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<document id="30CE46F9B97071CF0F7F44B4AFC6AF67" ID-DOI="10.11646/zootaxa.5415.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10692602" ID-ZooBank="4981E7AE-37FC-4619-BC55-1F89CB6A0F0B" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1708602063004" checkinUser="plazi" docAuthor="Muller, Tyler A. &amp; Simons, Andrew M." docDate="2024" docId="EF7587ECFFE3FD30FF464FFB13CDF840" docLanguage="en" docName="zootaxa.5415.1.3.pdf" docOrigin="Zootaxa 5415 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5415.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aphredoderus sayanus Gilliams 1824" docType="treatment" docVersion="3" lastPageNumber="84" masterDocId="134CFF94FFE5FD37FFD14F6C1278FF90" masterDocTitle="Taxonomic revision of the Pirate Perches, Aphredoderus, (Percopsiformes: Aphredoderidae) with descriptions of two new species" masterLastPageNumber="105" masterPageNumber="77" pageNumber="83" updateTime="1708957915222" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="4D38D3A01DEF3985B29FED7CD2B4B336">Taxonomic revision of the Pirate Perches, Aphredoderus, (Percopsiformes: Aphredoderidae) with descriptions of two new species</mods:title>
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<mods:namePart id="30B252FE946DFBFC4672AC8DB11B2230">Muller, Tyler A.</mods:namePart>
<mods:affiliation id="EC53D35510E0BDB598475593610B4353">Conservation Sciences Graduate Program, University of Minnesota, St. Paul, MN, USA 55108 &amp; Bell Museum of Natural History, St. Paul, MN, USA 55108</mods:affiliation>
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<mods:namePart id="333499EED3A2605C2E5C24A3A30071D5">Simons, Andrew M.</mods:namePart>
<mods:affiliation id="158B0AACCEF47734AF86A445868A5AD9">Department of Fisheries, Wildlife, and Conservation Biology, St. Paul, MN, USA 55108 &amp; Bell Museum of Natural History, St. Paul, MN, USA 55108</mods:affiliation>
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<mods:title id="6220E40CFAE6AAC3355268538BF73C7A">Zootaxa</mods:title>
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<mods:date id="DFB213C8B21394487E0FB02C0F02A45C">2024</mods:date>
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<treatment id="EF7587ECFFE3FD30FF464FFB13CDF840" ID-DOI="http://doi.org/10.5281/zenodo.10692612" ID-Zenodo-Dep="10692612" LSID="urn:lsid:plazi:treatment:EF7587ECFFE3FD30FF464FFB13CDF840" httpUri="http://treatment.plazi.org/id/EF7587ECFFE3FD30FF464FFB13CDF840" lastPageId="7" lastPageNumber="84" pageId="6" pageNumber="83">
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<paragraph id="676336FAFFE3FD31FF464FFB1027FF21" blockId="6.[151,607,151,249]" box="[151,607,151,178]" pageId="6" pageNumber="83">
<heading id="3C2B8196FFE3FD31FF464FFB1027FF21" bold="true" box="[151,607,151,178]" fontSize="11" level="1" pageId="6" pageNumber="83" reason="1">
<taxonomicName id="A0DC4D79FFE3FD31FF464FFB1027FF21" ID-CoL="FK24" authority="Gilliams 1824" authorityName="Gilliams" authorityYear="1824" box="[151,607,151,178]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="sayanus">
<emphasis id="55A8EAE8FFE3FD31FF464FFB1027FF21" bold="true" box="[151,607,151,178]" pageId="6" pageNumber="83">
<emphasis id="55A8EAE8FFE3FD31FF464FFB13D4FF21" bold="true" box="[151,428,151,178]" italics="true" pageId="6" pageNumber="83">Aphredoderus sayanus</emphasis>
<bibRefCitation id="034D4B0BFFE3FD31FE624FFB1027FF21" author="Gilliams, J." box="[435,607,151,177]" pageId="6" pageNumber="83" pagination="80 - 82" refId="ref18100" refString="Gilliams, J. (1824) Description of a new species of fish of the Linnean genus Perca. Journal of the Academy of Natural Sciences, 4 (Pt 1), 80 - 82." type="journal article" year="1824">Gilliams 1824</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="676336FAFFE3FD31FF464FD713D6FF45" blockId="6.[151,607,151,249]" box="[151,430,187,213]" pageId="6" pageNumber="83">Yellowbelly Pirate Perch</paragraph>
</subSubSection>
<subSubSection id="2FC66571FFE3FD31FF464FB31305FF69" box="[151,381,223,249]" pageId="6" pageNumber="83" type="description">
<paragraph id="676336FAFFE3FD31FF464FB31305FF69" blockId="6.[151,607,151,249]" box="[151,381,223,249]" pageId="6" pageNumber="83">
<figureCitation id="FFE72A7FFFE3FD31FF464FB31280FF69" box="[151,248,223,249]" captionStart="FIGURE 4" captionStartId="6.[152,255,1435,1459]" captionTargetBox="[159,1429,548,1403]" captionTargetId="figure-98@6.[151,1436,540,1412]" captionTargetPageId="6" captionText="FIGURE 4. A) Aphredoderus sayanus, ANSP 12690, lectotype from a fishpond in Philadelphia, Pennsylvania, U.S.A. B) JFBM 50405 A. sayanus from Little Creek, a tributary of the Delaware River in Burlington County, New Jersey, U.S.A. Photo credits: A—Amanda L. Labadie (ANSP), B—Tyler Muller." figureDoi="http://doi.org/10.5281/zenodo.10692610" httpUri="https://zenodo.org/record/10692610/files/figure.png" pageId="6" pageNumber="83">Figure 4</figureCitation>
;
<tableCitation id="2A5E0341FFE3FD31FED54FB3131AFF69" box="[260,354,223,249]" captionStart="TABLE 3" captionStartId="7.[152,245,764,788]" captionTargetPageId="7" captionText="TABLE 3. Meristic character distributions for five Aphredoderus species." pageId="6" pageNumber="83">Tables 3</tableCitation>
,
<tableCitation id="2A5E0341FFE3FD31FEBE4FB31305FF69" box="[367,381,223,249]" captionStart="TABLE 4" captionStartId="8.[152,245,152,176]" captionTargetPageId="8" captionText="TABLE 4. Morphometrics and meristics for A. sayanus." pageId="6" pageNumber="83">4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="2FC66571FFE3FD31FF464E481080FD94" pageId="6" pageNumber="83" type="reference_group">
<paragraph id="676336FAFFE3FD31FF464E48139FFECA" blockId="6.[151,1437,291,408]" pageId="6" pageNumber="83">
<treatmentCitationGroup id="47CC11D4FFE3FD31FF464E48139FFECA" pageId="6" pageNumber="83">
<treatmentCitation id="E67D10EBFFE3FD31FF464E48103BFEAC" author="Gilliams, J." box="[151,579,292,316]" page="82" pageId="6" pageNumber="83" year="1824">
<taxonomicName id="A0DC4D79FFE3FD31FF464E48103BFEAC" authority="Gilliams, 1824: 81 - 82" authorityName="Gilliams" authorityPageNumber="81 - 82" authorityYear="1824" box="[151,579,292,316]" family="Nemipteridae" genus="Scolopsis" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="sayanus">
<emphasis id="55A8EAE8FFE3FD31FF464E481329FEAB" box="[151,337,292,316]" italics="true" pageId="6" pageNumber="83">Scolopsis sayanus</emphasis>
<materialsCitation id="D7B43CA7FFE3FD31FE884E48103BFEAC" box="[345,579,292,316]" collectionCode="ANSP" country="United States of America" county="Philadelphia" location="Fish" municipality="Harrowgate" pageId="6" pageNumber="83" specimenCode="ANSP 12690" specimenCount="1" stateProvince="Pennsylvania" typeStatus="syntype">
<bibRefCitation id="034D4B0BFFE3FD31FE884E48103BFEAC" author="Gilliams, J." box="[345,579,292,316]" pageId="6" pageNumber="83" pagination="80 - 82" refId="ref18100" refString="Gilliams, J. (1824) Description of a new species of fish of the Linnean genus Perca. Journal of the Academy of Natural Sciences, 4 (Pt 1), 80 - 82." type="journal article" year="1824">Gilliams, 1824: 81–82</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="B8678858FFE3FD31FD9F4E4810DDFEAC" box="[590,677,292,316]" pageId="6" pageNumber="83" type="syntype">Syntype</typeStatus>
:
<specimenCode id="377A9E81FFE3FD31FD7E4E481141FEAB" box="[687,825,291,316]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="6" pageNumber="83">ANSP 12690</specimenCode>
,
<quantity id="A0249B1FFFE3FD31FC944E4811D8FEAB" box="[837,928,292,316]" metricMagnitude="-2" metricUnit="m" metricValue="4.91" pageId="6" pageNumber="83" unit="mm" value="49.1">49.1 mm</quantity>
SL,
<location id="62036021FFE3FD31FC044E481679FEAC" LSID="urn:lsid:plazi:treatment:EF7587ECFFE3FD30FF464FFB13CDF840:62036021FFE3FD31FC044E481679FEAC" box="[981,1025,292,316]" country="United States of America" county="Philadelphia" municipality="Harrowgate" name="Fish" pageId="6" pageNumber="83" stateProvince="Pennsylvania">Fish</location>
pond at
<collectingMunicipality id="8707AC80FFE3FD31FB8C4E4816AFFEAC" box="[1117,1239,292,316]" pageId="6" pageNumber="83">Harrowgate</collectingMunicipality>
, near
<collectingCounty id="8E024E76FFE3FD31FAC64E4817E0FEAC" box="[1303,1432,292,316]" pageId="6" pageNumber="83">Philadelphia</collectingCounty>
,
<collectingRegion id="A518F818FFE3FD31FF164E2E1329FECA" box="[199,337,322,346]" country="United States of America" name="Pennsylvania" pageId="6" pageNumber="83">Pennsylvania</collectingRegion>
,
<collectingCountry id="1FCB766AFFE3FD31FE8A4E2E13DBFECA" box="[347,419,322,346]" name="United States of America" pageId="6" pageNumber="83">U.S.A.</collectingCountry>
, 1824.
