160,1431,185,1919]" captionTargetId="figure-27@6.[151,1435,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Pselaphacus procerus Kuhnt, 1910. Male. (A) sternite IX; (B) tergite and sternite IX (C–G) aedeagus: (C–D) tegmen and parameres, ventral and lateral views; (E–F) penis, lateral and ventral views; (G) apex of accessory muscle. Scale bar: A, B, C, F = 0.5 mm, D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022041" httpUri="https://zenodo.org/record/14022041/files/figure.png" pageId="7" pageNumber="80">Fig. 4B</figureCitation>
) narrow, narrower than the width of tergum; gradually slightly narrowed apicad; distal margin widely rounded, marginated by fringe of long setae. Sternite IX (
<figureCitation id="13232A2DFFD6FD505903F8C1FBD545E1" box="[1032,1120,1899,1925]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[160,1431,185,1919]" captionTargetId="figure-27@6.[151,1435,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Pselaphacus procerus Kuhnt, 1910. Male. (A) sternite IX; (B) tergite and sternite IX (C–G) aedeagus: (C–D) tegmen and parameres, ventral and lateral views; (E–F) penis, lateral and ventral views; (G) apex of accessory muscle. Scale bar: A, B, C, F = 0.5 mm, D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022041" httpUri="https://zenodo.org/record/14022041/files/figure.png" pageId="7" pageNumber="80">Fig. 4A</figureCitation>
) elongate, subrectangular at base; apical region roundly narrowed apicad; apex trilobed; lateral lobes widened, extending dorsally and reaching tergite IX; with many long setae, more concentrate laterally; median lobe narrow, shorter than lateral, with distal margin strongly rounded and densely covered the setae. Aedeagus (
<figureCitation id="13232A2DFFD6FD505EA5F87CFB9B4594" box="[942,1070,2006,2032]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[160,1431,185,1919]" captionTargetId="figure-27@6.[151,1435,181,1919]" captionTargetPageId="6" captionText="FIGURE 4. Pselaphacus procerus Kuhnt, 1910. Male. (A) sternite IX; (B) tergite and sternite IX (C–G) aedeagus: (C–D) tegmen and parameres, ventral and lateral views; (E–F) penis, lateral and ventral views; (G) apex of accessory muscle. Scale bar: A, B, C, F = 0.5 mm, D, E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022041" httpUri="https://zenodo.org/record/14022041/files/figure.png" pageId="7" pageNumber="80">Fig. 4C–G</figureCitation>
): penis (median lobe) elongate, curved, with apex narrow; median strut longer than penis, with widened apex; flagellum (virga) longer than penis with head dilated. Parameres short, with many short setae; tegmen slightly expanded basad; tegmen arm short and bilobed.
</paragraph>
<caption id="DF676620FFD9FD5F5D93F8E1FED145E3" ID-DOI="http://doi.org/10.5281/zenodo.14022045" ID-Zenodo-Dep="14022045" httpUri="https://zenodo.org/record/14022045/files/figure.png" pageId="8" pageNumber="81" startId="8.[152,255,1867,1891]" targetBox="[151,1427,287,1838]" targetPageId="8" targetType="figure">
<paragraph id="8BA736A8FFD9FD5F5D93F8E1FED145E3" blockId="8.[151,1437,1867,1928]" pageId="8" pageNumber="81">
<emphasis id="B96CEABAFFD9FD5F5D93F8E1FEAC4500" bold="true" box="[152,281,1867,1892]" pageId="8" pageNumber="81">FIGURE 6.</emphasis>
<taxonomicName id="4C184D2BFFD9FD5F5C2AF8E6FD394500" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[289,652,1868,1892]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="81" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFD9FD5F5C2AF8E6FDB44507" box="[289,513,1868,1892]" italics="true" pageId="8" pageNumber="81">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFD9FD5F5F02F8E6FD394500" author="Kuhnt, P." box="[521,652,1868,1892]" pageId="8" pageNumber="81" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Mature larvae. 
<emphasis id="B96CEABAFFD9FD5F5E39F8E6FC374500" box="[818,898,1868,1892]" italics="true" pageId="8" pageNumber="81">Habitus</emphasis>
. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF676620FFD8FD5E5D93F8DEFD4F459C" ID-DOI="http://doi.org/10.5281/zenodo.14022049" ID-Zenodo-Dep="14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="9" pageNumber="82" startId="9.[152,255,1908,1932]" targetBox="[166,1422,186,1884]" targetPageId="9" targetType="figure">
<paragraph id="8BA736A8FFD8FD5E5D93F8DEFD4F459C" blockId="9.[151,1436,1908,2040]" pageId="9" pageNumber="82">
<emphasis id="B96CEABAFFD8FD5E5D93F8DEFEAC45E8" bold="true" box="[152,281,1908,1932]" pageId="9" pageNumber="82">FIGURE 7.</emphasis>
<taxonomicName id="4C184D2BFFD8FD5E5C2BF8DEFD3C45E8" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[288,649,1908,1932]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFD8FD5E5C2BF8DEFE4A45E8" box="[288,511,1908,1932]" italics="true" pageId="9" pageNumber="82">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFD8FD5E5F0DF8DEFD3C45E8" author="Kuhnt, P." box="[518,649,1908,1932]" pageId="9" pageNumber="82" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm.
</paragraph>
</caption>
<caption id="DF676620FFDBFD5D5D93F83DFAD34592" ID-DOI="http://doi.org/10.5281/zenodo.14022051" ID-Zenodo-Dep="14022051" httpUri="https://zenodo.org/record/14022051/files/figure.png" pageId="10" pageNumber="83" startId="10.[152,255,1943,1967]" targetBox="[153,1435,185,1919]" targetPageId="10" targetType="figure">
<paragraph id="8BA736A8FFDBFD5D5D93F83DFAD34592" blockId="10.[151,1437,1943,2039]" pageId="10" pageNumber="83">
<emphasis id="B96CEABAFFDBFD5D5D93F83DFEAD45CB" bold="true" box="[152,280,1943,1967]" pageId="10" pageNumber="83">FIGURE 8.</emphasis>
<taxonomicName id="4C184D2BFFDBFD5D5C15F83DFD3045CB" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[286,645,1943,1967]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFDBFD5D5C15F83DFE4845CB" box="[286,509,1943,1967]" italics="true" pageId="10" pageNumber="83">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFDBFD5D5F08F83DFD3045CB" author="Kuhnt, P." box="[515,645,1943,1967]" pageId="10" pageNumber="83" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Mature larva: (A) pro and mesonotum; (C) prothoracic leg, internal view; (D) pretarsus. Early larva: (B) pro and mesonotum; (E) prothoracic leg, internal view; (F) middle leg, external view; (G) thoracic spiracle; (H) segment IX, dorsal view; (I) urogomphi, apex. Scale bar: A, B = 1.0 mm, C, G, H = 0.1 mm, E, F = 0.5 mm.
</paragraph>
</caption>
<caption id="DF676620FFDAFD5C5D93F8DEFCE3459C" ID-DOI="http://doi.org/10.5281/zenodo.14022053" ID-Zenodo-Dep="14022053" httpUri="https://zenodo.org/record/14022053/files/figure.png" pageId="11" pageNumber="84" startId="11.[152,255,1908,1932]" targetBox="[164,1423,181,1884]" targetPageId="11" targetType="figure">
<paragraph id="8BA736A8FFDAFD5C5D93F8DEFCE3459C" blockId="11.[151,1437,1908,2040]" pageId="11" pageNumber="84">
<emphasis id="B96CEABAFFDAFD5C5D93F8DEFEAD45E8" bold="true" box="[152,280,1908,1932]" pageId="11" pageNumber="84">FIGURE 9.</emphasis>
<taxonomicName id="4C184D2BFFDAFD5C5C14F8DEFD3D45E8" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[287,648,1908,1932]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFDAFD5C5C14F8DEFE4A45E8" box="[287,511,1908,1932]" italics="true" pageId="11" pageNumber="84">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFDAFD5C5F0DF8DEFD3D45E8" author="Kuhnt, P." box="[518,648,1908,1932]" pageId="11" pageNumber="84" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm.
</paragraph>
</caption>
<caption id="DF676620FFDDFD5B5D93FA66FED1446C" ID-DOI="http://doi.org/10.5281/zenodo.14022057" ID-Zenodo-Dep="14022057" httpUri="https://zenodo.org/record/14022057/files/figure.png" pageId="12" pageNumber="85" startId="12.[152,255,1484,1508]" targetBox="[310,1278,181,1460]" targetPageId="12" targetType="figure">
<paragraph id="8BA736A8FFDDFD5B5D93FA66FED1446C" blockId="12.[151,1437,1484,1544]" pageId="12" pageNumber="85">
<emphasis id="B96CEABAFFDDFD5B5D93FA66FE934780" bold="true" box="[152,294,1484,1508]" pageId="12" pageNumber="85">FIGURE 10.</emphasis>
<taxonomicName id="4C184D2BFFDDFD5B5C25FA66FD2F4781" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[302,666,1484,1509]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="85" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFDDFD5B5C25FA66FDBA4780" box="[302,527,1484,1508]" italics="true" pageId="12" pageNumber="85">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFDDFD5B5F1CFA66FD2F4781" author="Kuhnt, P." box="[535,666,1484,1509]" pageId="12" pageNumber="85" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Early larvae. 
<emphasis id="B96CEABAFFDDFD5B5E24FA66FCCA4780" box="[815,895,1484,1508]" italics="true" pageId="12" pageNumber="85">Habitus</emphasis>
. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BA736A8FFDDFD5B5DCCF9F0FB664560" blockId="12.[151,1436,1626,2013]" pageId="12" pageNumber="85">
Female (
<figureCitation id="13232A2DFFDDFD5B5C23F9F0FEDE4411" box="[296,363,1626,1653]" captionStart="FIGURE 5" captionStartId="7.[152,255,1075,1099]" captionTargetBox="[153,1428,189,1051]" captionTargetId="figure-18@7.[151,1435,181,1051]" captionTargetPageId="7" captionText="FIGURE 5. Pselaphacus procerus Kuhnt, 1910. Female. (A) sternite VIII; (B) tergite VIII; (C) spermatheca; (D) ovipositor, dorsal view. Scale bar: A, B, D = 0.5 mm, C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022043" httpUri="https://zenodo.org/record/14022043/files/figure.png" pageId="12" pageNumber="85">Fig. 5</figureCitation>
): Tergite VIII (
<figureCitation id="13232A2DFFDDFD5B5F18F9F0FDDD4411" box="[531,616,1626,1653]" captionStart="FIGURE 5" captionStartId="7.[152,255,1075,1099]" captionTargetBox="[153,1428,189,1051]" captionTargetId="figure-18@7.[151,1435,181,1051]" captionTargetPageId="7" captionText="FIGURE 5. Pselaphacus procerus Kuhnt, 1910. Female. (A) sternite VIII; (B) tergite VIII; (C) spermatheca; (D) ovipositor, dorsal view. Scale bar: A, B, D = 0.5 mm, C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022043" httpUri="https://zenodo.org/record/14022043/files/figure.png" pageId="12" pageNumber="85">Fig. 5B</figureCitation>
) wider than long, slightly narrowed at apex; distal margin almost straight; marginated lateral and anteriorly by band of setae. Sternite VIII (
<figureCitation id="13232A2DFFDDFD5B5E70F9D4FC6644FD" box="[891,979,1662,1689]" captionStart="FIGURE 5" captionStartId="7.[152,255,1075,1099]" captionTargetBox="[153,1428,189,1051]" captionTargetId="figure-18@7.[151,1435,181,1051]" captionTargetPageId="7" captionText="FIGURE 5. Pselaphacus procerus Kuhnt, 1910. Female. (A) sternite VIII; (B) tergite VIII; (C) spermatheca; (D) ovipositor, dorsal view. Scale bar: A, B, D = 0.5 mm, C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022043" httpUri="https://zenodo.org/record/14022043/files/figure.png" pageId="12" pageNumber="85">Fig. 5A</figureCitation>
) transverse with lateral margins slightly rounded; basal prolongation 4.4x sternite length. Bursa elongate, membranous, with microspines. Coxites (
<figureCitation id="13232A2DFFDDFD5B5836F908FA2144D9" box="[1341,1428,1698,1725]" captionStart="FIGURE 5" captionStartId="7.[152,255,1075,1099]" captionTargetBox="[153,1428,189,1051]" captionTargetId="figure-18@7.[151,1435,181,1051]" captionTargetPageId="7" captionText="FIGURE 5. Pselaphacus procerus Kuhnt, 1910. Female. (A) sternite VIII; (B) tergite VIII; (C) spermatheca; (D) ovipositor, dorsal view. Scale bar: A, B, D = 0.5 mm, C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022043" httpUri="https://zenodo.org/record/14022043/files/figure.png" pageId="12" pageNumber="85">Fig. 5D</figureCitation>
) elongate and setose, surrounded by a ring formed by tergite and sternite IX; each coxite with one segmented stylus with several setae at apex. Spermatheca (
<figureCitation id="13232A2DFFDDFD5B5F68F940FD0C4561" box="[611,697,1770,1797]" captionStart="FIGURE 5" captionStartId="7.[152,255,1075,1099]" captionTargetBox="[153,1428,189,1051]" captionTargetId="figure-18@7.[151,1435,181,1051]" captionTargetPageId="7" captionText="FIGURE 5. Pselaphacus procerus Kuhnt, 1910. Female. (A) sternite VIII; (B) tergite VIII; (C) spermatheca; (D) ovipositor, dorsal view. Scale bar: A, B, D = 0.5 mm, C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022043" httpUri="https://zenodo.org/record/14022043/files/figure.png" pageId="12" pageNumber="85">Fig. 5C</figureCitation>
) large and oval; spermathecal ducts sclerotized.
