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<document id="FC0D327A534A3EEE98A908F05D00771F" ID-DOI="10.5281/zenodo.169460" ID-GBIF-Dataset="c6b6aaea-4e91-4f23-b4bd-c80a4f676e13" ID-ISSN="1175­5326" ID-Zenodo-Dep="169460" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461032056718" checkinUser="plazi" docAuthor="Johanson, Kjell Arne &amp; Holzenthal, Ralph W." docDate="2004" docId="03B1879DFFB5DE4EFE98FA397C2AF9EB" docLanguage="en" docName="zt00711.pdf" docOrigin="Zootaxa 711" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Helicopsyche (Feropsyche) sucrensis Johanson &amp; Holzenthal, 2004, new species" docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="FF88FFE5FFB7DE4AFF90FFD07F67FFB5" masterDocTitle="Thirteen new species and new distribution records of Helicopsyche (Feropsyche) Johanson from Venezuela (Trichoptera: Helicopsychidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="3" updateTime="1698573568728" updateUser="plazi">
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<mods:title id="688B4B06DCE5E9B6EEFE92E44F08AB6D">Thirteen new species and new distribution records of Helicopsyche (Feropsyche) Johanson from Venezuela (Trichoptera: Helicopsychidae)</mods:title>
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<mods:namePart id="3BA864856959477DE4910E50FEE8399B">Johanson, Kjell Arne</mods:namePart>
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<mods:namePart id="D476EACF7D4A8425EF19FECFA42B13F8">Holzenthal, Ralph W.</mods:namePart>
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<mods:date id="5C70487F70FECB6D764A72431DF00C46">2004</mods:date>
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<treatment id="03B1879DFFB5DE4EFE98FA397C2AF9EB" ID-DOI="http://doi.org/10.5281/zenodo.6271550" ID-GBIF-Taxon="119608239" ID-Zenodo-Dep="6271550" LSID="urn:lsid:plazi:treatment:03B1879DFFB5DE4EFE98FA397C2AF9EB" httpUri="http://treatment.plazi.org/id/03B1879DFFB5DE4EFE98FA397C2AF9EB" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="C3026500FFB5DE48FE98FA397EF3F993" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BA7368BFFB5DE48FE98FA397C01F9B6" blockId="2.[264,870,1513,1574]" box="[264,870,1513,1539]" pageId="2" pageNumber="3">
<heading id="D0EF81E7FFB5DE48FE98FA397C01F9B6" bold="true" box="[264,870,1513,1539]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B96CEA99FFB5DE48FE98FA397C01F9B6" bold="true" box="[264,870,1513,1539]" pageId="2" pageNumber="3">
<taxonomicName id="4C184D08FFB5DE48FE98FA397DA0F9B6" box="[264,711,1513,1539]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sucrensis" status="sp. nov." subGenus="Feropsyche">
<emphasis id="B96CEA99FFB5DE48FE98FA397ECEF9B6" bold="true" box="[264,425,1513,1539]" italics="true" pageId="2" pageNumber="3">Helicopsyche</emphasis>
(
<emphasis id="B96CEA99FFB5DE48FE2AFA397D21F9B6" bold="true" box="[442,582,1513,1539]" italics="true" pageId="2" pageNumber="3">Feropsyche</emphasis>
)
<emphasis id="B96CEA99FFB5DE48FDC7FA397DA0F9B6" bold="true" box="[599,711,1513,1539]" italics="true" pageId="2" pageNumber="3">sucrensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E2FFB5DE48FD47FA397C01F9B6" box="[727,870,1513,1539]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7368BFFB5DE48FE98F9DC7EF3F993" blockId="2.[264,870,1513,1574]" box="[264,404,1548,1574]" pageId="2" pageNumber="3">
<figureCitation id="13232A0EFFB5DE48FE98F9DC7E0CF993" box="[264,363,1548,1574]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="2" pageNumber="3">Fig. 17</figureCitation>
,
<figureCitation id="13232A0EFFB5DE48FEE7F9DC7EF3F993" box="[375,404,1548,1574]" captionStart="FIGURE 90" captionStartId="39.[264,367,1237,1261]" captionTargetBox="[276,1318,317,1202]" captionTargetId="figure@39.[264,1323,308,1212]" captionTargetPageId="39" captionText="FIGURE 90. Map of Venezuela showing distributions of H. merida (circle), H. laneblina (rectangle), H. sucrensis (cross), H. breviterga (diamond), and H. circulata (triangle)." httpUri="https://zenodo.org/record/169478/files/figure.png" pageId="2" pageNumber="3">90</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3026500FFB5DE4EFE98F98C7C86FADB" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="8BA7368BFFB5DE49FE98F98C7EF0FE63" blockId="2.[264,1323,1628,1894]" lastBlockId="3.[264,1323,284,470]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