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</paragraph>
<paragraph id="676336FAFFE3FD31FF464E0D1140FEE9" blockId="6.[151,1437,291,408]" box="[151,824,353,377]" pageId="6" pageNumber="83">
<taxonomicName id="A0DC4D79FFE3FD31FF464E0D13AEFEE8" ID-CoL="FK24" authority="Lesueur" authorityName="Lesueur" box="[151,470,353,377]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="sayanus">
<emphasis id="55A8EAE8FFE3FD31FF464E0D1306FEE9" box="[151,382,353,377]" italics="true" pageId="6" pageNumber="83">Aphredoderus sayanus</emphasis>
Lesueur
</taxonomicName>
in Cuvier and Valenciennes, 1833.
</paragraph>
<paragraph id="676336FAFFE3FD31FF464EEC10A3FE08" blockId="6.[151,1437,291,408]" box="[151,731,384,408]" pageId="6" pageNumber="83">
<taxonomicName id="A0DC4D79FFE3FD31FF464EEC10AFFE08" ID-CoL="FK24" authority="Boltz and Stauffer, 1993" authorityName="Boltz and Stauffer" authorityYear="1993" box="[151,727,384,408]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="6" pageNumber="83" phylum="Chordata" rank="subSpecies" species="sayanus" subSpecies="sayanus">
<emphasis id="55A8EAE8FFE3FD31FF464EEC13AEFE07" box="[151,470,384,408]" italics="true" pageId="6" pageNumber="83">Aphredoderus sayanus sayanus</emphasis>
<bibRefCitation id="034D4B0BFFE3FD31FE0D4EEC10AFFE08" author="Boltz, J. M. &amp; Stauffer, J. R." box="[476,727,384,408]" pageId="6" pageNumber="83" pagination="81" refId="ref17513" refString="Boltz, J. M. &amp; Stauffer, J. R. (1993) Systematics of Aphredoderus sayanus (Teleostei: Aphredoderidae). Copeia, 1993, 81. https: // doi. org / 10.2307 / 1446298" type="journal article" year="1993">Boltz and Stauffer, 1993</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="676336FAFFE3FD31FF464EA91080FD94" blockId="6.[151,1435,453,516]" pageId="6" pageNumber="83">
<materialsCitation id="D7B43CA7FFE3FD31FF464EA91728FE70" box="[151,1360,453,480]" collectionCode="ANSP" country="United States of America" county="Philadelphia" location="Fish" municipality="Harrowgate" pageId="6" pageNumber="83" specimenCode="ANSP 12690" specimenCount="1" stateProvince="Pennsylvania" typeStatus="syntype">
<emphasis id="55A8EAE8FFE3FD31FF464EA91379FE4F" bold="true" box="[151,257,453,479]" pageId="6" pageNumber="83">
<typeStatus id="B8678858FFE3FD31FF464EA91284FE4F" box="[151,252,453,479]" pageId="6" pageNumber="83" type="syntype">Syntype</typeStatus>
:
</emphasis>
<specimenCode id="377A9E81FFE3FD31FED74EAA13E2FE70" box="[262,410,454,480]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="6" pageNumber="83">ANSP 12690</specimenCode>
(1),
<quantity id="A0249B1FFFE3FD31FE014EAA104CFE70" box="[464,564,454,480]" metricMagnitude="-2" metricUnit="m" metricValue="4.91" pageId="6" pageNumber="83" unit="mm" value="49.1">49.1 mm</quantity>
SL,
<location id="62036021FFE3FD31FDB84EAA10E1FE70" LSID="urn:lsid:plazi:treatment:EF7587ECFFE3FD30FF464FFB13CDF840:62036021FFE3FD31FDB84EAA10E1FE70" box="[617,665,454,480]" country="United States of America" county="Philadelphia" municipality="Harrowgate" name="Fish" pageId="6" pageNumber="83" stateProvince="Pennsylvania">Fish</location>
pond at
<collectingMunicipality id="8707AC80FFE3FD31FD2B4EAA11F9FE70" box="[762,897,454,480]" pageId="6" pageNumber="83">Harrowgate</collectingMunicipality>
, near
<collectingCounty id="8E024E76FFE3FD31FC154EAA162BFE70" box="[964,1107,454,480]" pageId="6" pageNumber="83">Philadelphia</collectingCounty>
,
<collectingRegion id="A518F818FFE3FD31FB8F4EAA168EFE70" box="[1118,1270,454,480]" country="United States of America" name="Pennsylvania" pageId="6" pageNumber="83">Pennsylvania</collectingRegion>
,
<collectingCountry id="1FCB766AFFE3FD31FAD04EAA1728FE70" box="[1281,1360,454,480]" name="United States of America" pageId="6" pageNumber="83">U.S.A.</collectingCountry>
</materialsCitation>
, 1824.
<emphasis id="55A8EAE8FFE3FD31FF164E8513FCFD93" bold="true" box="[199,388,489,515]" pageId="6" pageNumber="83">Other material:</emphasis>
(
<specimenCount id="71DAFD73FFE3FD31FE424E861045FD94" box="[403,573,490,516]" count="133" pageId="6" pageNumber="83" type="generic">133 specimens</specimenCount>
,
<quantity id="A0249B1FFFE3FD31FD994E8610B9FD94" box="[584,705,490,516]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" metricValueMax="8.8" metricValueMin="1.9" pageId="6" pageNumber="83" unit="mm" value="53.5" valueMax="88.0" valueMin="19.0">19–88 mm</quantity>
SL).
</paragraph>
</subSubSection>
<caption id="33A36672FFE3FD31FF494AF71089FA6B" ID-DOI="http://doi.org/10.5281/zenodo.10692610" ID-Zenodo-Dep="10692610" httpUri="https://zenodo.org/record/10692610/files/figure.png" pageId="6" pageNumber="83" startId="6.[152,255,1435,1459]" targetBox="[159,1429,548,1403]" targetPageId="6" targetType="figure">
<paragraph id="676336FAFFE3FD31FF494AF71089FA6B" blockId="6.[151,1436,1435,1531]" pageId="6" pageNumber="83">
<emphasis id="55A8EAE8FFE3FD31FF494AF71346FA23" bold="true" box="[152,318,1435,1459]" pageId="6" pageNumber="83">FIGURE 4. A)</emphasis>
<taxonomicName id="A0DC4D79FFE3FD31FE994AF710F8FA23" authority=", ANSP" authorityName="ANSP" box="[328,640,1435,1459]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="sayanus">
<emphasis id="55A8EAE8FFE3FD31FE994AF7104AFA23" box="[328,562,1435,1459]" italics="true" pageId="6" pageNumber="83">Aphredoderus sayanus</emphasis>
, ANSP
</taxonomicName>
12690, lectotype from a fishpond in Philadelphia, Pennsylvania, U.S.A.
<emphasis id="55A8EAE8FFE3FD31FA524AF717E4FA23" bold="true" box="[1411,1436,1435,1459]" pageId="6" pageNumber="83">B)</emphasis>
JFBM 50405
<taxonomicName id="A0DC4D79FFE3FD31FEF24AD313EAFA47" box="[291,402,1471,1495]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="sayanus">
<emphasis id="55A8EAE8FFE3FD31FEF24AD313EAFA47" box="[291,402,1471,1495]" italics="true" pageId="6" pageNumber="83">A. sayanus</emphasis>
</taxonomicName>
from Little Creek, a tributary of the Delaware River in Burlington County, New Jersey, U.S.A. Photo credits: A—Amanda L. Labadie (ANSP), B—Tyler Muller.
</paragraph>
</caption>
<subSubSection id="2FC66571FFE3FD31FF164921102CF8F3" pageId="6" pageNumber="83" type="description">
<paragraph id="676336FAFFE3FD31FF164921102CF8F3" blockId="6.[151,1437,1613,2036]" pageId="6" pageNumber="83">
<emphasis id="55A8EAE8FFE3FD31FF164921133AF9F7" bold="true" box="[199,322,1613,1639]" pageId="6" pageNumber="83">
<collectingRegion id="A518F818FFE3FD31FF1649211344F9F7" box="[199,316,1613,1639]" country="United States of America" name="Delaware" pageId="6" pageNumber="83">Delaware</collectingRegion>
:
</emphasis>
Kent County: UF 20792 (1), UF 24206 (1). Sussex County: JFBM 50004 (12), JFBM 50005 (6).