</paragraph>
</subSubSection>
<subSubSection id="C3026523FFDDFD5A5DCCF8A4FB1242B1" lastPageId="13" lastPageNumber="86" pageId="12" pageNumber="85" type="materials_examined">
<paragraph id="8BA736A8FFDDFD5A5DCCF8A4FB1242B1" blockId="12.[151,1436,1626,2013]" lastBlockId="13.[151,1437,150,2013]" lastPageId="13" lastPageNumber="86" pageId="12" pageNumber="85">
<emphasis id="B96CEABAFFDDFD5B5DCCF8A4FE04454C" bold="true" box="[199,433,1806,1832]" pageId="12" pageNumber="85">Material examined.</emphasis>
<materialsCitation id="3B703CF5FFDDFD5B5CB3F8A5FC304529" collectingDate="1940-02" collectionCode="MZSP" collectorName="J. Guerin" country="Brazil" location="Benjamin Constant" municipality="Benjamin Constant" pageId="12" pageNumber="85" specimenCode="MZSP56285" specimenCount="1" stateProvince="Amazonas">
<collectingCountry id="F30F7638FFDDFD5B5CB3F8A5FD94454D" box="[440,545,1807,1833]" name="Brazil" pageId="12" pageNumber="85">BRAZIL</collectingCountry>
. 
<collectingRegion id="49DCF84AFFDDFD5B5F22F8A5FD11454D" box="[553,676,1807,1833]" country="Brazil" name="Amazonas" pageId="12" pageNumber="85">Amazonas</collectingRegion>
: 
<collectingMunicipality id="6BC3ACD2FFDDFD5B5FA5F8A4FC3D454D" box="[686,904,1806,1833]" pageId="12" pageNumber="85">Benjamin Constant</collectingMunicipality>
, 
<collectorName id="26ED537EFFDDFD5B5E9FF8A5FC6E454D" box="[916,987,1807,1833]" pageId="12" pageNumber="85">II.940</collectorName>
; ex-Coll J. Guérinn16383; 
<taxonomicName id="4C184D2BFFDDFD5B5807F8A4FA21454C" authorityName="Percheron" authorityYear="1835" box="[1292,1428,1806,1832]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="85" phylum="Arthropoda" rank="genus">Pselaphacus</taxonomicName>
/ 
<taxonomicName id="4C184D2BFFDDFD5B5D9CF898FE564528" authority="Lac., J. Guerin." authorityName="Lac., J. Guerin." box="[151,483,1842,1869]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="85" phylum="Arthropoda" rank="species" species="psecilosomus">
psecilosomus Lac., 
<collectorName id="26ED537EFFDDFD5B5C7FF899FE6B4528" box="[372,478,1842,1869]" pageId="12" pageNumber="85">J. Guérin</collectorName>
.
</taxonomicName>
det. 1944 (
<specimenCount id="9D1EFD21FFDDFD5B5F6FF899FD544528" box="[612,737,1842,1869]" count="1" pageId="12" pageNumber="85" type="generic">1 specimen</specimenCount>
<specimenCode id="DBBE9ED3FFDDFD5B5FE3F898FCC84529" box="[744,893,1842,1869]" pageId="12" pageNumber="85">MZSP56285</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="3B703CF5FFDDFD5B5E84F899FE1145F1" collectingDate="1993-12-21" collectionCode="MZSP" collectorName="Flechtmann" location="Horto Barra da Moeda" municipality="Tres Lagoas" pageId="12" pageNumber="85" specimenCode="MZSP56282" specimenCount="1" stateProvince="Mato Grosso do Sul">
<collectingRegion id="49DCF84AFFDDFD5B5E84F899FBC14528" box="[911,1140,1842,1869]" country="Brazil" name="Mato Grosso do Sul" pageId="12" pageNumber="85">Mato Grosso do Sul</collectingRegion>
: 
<collectingMunicipality id="6BC3ACD2FFDDFD5B5974F899FABE4529" box="[1151,1291,1843,1869]" pageId="12" pageNumber="85">Três Lagoas</collectingMunicipality>
, 
<location id="8EC76073FFDDFD5B581DF899FEB94514" LSID="urn:lsid:plazi:treatment:03B187BEFFD3FD465D9CF981FB22446D:8EC76073FFDDFD5B581DF899FEB94514" municipality="Tres Lagoas" name="Horto Barra da Moeda" pageId="12" pageNumber="85" stateProvince="Mato Grosso do Sul">Horto Barra da Moeda</location>
, 
<location id="8EC76073FFDDFD5B5C13F8FDFD8A4514" LSID="urn:lsid:plazi:treatment:03B187BEFFD3FD465D9CF981FB22446D:8EC76073FFDDFD5B5C13F8FDFD8A4514" box="[280,575,1878,1905]" municipality="Tres Lagoas" name="Tres Lagoas Agroflorestal" pageId="12" pageNumber="85" stateProvince="Mato Grosso do Sul">Três Lagoas Agroflorestal</location>
/ 
<date id="FFA61068FFDDFD5B5F5BF8FDFD6E4515" box="[592,731,1879,1905]" pageId="12" pageNumber="85" value="1993-12-21">
<collectingDate id="EFE2E980FFDDFD5B5F5BF8FDFD6E4515" box="[592,731,1879,1905]" pageId="12" pageNumber="85" value="1993-12-21">21.XII.1993</collectingDate>
</date>
, 
<collectorName id="26ED537EFFDDFD5B5FE3F8FCFCD94514" box="[744,876,1878,1904]" pageId="12" pageNumber="85">Flechtmann</collectorName>
col., Ethanol trap in 
<emphasis id="B96CEABAFFDDFD5B5969F8FCFA8D4514" box="[1122,1336,1878,1904]" italics="true" pageId="12" pageNumber="85">Eucaliptus grandis</emphasis>
stand (
<specimenCount id="9D1EFD21FFDDFD5B589BF8FDFEB445F0" count="1" pageId="12" pageNumber="85" type="generic">1 specimen</specimenCount>
<specimenCode id="DBBE9ED3FFDDFD5B5C0CF8D0FE2945F1" box="[263,412,1914,1941]" pageId="12" pageNumber="85">MZSP56282</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="3B703CF5FFDDFD5B5CA5F8D0FCF545DD" collectingDate="2018-11-25" collectionCode="MZSP" collectorName="Thiago Polizei &amp; Alice Sparvoli" location="Rancho Talisma Ana Gabriela. Rio Aguari" municipality="Uberlandia" pageId="12" pageNumber="85" specimenCode="MZSP 49098" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Minas Gerais">
<collectingRegion id="49DCF84AFFDDFD5B5CA5F8D0FDF245F0" box="[430,583,1914,1940]" country="Brazil" name="Minas Gerais" pageId="12" pageNumber="85">Minas Gerais</collectingRegion>
. 
<collectingMunicipality id="6BC3ACD2FFDDFD5B5F5AF8D0FD6545F0" box="[593,720,1914,1940]" pageId="12" pageNumber="85">Uberlândia</collectingMunicipality>
, 
<location id="8EC76073FFDDFD5B5FD0F8D0FB0445F0" LSID="urn:lsid:plazi:treatment:03B187BEFFD3FD465D9CF981FB22446D:8EC76073FFDDFD5B5FD0F8D0FB0445F0" box="[731,1201,1914,1941]" municipality="Uberlandia" name="Rancho Talisma Ana Gabriela. Rio Aguari" pageId="12" pageNumber="85" stateProvince="Minas Gerais">Rancho Talismã Ana Gabriela. Rio Aguari</location>
/ 
<date id="FFA61068FFDDFD5B59CBF8D0FAF145F1" box="[1216,1348,1914,1941]" pageId="12" pageNumber="85" value="2018-11-25">
<collectingDate id="EFE2E980FFDDFD5B59CBF8D0FAF145F1" box="[1216,1348,1914,1941]" pageId="12" pageNumber="85" value="2018-11-25">25.XI.2018</collectingDate>
</date>
. 
<collectorName id="26ED537EFFDDFD5B5846F8D0FF5045DC" pageId="12" pageNumber="85">Thiago Polizei</collectorName>
&amp; 
<collectorName id="26ED537EFFDDFD5B5C01F834FE1845DC" box="[266,429,1950,1976]" pageId="12" pageNumber="85">Alice Sparvoli</collectorName>
leg. (
<specimenCount id="9D1EFD21FFDDFD5B5CFAF835FDA045DD" box="[497,533,1951,1977]" count="1" pageId="12" pageNumber="85" type="male">1♂</specimenCount>
, 
<specimenCount id="9D1EFD21FFDDFD5B5F29F835FDF045DD" box="[546,581,1951,1977]" count="4" pageId="12" pageNumber="85" type="female">4♀</specimenCount>
<specimenCode id="DBBE9ED3FFDDFD5B5F46F835FD5245DD" box="[589,743,1951,1977]" pageId="12" pageNumber="85">MZSP 49098</specimenCode>
-49102)
</materialsCitation>
. 