In the male genitalia of the new species, the pointed process at the posterodorsal corner of the gonocoxite is longer and more prominent than in the group of similar American
<taxonomicName id="4C184D08FFB5DE48FB7EF9547E14F970" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEA99FFB5DE48FB7EF9547E14F970" italics="true" pageId="2" pageNumber="3">Helicopsyche</emphasis>
</taxonomicName>
species that includes
<taxonomicName id="4C184D08FFB5DE48FDE7F97C7C2CF973" authority="Flint" authorityName="Flint" box="[631,843,1708,1734]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="apicauda">
<emphasis id="B96CEA99FFB5DE48FDE7F97C7DEBF970" box="[631,652,1708,1733]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FD0CF97C7C6EF970" box="[668,777,1708,1733]" italics="true" pageId="2" pageNumber="3">apicauda</emphasis>
Flint
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FCC8F97C7B4AF973" authority="Johanson" authorityName="Johanson" box="[856,1069,1708,1734]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="flinti">
<emphasis id="B96CEA99FFB5DE48FCC8F97C7C0AF970" box="[856,877,1708,1733]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FCEDF97C7CD3F970" box="[893,948,1708,1733]" italics="true" pageId="2" pageNumber="3">flinti</emphasis>
Johanson
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FBA9F97C7E1AF95B" authority="Flint &amp; Sykora" authorityName="Flint &amp; Sykora" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="grenadensis">
<emphasis id="B96CEA99FFB5DE48FBA9F97C7B29F970" box="[1081,1102,1708,1733]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FBCEF97C7B8DF970" box="[1118,1258,1708,1733]" italics="true" pageId="2" pageNumber="3">grenadensis</emphasis>
Flint &amp; Sykora
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FE19F9047D5FF95B" authority="Ross" authorityName="Ross" box="[393,568,1748,1774]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="incisa">
<emphasis id="B96CEA99FFB5DE48FE19F9047EF9F958" box="[393,414,1748,1773]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FE3DF9047E94F958" box="[429,499,1748,1773]" italics="true" pageId="2" pageNumber="3">incisa</emphasis>
Ross
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FDD4F9047C05F95B" authority="Botosanuanu" authorityName="Botosanuanu" box="[580,866,1748,1774]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kalaom">
<emphasis id="B96CEA99FFB5DE48FDD4F9047D3EF958" box="[580,601,1748,1773]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FDF8F9047DD9F958" box="[616,702,1748,1773]" italics="true" pageId="2" pageNumber="3">kalaom</emphasis>
Botosanuanu
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FCFEF9047B4AF95B" authority="Flint" authorityName="Flint" box="[878,1069,1748,1774]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lambda">
<emphasis id="B96CEA99FFB5DE48FCFEF9047CE4F958" box="[878,899,1748,1773]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FC03F9047C8CF958" box="[915,1003,1748,1773]" italics="true" pageId="2" pageNumber="3">lambda</emphasis>
Flint
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FBA9F9047A65F95B" authority="Ross" authorityName="Ross" box="[1081,1282,1748,1774]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="limnella">
<emphasis id="B96CEA99FFB5DE48FBA9F9047B29F958" box="[1081,1102,1748,1773]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FBCEF9047BD9F958" box="[1118,1214,1748,1773]" italics="true" pageId="2" pageNumber="3">limnella</emphasis>
Ross
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FA9EF9047D3FF8A3" authority="Botosaneanu &amp; Flint" authorityName="Botosaneanu &amp; Flint" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="merida">