<emphasis id="55A8EAE8FFE3FD31FF46491D1353F91B" bold="true" box="[151,299,1649,1675]" pageId="6" pageNumber="83">
<collectingRegion id="A518F818FFE3FD31FF46491D135DF91B" box="[151,293,1649,1675]" country="United States of America" name="New Jersey" pageId="6" pageNumber="83">New Jersey</collectingRegion>
:
</emphasis>
Burlington County: JFBM 50207 (2), JFBM 50405 (7), NCSM 107824 (13), NCSM 107885 (12). Cumberland County: JFBM 50202 (7). Middlesex County: NCSM 63655 (2). Monmouth County: NCSM 62836 (12).
<emphasis id="55A8EAE8FFE3FD31FF0549D5132BF943" bold="true" box="[212,339,1721,1747]" pageId="6" pageNumber="83">
<collectingRegion id="A518F818FFE3FD31FF0549D51336F943" box="[212,334,1721,1747]" country="United States of America" name="New York" pageId="6" pageNumber="83">New York</collectingRegion>
:
</emphasis>
Suffolk County: JFBM 50407 (2) JFBM 50408 (2), JFBM 50472 (6).
<emphasis id="55A8EAE8FFE3FD31FBB949D516ACF943" bold="true" box="[1128,1236,1721,1747]" pageId="6" pageNumber="83">
<collectingRegion id="A518F818FFE3FD31FBB949D516B7F943" box="[1128,1231,1721,1747]" country="United States of America" name="Virginia" pageId="6" pageNumber="83">Virginia</collectingRegion>
:
</emphasis>
Caroline County: JFBM 50385 (1), NCSM 60660 (3), NCSM 62041 (2),. NCSM 62042 (1). Essex County: JFBM 50400 (5). King and Queen County: JFBM 46097 (6), 50389 (2), JFBM 50387 (8). Louisa County: JFBM 50382 (2), JFBM 50383 (1). Richmond County: JFBM 50391 (9). Spotsylvania County: NCSM 59063 (5), NCSM 60701 (1), NCSM 61827 (1). Stafford County: NCSM 60594 (1).
</paragraph>
</subSubSection>
<subSubSection id="2FC66571FFE3FD30FF1648011161FE8D" lastPageId="7" lastPageNumber="84" pageId="6" pageNumber="83" type="diagnosis">
<paragraph id="676336FAFFE3FD30FF1648011161FE8D" blockId="6.[151,1437,1613,2036]" lastBlockId="7.[151,1437,151,717]" lastPageId="7" lastPageNumber="84" pageId="6" pageNumber="83">
<emphasis id="55A8EAE8FFE3FD31FF164801133CF817" bold="true" box="[199,324,1901,1927]" pageId="6" pageNumber="83">Diagnosis:</emphasis>
<taxonomicName id="A0DC4D79FFE3FD31FE9D48011035F817" box="[332,589,1901,1927]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="sayanus">
<emphasis id="55A8EAE8FFE3FD31FE9D48011035F817" box="[332,589,1901,1927]" italics="true" pageId="6" pageNumber="83">Aphredoderus sayanus</emphasis>
</taxonomicName>
differs from all
<taxonomicName id="A0DC4D79FFE3FD31FCDA480111D1F817" box="[779,937,1901,1927]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="6" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis id="55A8EAE8FFE3FD31FCDA480111D1F817" box="[779,937,1901,1927]" italics="true" pageId="6" pageNumber="83">Aphredoderus</emphasis>
</taxonomicName>
by the large head size (mean 33.8% SL vs. 31.2%–33.4 %), and strongly contrasting abdominal color vs. slightly contrasting abdominal color. This species is further differentiated from
<taxonomicName id="A0DC4D79FFE3FD31FE1948DA1046F85F" authority="Muller &amp; Simons, 2024" authorityName="Muller &amp; Simons" authorityYear="2024" box="[456,574,1974,2000]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="ornatus" status="sp. nov">
<emphasis id="55A8EAE8FFE3FD31FE1948DA1046F85F" box="[456,574,1974,2000]" italics="true" pageId="6" pageNumber="83">A. ornatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4E9B5793FFE3FD31FD9448DA10EFF840" box="[581,663,1974,2000]" pageId="6" pageNumber="83" rank="species">sp. nov</taxonomicNameLabel>
by the absence of a lateral caudal peduncle stripe and higher caudal peduncle scale counts (mean 33 vs. 29). From remaining
<taxonomicName id="A0DC4D79FFE3FD31FCC648B511CDF863" box="[791,949,2009,2035]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="6" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis id="55A8EAE8FFE3FD31FCC648B511CDF863" box="[791,949,2009,2035]" italics="true" pageId="6" pageNumber="83">Aphredoderus</emphasis>
</taxonomicName>
this species differs by a higher dorsal spine count, four vs. three, occasionally higher anal spine count (almost always three vs. two or three), lower ray counts in pectoral fins (10 vs. 11), and a well-defined yellow pigment underneath the lateral line. From
<taxonomicName id="A0DC4D79FFE2FD30FB024FD71715FF44" box="[1235,1389,187,213]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="mesotrema">
<emphasis id="55A8EAE8FFE2FD30FB024FD71715FF44" box="[1235,1389,187,213]" italics="true" pageId="7" pageNumber="84">A. mesotrema</emphasis>
</taxonomicName>
this species differs by having caudal chevrons vs. a vertical caudal bar and a smaller eye. From
<taxonomicName id="A0DC4D79FFE2FD30FB424FB3176EFF69" authorityName="Lesueur" authorityYear="1833" box="[1171,1302,223,249]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="gibbosus">
<emphasis id="55A8EAE8FFE2FD30FB424FB3176EFF69" box="[1171,1302,223,249]" italics="true" pageId="7" pageNumber="84">A. gibbosus</emphasis>
</taxonomicName>
this species differs by having a larger eye (mean 25% HL vs. 23.9%).
</paragraph>
</subSubSection>
<subSubSection id="2FC66571FFE2FD30FF164E4A104DFD15" pageId="7" pageNumber="84" type="description">
<paragraph id="676336FAFFE2FD30FF164E4A12B2FE65" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="84">
<emphasis id="55A8EAE8FFE2FD30FF164E4A1323FED0" bold="true" box="[199,347,294,320]" pageId="7" pageNumber="84">Description:</emphasis>
Dorsal, IV:11 (IV:10–11). Anal, III:7 (III:6/7). Pectoral 10 (10–11). Lateral scales 44 (42–46) scale rows above lateral line 8 (8–9), caudal peduncle scales 33 (31–36). Deep bodied, 33% maximum body depth in SL and rarely less than 30%. Large head 34% of SL. Postorbital length long, typically over 16% of SL. Deep caudal peduncle, about 15% of SL, particularly in Southern portion of range. Predorsal length average, about 46% of SL. Eye size average, around 25% of head length. Interorbital space average, eye approximately 86% of interorbital size.
</paragraph>
<paragraph id="676336FAFFE2FD30FF164E92104DFD15" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="84">
<emphasis id="55A8EAE8FFE2FD30FF164E92132AFD88" bold="true" box="[199,338,510,536]" pageId="7" pageNumber="84">Coloration:</emphasis>
Dorsally black or brown, color sharply transitions to lighter yellow color on abdomen. Abdominal melanophores usually diffuse or absent leading to yellow or white abdomen. Two black triangular blotches over hypural on caudal peduncle, at times blends to form one large triangle or two connected triangles. Lateral line frequently outlined by light pigment.
</paragraph>
</subSubSection>
<subSubSection id="2FC66571FFE2FD30FF164DE211CEFD5D" pageId="7" pageNumber="84" type="distribution">
<paragraph id="676336FAFFE2FD30FF164DE211CEFD5D" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="84">
<emphasis id="55A8EAE8FFE2FD30FF164DE2131BFD38" bold="true" box="[199,355,654,680]" pageId="7" pageNumber="84">Distribution:</emphasis>
York River, VA north to coastal drainages in South Long Island, NY (
<figureCitation id="FFE72A7FFFE2FD30FB434DE316A0FD39" box="[1170,1240,655,681]" captionStart="FIGURE 9" captionStartId="18.[152,255,1719,1743]" captionTargetBox="[158,1430,752,1689]" captionTargetId="figure-60@18.[158,1430,752,1689]" captionTargetPageId="18" captionText="FIGURE 9. Range and sampling locations of Aphredoderus sayanus, A. gibbosus and A. ornatus specimens used in this study. Sampling locations with black outlines indicate genetic sampling. Type localities from A. sayanus georeferenced to the River Wards section of Philadelphia, PA (pers. comm. Mark Sabaj-Perez, pers. comm. Feb 8, 2022), A. gibbosus to the center of Lake Pontchartrain, LA." figureDoi="http://doi.org/10.5281/zenodo.10692629" httpUri="https://zenodo.org/record/10692629/files/figure.png" pageId="7" pageNumber="84">Fig. 9</figureCitation>
). Occurs with
<taxonomicName id="A0DC4D79FFE2FD30FA554DE31295FD5C" authorityName="Muller &amp; Simons" authorityYear="2024" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="55A8EAE8FFE2FD30FA554DE31295FD5C" italics="true" pageId="7" pageNumber="84">A. ornatus</emphasis>
</taxonomicName>
in the Rappahannock and Potomac River drainages in
<collectingRegion id="A518F818FFE2FD30FC824DDF11CAFD5D" box="[851,946,691,717]" country="United States of America" name="Virginia" pageId="7" pageNumber="84">Virginia</collectingRegion>
.