<materialsCitation id="3B703CF5FFDDFD5B5E40F835FE1045B9" collectingDate="1961-11" collectionCode="MZSP" collectorName="R. Reichardt" location="Piracicaba" municipality="Piracicaba" pageId="12" pageNumber="85" specimenCode="MZSP56283" specimenCount="1" stateProvince="Sao Paulo">
<collectingRegion id="49DCF84AFFDDFD5B5E40F835FC7745DC" box="[843,962,1950,1977]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="85">São Paulo</collectingRegion>
: 
<collectingMunicipality id="6BC3ACD2FFDDFD5B5EC6F834FBF045DC" box="[973,1093,1950,1976]" pageId="12" pageNumber="85">Piracicaba</collectingMunicipality>
, 
<date id="FFA61068FFDDFD5B595AF834FB0545DC" box="[1105,1200,1950,1976]" pageId="12" pageNumber="85" value="1961-11">
<collectingDate id="EFE2E980FFDDFD5B595AF834FB0545DC" box="[1105,1200,1950,1976]" pageId="12" pageNumber="85" value="1961-11">XI.1961</collectingDate>
</date>
, 
<collectorName id="26ED537EFFDDFD5B59B6F835FAFE45DC" box="[1213,1355,1950,1977]" pageId="12" pageNumber="85">R. Reichardt</collectorName>
col. (
<specimenCount id="9D1EFD21FFDDFD5B589BF835FEB445B8" count="1" pageId="12" pageNumber="85" type="generic">1 specimen</specimenCount>
<specimenCode id="DBBE9ED3FFDDFD5B5C02F868FE2845B9" box="[265,413,1986,2013]" pageId="12" pageNumber="85">MZSP56283</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="3B703CF5FFDDFD5B5CBBF868FB0C45B9" box="[432,1209,1986,2013]" collectingDate="2000-05-08" collectionCode="MZSP" collectorName="Santos-Silva" location="Salto Grande" municipality="Salto Grande" pageId="12" pageNumber="85" specimenCode="MZSP56284" specimenCount="1">
<collectingMunicipality id="6BC3ACD2FFDDFD5B5CBBF868FDF345B8" box="[432,582,1986,2012]" pageId="12" pageNumber="85">Salto Grande</collectingMunicipality>
, 
<date id="FFA61068FFDDFD5B5F59F869FD0F45B9" box="[594,698,1987,2013]" pageId="12" pageNumber="85" value="2000-05-08">
<collectingDate id="EFE2E980FFDDFD5B5F59F869FD0F45B9" box="[594,698,1987,2013]" pageId="12" pageNumber="85" value="2000-05-08">8.V.2000</collectingDate>
</date>
, 
<collectorName id="26ED537EFFDDFD5B5FCDF868FCE145B8" box="[710,852,1986,2012]" pageId="12" pageNumber="85">Santos-Silva</collectorName>
col. (
<specimenCount id="9D1EFD21FFDDFD5B5E9CF869FBA045B8" box="[919,1045,1986,2013]" count="1" pageId="12" pageNumber="85" type="generic">1 specimen</specimenCount>
<specimenCode id="DBBE9ED3FFDDFD5B5916F868FB0745B9" box="[1053,1202,1986,2013]" pageId="12" pageNumber="85">MZSP56284</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="3B703CF5FFDDFD5A59CFF868FA2C42D5" collectingDate="1945-02" collectionCode="MZSP" collectorName="J. Guerin" lastPageId="13" lastPageNumber="86" municipality="Marilia" pageId="12" pageNumber="85" specimenCode="MZSP 56286-56287" specimenCount="2">
<collectingMunicipality id="6BC3ACD2FFDDFD5B59CFF868FAAC45B8" box="[1220,1305,1986,2012]" pageId="12" pageNumber="85">Marilia</collectingMunicipality>
, 
<collectingDate id="EFE2E980FFDDFD5B582EF868FAD845B9" box="[1317,1389,1986,2013]" pageId="12" pageNumber="85" value="1945-02">
<date id="FFA61068FFDDFD5B582EF868FAD845B9" box="[1317,1389,1986,2013]" pageId="12" pageNumber="85" value="1945-02">II.945</date>
</collectingDate>
; ex-Coll. 
<collectorName id="26ED537EFFDCFD5A5DD3FF3DFEF442D5" box="[216,321,151,177]" pageId="13" pageNumber="86">J. Guérin</collectorName>
17413; 
<taxonomicName id="4C184D2BFFDCFD5A5CA9FF3DFC3C42D5" authority="Lac., J. Guerin." authorityName="Lac., J. Guerin." box="[418,905,151,177]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="psecilosomus">
<emphasis id="B96CEABAFFDCFD5A5CA9FF3DFD7B42D5" box="[418,718,151,177]" italics="true" pageId="13" pageNumber="86">Pselaphacus psecilosomus</emphasis>
Lac., 
<originalDetermination id="A8068E54FFDCFD5A5E12FF3DFC3042D5" box="[793,901,151,177]" pageId="13" pageNumber="86">J. Guérin</originalDetermination>
.
</taxonomicName>
det. 1945 (
<specimenCount id="9D1EFD21FFDCFD5A5904FF3DFB2942D5" box="[1039,1180,151,177]" count="2" pageId="13" pageNumber="86" type="generic">2 specimens</specimenCount>
<specimenCode id="DBBE9ED3FFDCFD5A59AEFF3DFA2442D5" box="[1189,1425,150,177]" pageId="13" pageNumber="86">MZSP 56286-56287</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="3B703CF5FFDCFD5A5D9CFF11FB1642B1" box="[151,1187,186,213]" collectingDate="1964-10" collectionCode="MZSP" location="Castilho" pageId="13" pageNumber="86" specimenCode="MZSP 56288-56289" specimenCount="2">
<location id="8EC76073FFDCFD5A5D9CFF11FF4242B1" LSID="urn:lsid:plazi:treatment:03B187BEFFD3FD465D9CF981FB22446D:8EC76073FFDCFD5A5D9CFF11FF4242B1" box="[151,247,187,213]" name="Castilho" pageId="13" pageNumber="86">Castilho</location>
, marg. esq. r. Paraná, 
<date id="FFA61068FFDCFD5A5CE6FF11FDF242B1" box="[493,583,187,213]" pageId="13" pageNumber="86" value="1964-10">
<collectingDate id="EFE2E980FFDCFD5A5CE6FF11FDF242B1" box="[493,583,187,213]" pageId="13" pageNumber="86" value="1964-10">X. 1964</collectingDate>
</date>
Exp. Depto Zool. (
<specimenCount id="9D1EFD21FFDCFD5A5E2BFF11FC1E42B1" box="[800,939,187,213]" count="2" pageId="13" pageNumber="86" type="generic">2 specimens</specimenCount>
<specimenCode id="DBBE9ED3FFDCFD5A5EBAFF11FB2E42B1" box="[945,1179,186,213]" pageId="13" pageNumber="86">MZSP 56288-56289</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3026523FFDCFD5A5DCCFF75FB2F41C1" pageId="13" pageNumber="86" type="description">
<paragraph id="8BA736A8FFDCFD5A5DCCFF75FF7F40A9" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="86">
<emphasis id="B96CEABAFFDCFD5A5DCCFF75FE8C429D" bold="true" box="[199,313,223,249]" italics="true" pageId="13" pageNumber="86">Variation.</emphasis>
The coloration is variable among the eleven examined specimens. All specimens are black with pronotum and elytra brownish-orange with black marks. On the pronotum the median basal black area is narrowed apically, reaching or not anterior margin. Sometimes the longitudinal prolongation is interrupted forming a black area starting at anterior margin. On the elytra, the sizes of black marks are variable. Variation in size was observed in specimens from different localities. Uberlândia specimens (n=5) total body length 10.4–11.0 mm; pronotum length 
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, greatest width 4.0– 
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and elytra length 7.75– 
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. Marília specimens (n=2) total body length 12.0– 
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; pronotum length 
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, greatest width 
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and elytra length 
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Castilho specimens (n=2) total body length 
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; pronotum length 
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, greatest width 
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and elytra length 
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. In Benjamin Constant specimen (n=1) total body length 
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; pronotum length 
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, greatest width 
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and elytra length 
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; Três Lagoas specimen (n=1) total body length 
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; pronotum length 
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, greatest width 
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and elytra length 
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; Piracicaba specimen (n=1) total body length 
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; pronotum length 
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, greatest width 
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and elytra length 
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; and Salto Grande specimen (n=1) total body length 
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; pronotum length 
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, greatest width 
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and elytra length 
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.
</paragraph>
<paragraph id="8BA736A8FFDCFD5A5DCCFD7DFCBA4139" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="86">
<emphasis id="B96CEABAFFDCFD5A5DCCFD7DFEAA4094" box="[199,287,727,753]" italics="true" pageId="13" pageNumber="86">
<emphasis id="B96CEABAFFDCFD5A5DCCFD7DFEAD4095" bold="true" box="[199,280,727,753]" italics="true" pageId="13" pageNumber="86">Larvae</emphasis>
.
</emphasis>
According to 
<bibRefCitation id="EF894B59FFDCFD5A5CCFFD7DFD0B4095" author="Wegrzynowicz, P." box="[452,702,727,753]" pageId="13" pageNumber="86" pagination="435 - 504" refId="ref10577" refString="Wegrzynowicz, P. (2002) Morphology, phylogeny and classification of the family Erotylidae based on adult characters (Coleoptera: Cucujoidea). Genus, 13, 435 - 504." type="journal article" year="2002">Wegrzynowicz (2002)</bibRefCitation>
the 
<taxonomicName id="4C184D2BFFDCFD5A5FFAFD7DFCD34095" box="[753,870,727,753]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="86" phylum="Arthropoda" rank="family">Erotylidae</taxonomicName>
larva undergoes three molts before pupation. The specimens we studied were probably in the last and penultimate instars. The description was based on the last instar larva (mature), and when there were differences in the structures of mature larva with those in the specimens from the early instar, the structure is described in both instars.
</paragraph>
<paragraph id="8BA736A8FFDCFD5A5DCCFCCDFB2F41C1" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="86">
Besides the different sizes, the larval instars are easily recognized by the dorsal sclerotized plates on abdomen (
<figureCitation id="13232A2DFFDCFD5A5D94FC21FEAC41C1" box="[159,281,907,933]" captionStart="FIGURE 6" captionStartId="8.[152,255,1867,1891]" captionTargetBox="[151,1427,287,1838]" captionTargetId="figure-62@8.[151,1435,272,1844]" captionTargetPageId="8" captionText="FIGURE 6. Pselaphacus procerus Kuhnt, 1910. Mature larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022045" httpUri="https://zenodo.org/record/14022045/files/figure.png" pageId="13" pageNumber="86">Fig. 6A–C</figureCitation>
) present only in last larval instar and missing (
<figureCitation id="13232A2DFFDCFD5A5E2FFC21FC1E41C1" box="[804,939,907,933]" captionStart="FIGURE 10" captionStartId="12.[152,255,1484,1508]" captionTargetBox="[310,1278,181,1460]" captionTargetId="figure-27@12.[310,1278,181,1460]" captionTargetPageId="12" captionText="FIGURE 10. Pselaphacus procerus Kuhnt, 1910. Early larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022057" httpUri="https://zenodo.org/record/14022057/files/figure.png" pageId="13" pageNumber="86">Fig. 10A–C</figureCitation>
) in the earlier larvae.
</paragraph>
</subSubSection>
<subSubSection id="C3026523FFDCFD5A5DCCFC04FAB84781" pageId="13" pageNumber="86" type="diagnosis">
<paragraph id="8BA736A8FFDCFD5A5DCCFC04FBA547A5" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="86">
<emphasis id="B96CEABAFFDCFD5A5DCCFC04FE2041AD" bold="true" box="[199,405,942,969]" italics="true" pageId="13" pageNumber="86">Larval diagnosis.</emphasis>
Body elongate, more or less parallel-sided, slightly narrowed anteriorly and at apex, subcylindrical, flattened ventrally. General coloration cream with dorsal transverse brown elliptical plates, yellow at middle and with some yellowish shadow outside; head yellowish, darker on early instar; area of stemmata darker, black in older instar; legs partially yellowish or yellowish-brown. Dorsally, pubescence short; setae inserted in darker punctures or tiny tubercles; ventrally apparently glabrous, with sparse tiny setae more concentrated near middle. Dorsally all segments with transverse, elliptical brown plates, yellow at middle; setae short inserted in darker punctures, densely distributed on darker contour and more concentrate laterally; plate of pronotum longer; plates on meso-and metanotum almost same size; plates on tergites I–VI almost same length; plates I–III slightly increasing in width; plates IV–VI slightly decreasing in width; plates VII–VIII slightly longer than previous, VII wider than VIII; plate IX larger, covering almost whole tergite and including urogomphi; urogomphi setose, brown, yellow dorsally. Early instar larvae with transverse sclerotized brown plates only on thorax and a small on first abdominal tergite; plate of pronotum longer; plates on meso- and metanotum almost same size; each plate of thorax, marginated by irregular band of short setae, inserted in darker punctures, and some sparse setae distributed by whole surface; setae on abdominal segments, short, inserted in darker punctures and concentrate transversal median on segments, same place of future plates; urogomphi setose, brown, clear dorsally.
