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<document id="1B22641F6D7CD4B2C1389C22E54D280E" ID-CLB-Dataset="7631" ID-DOI="10.1206/0003-0090-423.1.1" ID-GBIF-Dataset="2e886aea-b59a-45a6-aeaa-2427d584a894" ID-ISSN="0003-0090" ID-Zenodo-Dep="4613008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615995240537" checkinUser="felipe" docAuthor="Grimaldi, David A." docDate="2018" docId="03B187A8FFA8FFCEFE8D3F5E7C77FC99" docLanguage="en" docName="BulAmeMusNatHis.2018.423.1-97.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 2018 (423)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-423.1.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Microsania Zetterstedt 1837" docType="treatment" docVersion="8" lastPageNumber="97" masterDocId="FF88FFD0FF86FFFEFFAA39537F49FFF7" masterDocTitle="Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi" masterLastPageNumber="97" masterPageNumber="1" pageNumber="1" updateTime="1698933720305" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F297EA81A04C14A7F7BAD95E392F1B7E">Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi</mods:title>
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<treatment id="03B187A8FFA8FFCEFE8D3F5E7C77FC99" ID-DOI="http://doi.org/10.5281/zenodo.4631111" ID-GBIF-Taxon="180670811" ID-Zenodo-Dep="4631111" LSID="urn:lsid:plazi:treatment:03B187A8FFA8FFCEFE8D3F5E7C77FC99" httpUri="http://treatment.plazi.org/id/03B187A8FFA8FFCEFE8D3F5E7C77FC99" lastPageId="48" pageId="46">
<subSubSection id="C3026535FFA8FFD0FE8D3F5E7E8AF9D3" box="[295,451,1549,1572]" pageId="46" type="nomenclature">
<paragraph id="8BA736BEFFA8FFD0FE8D3F5E7E8AF9D3" blockId="46.[295,451,1549,1572]" box="[295,451,1549,1572]" pageId="46">
<heading id="D0EF81D2FFA8FFD0FE8D3F5E7E8AF9D3" box="[295,451,1549,1572]" centered="true" fontSize="9" level="2" pageId="46" reason="2">
<taxonomicName id="4C184D3DFFA8FFD0FE8D3F5E7ED6F9D3" ID-CoL="5SDF" ID-ENA="1445848" authorityName="Zetterstedt" authorityYear="1837" box="[295,415,1549,1572]" class="Insecta" family="Platypezidae" genus="Microsania" kingdom="Animalia" order="Diptera" pageId="46" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFA8FFD0FE8D3F5E7ED6F9D3" box="[295,415,1549,1572]" italics="true" pageId="46">Microsania</emphasis>
</taxonomicName>
sp.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFA8FFD0FE843F6D7EF3F9A1" box="[302,442,1598,1622]" pageId="46" type="description">
<paragraph id="8BA736BEFFA8FFD0FE843F6D7EF3F9A1" blockId="46.[302,442,1598,1622]" box="[302,442,1598,1622]" pageId="46">
<heading id="D0EF81D2FFA8FFD0FE843F6D7EF3F9A1" box="[302,442,1598,1622]" centered="true" fontSize="9" level="2" pageId="46" reason="2">
<figureCitation id="13232A3BFFA8FFD0FE843F6D7EDAF9A1" box="[302,403,1598,1622]" captionStart="FIG" captionStartId="12.[107,149,1705,1726]" captionTargetBox="[122,1198,224,1679]" captionTargetId="figure-9@12.[108,1212,224,1679]" captionTargetPageId="12" captionText="FIG. 5. Microsania sp. (Platypezidae), female, AMNH Ba502-14, in Eocene Baltic amber. A. Right lateral habitus. B. Frontal view of head." figureDoi="http://doi.org/10.5281/zenodo.4613020" httpUri="https://zenodo.org/record/4613020/files/figure.png" pageId="46">Figures 5</figureCitation>
,