</paragraph>
</subSubSection>
<caption id="33A36672FFE2FD30FF494D9011C9FC84" box="[152,945,764,788]" pageId="7" pageNumber="84" startId="7.[152,245,764,788]" targetBox="[163,1423,805,1070]" targetIsTable="true" targetPageId="7" targetType="table">
<paragraph id="676336FAFFE2FD30FF494D9011C9FC84" blockId="7.[152,945,764,788]" box="[152,945,764,788]" pageId="7" pageNumber="84">
<emphasis id="55A8EAE8FFE2FD30FF494D901377FC84" bold="true" box="[152,271,764,788]" pageId="7" pageNumber="84">TABLE 3.</emphasis>
Meristic character distributions for five
<taxonomicName id="A0DC4D79FFE2FD30FD6E4D9011C9FC84" box="[703,945,764,788]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="55A8EAE8FFE2FD30FD6E4D90112DFC84" box="[703,853,764,788]" italics="true" pageId="7" pageNumber="84">Aphredoderus</emphasis>
species.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="676336FAFFE2FD30FD034C4917F1FBBE" pageId="7" pageNumber="84">
<table id="15DCC45AFFE202C8FF724C4917F7FBBE" box="[163,1423,805,1070]" gridcols="22" gridrows="7" pageId="7" pageNumber="84" rowsContinueIn="7.[163,1423,1124,1390]">
<tr id="D9EC34B8FFE202C8FF724C4917F7FCAD" box="[163,1423,805,829]" gridrow="0" pageId="7" pageNumber="84">
<th id="9A3D5DC4FFE202C8FF724C4917F7FCAD" box="[163,1423,805,829]" colspan="22" colspanRight="21" gridcol="0" gridrow="0" pageId="7" pageNumber="84">Lateral Scales</th>
</tr>
<tr id="D9EC34B8FFE202C8FF724C2117F7FCF5" box="[163,1423,845,869]" gridrow="1" pageId="7" pageNumber="84" rowspan-0="1">
<td id="9A3D5DC4FFE202C8FE8B4C21130BFCF5" box="[346,371,845,869]" gridcol="1" gridrow="1" pageId="7" pageNumber="84">35</td>
<td id="9A3D5DC4FFE202C8FE5F4C2113DFFCF5" box="[398,423,845,869]" gridcol="2" gridrow="1" pageId="7" pageNumber="84">36</td>
<td id="9A3D5DC4FFE202C8FE154C2113A5FCF5" box="[452,477,845,869]" gridcol="3" gridrow="1" pageId="7" pageNumber="84">37</td>
<td id="9A3D5DC4FFE202C8FE2B4C21106BFCF5" box="[506,531,845,869]" gridcol="4" gridrow="1" pageId="7" pageNumber="84">38</td>
<td id="9A3D5DC4FFE202C8FDE34C211033FCF5" box="[562,587,845,869]" gridcol="5" gridrow="1" pageId="7" pageNumber="84">39</td>
<td id="9A3D5DC4FFE202C8FDBB4C2110FBFCF5" box="[618,643,845,869]" gridcol="6" gridrow="1" pageId="7" pageNumber="84">40</td>
<td id="9A3D5DC4FFE202C8FD734C2110C3FCF5" box="[674,699,845,869]" gridcol="7" gridrow="1" pageId="7" pageNumber="84">41</td>
<td id="9A3D5DC4FFE202C8FD084C21108AFCF5" box="[729,754,845,869]" gridcol="8" gridrow="1" pageId="7" pageNumber="84">42</td>
<td id="9A3D5DC4FFE202C8FCC14C211151FCF5" box="[784,809,845,869]" gridcol="9" gridrow="1" pageId="7" pageNumber="84">43</td>
<td id="9A3D5DC4FFE202C8FC974C211127FCF5" box="[838,863,845,869]" gridcol="10" gridrow="1" pageId="7" pageNumber="84">44</td>
<td id="9A3D5DC4FFE202C8FCAD4C2111EDFCF5" box="[892,917,845,869]" gridcol="11" gridrow="1" pageId="7" pageNumber="84">45</td>
<td id="9A3D5DC4FFE202C8FC614C2111B1FCF5" box="[944,969,845,869]" gridcol="12" gridrow="1" pageId="7" pageNumber="84">46</td>
<td id="9A3D5DC4FFE202C8FC354C211185FCF5" box="[996,1021,845,869]" gridcol="13" gridrow="1" pageId="7" pageNumber="84">47</td>
<td id="9A3D5DC4FFE202C8FBC44C211656FCF5" box="[1045,1070,845,869]" gridcol="14" gridrow="1" pageId="7" pageNumber="84">48</td>
<td id="9A3D5DC4FFE202C8FB944C211626FCF5" box="[1093,1118,845,869]" gridcol="15" gridrow="1" pageId="7" pageNumber="84">49</td>
<td id="9A3D5DC4FFE202C8FBA74C2116F7FCF5" box="[1142,1167,845,869]" gridcol="16" gridrow="1" pageId="7" pageNumber="84">50</td>
<td id="9A3D5DC4FFE202C8FB764C2116B8FCF5" box="[1191,1216,845,869]" gridcol="17" gridrow="1" pageId="7" pageNumber="84">51</td>
<td id="9A3D5DC4FFE202C8FB084C21168AFCF5" box="[1241,1266,845,869]" gridcol="18" gridrow="1" pageId="7" pageNumber="84">52</td>
<td id="9A3D5DC4FFE202C8FADC4C21175EFCF5" box="[1293,1318,845,869]" gridcol="19" gridrow="1" pageId="7" pageNumber="84">53</td>
<td id="9A3D5DC4FFE202C8FA904C211722FCF5" box="[1345,1370,845,869]" gridcol="20" gridrow="1" pageId="7" pageNumber="84">54</td>
<td id="9A3D5DC4FFE202C8FAA74C2117F7FCF5" box="[1398,1423,845,869]" gridcol="21" gridrow="1" pageId="7" pageNumber="84">55</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724C1917F7FC1D" box="[163,1423,885,909]" gridrow="2" pageId="7" pageNumber="84" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-20="1" rowspan-21="1">
<th id="9A3D5DC4FFE202C8FF724C191339FC1D" box="[163,321,885,909]" gridcol="0" gridrow="2" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724C1A1374FC1D" authorityName="Muller &amp; Simons" authorityYear="2024" box="[163,268,886,909]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="55A8EAE8FFE2FD30FF724C1A1374FC1D" box="[163,268,886,909]" italics="true" pageId="7" pageNumber="84">A. ornatus</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FE8B4C19130BFC1D" box="[346,371,885,909]" gridcol="1" gridrow="2" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FE5F4C1913DFFC1D" box="[398,423,885,909]" gridcol="2" gridrow="2" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FE154C1913A5FC1D" box="[452,477,885,909]" gridcol="3" gridrow="2" pageId="7" pageNumber="84">7</td>
<td id="9A3D5DC4FFE202C8FE2B4C19106BFC1D" box="[506,531,885,909]" gridcol="4" gridrow="2" pageId="7" pageNumber="84">13</td>
<td id="9A3D5DC4FFE202C8FDE34C191033FC1D" box="[562,587,885,909]" gridcol="5" gridrow="2" pageId="7" pageNumber="84">17</td>
<td id="9A3D5DC4FFE202C8FDBB4C1910FBFC1D" box="[618,643,885,909]" gridcol="6" gridrow="2" pageId="7" pageNumber="84">35</td>
<td id="9A3D5DC4FFE202C8FD734C1910C3FC1D" box="[674,699,885,909]" gridcol="7" gridrow="2" pageId="7" pageNumber="84">37</td>
<td id="9A3D5DC4FFE202C8FD084C19108AFC1D" box="[729,754,885,909]" gridcol="8" gridrow="2" pageId="7" pageNumber="84">29</td>
<td id="9A3D5DC4FFE202C8FCC14C191151FC1D" box="[784,809,885,909]" gridcol="9" gridrow="2" pageId="7" pageNumber="84">36</td>
<td id="9A3D5DC4FFE202C8FC974C191127FC1D" box="[838,863,885,909]" gridcol="10" gridrow="2" pageId="7" pageNumber="84">21</td>
<td id="9A3D5DC4FFE202C8FCAD4C1911EDFC1D" box="[892,917,885,909]" gridcol="11" gridrow="2" pageId="7" pageNumber="84">9</td>
<td id="9A3D5DC4FFE202C8FC614C1911B1FC1D" box="[944,969,885,909]" gridcol="12" gridrow="2" pageId="7" pageNumber="84">5</td>
<td id="9A3D5DC4FFE202C8FC354C191185FC1D" box="[996,1021,885,909]" gridcol="13" gridrow="2" pageId="7" pageNumber="84">5</td>
<td id="9A3D5DC4FFE202C8FBC44C191656FC1D" box="[1045,1070,885,909]" gridcol="14" gridrow="2" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FB944C191626FC1D" box="[1093,1118,885,909]" gridcol="15" gridrow="2" pageId="7" pageNumber="84">2</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724CF117F7FC26" box="[163,1423,925,950]" gridrow="3" pageId="7" pageNumber="84" rowspan-1="1" rowspan-17="1" rowspan-18="1" rowspan-19="1" rowspan-2="1" rowspan-20="1" rowspan-21="1" rowspan-3="1" rowspan-4="1">
<th id="9A3D5DC4FFE202C8FF724CF11339FC26" box="[163,321,925,950]" gridcol="0" gridrow="3" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724CF21369FC25" box="[163,273,926,949]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="sayanus">