</paragraph>
<paragraph id="8BA736A8FFDCFD5A5DCCFA60FAB84781" blockId="13.[151,1437,150,2013]" box="[199,1293,1482,1509]" pageId="13" pageNumber="86">
<emphasis id="B96CEABAFFDCFD5A5DCCFA60FE144781" bold="true" box="[199,417,1482,1509]" italics="true" pageId="13" pageNumber="86">Larval description.</emphasis>
(
<figureCitation id="13232A2DFFDCFD5A5CBBFA61FD964781" box="[432,547,1483,1509]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStart-5="FIGURE 11" captionStartId-0="8.[152,255,1867,1891]" captionStartId-1="9.[152,255,1908,1932]" captionStartId-2="10.[152,255,1943,1967]" captionStartId-3="11.[152,255,1908,1932]" captionStartId-4="12.[152,255,1484,1508]" captionStartId-5="15.[152,255,1908,1932]" captionTargetBox-0="[151,1427,287,1838]" captionTargetBox-1="[166,1422,186,1884]" captionTargetBox-2="[153,1435,185,1919]" captionTargetBox-3="[164,1423,181,1884]" captionTargetBox-4="[310,1278,181,1460]" captionTargetBox-5="[166,1419,184,1878]" captionTargetId-0="figure-62@8.[151,1435,272,1844]" captionTargetId-1="figure-18@9.[164,1423,181,1884]" captionTargetId-2="figure-27@10.[151,1435,181,1919]" captionTargetId-3="figure-18@11.[164,1423,181,1884]" captionTargetId-4="figure-27@12.[310,1278,181,1460]" captionTargetId-5="figure-18@15.[164,1423,181,1884]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="15" captionText-0="FIGURE 6. Pselaphacus procerus Kuhnt, 1910. Mature larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." captionText-1="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." captionText-2="FIGURE 8. Pselaphacus procerus Kuhnt, 1910. Mature larva: (A) pro and mesonotum; (C) prothoracic leg, internal view; (D) pretarsus. Early larva: (B) pro and mesonotum; (E) prothoracic leg, internal view; (F) middle leg, external view; (G) thoracic spiracle; (H) segment IX, dorsal view; (I) urogomphi, apex. Scale bar: A, B = 1.0 mm, C, G, H = 0.1 mm, E, F = 0.5 mm." captionText-3="FIGURE 9. Pselaphacus procerus Kuhnt, 1910. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm." captionText-4="FIGURE 10. Pselaphacus procerus Kuhnt, 1910. Early larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." captionText-5="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14022045" figureDoi-1="http://doi.org/10.5281/zenodo.14022049" figureDoi-2="http://doi.org/10.5281/zenodo.14022051" figureDoi-3="http://doi.org/10.5281/zenodo.14022053" figureDoi-4="http://doi.org/10.5281/zenodo.14022057" figureDoi-5="http://doi.org/10.5281/zenodo.14022059" httpUri-0="https://zenodo.org/record/14022045/files/figure.png" httpUri-1="https://zenodo.org/record/14022049/files/figure.png" httpUri-2="https://zenodo.org/record/14022051/files/figure.png" httpUri-3="https://zenodo.org/record/14022053/files/figure.png" httpUri-4="https://zenodo.org/record/14022057/files/figure.png" httpUri-5="https://zenodo.org/record/14022059/files/figure.png" pageId="13" pageNumber="86">Figs 6–11</figureCitation>
). Mature larvae: length: 
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. Early larvae: length: 
<quantity id="4CE09B4DFFDCFD5A59A2FA60FABC4781" box="[1193,1289,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="13" pageNumber="86" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3026523FFDCFD465DCCFA45FAF94391" lastPageId="17" lastPageNumber="90" pageId="13" pageNumber="86" type="description">
<paragraph id="8BA736A8FFDCFD595DCCFA45FE01446D" blockId="13.[151,1437,150,2013]" lastBlockId="14.[151,1437,151,2013]" lastPageId="14" lastPageNumber="87" pageId="13" pageNumber="86">
<emphasis id="B96CEABAFFDCFD5A5DCCFA45FEB1446D" box="[199,260,1519,1545]" italics="true" pageId="13" pageNumber="86">Head</emphasis>
(
<figureCitation id="13232A2DFFDCFD5A5C1DFA45FE22446D" box="[278,407,1519,1545]" captionStart="FIGURE 1" captionStartId="3.[152,255,1145,1169]" captionTargetBox="[181,1410,310,1115]" captionTargetId="figure-70@3.[176,1410,306,1119]" captionTargetPageId="3" captionText="FIGURE 1. Map showing the known distribution of the Pselaphacus procerus Kuhnt, 1910. (A–G) Brazil: (A) Benjamin Constant, Amazonas state: 4° 23′ 0″ S, 70° 1′ 53″W. (B) Uberlândia, Minas Gerais state: 18° 54′41″ S, 48° 15′ 44″W. (C) Três Lagoas, Mato Grosso do Sul state: 20° 45′ 35″ S, 51° 41′ 42″W. (D) Castilho, São Paulo state: 20° 52′ 12″ S, 51° 29′ 13″W. (E) Marília, São Paulo state: 22° 13′ 15″ S, 49° 56′ 55″W. (F) Salto Grande, São Paulo state: 22° 52′ 53″ S, 49° 57′ 44″W. (G) Piracicaba, São Paulo state: 22° 43’ 30” S, 47° 38’ 56” W. (H) Misiones, Argentina: 26° 55′ S, 54° 31′ W, [type locality]." figureDoi="http://doi.org/10.5281/zenodo.14022033" httpUri="https://zenodo.org/record/14022033/files/figure.png" pageId="13" pageNumber="86">Figs 1A–B</figureCitation>
, 
<figureCitation id="13232A2DFFDCFD5A5CACFA45FE59446D" box="[423,492,1519,1545]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="13" pageNumber="86">7A–B</figureCitation>
, 
<figureCitation id="13232A2DFFDCFD5A5CF7FA45FDD2446D" box="[508,615,1519,1545]" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-18@11.[164,1423,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Pselaphacus procerus Kuhnt, 1910. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022053" httpUri="https://zenodo.org/record/14022053/files/figure.png" pageId="13" pageNumber="86">9A, C–D</figureCitation>
, 
<figureCitation id="13232A2DFFDCFD5A5F7DFA45FD7C446D" box="[630,713,1519,1545]" captionStart="FIGURE 10" captionStartId="12.[152,255,1484,1508]" captionTargetBox="[310,1278,181,1460]" captionTargetId="figure-27@12.[310,1278,181,1460]" captionTargetPageId="12" captionText="FIGURE 10. Pselaphacus procerus Kuhnt, 1910. Early larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022057" httpUri="https://zenodo.org/record/14022057/files/figure.png" pageId="13" pageNumber="86">10A–B</figureCitation>
, 
<figureCitation id="13232A2DFFDCFD5A5FD3FA45FC9C446D" box="[728,809,1519,1545]" captionStart="FIGURE 11" captionStartId="15.[152,255,1908,1932]" captionTargetBox="[166,1419,184,1878]" captionTargetId="figure-18@15.[164,1423,181,1884]" captionTargetPageId="15" captionText="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="13" pageNumber="86">11A–B</figureCitation>
) protracted and moderately declined, almost globular. Epicranial sutures (
<figureCitation id="13232A2DFFDCFD5A5C79F9B9FE7E4449" box="[370,459,1555,1581]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="13" pageNumber="86">Fig. 7A</figureCitation>
) U-Shaped; coronal suture absent. Frons (
<figureCitation id="13232A2DFFDCFD5A5EA0F9B9FBB14449" box="[939,1028,1555,1581]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="13" pageNumber="86">Fig. 7A</figureCitation>
) with two rounded slightly concave areas, and with four pair of setae. Each epicranial half with seven setae inserted in small tubercles; five stemmata below each antenna and three setae near them. Ventrally (
<figureCitation id="13232A2DFFDCFD5A5E2BF9F1FCCD4411" box="[800,888,1627,1653]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="13" pageNumber="86">Fig. 7B</figureCitation>
), each side with four setae; two below antennae, each inserted in a small tubercle. In early instar, epicranial suture (
<figureCitation id="13232A2DFFDCFD5A5E79F9D5FC6344FD" box="[882,982,1663,1689]" captionStart="FIGURE 11" captionStartId="15.[152,255,1908,1932]" captionTargetBox="[166,1419,184,1878]" captionTargetId="figure-18@15.[164,1423,181,1884]" captionTargetPageId="15" captionText="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="13" pageNumber="86">Fig. 11A</figureCitation>
) indistinct; dorsal area with two rounded slightly concave areas near middle, 14 pair of setae of varied sizes, not inserted in tiny tubercle; five stemmata below each antenna and three setae near them and one below; a group of about five short setae near each posterior angle; ventrally (
<figureCitation id="13232A2DFFDCFD5A5C54F941FE0E4561" box="[351,443,1771,1797]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="13" pageNumber="86">Fig. 7B</figureCitation>
), each side with seven setae. Antennae (
<figureCitation id="13232A2DFFDCFD5A5E95F941FBAF4561" box="[926,1050,1771,1797]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="13" pageNumber="86">Fig. 7A–E</figureCitation>
) short, with three antennomeres; antennifer wider, membranous and microspined; antennomere one transverse with two sensilla, and microspined at apex; antennomere two elongate, narrower than previous, at apex with antennomere three and four setae, and two ventral membranous sensoria, one very smaller; antennomere three elongate, reduced with two short setae at apex, and one dorsal campaniform sensillum near apex; apex with some small sensoria. In early instar (
<figureCitation id="13232A2DFFDCFD5A580EF8D1FA2545F1" box="[1285,1424,1915,1941]" captionStart="FIGURE 11" captionStartId="15.[152,255,1908,1932]" captionTargetBox="[166,1419,184,1878]" captionTargetId="figure-18@15.[164,1423,181,1884]" captionTargetPageId="15" captionText="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="13" pageNumber="86">Fig. 11C–D</figureCitation>
), basal antennomere transverse, dorsally with two campaniform sensilla and microspines near apex and ventrally with microspines laterally near apex; antennomere two elongate, dorsally with one campaniform sensillum and one short seta near apex, and ventrally with three short setae near apex, and at apex, with antennomere three and membranous apical sensorium; distal antennomere ventrally with one campaniform sensillum near apex, and three wide and long and three wide and short setae at apex. Frontoclypeal suture (
<figureCitation id="13232A2DFFDFFD595EF6FF75FBE3429D" box="[1021,1110,223,249]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="14" pageNumber="87">Fig. 7A</figureCitation>