<emphasis id="B96CEA99FFB5DE48FA9EF9047A44F958" box="[1294,1315,1748,1773]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FE98F92C7E3CF8A0" box="[264,347,1788,1813]" italics="true" pageId="2" pageNumber="3">merida</emphasis>
Botosaneanu &amp; Flint
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FDF3F92C7CE4F8A3" authority="Botosaneanu" authorityName="Botosaneanu" box="[611,899,1788,1814]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="molesta">
<emphasis id="B96CEA99FFB5DE48FDF3F92C7D1FF8A0" box="[611,632,1788,1813]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FD16F92C7D86F8A0" box="[646,737,1788,1813]" italics="true" pageId="2" pageNumber="3">molesta</emphasis>
Botosaneanu
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FC1DF92C7B26F8A3" authority="Flint" authorityName="Flint" box="[909,1089,1788,1814]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="monda">
<emphasis id="B96CEA99FFB5DE48FC1DF92C7CC5F8A0" box="[909,930,1788,1813]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FC20F92C7B67F8A0" box="[944,1024,1788,1813]" italics="true" pageId="2" pageNumber="3">monda</emphasis>
Flint
</taxonomicName>
,
<taxonomicName id="4C184D08FFB5DE48FBDCF92C7EBDF88B" authority="Botosaneanu &amp; Flint" authorityName="Botosaneanu &amp; Flint" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="paucispina">
<emphasis id="B96CEA99FFB5DE48FBDCF92C7B06F8A0" box="[1100,1121,1788,1813]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FBFFF92C7B97F8A0" box="[1135,1264,1788,1813]" italics="true" pageId="2" pageNumber="3">paucispina</emphasis>
Botosaneanu &amp; Flint
</taxonomicName>
, and
<taxonomicName id="4C184D08FFB5DE48FD8EF8F47D90F88B" authority="Flint." authorityName="Flint." box="[542,759,1828,1854]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="valligera">
<emphasis id="B96CEA99FFB5DE48FD8EF8F47D54F888" box="[542,563,1828,1853]" italics="true" pageId="2" pageNumber="3">H</emphasis>
.
<emphasis id="B96CEA99FFB5DE48FDD5F8F47DC8F888" box="[581,687,1828,1853]" italics="true" pageId="2" pageNumber="3">valligera</emphasis>
Flint.
</taxonomicName>
<taxonomicName id="4C184D08FFB5DE48FC9CF8F47B47F888" box="[780,1056,1828,1853]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sucrensis">
<emphasis id="B96CEA99FFB5DE48FC9CF8F47B47F888" box="[780,1056,1828,1853]" italics="true" pageId="2" pageNumber="3">Helicopsyche sucrensis</emphasis>
</taxonomicName>
can also be separated from these species by the anterior and posterior margins of the gonocoxite that run nearly parallel. It can be further distinguished from
<taxonomicName id="4C184D08FFB4DE49FCBFFECC7CA5FE80" box="[815,962,284,310]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="apicauda">
<emphasis id="B96CEA99FFB4DE49FCBFFECC7C23FE80" box="[815,836,284,309]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FCC5FECC7CA5FE80" box="[853,962,284,309]" italics="true" pageId="3" pageNumber="4">apicauda</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D08FFB4DE49FC43FECC7B57FE80" box="[979,1072,284,310]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="flinti">
<emphasis id="B96CEA99FFB4DE49FC43FECC7C8FFE80" box="[979,1000,284,309]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FC69FECC7B57FE80" box="[1017,1072,284,309]" italics="true" pageId="3" pageNumber="4">flinti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D08FFB4DE49FBDBFECC7B9AFE80" box="[1099,1277,284,310]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="grenadensis">
<emphasis id="B96CEA99FFB4DE49FBDBFECC7B07FE80" box="[1099,1120,284,309]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FBE1FECC7B9AFE80" box="[1137,1277,284,309]" italics="true" pageId="3" pageNumber="4">grenadensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D08FFB4DE49FA9EFECC7E38FEE8" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lambda">