<emphasis id="55A8EAE8FFE2FD30FF724CF21369FC25" box="[163,273,926,949]" italics="true" pageId="7" pageNumber="84">A. sayanus</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FDE34CF11033FC26" box="[562,587,925,950]" gridcol="5" gridrow="3" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FDBB4CF110FBFC26" box="[618,643,925,950]" gridcol="6" gridrow="3" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FD734CF110C3FC26" box="[674,699,925,950]" gridcol="7" gridrow="3" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FD084CF1108AFC26" box="[729,754,925,950]" gridcol="8" gridrow="3" pageId="7" pageNumber="84">15</td>
<td id="9A3D5DC4FFE202C8FCC14CF11151FC26" box="[784,809,925,950]" gridcol="9" gridrow="3" pageId="7" pageNumber="84">21</td>
<td id="9A3D5DC4FFE202C8FC974CF11127FC26" box="[838,863,925,950]" gridcol="10" gridrow="3" pageId="7" pageNumber="84">24</td>
<td id="9A3D5DC4FFE202C8FCAD4CF111EDFC26" box="[892,917,925,950]" gridcol="11" gridrow="3" pageId="7" pageNumber="84">17</td>
<td id="9A3D5DC4FFE202C8FC614CF111B1FC26" box="[944,969,925,950]" gridcol="12" gridrow="3" pageId="7" pageNumber="84">12</td>
<td id="9A3D5DC4FFE202C8FC354CF11185FC26" box="[996,1021,925,950]" gridcol="13" gridrow="3" pageId="7" pageNumber="84">13</td>
<td id="9A3D5DC4FFE202C8FBC44CF11656FC26" box="[1045,1070,925,950]" gridcol="14" gridrow="3" pageId="7" pageNumber="84">6</td>
<td id="9A3D5DC4FFE202C8FB944CF11626FC26" box="[1093,1118,925,950]" gridcol="15" gridrow="3" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FBA74CF116F7FC26" box="[1142,1167,925,950]" gridcol="16" gridrow="3" pageId="7" pageNumber="84">1</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724CAA17F7FC4E" box="[163,1423,966,990]" gridrow="4" pageId="7" pageNumber="84" rowspan-1="1" rowspan-19="1" rowspan-2="1" rowspan-20="1" rowspan-21="1" rowspan-3="1" rowspan-5="1">
<th id="9A3D5DC4FFE202C8FF724CAA1339FC4E" box="[163,321,966,990]" gridcol="0" gridrow="4" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724CAA1339FC4E" authorityName="Muller &amp; Simons" authorityYear="2024" box="[163,321,966,990]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="retrodorsalis">
<emphasis id="55A8EAE8FFE2FD30FF724CAA1339FC4E" box="[163,321,966,990]" italics="true" pageId="7" pageNumber="84">A. retrodorsalis</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FE2B4CAA106BFC4E" box="[506,531,966,990]" gridcol="4" gridrow="4" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FDBB4CAA10FBFC4E" box="[618,643,966,990]" gridcol="6" gridrow="4" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FD734CAA10C3FC4E" box="[674,699,966,990]" gridcol="7" gridrow="4" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FD084CAA108AFC4E" box="[729,754,966,990]" gridcol="8" gridrow="4" pageId="7" pageNumber="84">6</td>
<td id="9A3D5DC4FFE202C8FCC14CAA1151FC4E" box="[784,809,966,990]" gridcol="9" gridrow="4" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FC974CAA1127FC4E" box="[838,863,966,990]" gridcol="10" gridrow="4" pageId="7" pageNumber="84">6</td>
<td id="9A3D5DC4FFE202C8FCAD4CAA11EDFC4E" box="[892,917,966,990]" gridcol="11" gridrow="4" pageId="7" pageNumber="84">9</td>
<td id="9A3D5DC4FFE202C8FC614CAA11B1FC4E" box="[944,969,966,990]" gridcol="12" gridrow="4" pageId="7" pageNumber="84">8</td>
<td id="9A3D5DC4FFE202C8FC354CAA1185FC4E" box="[996,1021,966,990]" gridcol="13" gridrow="4" pageId="7" pageNumber="84">16</td>
<td id="9A3D5DC4FFE202C8FBC44CAA1656FC4E" box="[1045,1070,966,990]" gridcol="14" gridrow="4" pageId="7" pageNumber="84">12</td>
<td id="9A3D5DC4FFE202C8FB944CAA1626FC4E" box="[1093,1118,966,990]" gridcol="15" gridrow="4" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FBA74CAA16F7FC4E" box="[1142,1167,966,990]" gridcol="16" gridrow="4" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FB764CAA16B8FC4E" box="[1191,1216,966,990]" gridcol="17" gridrow="4" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FB084CAA168AFC4E" box="[1241,1266,966,990]" gridcol="18" gridrow="4" pageId="7" pageNumber="84">1</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724C8217F7FB96" box="[163,1423,1006,1030]" gridrow="5" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1" rowspan-21="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="9A3D5DC4FFE202C8FF724C821339FB96" box="[163,321,1006,1030]" gridcol="0" gridrow="5" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724C821361FB96" authorityName="Lesueur" authorityYear="1833" box="[163,281,1006,1030]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="gibbosus">
<emphasis id="55A8EAE8FFE2FD30FF724C821361FB96" box="[163,281,1006,1030]" italics="true" pageId="7" pageNumber="84">A. gibbosus</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FD734C8210C3FB96" box="[674,699,1006,1030]" gridcol="7" gridrow="5" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FD084C82108AFB96" box="[729,754,1006,1030]" gridcol="8" gridrow="5" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FCC14C821151FB96" box="[784,809,1006,1030]" gridcol="9" gridrow="5" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FC974C821127FB96" box="[838,863,1006,1030]" gridcol="10" gridrow="5" pageId="7" pageNumber="84">23</td>
<td id="9A3D5DC4FFE202C8FCAD4C8211EDFB96" box="[892,917,1006,1030]" gridcol="11" gridrow="5" pageId="7" pageNumber="84">20</td>
<td id="9A3D5DC4FFE202C8FC614C8211B1FB96" box="[944,969,1006,1030]" gridcol="12" gridrow="5" pageId="7" pageNumber="84">37</td>
<td id="9A3D5DC4FFE202C8FC354C821185FB96" box="[996,1021,1006,1030]" gridcol="13" gridrow="5" pageId="7" pageNumber="84">51</td>
<td id="9A3D5DC4FFE202C8FBC44C821656FB96" box="[1045,1070,1006,1030]" gridcol="14" gridrow="5" pageId="7" pageNumber="84">34</td>
<td id="9A3D5DC4FFE202C8FB944C821626FB96" box="[1093,1118,1006,1030]" gridcol="15" gridrow="5" pageId="7" pageNumber="84">36</td>
<td id="9A3D5DC4FFE202C8FBA74C8216F7FB96" box="[1142,1167,1006,1030]" gridcol="16" gridrow="5" pageId="7" pageNumber="84">17</td>
<td id="9A3D5DC4FFE202C8FB764C8216B8FB96" box="[1191,1216,1006,1030]" gridcol="17" gridrow="5" pageId="7" pageNumber="84">13</td>
<td id="9A3D5DC4FFE202C8FB084C82168AFB96" box="[1241,1266,1006,1030]" gridcol="18" gridrow="5" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FADC4C82175EFB96" box="[1293,1318,1006,1030]" gridcol="19" gridrow="5" pageId="7" pageNumber="84">5</td>
<td id="9A3D5DC4FFE202C8FA904C821722FB96" box="[1345,1370,1006,1030]" gridcol="20" gridrow="5" pageId="7" pageNumber="84">2</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724B7A17F7FBBE" box="[163,1423,1046,1070]" gridrow="6" pageId="7" pageNumber="84" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="9A3D5DC4FFE202C8FF724B7A1339FBBE" box="[163,321,1046,1070]" gridcol="0" gridrow="6" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724B7B1355FBBE" box="[163,301,1047,1070]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="mesotrema">
<emphasis id="55A8EAE8FFE2FD30FF724B7B1355FBBE" box="[163,301,1047,1070]" italics="true" pageId="7" pageNumber="84">A. mesotrema</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FCC14B7A1151FBBE" box="[784,809,1046,1070]" gridcol="9" gridrow="6" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FC974B7A1127FBBE" box="[838,863,1046,1070]" gridcol="10" gridrow="6" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FCAD4B7A11EDFBBE" box="[892,917,1046,1070]" gridcol="11" gridrow="6" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FC614B7A11B1FBBE" box="[944,969,1046,1070]" gridcol="12" gridrow="6" pageId="7" pageNumber="84">10</td>