) present. Clypeus transverse with anterior margin emarginate and posterior prominent at middle; each side with two setae and two campaniform sensilla. In early instar (
<figureCitation id="13232A2DFFDFFD595CAFFE8DFDBD4325" box="[420,520,295,321]" captionStart="FIGURE 11" captionStartId="15.[152,255,1908,1932]" captionTargetBox="[166,1419,184,1878]" captionTargetId="figure-18@15.[164,1423,181,1884]" captionTargetPageId="15" captionText="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="14" pageNumber="87">Fig. 11E</figureCitation>
), with one seta on each fore angle and each side with two setae near base and one campaniform sensillum. Labrum band-like with anterior margin emarginate; two short setae. In early instar also with four campaniform sensilla. Epipharynx (
<figureCitation id="13232A2DFFDFFD595F68FEC5FD0943ED" box="[611,700,367,393]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="14" pageNumber="87">Fig. 7G</figureCitation>
) marginate anteriorly by row of short and stout setae midwardly directed, interrupted at middle; median anterior area with group of setae backwardly directed, with a group of about four sensilla each side; subapical area verrucose on each side and setae backwardly directed at middle. In early instar (
<figureCitation id="13232A2DFFDFFD595D94FE71FEB74391" box="[159,258,475,501]" captionStart="FIGURE 11" captionStartId="15.[152,255,1908,1932]" captionTargetBox="[166,1419,184,1878]" captionTargetId="figure-18@15.[164,1423,181,1884]" captionTargetPageId="15" captionText="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="14" pageNumber="87">Fig. 11F</figureCitation>
), the anterior margin of epipharynx emarginate; each side marginate anteriorly by row of long and thin setae medially directed, interrupted at the middle; the subapical area laterally verrucose; groups of short setae near middle. Mandibles (
<figureCitation id="13232A2DFFDFFD595C72FD89FE5D4059" box="[377,488,547,573]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="14" pageNumber="87">Fig. 7H–I</figureCitation>
) symmetrical, slightly longer than wide, wider at basal half; apex blackish in mature larvae; apex concave and tridentate; without accessory ventral process; mola absent; molar area with membranous lobe armed with asperites; one lateroexternal seta. In early instar, mandibles (
<figureCitation id="13232A2DFFDFFD595916FDC1FB2940E1" box="[1053,1180,619,645]" captionStart="FIGURE 11" captionStartId="15.[152,255,1908,1932]" captionTargetBox="[166,1419,184,1878]" captionTargetId="figure-18@15.[164,1423,181,1884]" captionTargetPageId="15" captionText="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="14" pageNumber="87">Fig. 11G–I</figureCitation>
) were almost as wide as long. Ventral mouthparts (
<figureCitation id="13232A2DFFDFFD595CE9FD25FD8440CD" box="[482,561,655,681]" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-18@11.[164,1423,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Pselaphacus procerus Kuhnt, 1910. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022053" httpUri="https://zenodo.org/record/14022053/files/figure.png" pageId="14" pageNumber="87">Fig. 9I</figureCitation>
) retracted. Maxilla (
<figureCitation id="13232A2DFFDFFD595E17FD25FC0040CD" box="[796,949,655,681]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="14" pageNumber="87">Fig. 7B, J–K</figureCitation>
). Cardo subtriangular with one seta; stipes elongate, ventrally with six setae (one basal shorter); dorsally with one or two campaniform sensilla, microspined at base galea and lacinia; galea and lacinia fused; mala (
<figureCitation id="13232A2DFFDFFD595E06FD7DFC174095" box="[781,930,727,753]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="14" pageNumber="87">Fig. 7J, L–M</figureCitation>
) bilobed; internal lobe smaller, with rounded apex; ventrally with one seta (with irregular apex in early instar) and one campaniform sensillum; dorsally with teeth, smaller, sharpened and in larger number in early instar, and rounded in mature; external lobe wider with apex almost straight; apex ventrally with small teeth of varied sizes, and dorsally with 5 stout setae; some setae in earlier instar with bifurcate apex. Palpi trimerous: basal palpomere transverse, ventrally with two campaniform sensilla and dorsally with one lateroexternal seta; second transverse, ventrally with one campaniform sensillum and one lateroventral seta; distal elongate, ventrally with one laterointernal seta, and dorsally with one wide lateroexternal seta inserted in groove, and one simple seta, and some small peg-like sensilla at apex. In early instar, maxilla (
<figureCitation id="13232A2DFFDFFD595864FC79FEB74675" captionStart="FIGURE 11" captionStartId="15.[152,255,1908,1932]" captionTargetBox="[166,1419,184,1878]" captionTargetId="figure-18@15.[164,1423,181,1884]" captionTargetPageId="15" captionText="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="14" pageNumber="87">Fig. 11B, J–K</figureCitation>
) with cardo transverse, subtriangular with one seta; stipes with six setae and three campaniform sensilla; apex of mala (
<figureCitation id="13232A2DFFDFFD595C3FFBB1FE0E4651" box="[308,443,1051,1077]" captionStart="FIGURE 11" captionStartId="15.[152,255,1908,1932]" captionTargetBox="[166,1419,184,1878]" captionTargetId="figure-18@15.[164,1423,181,1884]" captionTargetPageId="15" captionText="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="14" pageNumber="87">Fig. 11L–M</figureCitation>
) almost truncate with weak rounded lobe internally; ventrally with two teeth and six wide setae (three with irregular apex); dorsally with many small and irregular teeth. Palpomere basal transverse, ventrally with two campaniform sensilla; second transverse, ventrally with one campaniform sensillum and one laterointernal short seta; distal, ventrally with laterointernal short seta, dorsally with one seta in groove, and many apical peg-like sensilla at apex. Labium (
<figureCitation id="13232A2DFFDFFD595CBEFB01FD8746A1" box="[437,562,1195,1221]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="14" pageNumber="87">Figs 7B, O</figureCitation>
; 
<figureCitation id="13232A2DFFDFFD595F35FB01FD2446A1" box="[574,657,1195,1221]" captionStart="FIGURE 11" captionStartId="15.[152,255,1908,1932]" captionTargetBox="[166,1419,184,1878]" captionTargetId="figure-18@15.[164,1423,181,1884]" captionTargetPageId="15" captionText="FIGURE 11. Pselaphacus procerus Kuhnt, 1910. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="14" pageNumber="87">11B, N</figureCitation>
): prementum transverse, ventrally with one long seta below each palp and two short setae and one campaniform sensillum on each side; postmentum elongate, rectangular with two pair of setae and one pair of campaniform sensilla; ligula indistinct. Labial palpi with two palpomeres; basal transverse with one lateroexternal seta; distal elongate with one dorsal campaniform sensillum and peg-like sensilla at apex. Hypopharynx (
<figureCitation id="13232A2DFFDFFD595C4BFA91FE224731" box="[320,407,1339,1365]" captionStart="FIGURE 7" captionStartId="9.[152,255,1908,1932]" captionTargetBox="[166,1422,186,1884]" captionTargetId="figure-18@9.[164,1423,181,1884]" captionTargetPageId="9" captionText="FIGURE 7. Pselaphacus procerus Kuhnt, 1910. Mature larvae. (A–B) head, dorsal and ventral views; (C–E) antenna, dorsal, ventral and frontal views; (F) clypeus and labrum; (G) epipharynx; (H–I) mandible, ventral and internal views; (J–K) maxilla, apex dorsal and ventral views; (L–M) apex of mala, ventral view, with and without setae; (N) hypopharynx; (O) labium. Scale bar: A–B = 0.5 mm, C–K, N–O = 0.1 mm, L–M = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.14022049" httpUri="https://zenodo.org/record/14022049/files/figure.png" pageId="14" pageNumber="87">Fig. 7N</figureCitation>
) raised in a median rounded area with setae on each side, inwardly directed; median anterior region with four short setae and one longer on each side. Hypopharyngeal scleroma slightly elongated with base curved. Hypostomal rods absent. Ventral epicranial ridges and gula absent. Each segment of thorax, dorsally with sclerotized, transverse, oval-shaped plate, margined by brown and yellow at middle; brown band with dense setae, inserted in small tubercles. In early instar the plates are weakly sclerotized or lacking, and setae are longer and without tubercles at base.
</paragraph>
<paragraph id="8BA736A8FFDFFD595DCCF9B9FBEF45B9" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="87">
<emphasis id="B96CEABAFFDFFD595DCCF9B9FEAD4449" box="[199,280,1555,1581]" italics="true" pageId="14" pageNumber="87">Thorax</emphasis>
(
<figureCitation id="13232A2DFFDFFD595C21F9B9FE124449" box="[298,423,1555,1581]" captionStart="FIGURE 8" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[153,1435,185,1919]" captionTargetId="figure-27@10.[151,1435,181,1919]" captionTargetPageId="10" captionText="FIGURE 8. Pselaphacus procerus Kuhnt, 1910. Mature larva: (A) pro and mesonotum; (C) prothoracic leg, internal view; (D) pretarsus. Early larva: (B) pro and mesonotum; (E) prothoracic leg, internal view; (F) middle leg, external view; (G) thoracic spiracle; (H) segment IX, dorsal view; (I) urogomphi, apex. Scale bar: A, B = 1.0 mm, C, G, H = 0.1 mm, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022051" httpUri="https://zenodo.org/record/14022051/files/figure.png" pageId="14" pageNumber="87">Fig. 8A–B</figureCitation>
, 
<figureCitation id="13232A2DFFDFFD595CBEF9B9FE4E4449" box="[437,507,1555,1581]" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-18@11.[164,1423,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Pselaphacus procerus Kuhnt, 1910. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022053" httpUri="https://zenodo.org/record/14022053/files/figure.png" pageId="14" pageNumber="87">9B, G</figureCitation>