<emphasis id="B96CEA99FFB4DE49FA9EFECC7A44FE80" box="[1294,1315,284,309]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FE98FE947E38FEE8" box="[264,351,324,349]" italics="true" pageId="3" pageNumber="4">lambda</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C184D08FFB4DE49FE33FE947D55FEE8" box="[419,562,324,350]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="valligera">
<emphasis id="B96CEA99FFB4DE49FE33FE947EDFFEE8" box="[419,440,324,349]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FE58FE947D55FEE8" box="[456,562,324,349]" italics="true" pageId="3" pageNumber="4">valligera</emphasis>
</taxonomicName>
by the absence of small lobes along the posterior margin of the gonocoxite. Finally,
<taxonomicName id="4C184D08FFB4DE49FD84FEBC7DCCFE30" box="[532,683,364,389]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sucrensis">
<emphasis id="B96CEA99FFB4DE49FD84FEBC7DCCFE30" box="[532,683,364,389]" italics="true" pageId="3" pageNumber="4">H. sucrensis</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C184D08FFB4DE49FC4CFEBC7B01FE30" box="[988,1126,364,390]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="limnella">
<emphasis id="B96CEA99FFB4DE49FC4CFEBC7C96FE30" box="[988,1009,364,389]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FB96FEBC7B01FE30" box="[1030,1126,364,389]" italics="true" pageId="3" pageNumber="4">limnella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D08FFB4DE49FBEBFEBC7B9EFE30" box="[1147,1273,364,390]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="merida">
<emphasis id="B96CEA99FFB4DE49FBEBFEBC7BF7FE30" box="[1147,1168,364,389]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FB36FEBC7B9EFE30" box="[1190,1273,364,389]" italics="true" pageId="3" pageNumber="4">merida</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D08FFB4DE49FA9EFEBC7E04FE18" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="molesta">
<emphasis id="B96CEA99FFB4DE49FA9EFEBC7A44FE30" box="[1294,1315,364,389]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FE98FE447E04FE18" box="[264,355,404,429]" italics="true" pageId="3" pageNumber="4">molesta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D08FFB4DE49FEE2FE447E82FE18" box="[370,485,404,430]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="monda">
<emphasis id="B96CEA99FFB4DE49FEE2FE447EE0FE18" box="[370,391,404,429]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FE05FE447E82FE18" box="[405,485,404,429]" italics="true" pageId="3" pageNumber="4">monda</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C184D08FFB4DE49FDB5FE447DADFE18" box="[549,714,404,430]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="paucispina">
<emphasis id="B96CEA99FFB4DE49FDB5FE447D5DFE18" box="[549,570,404,429]" italics="true" pageId="3" pageNumber="4">H</emphasis>
.
<emphasis id="B96CEA99FFB4DE49FDD9FE447DADFE18" box="[585,714,404,429]" italics="true" pageId="3" pageNumber="4">paucispina</emphasis>
</taxonomicName>
by the slightly wider anterior lobe of segment IX in lateral view.
</paragraph>
<caption id="DF676603FFB4DE49FE98F9B47C17F974" httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="3" pageNumber="4" targetBox="[270,1298,537,1635]" targetPageId="3">
<paragraph id="8BA7368BFFB4DE49FE98F9B47C17F974" blockId="3.[264,1323,1636,1729]" pageId="3" pageNumber="4">
<emphasis id="B96CEA99FFB4DE49FE98F9B47ED2F9C9" bold="true" box="[264,437,1636,1660]" pageId="3" pageNumber="4">FIGURES 17.</emphasis>
<taxonomicName id="4C184D08FFB4DE49FE56F9B47DA6F9CE" box="[454,705,1636,1659]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sucrensis" status="sp. nov.">
<emphasis id="B96CEA99FFB4DE49FE56F9B47DA6F9CE" box="[454,705,1636,1659]" italics="true" pageId="3" pageNumber="4">Helicopsyche sucrensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E2FFB4DE49FD40F9B47C32F9CE" box="[720,853,1636,1659]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
, holotype. 1—sternum VI process, lateral; 2—sternum VI process, ventral; 3—male genitalia, lateral; 4— male genitalia, dorsal; 5—male genitalia, ventral; 6—phallus, lateral; 7—phallus, ventral.