<td id="9A3D5DC4FFE202C8FC354B7A1185FBBE" box="[996,1021,1046,1070]" gridcol="13" gridrow="6" pageId="7" pageNumber="84">20</td>
<td id="9A3D5DC4FFE202C8FBC44B7A1656FBBE" box="[1045,1070,1046,1070]" gridcol="14" gridrow="6" pageId="7" pageNumber="84">28</td>
<td id="9A3D5DC4FFE202C8FB944B7A1626FBBE" box="[1093,1118,1046,1070]" gridcol="15" gridrow="6" pageId="7" pageNumber="84">15</td>
<td id="9A3D5DC4FFE202C8FBA74B7A16F7FBBE" box="[1142,1167,1046,1070]" gridcol="16" gridrow="6" pageId="7" pageNumber="84">20</td>
<td id="9A3D5DC4FFE202C8FB764B7A16B8FBBE" box="[1191,1216,1046,1070]" gridcol="17" gridrow="6" pageId="7" pageNumber="84">13</td>
<td id="9A3D5DC4FFE202C8FB084B7A168AFBBE" box="[1241,1266,1046,1070]" gridcol="18" gridrow="6" pageId="7" pageNumber="84">8</td>
<td id="9A3D5DC4FFE202C8FADC4B7A175EFBBE" box="[1293,1318,1046,1070]" gridcol="19" gridrow="6" pageId="7" pageNumber="84">5</td>
<td id="9A3D5DC4FFE202C8FA904B7A1722FBBE" box="[1345,1370,1046,1070]" gridcol="20" gridrow="6" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FAA74B7A17F7FBBE" box="[1398,1423,1046,1070]" gridcol="21" gridrow="6" pageId="7" pageNumber="84">1</td>
</tr>
</table>
</paragraph>
<paragraph id="676336FAFFE2FD30FF204B081726FAFE" pageId="7" pageNumber="84">
<table id="15DCC45AFFE202C8FF724B0817F7FAFE" box="[163,1423,1124,1390]" gridcols="20" gridrows="7" pageId="7" pageNumber="84" rowsContinueFrom="7.[163,1423,805,1070]">
<tr id="D9EC34B8FFE202C8FF724B0817F7FBEC" box="[163,1423,1124,1148]" gridrow="0" pageId="7" pageNumber="84" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-19="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="9A3D5DC4FFE202C8FF724B081035FBEC" box="[163,589,1124,1148]" colspan="5" colspanRight="4" gridcol="0" gridrow="0" pageId="7" pageNumber="84">Scale Rows Above Lateral Line</th>
<th id="9A3D5DC4FFE202C8FCC94B081677FBEC" box="[792,1039,1124,1148]" colspan="5" colspanRight="4" gridcol="8" gridrow="0" pageId="7" pageNumber="84">Caudal Peduncle Scales</th>
<th id="9A3D5DC4FFE202C8FB194B081727FBEC" box="[1224,1375,1124,1148]" colspan="3" colspanRight="2" gridcol="16" gridrow="0" pageId="7" pageNumber="84">Pectoral Rays</th>
</tr>
<tr id="D9EC34B8FFE202C8FF724BE017F7FB34" box="[163,1423,1164,1188]" gridrow="1" pageId="7" pageNumber="84" rowspan-0="1">
<td id="9A3D5DC4FFE202C8FEBC4BE013FEFB34" box="[365,390,1164,1188]" gridcol="1" gridrow="1" pageId="7" pageNumber="84">7</td>
<td id="9A3D5DC4FFE202C8FE744BE013B3FB34" box="[421,459,1164,1188]" gridcol="2" gridrow="1" pageId="7" pageNumber="84">8</td>
<td id="9A3D5DC4FFE202C8FE374BE01074FB34" box="[486,524,1164,1188]" gridcol="3" gridrow="1" pageId="7" pageNumber="84">9</td>
<td id="9A3D5DC4FFE202C8FDF64BE01035FB34" box="[551,589,1164,1188]" gridcol="4" gridrow="1" pageId="7" pageNumber="84">10</td>
<td id="9A3D5DC4FFE202C8FDB64BE01007FB34" box="[615,639,1164,1188]" gridcol="5" gridrow="1" pageId="7" pageNumber="84">11</td>
<td id="9A3D5DC4FFE202C8FD7C4BE010BEFB34" box="[685,710,1164,1188]" gridcol="6" gridrow="1" pageId="7" pageNumber="84">24</td>
<td id="9A3D5DC4FFE202C8FD354BE01085FB34" box="[740,765,1164,1188]" gridcol="7" gridrow="1" pageId="7" pageNumber="84">26</td>
<td id="9A3D5DC4FFE202C8FCC94BE0114DFB34" box="[792,821,1164,1188]" gridcol="8" gridrow="1" pageId="7" pageNumber="84">28</td>
<td id="9A3D5DC4FFE202C8FC804BE01112FB34" box="[849,874,1164,1188]" gridcol="9" gridrow="1" pageId="7" pageNumber="84">30</td>
<td id="9A3D5DC4FFE202C8FC574BE011E7FB34" box="[902,927,1164,1188]" gridcol="10" gridrow="1" pageId="7" pageNumber="84">32</td>
<td id="9A3D5DC4FFE202C8FC6F4BE011AFFB34" box="[958,983,1164,1188]" gridcol="11" gridrow="1" pageId="7" pageNumber="84">34</td>
<td id="9A3D5DC4FFE202C8FC274BE01677FB34" box="[1014,1039,1164,1188]" gridcol="12" gridrow="1" pageId="7" pageNumber="84">36</td>
<td id="9A3D5DC4FFE202C8FBFA4BE0163CFB34" box="[1067,1092,1164,1188]" gridcol="13" gridrow="1" pageId="7" pageNumber="84">38</td>
<td id="9A3D5DC4FFE202C8FB8F4BE0160FFB34" box="[1118,1143,1164,1188]" gridcol="14" gridrow="1" pageId="7" pageNumber="84">40</td>
<td id="9A3D5DC4FFE202C8FB704BE016D6FB34" box="[1185,1198,1164,1188]" gridcol="15" gridrow="1" pageId="7" pageNumber="84">9</td>
<td id="9A3D5DC4FFE202C8FB194BE01696FB34" box="[1224,1262,1164,1188]" gridcol="16" gridrow="1" pageId="7" pageNumber="84">10</td>
<td id="9A3D5DC4FFE202C8FAD74BE01754FB34" box="[1286,1324,1164,1188]" gridcol="17" gridrow="1" pageId="7" pageNumber="84">11</td>
<td id="9A3D5DC4FFE202C8FA944BE01727FB34" box="[1349,1375,1164,1188]" gridcol="18" gridrow="1" pageId="7" pageNumber="84">12</td>
<td id="9A3D5DC4FFE202C8FAA74BE017F7FB34" box="[1398,1423,1164,1188]" gridcol="19" gridrow="1" pageId="7" pageNumber="84">13</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724BD817F7FB5C" box="[163,1423,1204,1228]" gridrow="2" pageId="7" pageNumber="84" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-18="1" rowspan-19="1" rowspan-5="1">
<th id="9A3D5DC4FFE202C8FF724BD81339FB5C" box="[163,321,1204,1228]" gridcol="0" gridrow="2" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724BD91374FB5C" authorityName="Muller &amp; Simons" authorityYear="2024" box="[163,268,1205,1228]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="55A8EAE8FFE2FD30FF724BD91374FB5C" box="[163,268,1205,1228]" italics="true" pageId="7" pageNumber="84">A. ornatus</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FEBC4BD813FEFB5C" box="[365,390,1204,1228]" gridcol="1" gridrow="2" pageId="7" pageNumber="84">39</td>
<td id="9A3D5DC4FFE202C8FE744BD813B3FB5C" box="[421,459,1204,1228]" gridcol="2" gridrow="2" pageId="7" pageNumber="84">215</td>
<td id="9A3D5DC4FFE202C8FE374BD81074FB5C" box="[486,524,1204,1228]" gridcol="3" gridrow="2" pageId="7" pageNumber="84">38</td>
<td id="9A3D5DC4FFE202C8FDF64BD81035FB5C" box="[551,589,1204,1228]" gridcol="4" gridrow="2" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FD7C4BD810BEFB5C" box="[685,710,1204,1228]" gridcol="6" gridrow="2" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FD354BD81085FB5C" box="[740,765,1204,1228]" gridcol="7" gridrow="2" pageId="7" pageNumber="84">25</td>
<td id="9A3D5DC4FFE202C8FCC94BD8114DFB5C" box="[792,821,1204,1228]" gridcol="8" gridrow="2" pageId="7" pageNumber="84">47</td>
<td id="9A3D5DC4FFE202C8FC804BD81112FB5C" box="[849,874,1204,1228]" gridcol="9" gridrow="2" pageId="7" pageNumber="84">50</td>
<td id="9A3D5DC4FFE202C8FC574BD811E7FB5C" box="[902,927,1204,1228]" gridcol="10" gridrow="2" pageId="7" pageNumber="84">14</td>
<td id="9A3D5DC4FFE202C8FC6F4BD811AFFB5C" box="[958,983,1204,1228]" gridcol="11" gridrow="2" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FB704BD816D6FB5C" box="[1185,1198,1204,1228]" gridcol="15" gridrow="2" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FB194BD81696FB5C" box="[1224,1262,1204,1228]" gridcol="16" gridrow="2" pageId="7" pageNumber="84">102</td>