) Prothorax (
<figureCitation id="13232A2DFFDFFD595F85F9B9FCBC4449" box="[654,777,1555,1581]" captionStart="FIGURE 6" captionStartId="8.[152,255,1867,1891]" captionTargetBox="[151,1427,287,1838]" captionTargetId="figure-62@8.[151,1435,272,1844]" captionTargetPageId="8" captionText="FIGURE 6. Pselaphacus procerus Kuhnt, 1910. Mature larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022045" httpUri="https://zenodo.org/record/14022045/files/figure.png" pageId="14" pageNumber="87">Fig. 6A–C</figureCitation>
) longer and narrower than mesothorax, shorter ventrally; dorsally convex with large sclerotized plate; plate ventrally glabrous; prosternum with one anterior sclerite on each side; episternum with two small flattened lobes near each coxa, each with one clear sclerite and one short setae; anterior larger. Ventrally with two small plates on each side, clearer or only one each side on early instar. Meso and metathorax of almost same size, each with one short elliptical sclerotized plate with setae; mesothorax anterolaterally with one anullar-biforous spiracle (
<figureCitation id="13232A2DFFDFFD595FCCF96DFCAB4485" box="[711,798,1735,1761]" captionStart="FIGURE 8" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[153,1435,185,1919]" captionTargetId="figure-27@10.[151,1435,181,1919]" captionTargetPageId="10" captionText="FIGURE 8. Pselaphacus procerus Kuhnt, 1910. Mature larva: (A) pro and mesonotum; (C) prothoracic leg, internal view; (D) pretarsus. Early larva: (B) pro and mesonotum; (E) prothoracic leg, internal view; (F) middle leg, external view; (G) thoracic spiracle; (H) segment IX, dorsal view; (I) urogomphi, apex. Scale bar: A, B = 1.0 mm, C, G, H = 0.1 mm, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022051" httpUri="https://zenodo.org/record/14022051/files/figure.png" pageId="14" pageNumber="87">Fig. 8G</figureCitation>
) on each side, marginated by a crescent sclerotized patch and a lateroposterior brown patch on each side; meso and metacoxae marginate externally by two tubercles on each side, anterior yellowish and posterior brown. In early instar (
<figureCitation id="13232A2DFFDFFD595E9EF8A5FB95454D" box="[917,1056,1807,1833]" captionStart="FIGURE 10" captionStartId="12.[152,255,1484,1508]" captionTargetBox="[310,1278,181,1460]" captionTargetId="figure-27@12.[310,1278,181,1460]" captionTargetPageId="12" captionText="FIGURE 10. Pselaphacus procerus Kuhnt, 1910. Early larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022057" httpUri="https://zenodo.org/record/14022057/files/figure.png" pageId="14" pageNumber="87">Fig. 10A–C</figureCitation>
), metathorax laterally sometimes bearing two brown patches. Legs (
<figureCitation id="13232A2DFFDFFD595F11F899FD224529" box="[538,663,1843,1869]" captionStart="FIGURE 6" captionStartId="8.[152,255,1867,1891]" captionTargetBox="[151,1427,287,1838]" captionTargetId="figure-62@8.[151,1435,272,1844]" captionTargetPageId="8" captionText="FIGURE 6. Pselaphacus procerus Kuhnt, 1910. Mature larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022045" httpUri="https://zenodo.org/record/14022045/files/figure.png" pageId="14" pageNumber="87">Figs 6B–C</figureCitation>
, 
<figureCitation id="13232A2DFFDFFD595FA8F899FD524529" box="[675,743,1843,1869]" captionStart="FIGURE 8" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[153,1435,185,1919]" captionTargetId="figure-27@10.[151,1435,181,1919]" captionTargetPageId="10" captionText="FIGURE 8. Pselaphacus procerus Kuhnt, 1910. Mature larva: (A) pro and mesonotum; (C) prothoracic leg, internal view; (D) pretarsus. Early larva: (B) pro and mesonotum; (E) prothoracic leg, internal view; (F) middle leg, external view; (G) thoracic spiracle; (H) segment IX, dorsal view; (I) urogomphi, apex. Scale bar: A, B = 1.0 mm, C, G, H = 0.1 mm, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022051" httpUri="https://zenodo.org/record/14022051/files/figure.png" pageId="14" pageNumber="87">8C–D</figureCitation>
) long, increasing in size from anterior to posterior ones; coxa inserted distant from midline of thorax; coxae robust, sclerotized with short setae inserted in sclerotized punctures; setae more concentrate internally; trochanter with setae more concentrate ventrally; trochanter and femur darker, elongated, and setose; pretarsus wide, with two basal setae. In early instar (
<figureCitation id="13232A2DFFDFFD595ED8F835FBFE45DD" box="[979,1099,1951,1977]" captionStart="FIGURE 8" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[153,1435,185,1919]" captionTargetId="figure-27@10.[151,1435,181,1919]" captionTargetPageId="10" captionText="FIGURE 8. Pselaphacus procerus Kuhnt, 1910. Mature larva: (A) pro and mesonotum; (C) prothoracic leg, internal view; (D) pretarsus. Early larva: (B) pro and mesonotum; (E) prothoracic leg, internal view; (F) middle leg, external view; (G) thoracic spiracle; (H) segment IX, dorsal view; (I) urogomphi, apex. Scale bar: A, B = 1.0 mm, C, G, H = 0.1 mm, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022051" httpUri="https://zenodo.org/record/14022051/files/figure.png" pageId="14" pageNumber="87">Figs 8E–F</figureCitation>
; 
<figureCitation id="13232A2DFFDFFD59595DF835FB1D45DD" box="[1110,1192,1951,1977]" captionStart="FIGURE 10" captionStartId="12.[152,255,1484,1508]" captionTargetBox="[310,1278,181,1460]" captionTargetId="figure-27@12.[310,1278,181,1460]" captionTargetPageId="12" captionText="FIGURE 10. Pselaphacus procerus Kuhnt, 1910. Early larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022057" httpUri="https://zenodo.org/record/14022057/files/figure.png" pageId="14" pageNumber="87">10B–C</figureCitation>
; 
<figureCitation id="13232A2DFFDFFD5959B8F835FABC45DD" box="[1203,1289,1951,1977]" captionStart="FIGURE 12" captionStartId="16.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-27@16.[164,1423,181,1884]" captionTargetPageId="16" captionText="FIGURE 12. Pselaphacus procerus Kuhnt, 1910. Early larvae under SEM. (A–B) head, ventral and lateral views; (C) mouthparts, ventral; (D) antenna; (E) mesothoracic tubercle near coxa; (F) mesonotum; (G–H) thorax, lateral and ventral views; (I) thoracic spiracle; (J) abdominal spiracle; (K) segments VIII–X, lateral view. Scale bar: A, B, F, G, H, K = 0.1 mm; C, D, E = 0.03 mm; I, J = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022063" httpUri="https://zenodo.org/record/14022063/files/figure.png" pageId="14" pageNumber="87">12G–H</figureCitation>
), legs similar to mature, apparently setae a little longer. Legs clearer and more setose in early instar.
</paragraph>
<caption id="DF676620FFDEFD585D93F8DEFEF3459C" ID-DOI="http://doi.org/10.5281/zenodo.14022059" ID-Zenodo-Dep="14022059" httpUri="https://zenodo.org/record/14022059/files/figure.png" pageId="15" pageNumber="88" startId="15.[152,255,1908,1932]" targetBox="[166,1419,184,1878]" targetPageId="15" targetType="figure">
<paragraph id="8BA736A8FFDEFD585D93F8DEFEF3459C" blockId="15.[151,1436,1908,2040]" pageId="15" pageNumber="88">
<emphasis id="B96CEABAFFDEFD585D93F8DEFE9045E8" bold="true" box="[152,293,1908,1932]" pageId="15" pageNumber="88">FIGURE 11.</emphasis>
<taxonomicName id="4C184D2BFFDEFD585C27F8DEFD2245E8" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[300,663,1908,1932]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFDEFD585C27F8DEFDB845E8" box="[300,525,1908,1932]" italics="true" pageId="15" pageNumber="88">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFDEFD585F1FF8DEFD2245E8" author="Kuhnt, P." box="[532,663,1908,1932]" pageId="15" pageNumber="88" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Early larvae. (A–B) head, dorsal and ventral views; (C–D) antenna, dorsal and ventral views; (E) clypeus and labrum; (F) epipharynx; (G–I) mandible, dorsal, external and internal views; (J–K) maxilla, dorsal and ventral views; (L–M) apex of mala, dorsal and ventral views; (N) labium. Scale bar: A = 0,5 mm, B–K, N = 0.1 mm, L, M = 0.05 mm.
</paragraph>
</caption>
<caption id="DF676620FFC1FD475D93F8DEFE1A459C" ID-DOI="http://doi.org/10.5281/zenodo.14022063" ID-Zenodo-Dep="14022063" httpUri="https://zenodo.org/record/14022063/files/figure.png" pageId="16" pageNumber="89" startId="16.[152,255,1908,1932]" targetBox="[164,1423,181,1884]" targetPageId="16" targetType="figure">
<paragraph id="8BA736A8FFC1FD475D93F8DEFE1A459C" blockId="16.[151,1436,1908,2040]" pageId="16" pageNumber="89">
<emphasis id="B96CEABAFFC1FD475D93F8DEFE9F45E8" bold="true" box="[152,298,1908,1932]" pageId="16" pageNumber="89">FIGURE 12.</emphasis>
<taxonomicName id="4C184D2BFFC1FD475C3DF8DEFD1845E8" authority="Kuhnt, 1910" authorityName="Kuhnt" authorityYear="1910" box="[310,685,1908,1932]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFC1FD475C3DF8DEFDAF45E8" box="[310,538,1908,1932]" italics="true" pageId="16" pageNumber="89">Pselaphacus procerus</emphasis>
<bibRefCitation id="EF894B59FFC1FD475F2DF8DEFD1845E8" author="Kuhnt, P." box="[550,685,1908,1932]" pageId="16" pageNumber="89" pagination="219 - 270" refId="ref9534" refString="Kuhnt, P. (1910) Neue Erotylidae. (Col.). Deutsche entomologische Zeitschrift 219 - 270. Available from: https: // www. biodiversitylibrary. org / page / 35571530 (accessed 14 April 2023) https: // doi. org / 10.1002 / mmnd. 4801910301" type="book chapter" year="1910">Kuhnt, 1910</bibRefCitation>
</taxonomicName>
. Early larvae under SEM. (A–B) head, ventral and lateral views; (C) mouthparts, ventral; (D) antenna; (E) mesothoracic tubercle near coxa; (F) mesonotum; (G–H) thorax, lateral and ventral views; (I) thoracic spiracle; (J) abdominal spiracle; (K) segments VIII–X, lateral view. Scale bar: A, B, F, G, H, K = 0.1 mm; C, D, E = 0.03 mm; I, J = 0.01 mm.