</paragraph>
</caption>
<paragraph id="8BA7368BFFB4DE4EFEA8F8CC7C86FADB" blockId="3.[264,1323,1820,1886]" lastBlockId="4.[264,1325,284,1630]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Male. Head: Antennal scape slightly shorter than eye diameter. Maxillary palp distal segment slightly shorter than proximal segment and scape. Cephalic warts slightly oval, about 1/2 as long as eye diameter, with pale grey and dark brown setae. Forewing golden brownish, length
<quantity id="4CE09B6EFFB3DE4EFE49FE947D5EFEEB" box="[473,569,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="5" unit="mm" value="3.3">3.3 mm</quantity>
. Sternum VI process (
<figureCitation id="13232A0EFFB3DE4EFCDFFE947CD0FEEB" box="[847,951,324,350]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 1, 2</figureCitation>
) nearly 2/3 its segment length, apparently without microtrichiae; nearly straight in lateral view (
<figureCitation id="13232A0EFFB3DE4EFB9EFEBC7B33FE33" box="[1038,1108,364,390]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
), oriented posteroventrally, tubular along its length, except slightly wider at apex; apex lamellate (
<figureCitation id="13232A0EFFB3DE4EFB54FE447A6CFE1B" box="[1220,1291,404,430]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
). Male genitalia (
<figureCitation id="13232A0EFFB3DE4EFE55FE6C7D4CFE63" box="[453,555,444,470]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 37</figureCitation>
). Segment IX, in lateral view (
<figureCitation id="13232A0EFFB3DE4EFC3CFE6C7C92FE63" box="[940,1013,444,470]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
), with anterior lobe ellipsoid, oriented anteriad, present midlaterally; anterodorsal margin slightly concave; anteroventral margin concave; in dorsal view (
<figureCitation id="13232A0EFFB3DE4EFCA1FDDC7C1CFD93" box="[817,891,524,550]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
), with inner margin uniformly ellipsoid; in ventral view (
<figureCitation id="13232A0EFFB3DE4EFD98FDE47D28FDFB" box="[520,591,564,590]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
), with posterior process midventrally; lateral apodeme present as horizontal, nearly straight line (
<figureCitation id="13232A0EFFB3DE4EFD0DFD8C7D82FDC3" box="[669,741,604,630]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
), tapering anteriorly, diminishing before meeting anterior margin; sub­marginal line present; tergal transverse apodeme apparently absent; sternal transverse apodeme absent. Segment X, in lateral view (
<figureCitation id="13232A0EFFB3DE4EFB9DFD7C7B30FD73" box="[1037,1111,684,710]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
), oriented posteroventrad, nearly straight; dorsally slightly produced; apex truncate; in dorsal view (
<figureCitation id="13232A0EFFB3DE4EFB45FD047A7BFD5B" box="[1237,1308,724,750]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
), parallel along its length, apical part weakly notched; with about 9 pairs of megasetae in lateral group starting midway on segment, the lateral megasetae somewhat shorter than the others. Superior appendage (
<figureCitation id="13232A0EFFB3DE4EFDFBFC9C7DD3FCD3" box="[619,692,844,870]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
) strongly club­shaped, oriented posterolaterad. Primary branch of gonocoxite, in lateral view (
<figureCitation id="13232A0EFFB3DE4EFC82FCA47C3DFC3B" box="[786,858,884,910]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
), generally parallel­sided at midlength, with strongly undulate dorsal margin due to slightly erect setal bases; apex strongly produced posteriorly and curving medially; central part of primary branch slightly wider than height of central part of tergum X (
<figureCitation id="13232A0EFFB3DE4EFD25FC3C7D99FBB3" box="[693,766,1004,1030]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
); anterodorsal margin nearly straight, smooth; posteroventral margin without undulations; basimesal lobe visible in lateral view (
<figureCitation id="13232A0EFFB3DE4EFB43FBC47A7CFB9B" box="[1235,1307,1044,1070]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
), wide, rounded in ventral view (
<figureCitation id="13232A0EFFB3DE4EFDEEFBEC7DA0FBE3" box="[638,711,1084,1110]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
), with weakly undulating median margins, smooth; with about 7 megasetae on dorsal surface; basal plate, in lateral view (
<figureCitation id="13232A0EFFB3DE4EFBE9FBB47BA2FBCB" box="[1145,1221,1124,1150]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
), nearly straight, wide along its length; in ventral view (
<figureCitation id="13232A0EFFB3DE4EFCA8FB5C7CE7FB13" box="[824,896,1164,1190]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
), triangular, apex rounded. Phallus, in lateral view (
<figureCitation id="13232A0EFFB3DE4EFE4BFB647D4BFB7B" box="[475,556,1204,1230]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
), with anterior half nearly straight, sharply bent ventrad at midlength, dorsal margin weakly undulating; ventral margin strongly angled; anterior 1/4 twice as broad as its central part (
<figureCitation id="13232A0EFFB3DE4EFD30FAD47D8DFAAB" box="[672,746,1284,1310]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
); in ventral view (
<figureCitation id="13232A0EFFB3DE4EFC5AFAD47B73FAAB" box="[970,1044,1284,1310]" captionStart="FIGURES 1 7" captionStartId="3.[264,382,1636,1660]" captionTargetBox="[270,1298,537,1635]" captionTargetId="figure@3.[270,1317,529,1635]" captionTargetPageId="3" captionText="FIGURES 1 7. Helicopsyche sucrensis, new species, holotype. 1 — sternum VI process, lateral; 2 — sternum VI process, ventral; 3 — male genitalia, lateral; 4 — male genitalia, dorsal; 5 — male genitalia, ventral; 6 — phallus, lateral; 7 — phallus, ventral." httpUri="https://zenodo.org/record/169461/files/figure.png" pageId="4" pageNumber="5">Fig. 7</figureCitation>
), phallobase present as narrow lobe; endotheca weakly produced; sperm channel divided into thick posterior and slender anterior parts; posteroventral part strongly sclerotized.