<td id="9A3D5DC4FFE202C8FAD74BD81754FB5C" box="[1286,1324,1204,1228]" gridcol="17" gridrow="2" pageId="7" pageNumber="84">32</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724BB017F7FB65" box="[163,1423,1244,1269]" gridrow="3" pageId="7" pageNumber="84" rowspan-19="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="9A3D5DC4FFE202C8FF724BB01339FB65" box="[163,321,1244,1269]" gridcol="0" gridrow="3" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724BB11369FB64" box="[163,273,1245,1268]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="sayanus">
<emphasis id="55A8EAE8FFE2FD30FF724BB11369FB64" box="[163,273,1245,1268]" italics="true" pageId="7" pageNumber="84">A. sayanus</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FEBC4BB013FEFB65" box="[365,390,1244,1269]" gridcol="1" gridrow="3" pageId="7" pageNumber="84">10</td>
<td id="9A3D5DC4FFE202C8FE744BB013B3FB65" box="[421,459,1244,1269]" gridcol="2" gridrow="3" pageId="7" pageNumber="84">89</td>
<td id="9A3D5DC4FFE202C8FE374BB01074FB65" box="[486,524,1244,1269]" gridcol="3" gridrow="3" pageId="7" pageNumber="84">30</td>
<td id="9A3D5DC4FFE202C8FCC94BB0114DFB65" box="[792,821,1244,1269]" gridcol="8" gridrow="3" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FC804BB01112FB65" box="[849,874,1244,1269]" gridcol="9" gridrow="3" pageId="7" pageNumber="84">13</td>
<td id="9A3D5DC4FFE202C8FC574BB011E7FB65" box="[902,927,1244,1269]" gridcol="10" gridrow="3" pageId="7" pageNumber="84">34</td>
<td id="9A3D5DC4FFE202C8FC6F4BB011AFFB65" box="[958,983,1244,1269]" gridcol="11" gridrow="3" pageId="7" pageNumber="84">22</td>
<td id="9A3D5DC4FFE202C8FC274BB01677FB65" box="[1014,1039,1244,1269]" gridcol="12" gridrow="3" pageId="7" pageNumber="84">15</td>
<td id="9A3D5DC4FFE202C8FBFA4BB0163CFB65" box="[1067,1092,1244,1269]" gridcol="13" gridrow="3" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FB8F4BB0160FFB65" box="[1118,1143,1244,1269]" gridcol="14" gridrow="3" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FB704BB016D6FB65" box="[1185,1198,1244,1269]" gridcol="15" gridrow="3" pageId="7" pageNumber="84">5</td>
<td id="9A3D5DC4FFE202C8FB194BB01696FB65" box="[1224,1262,1244,1269]" gridcol="16" gridrow="3" pageId="7" pageNumber="84">65</td>
<td id="9A3D5DC4FFE202C8FAD74BB01754FB65" box="[1286,1324,1244,1269]" gridcol="17" gridrow="3" pageId="7" pageNumber="84">17</td>
<td id="9A3D5DC4FFE202C8FA944BB01727FB65" box="[1349,1375,1244,1269]" gridcol="18" gridrow="3" pageId="7" pageNumber="84">1</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724A6917F7FA8D" box="[163,1423,1285,1309]" gridrow="4" pageId="7" pageNumber="84" rowspan-1="1" rowspan-15="1" rowspan-19="1" rowspan-6="1" rowspan-7="1">
<th id="9A3D5DC4FFE202C8FF724A691339FA8D" box="[163,321,1285,1309]" gridcol="0" gridrow="4" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724A691339FA8D" authorityName="Muller &amp; Simons" authorityYear="2024" box="[163,321,1285,1309]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="retrodorsalis">
<emphasis id="55A8EAE8FFE2FD30FF724A691339FA8D" box="[163,321,1285,1309]" italics="true" pageId="7" pageNumber="84">A. retrodorsalis</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FE744A6913B3FA8D" box="[421,459,1285,1309]" gridcol="2" gridrow="4" pageId="7" pageNumber="84">19</td>
<td id="9A3D5DC4FFE202C8FE374A691074FA8D" box="[486,524,1285,1309]" gridcol="3" gridrow="4" pageId="7" pageNumber="84">35</td>
<td id="9A3D5DC4FFE202C8FDF64A691035FA8D" box="[551,589,1285,1309]" gridcol="4" gridrow="4" pageId="7" pageNumber="84">19</td>
<td id="9A3D5DC4FFE202C8FDB64A691007FA8D" box="[615,639,1285,1309]" gridcol="5" gridrow="4" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FCC94A69114DFA8D" box="[792,821,1285,1309]" gridcol="8" gridrow="4" pageId="7" pageNumber="84">6</td>
<td id="9A3D5DC4FFE202C8FC804A691112FA8D" box="[849,874,1285,1309]" gridcol="9" gridrow="4" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FC574A6911E7FA8D" box="[902,927,1285,1309]" gridcol="10" gridrow="4" pageId="7" pageNumber="84">9</td>
<td id="9A3D5DC4FFE202C8FC6F4A6911AFFA8D" box="[958,983,1285,1309]" gridcol="11" gridrow="4" pageId="7" pageNumber="84">15</td>
<td id="9A3D5DC4FFE202C8FC274A691677FA8D" box="[1014,1039,1285,1309]" gridcol="12" gridrow="4" pageId="7" pageNumber="84">9</td>
<td id="9A3D5DC4FFE202C8FBFA4A69163CFA8D" box="[1067,1092,1285,1309]" gridcol="13" gridrow="4" pageId="7" pageNumber="84">5</td>
<td id="9A3D5DC4FFE202C8FB8F4A69160FFA8D" box="[1118,1143,1285,1309]" gridcol="14" gridrow="4" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FB194A691696FA8D" box="[1224,1262,1285,1309]" gridcol="16" gridrow="4" pageId="7" pageNumber="84">15</td>
<td id="9A3D5DC4FFE202C8FAD74A691754FA8D" box="[1286,1324,1285,1309]" gridcol="17" gridrow="4" pageId="7" pageNumber="84">33</td>
<td id="9A3D5DC4FFE202C8FA944A691727FA8D" box="[1349,1375,1285,1309]" gridcol="18" gridrow="4" pageId="7" pageNumber="84">1</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724A4117F7FAD5" box="[163,1423,1325,1349]" gridrow="5" pageId="7" pageNumber="84" rowspan-1="1" rowspan-6="1" rowspan-7="1">
<th id="9A3D5DC4FFE202C8FF724A411339FAD5" box="[163,321,1325,1349]" gridcol="0" gridrow="5" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724A411361FAD5" authorityName="Lesueur" authorityYear="1833" box="[163,281,1325,1349]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="gibbosus">
<emphasis id="55A8EAE8FFE2FD30FF724A411361FAD5" box="[163,281,1325,1349]" italics="true" pageId="7" pageNumber="84">A. gibbosus</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FE744A4113B3FAD5" box="[421,459,1325,1349]" gridcol="2" gridrow="5" pageId="7" pageNumber="84">31</td>
<td id="9A3D5DC4FFE202C8FE374A411074FAD5" box="[486,524,1325,1349]" gridcol="3" gridrow="5" pageId="7" pageNumber="84">160</td>
<td id="9A3D5DC4FFE202C8FDF64A411035FAD5" box="[551,589,1325,1349]" gridcol="4" gridrow="5" pageId="7" pageNumber="84">100</td>
<td id="9A3D5DC4FFE202C8FDB64A411007FAD5" box="[615,639,1325,1349]" gridcol="5" gridrow="5" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FCC94A41114DFAD5" box="[792,821,1325,1349]" gridcol="8" gridrow="5" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FC804A411112FAD5" box="[849,874,1325,1349]" gridcol="9" gridrow="5" pageId="7" pageNumber="84">6</td>
<td id="9A3D5DC4FFE202C8FC574A4111E7FAD5" box="[902,927,1325,1349]" gridcol="10" gridrow="5" pageId="7" pageNumber="84">23</td>
<td id="9A3D5DC4FFE202C8FC6F4A4111AFFAD5" box="[958,983,1325,1349]" gridcol="11" gridrow="5" pageId="7" pageNumber="84">58</td>
<td id="9A3D5DC4FFE202C8FC274A411677FAD5" box="[1014,1039,1325,1349]" gridcol="12" gridrow="5" pageId="7" pageNumber="84">51</td>
<td id="9A3D5DC4FFE202C8FBFA4A41163CFAD5" box="[1067,1092,1325,1349]" gridcol="13" gridrow="5" pageId="7" pageNumber="84">29</td>
<td id="9A3D5DC4FFE202C8FB8F4A41160FFAD5" box="[1118,1143,1325,1349]" gridcol="14" gridrow="5" pageId="7" pageNumber="84">9</td>
<td id="9A3D5DC4FFE202C8FB704A4116D6FAD5" box="[1185,1198,1325,1349]" gridcol="15" gridrow="5" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FB194A411696FAD5" box="[1224,1262,1325,1349]" gridcol="16" gridrow="5" pageId="7" pageNumber="84">18</td>
<td id="9A3D5DC4FFE202C8FAD74A411754FAD5" box="[1286,1324,1325,1349]" gridcol="17" gridrow="5" pageId="7" pageNumber="84">104</td>
<td id="9A3D5DC4FFE202C8FA944A411727FAD5" box="[1349,1375,1325,1349]" gridcol="18" gridrow="5" pageId="7" pageNumber="84">40</td>
<td id="9A3D5DC4FFE202C8FAA74A4117F7FAD5" box="[1398,1423,1325,1349]" gridcol="19" gridrow="5" pageId="7" pageNumber="84">1</td>