</paragraph>
</caption>
<paragraph id="8BA736A8FFC0FD465DCCFF3DFAF94391" blockId="17.[151,1437,151,1545]" pageId="17" pageNumber="90">
<emphasis id="B96CEABAFFC0FD465DCCFF3DFE8442D5" box="[199,305,151,177]" italics="true" pageId="17" pageNumber="90">Abdomen</emphasis>
(
<figureCitation id="13232A2DFFC0FD465C4AFF3DFE0042D5" box="[321,437,151,177]" captionStart="FIGURE 6" captionStartId="8.[152,255,1867,1891]" captionTargetBox="[151,1427,287,1838]" captionTargetId="figure-62@8.[151,1435,272,1844]" captionTargetPageId="8" captionText="FIGURE 6. Pselaphacus procerus Kuhnt, 1910. Mature larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022045" httpUri="https://zenodo.org/record/14022045/files/figure.png" pageId="17" pageNumber="90">Fig.6A–C</figureCitation>
), segments I–III slightly increasing in length and width; segments IV–VIII increasing in length and decreasing in width; segments I–VIII, each with a dorsal elliptical plate, brown and yellow at middle, with setae inserted in small sclerotized tubercles, irregularly distributed; dorsal plates slightly decreasing in width and increasing in length to apex direction; each segment with one laterodorsal annular-biforous spiracle (
<figureCitation id="13232A2DFFC0FD465832FEA9FA3A4379" box="[1337,1423,259,285]" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-18@11.[164,1423,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Pselaphacus procerus Kuhnt, 1910. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022053" httpUri="https://zenodo.org/record/14022053/files/figure.png" pageId="17" pageNumber="90">Fig. 9F</figureCitation>
), smaller than thoracic ones and one pleural rounded lobe on each side, with several tiny setae; segment IX (
<figureCitation id="13232A2DFFC0FD46584AFE8DFF1C4301" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-18@11.[164,1423,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Pselaphacus procerus Kuhnt, 1910. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022053" httpUri="https://zenodo.org/record/14022053/files/figure.png" pageId="17" pageNumber="90">Fig. 9J– L</figureCitation>
) narrower, gradually narrowed apicad, with dorsal plate almost covering whole tergite and fused with urogomphi; each side with one vestigial spiracle. Urogomphi (
<figureCitation id="13232A2DFFC0FD465FD0FEC5FCFE43ED" box="[731,843,367,393]" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-18@11.[164,1423,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Pselaphacus procerus Kuhnt, 1910. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022053" httpUri="https://zenodo.org/record/14022053/files/figure.png" pageId="17" pageNumber="90">Fig. 9J–L</figureCitation>
) fix and strongly sclerotized, with several flattened tubercles with one seta at base. Segment X (
<figureCitation id="13232A2DFFC0FD465F8BFE39FD5943C9" box="[640,748,403,429]" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-18@11.[164,1423,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Pselaphacus procerus Kuhnt, 1910. Mature larvae under SEM. (A, C–D) head, lateral, frontal and ventral views; (B) head, thorax and abdominal segment I, lateral view; (E) ventral mouthparts; (F) abdominal spiracle; (G) pro and mesonotum; (H) prothoracic leg; (I) integument of coxa; (J–L) segments IX–X, lateral and ventral views; (K) segment X, dorsal view. Scale bar: A, C, D, E, G, H, J, K, L = 0.1 mm. B = 0.3 mm; F, I = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022053" httpUri="https://zenodo.org/record/14022053/files/figure.png" pageId="17" pageNumber="90">Fig. 9J, L</figureCitation>
) ventral and tubular, apicoventral, apex forming two lobes. In early instar (
<figureCitation id="13232A2DFFC0FD465C28FE1DFE2C43B5" box="[291,409,439,465]" captionStart="FIGURE 8" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[153,1435,185,1919]" captionTargetId="figure-27@10.[151,1435,181,1919]" captionTargetPageId="10" captionText="FIGURE 8. Pselaphacus procerus Kuhnt, 1910. Mature larva: (A) pro and mesonotum; (C) prothoracic leg, internal view; (D) pretarsus. Early larva: (B) pro and mesonotum; (E) prothoracic leg, internal view; (F) middle leg, external view; (G) thoracic spiracle; (H) segment IX, dorsal view; (I) urogomphi, apex. Scale bar: A, B = 1.0 mm, C, G, H = 0.1 mm, E, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022051" httpUri="https://zenodo.org/record/14022051/files/figure.png" pageId="17" pageNumber="90">Figs 8H–I</figureCitation>
; 
<figureCitation id="13232A2DFFC0FD465CAFFE1DFE4D43B5" box="[420,504,439,465]" captionStart="FIGURE 10" captionStartId="12.[152,255,1484,1508]" captionTargetBox="[310,1278,181,1460]" captionTargetId="figure-27@12.[310,1278,181,1460]" captionTargetPageId="12" captionText="FIGURE 10. Pselaphacus procerus Kuhnt, 1910. Early larvae. Habitus. (A) dorsal view; (B) ventral view; (C) lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14022057" httpUri="https://zenodo.org/record/14022057/files/figure.png" pageId="17" pageNumber="90">10A–C</figureCitation>
; 
<figureCitation id="13232A2DFFC0FD465F08FE1DFD8243B5" box="[515,567,439,465]" captionStart="FIGURE 12" captionStartId="16.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-27@16.[164,1423,181,1884]" captionTargetPageId="16" captionText="FIGURE 12. Pselaphacus procerus Kuhnt, 1910. Early larvae under SEM. (A–B) head, ventral and lateral views; (C) mouthparts, ventral; (D) antenna; (E) mesothoracic tubercle near coxa; (F) mesonotum; (G–H) thorax, lateral and ventral views; (I) thoracic spiracle; (J) abdominal spiracle; (K) segments VIII–X, lateral view. Scale bar: A, B, F, G, H, K = 0.1 mm; C, D, E = 0.03 mm; I, J = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022063" httpUri="https://zenodo.org/record/14022063/files/figure.png" pageId="17" pageNumber="90">12K</figureCitation>
), dorsal plates are absent or small and present only on first or first and second segments; urogomphi (
<figureCitation id="13232A2DFFC0FD465C93FE71FDB54391" box="[408,512,475,501]" captionStart="FIGURE 12" captionStartId="16.[152,255,1908,1932]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-27@16.[164,1423,181,1884]" captionTargetPageId="16" captionText="FIGURE 12. Pselaphacus procerus Kuhnt, 1910. Early larvae under SEM. (A–B) head, ventral and lateral views; (C) mouthparts, ventral; (D) antenna; (E) mesothoracic tubercle near coxa; (F) mesonotum; (G–H) thorax, lateral and ventral views; (I) thoracic spiracle; (J) abdominal spiracle; (K) segments VIII–X, lateral view. Scale bar: A, B, F, G, H, K = 0.1 mm; C, D, E = 0.03 mm; I, J = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14022063" httpUri="https://zenodo.org/record/14022063/files/figure.png" pageId="17" pageNumber="90">Fig. 12K</figureCitation>
), sclerotized only at the apex, and setae of the abdomen are anchor-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C3026523FFC0FD465DCCFE54FB5D4171" pageId="17" pageNumber="90" type="materials_examined">
<paragraph id="8BA736A8FFC0FD465DCCFE54FC4A4059" blockId="17.[151,1437,151,1545]" pageId="17" pageNumber="90">
<emphasis id="B96CEABAFFC0FD465DCCFE54FE13407D" bold="true" box="[199,422,510,537]" italics="true" pageId="17" pageNumber="90">Material examined.</emphasis>
<materialsCitation id="3B703CF5FFC0FD465CA7FE55FC4F4059" collectingDate="2018-11-25" collectionCode="T" collectorName="T. Polizei &amp; A. Sparvoli" country="Brazil" county="Uberlandia" location="Rio Araguari" municipality="Uberlandia" pageId="17" pageNumber="90" specimenCode="MZSP 10375" specimenCount="1" stateProvince="Minas Gerais">
<collectingCountry id="F30F7638FFC0FD465CA7FE55FE41407D" box="[428,500,511,537]" name="Brazil" pageId="17" pageNumber="90">Brazil</collectingCountry>
. 
<collectingRegion id="49DCF84AFFC0FD465CF7FE55FD21407D" box="[508,660,511,537]" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="90">Minas Gerais</collectingRegion>
: 
<collectingMunicipality id="6BC3ACD2FFC0FD465F96FE55FCA9407D" box="[669,796,511,537]" pageId="17" pageNumber="90">Uberlândia</collectingMunicipality>
, 
<location id="8EC76073FFC0FD465E2EFE55FC02407D" LSID="urn:lsid:plazi:treatment:03B187BEFFD3FD465D9CF981FB22446D:8EC76073FFC0FD465E2EFE55FC02407D" box="[805,951,511,537]" country="Brazil" county="Uberlandia" municipality="Uberlandia" name="Rio Araguari" pageId="17" pageNumber="90" stateProvince="Minas Gerais">Rio Araguari</location>
, 
<location id="8EC76073FFC0FD465ECBFE55FAB9407D" LSID="urn:lsid:plazi:treatment:03B187BEFFD3FD465D9CF981FB22446D:8EC76073FFC0FD465ECBFE55FAB9407D" box="[960,1292,511,537]" country="Brazil" county="Uberlandia" municipality="Uberlandia" name="Rancho Talisma Ana Gabriela" pageId="17" pageNumber="90" stateProvince="Minas Gerais">Rancho Talismã Ana Gabriela</location>
, 
<date id="FFA61068FFC0FD46581DFE54FA22407D" box="[1302,1431,510,537]" pageId="17" pageNumber="90" value="2018-11-25">
<collectingDate id="EFE2E980FFC0FD46581DFE54FA22407D" box="[1302,1431,510,537]" pageId="17" pageNumber="90" value="2018-11-25">25.XI.2018</collectingDate>
</date>
, 
<collectorName id="26ED537EFFC0FD465D9CFD89FEB74059" box="[151,258,547,573]" pageId="17" pageNumber="90">T. Polizei</collectorName>
&amp; 
<collectorName id="26ED537EFFC0FD465C2FFD89FE1D4059" box="[292,424,547,573]" pageId="17" pageNumber="90">A. Sparvoli</collectorName>
, (ca. 300 early and 36 mature larvae) (
<specimenCode id="DBBE9ED3FFC0FD465E5CFD89FC464059" box="[855,1011,546,573]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="17" pageNumber="90" type="Museum">MZSP 10375</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA736A8FFC0FD465DCCFDECFB5D4171" blockId="17.[151,1437,151,1545]" pageId="17" pageNumber="90">
<emphasis id="B96CEABAFFC0FD465DCCFDECFE904005" bold="true" box="[199,293,582,609]" italics="true" pageId="17" pageNumber="90">Habitat.</emphasis>
The adults associated with the immatures were collected in the summer of 
<quantity id="4CE09B4DFFC0FD46599FFDEDFB594005" box="[1172,1260,583,609]" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" pageId="17" pageNumber="90" unit="in" value="2018.0">2018 in</quantity>
fruiting bodies of 
<taxonomicName id="4C184D2BFFC0FD465DBCFDC1FEEF40E1" authority="Fries." authorityName="Fries." box="[183,346,619,645]" class="Agaricomycetes" family="Polyporaceae" genus="Favolus" kingdom="Fungi" order="Polyporales" pageId="17" pageNumber="90" phylum="Basidiomycota" rank="genus">
<emphasis id="B96CEABAFFC0FD465DBCFDC1FEA440E1" box="[183,273,619,645]" italics="true" pageId="17" pageNumber="90">Favolus</emphasis>
Fries.
</taxonomicName>
(
<taxonomicName id="4C184D2BFFC0FD465C61FDC1FDBD40E1" box="[362,520,619,645]" class="Agaricomycetes" family="Polyporaceae" kingdom="Fungi" order="Polyporales" pageId="17" pageNumber="90" phylum="Basidiomycota" rank="family">Polyporaceae</taxonomicName>
). The specimens were found by the last author (
<collectionCode id="ED09AE6DFFC0FD46593BFDC1FBC040E1" box="[1072,1141,619,645]" pageId="17" pageNumber="90">TTSP</collectionCode>
) forming gregarious balls over the fruiting of fungi in a rotten wood trunk, in a ranch nearby Aguari river, Uberlândia municipality, 
<collectingRegion id="49DCF84AFFC0FD46585DFD25FEAA40A9" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="90">Minas Gerais state</collectingRegion>
, 
<collectingCountry id="F30F7638FFC0FD465C27FD19FEC140A9" box="[300,372,691,717]" name="Brazil" pageId="17" pageNumber="90">Brazil</collectingCountry>
. The hundreds of larvae were feeding on fungus and taken care by 
<specimenCount id="9D1EFD21FFC0FD465971FD19FB4440A9" box="[1146,1265,691,717]" count="5" pageId="17" pageNumber="90" type="adult">five adults</specimenCount>
(
<specimenCount id="9D1EFD21FFC0FD465809FD19FADE40A9" box="[1282,1387,691,717]" count="1" pageId="17" pageNumber="90" type="male">one male</specimenCount>
and 
<specimenCount id="9D1EFD21FFC0FD465D9CFD7DFE9C4095" box="[151,297,727,753]" count="4" pageId="17" pageNumber="90" type="female">four females</specimenCount>
). This kind of behaviour has been reported in the 
<taxonomicName id="4C184D2BFFC0FD465E5CFD7DFC794095" box="[855,972,727,753]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="family">Erotylidae</taxonomicName>
family, especially in 
<taxonomicName id="4C184D2BFFC0FD4659B7FD7DFAFF4095" authorityName="Percheron" authorityYear="1835" box="[1212,1354,727,753]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEABAFFC0FD4659B7FD7DFAFF4095" box="[1212,1354,727,753]" italics="true" pageId="17" pageNumber="90">Pselaphacus</emphasis>
</taxonomicName>
, where another seven species have exhibited maternal care of the brood mass (
<bibRefCitation id="EF894B59FFC0FD465EBAFD51FB694171" author="Chaboo, C. S. &amp; McHugh, J. V." box="[945,1244,763,789]" pageId="17" pageNumber="90" pagination="116 - 118" refId="ref8967" refString="Chaboo, C. S. &amp; McHugh, J. V. (2010) Maternal care by a species of Pselaphacus Percheron (Coleoptera: Erotylidae: Erorylinae) from Peru. The Coleopterists Bulletin, 64 (2), 116 - 118. https: // doi. org / 10.1649 / 072.064.0204" type="journal article" year="2010">Chaboo &amp; McHugh, 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3026523FFC0FD465DCCFCB4FADA4651" pageId="17" pageNumber="90" type="distribution">
<paragraph id="8BA736A8FFC0FD465DCCFCB4FADA4651" blockId="17.[151,1437,151,1545]" pageId="17" pageNumber="90">
<emphasis id="B96CEABAFFC0FD465DCCFCB4FEEF415D" bold="true" box="[199,346,798,825]" italics="true" pageId="17" pageNumber="90">Distribution.</emphasis>
<collectingCountry id="F30F7638FFC0FD465C68FCB5FE12415D" box="[355,423,799,825]" name="Brazil" pageId="17" pageNumber="90">Brazil</collectingCountry>
(first country record): Amazonas (?), 
<collectingRegion id="49DCF84AFFC0FD465E55FCB5FC4F415D" box="[862,1018,799,825]" country="Brazil" name="Minas Gerais" pageId="17" pageNumber="90">Minas Gerais</collectingRegion>
, 
<collectingRegion id="49DCF84AFFC0FD46590CFCB5FB5B415D" box="[1031,1262,799,825]" country="Brazil" name="Mato Grosso do Sul" pageId="17" pageNumber="90">Mato Grosso do Sul</collectingRegion>
and 
<collectingRegion id="49DCF84AFFC0FD465822FCB5FA29415D" box="[1321,1436,799,825]" country="Brazil" name="Sao Paulo" pageId="17" pageNumber="90">São Paulo</collectingRegion>
States. 