</paragraph>
</subSubSection>
<subSubSection id="C3026500FFB3DE4EFEA8FAAC7C2AF9EB" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BA7368BFFB3DE4EFEA8FAAC7DDAFA53" blockId="4.[264,1325,284,1630]" pageId="4" pageNumber="5">
<emphasis id="B96CEA99FFB3DE4EFEA8FAAC7C49FA23" bold="true" box="[312,814,1404,1430]" pageId="4" pageNumber="5">
<typeStatus id="54A38829FFB3DE4EFEA8FAAC7ECDFA23" box="[312,426,1404,1430]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F30F761BFFB3DE4EFD9CFAAC7DA3FA23" box="[524,708,1404,1430]" name="Venezuela" pageId="4" pageNumber="5">VENEZUELA</collectingCountry>
: Sucre:
</emphasis>
Peninsula de Paria, Puerto Viejo, &quot;Rio el Pozo&quot;,
<geoCoordinate id="EE2C504CFFB3DE4EFEFCFA747D64FA0B" box="[364,515,1444,1470]" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="10.717883">10°43.073'N</geoCoordinate>
,
<geoCoordinate id="EE2C504CFFB3DE4EFD8EFA747DDFFA0B" box="[542,696,1444,1470]" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="-62.47615">62°28.569'W</geoCoordinate>
,
<quantity id="4CE09B6EFFB3DE4EFD43FA747C46FA0B" box="[723,801,1444,1470]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="m" value="20.0">20 m</quantity>
,
<date id="FFA6104BFFB3DE4EFCABFA747CCDFA0B" box="[827,938,1444,1470]" pageId="4" pageNumber="5" value="1995-04-03">3.iv.1995</date>
, Holzenthal, Flint &amp; Cressa (
<collectionCode id="ED09AE4EFFB3DE4EFE80FA1C7E06FA53" box="[272,353,1484,1510]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="4" pageNumber="5">UMSP</collectionCode>
000022229) (
<collectionCode id="ED09AE4EFFB3DE4EFE6FFA1C7D37FA53" box="[511,592,1484,1510]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="4" pageNumber="5">UMSP</collectionCode>
, pinned).
</paragraph>
<paragraph id="8BA7368BFFB3DE4EFEA8FA247DDFF9BB" blockId="4.[264,1325,284,1630]" box="[312,696,1524,1550]" pageId="4" pageNumber="5">
Distribution.
<collectingCountry id="F30F761BFFB3DE4EFE4BFA247D34F9BB" box="[475,595,1524,1550]" name="Venezuela" pageId="4" pageNumber="5">Venezuela</collectingCountry>
(Sucre).
</paragraph>
<paragraph id="8BA7368BFFB3DE4EFEA8F9CC7C2AF9EB" blockId="4.[264,1325,284,1630]" pageId="4" pageNumber="5">
Etymology. The name is derived from the
<typeStatus id="54A38829FFB3DE4EFCBEF9CC7C06F983" box="[814,865,1564,1590]" pageId="4" pageNumber="5">type</typeStatus>
locality, the Venezuelan state of Sucre. The name is to be treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>