<figureCitation id="13232A3BFFA8FFD0FE0A3F6D7EF3F9A1" box="[416,442,1598,1622]" captionStart="FIG" captionStartId="47.[111,152,1521,1542]" captionTargetBox="[152,1168,233,1488]" captionTargetId="figure-7@47.[108,1212,224,1494]" captionTargetPageId="47" captionText="FIG. 31. Microsania sp. (Platypezidae), female, AMNH Ba502-14, in Eocene Baltic amber. A. Wing. B. Thorax, lateral view. C. Head, frontal. D. Antenna. E. Thorax, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4613088" httpUri="https://zenodo.org/record/4613088/files/figure.png" pageId="46">31</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFA8FFD0FF263F237CD9F9D4" pageId="46" type="discussion">
<paragraph id="8BA736BEFFA8FFD0FF263F237CD9F9D4" blockId="46.[108,636,1647,1738]" lastBlockId="46.[684,1212,1283,1737]" pageId="46">
SUMMARY: Complete female in Baltic amber; with typical array of
<taxonomicName id="4C184D3DFFA8FFD0FEF23FC27E99F95F" authorityName="Zetterstedt" authorityYear="1837" box="[344,464,1681,1704]" class="Insecta" family="Platypezidae" genus="Microsania" kingdom="Animalia" order="Diptera" pageId="46" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFA8FFD0FEF23FC27E99F95F" box="[344,464,1681,1704]" italics="true" pageId="46">Microsania</emphasis>
</taxonomicName>
characters (see generic diagnosis above). Additional details include the following: M, Cu veins with minute gap between apices and wing margin; bm cell and vein CuA2 appear incomplete (the former possibly an artifact of preservation); 2 pairs fronto-orbital setae, anterior pair inclinate, posterior pair latero-proclinate; strong pair of interfrontals present; pair of long, fine presutural setae (probably dorsocentrals) present; prescutellum appears present.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFA8FFCEFD673F7E7CDEFDB3" lastPageId="48" pageId="46" type="description">
<paragraph id="8BA736BEFFA8FFCEFD673F7E7EFFFE89" blockId="46.[684,1212,1283,1737]" lastBlockId="48.[108,638,226,1738]" lastPageId="48" pageId="46">
DESCRIPTION:
<emphasis id="B96CEAACFFA8FFD0FCC53F7E7CFAF9B2" bold="true" box="[879,947,1581,1605]" pageId="46">Head:</emphasis>
Eyes bare, red; dichoptic. Frons with faint median furrow on anterior half; pair of interfrontals present, well developed, proclinate; 2 pairs of fronto-orbitals, anterior pair inclinate, slightly more posterior pair latero- proclinate. Pair of ocellar setae present, long, divergent, smaller pair posteriad. Lateroclinate pair of outer verticals present; inclinate pair of inner verticals, plus other setae on vertex; postoccipital setae well developed.
</paragraph>
<caption id="DF676636FFA9FFD1FFC53CA27D93F9D3" ID-DOI="http://doi.org/10.5281/zenodo.4613088" ID-Zenodo-Dep="4613088" httpUri="https://zenodo.org/record/4613088/files/figure.png" pageId="47" startId="47.[111,152,1521,1542]" targetBox="[152,1168,233,1488]" targetPageId="47">
<paragraph id="8BA736BEFFA9FFD1FFC53CA27D93F9D3" blockId="47.[111,1214,1518,1572]" pageId="47">
FIG. 31.
<taxonomicName id="4C184D3DFFA9FFD1FF6F3CBC7E7DF9F1" authorityName="Zetterstedt" authorityYear="1837" box="[197,308,1519,1542]" class="Insecta" family="Platypezidae" genus="Microsania" kingdom="Animalia" order="Diptera" pageId="47" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFA9FFD1FF6F3CBC7E7DF9F1" box="[197,308,1519,1542]" italics="true" pageId="47">Microsania</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C184D3DFFA9FFD1FECB3CBC7EAFF9F0" box="[353,486,1519,1543]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="47" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
), female, AMNH Ba502-14, in Eocene Baltic amber.