<collectingCountry id="F30F7638FFC0FD465DECFCE9FEEE4139" box="[231,347,835,861]" name="Argentina" pageId="17" pageNumber="90">Argentina</collectingCountry>
: 
<collectingRegion id="49DCF84AFFC0FD465C6DFCE9FE784139" box="[358,461,835,861]" country="Argentina" name="Misiones" pageId="17" pageNumber="90">Misiones</collectingRegion>
(
<figureCitation id="13232A2DFFC0FD465CD6FCE9FDAF4139" box="[477,538,835,861]" captionStart="FIGURE 1" captionStartId="3.[152,255,1145,1169]" captionTargetBox="[181,1410,310,1115]" captionTargetId="figure-70@3.[176,1410,306,1119]" captionTargetPageId="3" captionText="FIGURE 1. Map showing the known distribution of the Pselaphacus procerus Kuhnt, 1910. (A–G) Brazil: (A) Benjamin Constant, Amazonas state: 4° 23′ 0″ S, 70° 1′ 53″W. (B) Uberlândia, Minas Gerais state: 18° 54′41″ S, 48° 15′ 44″W. (C) Três Lagoas, Mato Grosso do Sul state: 20° 45′ 35″ S, 51° 41′ 42″W. (D) Castilho, São Paulo state: 20° 52′ 12″ S, 51° 29′ 13″W. (E) Marília, São Paulo state: 22° 13′ 15″ S, 49° 56′ 55″W. (F) Salto Grande, São Paulo state: 22° 52′ 53″ S, 49° 57′ 44″W. (G) Piracicaba, São Paulo state: 22° 43’ 30” S, 47° 38’ 56” W. (H) Misiones, Argentina: 26° 55′ S, 54° 31′ W, [type locality]." figureDoi="http://doi.org/10.5281/zenodo.14022033" httpUri="https://zenodo.org/record/14022033/files/figure.png" pageId="17" pageNumber="90">Fig.1</figureCitation>
). The original description of 
<taxonomicName id="4C184D2BFFC0FD465E68FCE9FC574138" authorityName="Kuhnt" authorityYear="1910" box="[867,994,835,860]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFC0FD465E68FCE9FC574138" box="[867,994,835,860]" italics="true" pageId="17" pageNumber="90">P. procerus</emphasis>
</taxonomicName>
was based on material from 
<collectingCountry id="F30F7638FFC0FD46582DFCE9FA2C4139" box="[1318,1433,835,861]" name="Argentina" pageId="17" pageNumber="90">Argentina</collectingCountry>
. During the study of 
<taxonomicName id="4C184D2BFFC0FD465C70FCCDFE4541E5" box="[379,496,871,897]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="family">Erotylidae</taxonomicName>
collection of MZSP, it was found 
<specimenCount id="9D1EFD21FFC0FD465E78FCCDFBB841E5" box="[883,1037,871,897]" count="1" pageId="17" pageNumber="90" type="generic">one specimen</specimenCount>
from Amazonas state, 
<collectingCountry id="F30F7638FFC0FD46581BFCCDFAED41E5" box="[1296,1368,871,897]" name="Brazil" pageId="17" pageNumber="90">Brazil</collectingCountry>
. After the comparison of this specimen with the original description and remainder specimens, there is no doubt that the specimen from Amazonas state (Amazon region) also corresponds to 
<taxonomicName id="4C184D2BFFC0FD465EB1FC05FB8541AC" box="[954,1072,943,968]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="proceru">
<emphasis id="B96CEABAFFC0FD465EB1FC05FB8541AC" box="[954,1072,943,968]" italics="true" pageId="17" pageNumber="90">P. proceru</emphasis>
</taxonomicName>
s. However, the material of this region, including this specimen, is very old and came from a private collection (J. Guérin), incorporated at the MZSP a few years ago. Based on the distribution of the other specimens of this species, collected in Central and Southeast 
<collectingCountry id="F30F7638FFC0FD465C00FBB1FEE74651" box="[267,338,1051,1077]" name="Brazil" pageId="17" pageNumber="90">Brazil</collectingCountry>
, and the 
<typeStatus id="54A3880AFFC0FD465CBDFBB1FE534651" box="[438,486,1051,1077]" pageId="17" pageNumber="90">type</typeStatus>
locality in 
<collectingCountry id="F30F7638FFC0FD465F6FFBB1FD664651" box="[612,723,1051,1077]" name="Argentina" pageId="17" pageNumber="90">Argentina</collectingCountry>
(
<figureCitation id="13232A2DFFC0FD465FE9FBB1FC914651" box="[738,804,1051,1077]" captionStart="FIGURE 1" captionStartId="3.[152,255,1145,1169]" captionTargetBox="[181,1410,310,1115]" captionTargetId="figure-70@3.[176,1410,306,1119]" captionTargetPageId="3" captionText="FIGURE 1. Map showing the known distribution of the Pselaphacus procerus Kuhnt, 1910. (A–G) Brazil: (A) Benjamin Constant, Amazonas state: 4° 23′ 0″ S, 70° 1′ 53″W. (B) Uberlândia, Minas Gerais state: 18° 54′41″ S, 48° 15′ 44″W. (C) Três Lagoas, Mato Grosso do Sul state: 20° 45′ 35″ S, 51° 41′ 42″W. (D) Castilho, São Paulo state: 20° 52′ 12″ S, 51° 29′ 13″W. (E) Marília, São Paulo state: 22° 13′ 15″ S, 49° 56′ 55″W. (F) Salto Grande, São Paulo state: 22° 52′ 53″ S, 49° 57′ 44″W. (G) Piracicaba, São Paulo state: 22° 43’ 30” S, 47° 38’ 56” W. (H) Misiones, Argentina: 26° 55′ S, 54° 31′ W, [type locality]." figureDoi="http://doi.org/10.5281/zenodo.14022033" httpUri="https://zenodo.org/record/14022033/files/figure.png" pageId="17" pageNumber="90">Fig. 1</figureCitation>
), this specimen from Amazonas may be mislabeled.
</paragraph>
</subSubSection>
<subSubSection id="C3026523FFC0FD465DCCFB94FB22446D" pageId="17" pageNumber="90" type="discussion">
<paragraph id="8BA736A8FFC0FD465DCCFB94FE2846A1" blockId="17.[151,1437,151,1545]" pageId="17" pageNumber="90">
<emphasis id="B96CEABAFFC0FD465DCCFB94FE04463D" bold="true" box="[199,433,1086,1113]" pageId="17" pageNumber="90">Comparative notes.</emphasis>
Based on adults, 
<taxonomicName id="4C184D2BFFC0FD465F73FB95FD42463C" authorityName="Kuhnt" authorityYear="1910" box="[632,759,1087,1112]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFC0FD465F73FB95FD42463C" box="[632,759,1087,1112]" italics="true" pageId="17" pageNumber="90">P. procerus</emphasis>
</taxonomicName>
is similar both in size and coloration to other two species of the genus: 
<taxonomicName id="4C184D2BFFC0FD465C1AFBC9FD394619" authority="Lacordaire, 1942" authorityName="Lacordaire" authorityYear="1942" box="[273,652,1123,1149]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="poecilosomus">
<emphasis id="B96CEABAFFC0FD465C1AFBC9FE704619" box="[273,453,1123,1149]" italics="true" pageId="17" pageNumber="90">P. poecilosomus</emphasis>
Lacordaire, 1942
</taxonomicName>
and 
<taxonomicName id="4C184D2BFFC0FD465FC8FBC9FC514619" authority="Crotch, 1973" authorityName="Crotch" authorityYear="1973" box="[707,996,1123,1149]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="vitticollis">
<emphasis id="B96CEABAFFC0FD465FC8FBC9FCF24619" box="[707,839,1123,1149]" italics="true" pageId="17" pageNumber="90">P. vitticollis</emphasis>
Crotch, 1973
</taxonomicName>
. It can be distinguished from these two by the absence of a black mark at the humeral angle of the elytron and by the separation of the longitudinal strips from the sutural marks.
</paragraph>
<paragraph id="8BA736A8FFC0FD465DCCFB65FDDF47F9" blockId="17.[151,1437,151,1545]" pageId="17" pageNumber="90">
Comparing the larvae of 
<taxonomicName id="4C184D2BFFC0FD465CE2FB65FD56468C" authorityName="Kuhnt" authorityYear="1910" box="[489,739,1231,1257]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFC0FD465CE2FB65FD56468C" box="[489,739,1231,1257]" italics="true" pageId="17" pageNumber="90">Pselaphacus procerus</emphasis>
</taxonomicName>
with that of 
<taxonomicName id="4C184D2BFFC0FD465E70FB65FBF7468C" authorityName="Percheron" authorityYear="1835" box="[891,1090,1231,1256]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="nigropunctatus">
<emphasis id="B96CEABAFFC0FD465E70FB65FBF7468C" box="[891,1090,1231,1256]" italics="true" pageId="17" pageNumber="90">P. nigropunctatus</emphasis>
</taxonomicName>
it was verified that the larvae of both species are different, especially in coloration. In the first species, there is darker dorsal plates, yellow with contour dark-brown, and in second, the plates are brownish, but in both the setae are inserted in small sclerotized tubercles. Both species present epicranial suture U-shaped, antennae with three antennomeres, antennomere 2 with membranous sensorium, mandibles concave with apex 3-toothed, prostheca membranous, lobate and microspined, and urogomphi fixed and one-segmented.
</paragraph>
<paragraph id="8BA736A8FFC0FD465DCCFA0DFB22446D" blockId="17.[151,1437,151,1545]" pageId="17" pageNumber="90">
In a more detailed comparison of larvae of both species, it was verified that 
<taxonomicName id="4C184D2BFFC0FD465EE7FA0DFB7347A4" box="[1004,1222,1447,1473]" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B96CEABAFFC0FD465EE7FA0DFB7347A4" box="[1004,1222,1447,1473]" italics="true" pageId="17" pageNumber="90">Pselaphus procerus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C184D2BFFC0FD4659D9FA0DFA2647A4" authorityName="Percheron" authorityYear="1835" box="[1234,1427,1447,1472]" class="Insecta" family="Erotylidae" genus="Pselaphacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="nigropunctatus">
<emphasis id="B96CEABAFFC0FD4659D9FA0DFA2647A4" box="[1234,1427,1447,1472]" italics="true" pageId="17" pageNumber="90">P.nigropunctatus</emphasis>
</taxonomicName>
) has clypeus and labrum separated (partially fused); gular sutures absent (gular sutures short); some marginal areas of mandibular teeth micro-toothed (straight); urogomphi with a distal tooth (without tooth).
</paragraph>
</subSubSection>
</treatment>
</document>