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<document id="2620799215994D4815084683DE9C5C23" ID-CLB-Dataset="7631" ID-DOI="10.1206/0003-0090-423.1.1" ID-GBIF-Dataset="2e886aea-b59a-45a6-aeaa-2427d584a894" ID-ISSN="0003-0090" ID-Zenodo-Dep="4613008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615995240537" checkinUser="felipe" docAuthor="Grimaldi, David A." docDate="2018" docId="03B187A8FFB5FFCBFF0B3DB17E6BFBC1" docLanguage="en" docName="BulAmeMusNatHis.2018.423.1-97.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 2018 (423)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-423.1.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Chandleromyia anomala Grimaldi 2018, new species" docType="treatment" docVersion="7" lastPageNumber="97" masterDocId="FF88FFD0FF86FFFEFFAA39537F49FFF7" masterDocTitle="Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi" masterLastPageNumber="97" masterPageNumber="1" pageNumber="1" updateTime="1698933720305" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CE98FD768FA129B731BF8778F36B1D80">Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi</mods:title>
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<mods:name id="01E4F910A6C33D51B23E6B0D134C18A3" type="personal">
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<mods:namePart id="2E41BC493ADA2785DB671DC3855F04E9">Grimaldi, David A.</mods:namePart>
<mods:affiliation id="165D50A8A414AAE26F1E063E2658EC8D">Division of Invertebrate Zoology American Museum of Natural History, New York</mods:affiliation>
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<mods:date id="DA79CB3CB12C620B682FE4D47DC2F492">2018</mods:date>
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<mods:number id="CD3F9D4691E40D73B6905E8823F4401E">2018-10-24</mods:number>
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<treatment id="03B187A8FFB5FFCBFF0B3DB17E6BFBC1" ID-DOI="http://doi.org/10.5281/zenodo.4631127" ID-GBIF-Taxon="180670799" ID-Zenodo-Dep="4631127" LSID="urn:lsid:plazi:treatment:03B187A8FFB5FFCBFF0B3DB17E6BFBC1" httpUri="http://treatment.plazi.org/id/03B187A8FFB5FFCBFF0B3DB17E6BFBC1" lastPageId="53" pageId="51">
<subSubSection id="C3026535FFB5FFCDFF0B3DB17D0EFB0D" box="[161,583,1250,1274]" pageId="51" type="nomenclature">
<paragraph id="8BA736BEFFB5FFCDFF0B3DB17D0EFB0D" blockId="51.[161,583,1250,1274]" box="[161,583,1250,1274]" pageId="51">
<heading id="D0EF81D2FFB5FFCDFF0B3DB17D0EFB0D" box="[161,583,1250,1274]" centered="true" fontSize="9" level="2" pageId="51" reason="2">
<taxonomicName id="4C184D3DFFB5FFCDFF0B3DB17EF0FB0D" ID-CoL="8BFQC" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[161,441,1250,1274]" class="Insecta" family="Platypezidae" genus="Chandleromyia" kingdom="Animalia" order="Diptera" pageId="51" phylum="Arthropoda" rank="species" species="anomala" status="sp. nov.">
<emphasis id="B96CEAACFFB5FFCDFF0B3DB17EF0FB0D" bold="true" box="[161,441,1250,1274]" italics="true" pageId="51">Chandleromyia anomala</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFB5FFCDFE6D3DB07D0EFB0D" box="[455,583,1250,1274]" pageId="51" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB5FFCDFE843C477EF3FADB" box="[302,442,1300,1324]" pageId="51" type="description">
<paragraph id="8BA736BEFFB5FFCDFE843C477EF3FADB" blockId="51.[302,442,1300,1324]" box="[302,442,1300,1324]" pageId="51">
<heading id="D0EF81D2FFB5FFCDFE843C477EF3FADB" box="[302,442,1300,1324]" centered="true" fontSize="9" level="2" pageId="51" reason="2">