<emphasis id="B96CEAACFFA9FFD1FC593CBD7B45F9F1" bold="true" box="[1011,1036,1518,1542]" pageId="47">A.</emphasis>
Wing.
<emphasis id="B96CEAACFFA9FFD1FBFE3CBC7B23F9F0" bold="true" box="[1108,1130,1519,1543]" pageId="47">B.</emphasis>
Thorax, lateral view.
<emphasis id="B96CEAACFFA9FFD1FF453F5F7E4EF9D3" bold="true" box="[239,263,1548,1572]" pageId="47">C.</emphasis>
Head, frontal.
<emphasis id="B96CEAACFFA9FFD1FE0B3F5F7EF3F9D3" bold="true" box="[417,442,1548,1572]" pageId="47">D.</emphasis>
Antenna.
<emphasis id="B96CEAACFFA9FFD1FD8F3F5F7D73F9D3" bold="true" box="[549,570,1548,1572]" pageId="47">E.</emphasis>
Thorax, dorsal.
</paragraph>
</caption>
<paragraph id="8BA736BEFFB6FFCEFF2638D47D35FE37" blockId="48.[108,638,226,1738]" pageId="48">Clypeus (face) bulging; vibrissa, genal setae absent. Proboscis relatively broad but short.</paragraph>
<paragraph id="8BA736BEFFB6FFCEFFC6389A7E68FD71" blockId="48.[108,638,226,1738]" pageId="48">
<emphasis id="B96CEAACFFB6FFCEFFC6389A7F9CFE16" bold="true" box="[108,213,457,481]" pageId="48">Antenna:</emphasis>
Scape and pedicel short, ringlike; pedicel with setulae; arista terminal, basal flagellomere ovoid, with sensory/glandular pit and seta on dorsal surface; basal aristomere minute, short; apical aristomere long, not particularly fine, micropubescent.
</paragraph>
<paragraph id="8BA736BEFFB6FFCEFF263BC37CDEFDB3" blockId="48.[108,638,226,1738]" lastBlockId="48.[684,1215,226,878]" pageId="48">
<emphasis id="B96CEAACFFB6FFCEFF263BC37FAAFD5F" bold="true" box="[140,227,656,680]" pageId="48">Thorax:</emphasis>
Short, deep in lateral view, rounded and slightly arched. Acrostichals uniserial, ~
<quantity id="4CE09B5BFFB6FFCEFDE53BE27D35FD3F" box="[591,636,688,712]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="48" unit="in" value="7.0">7 in</quantity>
row; dorsocentrals in pair of rows, ~78 setae each row, anteriormost pair long, fine, ~3× length of posterior dcs, ~ 5 scattered setae lateral to dcs. 1 large supraalar seta; 1 large postalar; 6 scattered notopleural setae. Anepisternum, katepisternum, anepimeron bare. Scutellum short, broad; with two pairs setae, apical pair cruciate; prescutellum present.