<figureCitation id="13232A3BFFB5FFCDFE843C477EDAFADB" box="[302,403,1300,1324]" captionStart="FIG" captionStartId="15.[108,150,1420,1441]" captionTargetBox="[108,1212,227,1393]" captionTargetId="figure-7@15.[108,1212,227,1393]" captionTargetPageId="15" captionText="FIG. 8. Chandleromyia anomala, n. gen., n. sp. (Platypezidae), male holotype in Burmese amber, AMNH Bu-KL30-27. A. Dorsolateral habitus. B. Head, ventrolateral view. C. Dorsolateral view. D. Metatarsus. E. Male terminalia, left posterolateral view." figureDoi="http://doi.org/10.5281/zenodo.4613026" httpUri="https://zenodo.org/record/4613026/files/figure.png" pageId="51">Figures 8</figureCitation>
,
<figureCitation id="13232A3BFFB5FFCDFE0A3C477EF3FADB" box="[416,442,1300,1324]" captionStart="FIG" captionStartId="52.[108,150,1501,1522]" captionTargetBox="[132,1181,228,1470]" captionTargetId="figure-9@52.[108,1212,224,1475]" captionTargetPageId="52" captionText="FIG. 33. Chandleromyia anomala, n. gen., n. sp. (Platypezidae), male holotype in Burmese amber, AMNH Bu-KL30-27. A. Wing. B. Antenna. C. Metatarsus, lateral. D. Thorax, lateral." figureDoi="http://doi.org/10.5281/zenodo.4613092" httpUri="https://zenodo.org/record/4613092/files/figure.png" pageId="51">33</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB5FFCDFF263C157D77FAA9" box="[140,574,1350,1374]" pageId="51" type="diagnosis">
<paragraph id="8BA736BEFFB5FFCDFF263C157D77FAA9" blockId="51.[108,636,1350,1738]" box="[140,574,1350,1374]" pageId="51">DIAGNOSIS: As for genus, by monotypy.</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB5FFCBFF263C347E6BFC24" lastPageId="53" pageId="51" type="description">
<paragraph id="8BA736BEFFB5FFCDFF263C347BF4FD71" blockId="51.[108,636,1350,1738]" lastBlockId="51.[684,1214,226,1738]" pageId="51">
DESCRIPTION: (male only).
<emphasis id="B96CEAACFFB5FFCDFE4B3C347D6FFA88" bold="true" box="[481,550,1383,1407]" pageId="51">Head:</emphasis>
Large, broader than thorax. Eyes bare, large, holoptic for ~30 facets (virtually entire length of frons except short segment above antennae). Dorsal facets (~ 70% of eye) large, 2× diameter of ventral facets. Frontal margins of eyes emarginate around bases of antennae, margins slightly divergent ventral to this. Median furrow of frons, interfrontal, fronto-orbital setae (presence/ absence) not observable. Face entirely bare. Ocelli on tubercle, with pair of short posterior setae (length shorter than diameter of ocellus). Vertical setae absent; postoculars minute, except for short row of 45 short, thick seate on dorsal portion. Occiput with fine, short, scattered, light setae (lengths ca. equal to diameter of large eye facets); occipital setae do not extend to gena. Vibrissa, genal setae absent.
<emphasis id="B96CEAACFFB5FFCDFBA938FB7B26FE37" bold="true" box="[1027,1135,424,448]" pageId="51">Antenna:</emphasis>
Scape small, barely exposed, apparently bare; pedicel small, conical, with apical circlet of setulae; arista terminal; basal flagellomere small, tapered, with incised portion mesally; two small basal aristomeres (lengths ~3× the diameter); apical aristomere long, fine, bare of any micropubescence.
</paragraph>
<paragraph id="8BA736BEFFB5FFCDFD063BC37B36FCFC" blockId="51.[684,1214,226,1738]" pageId="51">
<emphasis id="B96CEAACFFB5FFCDFD063BC37C74FD5F" bold="true" box="[684,829,656,680]" pageId="51">Mouthparts:</emphasis>
Oral margin squared; clypeus a broad triangle; labrum broad, apex pointed and sclerotized; labellar lobes deeply separated anteriorly, ~0.65× lobe length; palp very small.
</paragraph>
<paragraph id="8BA736BEFFB5FFCDFD673A477BF5FBA2" blockId="51.[684,1214,226,1738]" pageId="51">
<emphasis id="B96CEAACFFB5FFCDFD673A477C6DFCDB" bold="true" box="[717,804,788,812]" pageId="51">Thorax:</emphasis>
Scutum convexed but not arched, flatter anteriorly. Acrostichals completely absent, even minute ones. Only 1 pair of dorsocentrals present, short, prescutellar, no smaller, graded dorsocentrals present. Supraalar setae absent; 45 stout notopleural setae present (4 on one side of specimen, 5 on other side); anepisternum and katepisternum bare. One pair of short, stiff postalar setae present. Prescutellum absent. Scutellum with one pair of apical setae only; setae short, stout.
</paragraph>
<paragraph id="8BA736BEFFB5FFCBFD063D0D7E28FDD3" blockId="51.[684,1214,226,1738]" lastBlockId="53.[108,637,226,1078]" lastPageId="53" pageId="51">
<emphasis id="B96CEAACFFB5FFCDFD063D0D7DB9FB81" bold="true" box="[684,752,1118,1142]" pageId="51">Wing:</emphasis>
Length/width 2.4; hyaline, except for faint tint between Sc and R
<subScript id="179C34FBFFB5FFCDFC0B3DD87CE0FB6B" attach="left" box="[929,937,1163,1180]" fontSize="7" pageId="51">1</subScript>
. Membrane microtrichia well developed over entire surface of wing, even in cell Sc. Two prehumeral costal setae at base of C; C with spinules, end near apex of R
<subScript id="179C34FBFFB5FFCDFBEF3DBD7B29FB08" attach="left" box="[1093,1120,1262,1279]" fontSize="7" pageId="51">2+3</subScript>
. Sc relatively short, apex slightly beyond level of crossvein r-m; R
<subScript id="179C34FBFFB5FFCDFD503C637C4AFAB6" attach="left" box="[762,771,1328,1345]" fontSize="7" pageId="51">1</subScript>
straight, bare, length 0.72× the wing length; bases of Sc and R
<subScript id="179C34FBFFB5FFCDFC1E3C027CF4FA95" attach="left" box="[948,957,1361,1378]" fontSize="7" pageId="51">1</subScript>
close, divergent for apical half. Pterostigma absent. Vein R
<subScript id="179C34FBFFB5FFCDFBEE3C217B16FA74" attach="left" box="[1092,1119,1394,1411]" fontSize="7" pageId="51">4+5</subScript>
ends at wing tip; cells rm and dm closed; crossvein br-m slightly basal to level of Sc apex; crossvein dm-cu close to wing margin, separated from margin 0.75× its length. M not forked (M
<subScript id="179C34FBFFB5FFCDFBB23CA47B68F9FF" attach="left" box="[1048,1057,1527,1544]" fontSize="7" pageId="51">2</subScript>
lost); M
<subScript id="179C34FBFFB5FFCDFBD73CA47BCFF9FF" attach="left" box="[1149,1158,1527,1544]" fontSize="7" pageId="51">1</subScript>
bent in middle, deflected anteriad. Tip of CuA
<subScript id="179C34FBFFB5FFCDFBCA3F4B7B20F9DE" attach="left" box="[1120,1129,1560,1577]" fontSize="7" pageId="51">1</subScript>
incomplete; cell cup longer than vein at apex, cell not extremely acute or narrow; CuP slightly curved, not straight; CuP+A
<subScript id="179C34FBFFB5FFCDFC3B3F287CD3F97B" attach="left" box="[913,922,1659,1676]" fontSize="7" pageId="51">1</subScript>
tip incomplete.
<emphasis id="B96CEAACFFB5FFCDFBFC3F237BD8F97F" bold="true" box="[1110,1169,1648,1672]" pageId="51">Legs:</emphasis>
All femora without macrosetae, spines or tubercles. Dark bifid scales on all tibiae and tarsi: scattered on partial, lateral surface of tibiae, encircling ventral portion of tibial apex; in dense lateral row on tarsi. Metatarsomeres 14 flattened laterally, expanded; tarsomere 2 with blunt spine on lateroapical corner; tarsomere 3 with small, light, clavate seta preapically near row of bifid scales. Mesotibia with 2 ventroapical spurs, large one mesally (its length ca. diameter of tibia). Pulvilli well developed; empodium not observable (probably minute, setiform).