<emphasis id="B96CEAACFFB6FFCEFF313AEA7F96FC26" bold="true" box="[155,223,953,977]" pageId="48">Wing:</emphasis>
Relatively elongate, length 2.3× width; darkened between Sc and R
<subScript id="179C34FBFFB6FFCEFE323AB57EE9FC00" attach="left" box="[408,416,998,1015]" fontSize="7" pageId="48">1</subScript>
, with darkened area extending past apex of R
<subScript id="179C34FBFFB6FFCEFEC73D547E3CFBEF" attach="left" box="[365,373,1031,1048]" fontSize="7" pageId="48">1</subScript>
; membrane entirely covered with well-developed microtrichia. Vein C with spinules, longer on portion from apex of R
<subScript id="179C34FBFFB6FFCEFDF03D1A7D2AFBAD" attach="left" box="[602,611,1097,1114]" fontSize="7" pageId="48">1</subScript>
to base; C terminates at apex of R
<subScript id="179C34FBFFB6FFCEFE003D397E8DFB8C" attach="right" box="[426,452,1130,1147]" fontSize="7" pageId="48">4+5</subScript>
, portion between terminus and apex of R
<subScript id="179C34FBFFB6FFCEFED13DD87ECDFB6B" attach="left" box="[379,388,1163,1180]" fontSize="7" pageId="48">1</subScript>
hardly sclerotized. Sc about half the length of R
<subScript id="179C34FBFFB6FFCEFEDD3DFF7E36FB4A" attach="right" box="[375,383,1196,1213]" fontSize="7" pageId="48">1</subScript>
; R
<subScript id="179C34FBFFB6FFCEFE363DFF7EFEFB4A" attach="left" box="[412,439,1196,1213]" fontSize="7" pageId="48">4+5</subScript>
terminates at apex of wing. Cell rm absent; cell bm open apically (bmcu appears incomplete), crossvein dm-cu (cell dm) absent. M
<subScript id="179C34FBFFB6FFCEFF4A3C5C7FA0FAD7" attach="left" box="[224,233,1295,1312]" fontSize="7" pageId="48">1</subScript>
basally incomplete; M
<subScript id="179C34FBFFB6FFCEFE503C5C7D4AFAD7" attach="left" box="[506,515,1295,1312]" fontSize="7" pageId="48">2</subScript>
complete; between apices of M and Cu veins and wing margin with a minute gap. CuA
<subScript id="179C34FBFFB6FFCEFE213C027EDDFA95" attach="left" box="[395,404,1361,1378]" fontSize="7" pageId="48">2</subScript>
short, apically incomplete(?), cell cup small, possibly open; A
<subScript id="179C34FBFFB6FFCEFDB63C217D6CFA74" attach="left" box="[540,549,1394,1411]" fontSize="7" pageId="48">1</subScript>
apically evanescent; anal lobe well developed, alula with long flattened setae.
<emphasis id="B96CEAACFFB6FFCEFEE03CFA7ECAFA36" bold="true" box="[330,387,1449,1473]" pageId="48">Legs:</emphasis>
Coxa laterally attached to thorax; length of femur ≊ tibia on all legs; all podomeres setulose. Largest setae on legs stout, spine like; 2 on dorsal surface mesotibia (at 0.3× length of tibia, plus preapically); 1 dorsally on metatibia and metabasitarsomere, both preapical. Setae and setulae on all tibiae and tarsi in longitudinal rows. Mesotibia and tarsus with 2 ventral, seamlike rows of stout, short setulae. Metatarso- meres 1 and 2 with transverse rows setulae. Claws and pulvilli well developed. Metabasitarsomere barely expanded in lateral view.
<emphasis id="B96CEAACFFB6FFCEFC5738707B3BFECC" bold="true" box="[1021,1138,291,315]" pageId="48">Abdomen:</emphasis>
Abdomen relatively short, stout, distinctly shorter than wings; 8 well-developed tergites, 6 well-developed sternites visible. Tergites with short, scattered setae (no macrosetae), sparser on sternites; sternites nearly equal in size to tergites on same segment. Pleural membrane wrinkled and exposed between lateral margins of abdominal sclerites. Cerci small, 1-segmented, suboval.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB6FFCEFD663B1E7B29FD1D" pageId="48" type="materials_examined">
<paragraph id="8BA736BEFFB6FFCEFD663B1E7B29FD1D" blockId="48.[684,1215,226,878]" pageId="48">SPECIMEN EXAMINED: Female, AMNH Ba-502-14, in Eocene Baltic amber. Specimen is complete and well preserved (the eyes have even retained a reddish color), but portions of the cuticle are obscured by a milky coating.</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB6FFCEFD663BA07C77FC99" pageId="48" type="discussion">
<paragraph id="8BA736BEFFB6FFCEFD663BA07C77FC99" blockId="48.[684,1215,226,878]" pageId="48">COMMENTS: I chose not to name this fossil because I could find no diagnostic characters to separate it from extant species, one reason being it is a female.</paragraph>
</subSubSection>
</treatment>
</document>