</paragraph>
<caption id="DF676636FFB2FFCAFFC63C8E7C32F9E7" ID-DOI="http://doi.org/10.5281/zenodo.4613092" ID-Zenodo-Dep="4613092" httpUri="https://zenodo.org/record/4613092/files/figure.png" pageId="52" startId="52.[108,150,1501,1522]" targetBox="[132,1181,228,1470]" targetPageId="52">
<paragraph id="8BA736BEFFB2FFCAFFC63C8E7C32F9E7" blockId="52.[108,1212,1499,1553]" pageId="52">
FIG. 33.
<taxonomicName id="4C184D3DFFB2FFCAFF623C887E8BFA04" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[200,450,1499,1523]" class="Insecta" family="Platypezidae" genus="Chandleromyia" kingdom="Animalia" order="Diptera" pageId="52" phylum="Arthropoda" rank="species" species="anomala" status="n. gen., n. sp.">
<emphasis id="B96CEAACFFB2FFCAFF623C887E8BFA04" box="[200,450,1499,1523]" italics="true" pageId="52">Chandleromyia anomala</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFB2FFCAFE7B3C8F7D17FA04" box="[465,606,1500,1524]" pageId="52" rank="species">n. gen., n. sp.</taxonomicNameLabel>
(
<taxonomicName id="4C184D3DFFB2FFCAFDC73C887DBAFA04" box="[621,755,1499,1523]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="52" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
), male holotype in Burmese amber, AMNH Bu-KL30-27.
<emphasis id="B96CEAACFFB2FFCAFF5C3CAB7E46F9E7" bold="true" box="[246,271,1528,1552]" pageId="52">A.</emphasis>
Wing.
<emphasis id="B96CEAACFFB2FFCAFEF13CAA7E3BF9E6" bold="true" box="[347,370,1529,1553]" pageId="52">B.</emphasis>
Antenna.
<emphasis id="B96CEAACFFB2FFCAFE773CAB7EBCF9E7" bold="true" box="[477,501,1528,1552]" pageId="52">C.</emphasis>
Metatarsus, lateral.
<emphasis id="B96CEAACFFB2FFCAFD683CAA7D92F9E6" bold="true" box="[706,731,1529,1553]" pageId="52">D.</emphasis>
Thorax, lateral.
</paragraph>
</caption>
<paragraph id="8BA736BEFFB3FFCBFF263B7E7D4EFCDA" blockId="53.[108,637,226,1078]" pageId="53">
<emphasis id="B96CEAACFFB3FFCBFF263B7E7E4FFDB2" bold="true" box="[140,262,557,581]" pageId="53">Abdomen:</emphasis>
Tergites without macrosetae or setulae; 8 tergites visible; anterior 6 broad (14 broadest); tergite 7 narrow, longer than wide, tubular, ventroflexed (presumably circumverted); epandrium small, capsular. Male genitalia not observable (preservation insufficient). Sternites well developed, but lateral margins do not meet tergites (pleural membrane exposed).
</paragraph>
<paragraph id="8BA736BEFFB3FFCBFF273A657E6BFC24" blockId="53.[108,637,226,1078]" pageId="53">
<materialsCitation id="3B703CE3FFB3FFCBFF273A657E6BFC24" ID-GBIF-Occurrence="3059949307" collectionCode="AMNH" pageId="53" specimenCount="1" typeStatus="holotype">
SPECIMENS EXAMINED:
<typeStatus id="54A3881CFFB3FFCBFE123A657D6CFCB9" box="[440,549,822,846]" pageId="53" type="holotype">Holotype</typeStatus>
, male,
<collectionCode id="ED09AE7BFFB3FFCBFFC63A0B7F89FC87" box="[108,192,856,880]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="53">AMNH</collectionCode>
Bu-KL30-27, in Burmese amber. The fly is extremely well preserved, complete, though some details are obscured (e.g., small structures of the genitalia).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB3FFCBFF263A8F7E6BFBC1" pageId="53" type="etymology">
<paragraph id="8BA736BEFFB3FFCBFF263A8F7E6BFBC1" blockId="53.[108,637,226,1078]" pageId="53">ETYMOLOGY: In reference to the anomalously early appearance of a platypezine in the mid-Cretaceous.</paragraph>
</subSubSection>
</treatment>
</document>