</tr>
<tr id="D9EC34B8FFE202C8FF724A3917F7FAFE" box="[163,1423,1365,1390]" gridrow="6" pageId="7" pageNumber="84" rowspan-1="1" rowspan-14="1" rowspan-15="1" rowspan-19="1" rowspan-2="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="9A3D5DC4FFE202C8FF724A391339FAFE" box="[163,321,1365,1390]" gridcol="0" gridrow="6" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF724A3A1355FAFD" box="[163,301,1366,1389]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="mesotrema">
<emphasis id="55A8EAE8FFE2FD30FF724A3A1355FAFD" box="[163,301,1366,1389]" italics="true" pageId="7" pageNumber="84">A. mesotrema</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FE374A391074FAFE" box="[486,524,1365,1390]" gridcol="3" gridrow="6" pageId="7" pageNumber="84">51</td>
<td id="9A3D5DC4FFE202C8FDF64A391035FAFE" box="[551,589,1365,1390]" gridcol="4" gridrow="6" pageId="7" pageNumber="84">96</td>
<td id="9A3D5DC4FFE202C8FDB64A391007FAFE" box="[615,639,1365,1390]" gridcol="5" gridrow="6" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FC804A391112FAFE" box="[849,874,1365,1390]" gridcol="9" gridrow="6" pageId="7" pageNumber="84">13</td>
<td id="9A3D5DC4FFE202C8FC574A3911E7FAFE" box="[902,927,1365,1390]" gridcol="10" gridrow="6" pageId="7" pageNumber="84">22</td>
<td id="9A3D5DC4FFE202C8FC6F4A3911AFFAFE" box="[958,983,1365,1390]" gridcol="11" gridrow="6" pageId="7" pageNumber="84">27</td>
<td id="9A3D5DC4FFE202C8FC274A391677FAFE" box="[1014,1039,1365,1390]" gridcol="12" gridrow="6" pageId="7" pageNumber="84">13</td>
<td id="9A3D5DC4FFE202C8FBFA4A39163CFAFE" box="[1067,1092,1365,1390]" gridcol="13" gridrow="6" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FB194A391696FAFE" box="[1224,1262,1365,1390]" gridcol="16" gridrow="6" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FAD74A391754FAFE" box="[1286,1324,1365,1390]" gridcol="17" gridrow="6" pageId="7" pageNumber="84">47</td>
<td id="9A3D5DC4FFE202C8FA944A391727FAFE" box="[1349,1375,1365,1390]" gridcol="18" gridrow="6" pageId="7" pageNumber="84">18</td>
</tr>
</table>
</paragraph>
<paragraph id="676336FAFFE2FD30FEC24AD417F1F90A" pageId="7" pageNumber="84">
<table id="15DCC45AFFE202C8FF724AD417F1F90A" box="[163,1417,1464,1690]" gridcols="16" gridrows="6" pageId="7" pageNumber="84">
<tr id="D9EC34B8FFE202C8FF724AD417F1FA41" box="[163,1417,1464,1489]" gridrow="0" pageId="7" pageNumber="84" rowspan-11="1" rowspan-15="1" rowspan-2="1" rowspan-3="1" rowspan-7="1">
<th id="9A3D5DC4FFE202C8FF724AD413D7FA41" box="[163,431,1464,1489]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="7" pageNumber="84">Dorsal Spines</th>
<th id="9A3D5DC4FFE202C8FD594AD4115CFA41" box="[648,804,1464,1489]" colspan="3" colspanRight="2" gridcol="4" gridrow="0" pageId="7" pageNumber="84">Dorsal Rays</th>
<th id="9A3D5DC4FFE202C8FC7A4AD41645FA41" box="[939,1085,1464,1489]" colspan="3" colspanRight="2" gridcol="8" gridrow="0" pageId="7" pageNumber="84">Anal Spines</th>
<th id="9A3D5DC4FFE202C8FB644AD41737FA41" box="[1205,1359,1464,1489]" colspan="3" colspanRight="2" gridcol="12" gridrow="0" pageId="7" pageNumber="84">Anal Rays</th>
</tr>
<tr id="D9EC34B8FFE202C8FF724A8C17F1FA69" box="[163,1417,1504,1529]" gridrow="1" pageId="7" pageNumber="84" rowspan-0="1">
<td id="9A3D5DC4FFE202C8FE584A8C13D7FA69" box="[393,431,1504,1529]" gridcol="1" gridrow="1" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FE0E4A8C107DFA69" box="[479,517,1504,1529]" gridcol="2" gridrow="1" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FD894A8C101DFA69" box="[600,613,1504,1529]" gridcol="3" gridrow="1" pageId="7" pageNumber="84">9</td>
<td id="9A3D5DC4FFE202C8FD594A8C10D9FA69" box="[648,673,1504,1529]" gridcol="4" gridrow="1" pageId="7" pageNumber="84">10</td>
<td id="9A3D5DC4FFE202C8FD154A8C1092FA69" box="[708,746,1504,1529]" gridcol="5" gridrow="1" pageId="7" pageNumber="84">11</td>
<td id="9A3D5DC4FFE202C8FCDA4A8C115CFA69" box="[779,804,1504,1529]" gridcol="6" gridrow="1" pageId="7" pageNumber="84">12</td>
<td id="9A3D5DC4FFE202C8FC944A8C1126FA69" box="[837,862,1504,1529]" gridcol="7" gridrow="1" pageId="7" pageNumber="84">13</td>
<td id="9A3D5DC4FFE202C8FC7A4A8C11BCFA69" box="[939,964,1504,1529]" gridcol="8" gridrow="1" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FC3B4A8C1668FA69" box="[1002,1040,1504,1529]" gridcol="9" gridrow="1" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FBE14A8C1645FA69" box="[1072,1085,1504,1529]" gridcol="10" gridrow="1" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FB564A8C16ECFA69" box="[1159,1172,1504,1529]" gridcol="11" gridrow="1" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FB644A8C16B6FA69" box="[1205,1230,1504,1529]" gridcol="12" gridrow="1" pageId="7" pageNumber="84">5</td>
<td id="9A3D5DC4FFE202C8FB3B4A8C1768FA69" box="[1258,1296,1504,1529]" gridcol="13" gridrow="1" pageId="7" pageNumber="84">6</td>
<td id="9A3D5DC4FFE202C8FAE74A8C1737FA69" box="[1334,1359,1504,1529]" gridcol="14" gridrow="1" pageId="7" pageNumber="84">7</td>
<td id="9A3D5DC4FFE202C8FAAD4A8C17F1FA69" box="[1404,1417,1504,1529]" gridcol="15" gridrow="1" pageId="7" pageNumber="84">8</td>
</tr>
<tr id="D9EC34B8FFE202C8FF72496517F1F9B1" box="[163,1417,1545,1569]" gridrow="2" pageId="7" pageNumber="84" rowspan-10="1" rowspan-11="1">
<th id="9A3D5DC4FFE202C8FF7249651339F9B1" box="[163,321,1545,1569]" gridcol="0" gridrow="2" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF7249651374F9B1" authorityName="Muller &amp; Simons" authorityYear="2024" box="[163,268,1545,1569]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="55A8EAE8FFE2FD30FF7249651374F9B1" box="[163,268,1545,1569]" italics="true" pageId="7" pageNumber="84">A. ornatus</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FE58496513D7F9B1" box="[393,431,1545,1569]" gridcol="1" gridrow="2" pageId="7" pageNumber="84">44</td>
<td id="9A3D5DC4FFE202C8FE0E4965107DF9B1" box="[479,517,1545,1569]" gridcol="2" gridrow="2" pageId="7" pageNumber="84">262</td>
<td id="9A3D5DC4FFE202C8FD894965101DF9B1" box="[600,613,1545,1569]" gridcol="3" gridrow="2" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FD59496510D9F9B1" box="[648,673,1545,1569]" gridcol="4" gridrow="2" pageId="7" pageNumber="84">94</td>
<td id="9A3D5DC4FFE202C8FD1549651092F9B1" box="[708,746,1545,1569]" gridcol="5" gridrow="2" pageId="7" pageNumber="84">163</td>
<td id="9A3D5DC4FFE202C8FCDA4965115CF9B1" box="[779,804,1545,1569]" gridcol="6" gridrow="2" pageId="7" pageNumber="84">26</td>
<td id="9A3D5DC4FFE202C8FC9449651126F9B1" box="[837,862,1545,1569]" gridcol="7" gridrow="2" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FC7A496511BCF9B1" box="[939,964,1545,1569]" gridcol="8" gridrow="2" pageId="7" pageNumber="84">15</td>
<td id="9A3D5DC4FFE202C8FC3B49651668F9B1" box="[1002,1040,1545,1569]" gridcol="9" gridrow="2" pageId="7" pageNumber="84">298</td>
<td id="9A3D5DC4FFE202C8FB64496516B6F9B1" box="[1205,1230,1545,1569]" gridcol="12" gridrow="2" pageId="7" pageNumber="84">19</td>
<td id="9A3D5DC4FFE202C8FB3B49651768F9B1" box="[1258,1296,1545,1569]" gridcol="13" gridrow="2" pageId="7" pageNumber="84">204</td>
<td id="9A3D5DC4FFE202C8FAE749651737F9B1" box="[1334,1359,1545,1569]" gridcol="14" gridrow="2" pageId="7" pageNumber="84">74</td>
<td id="9A3D5DC4FFE202C8FAAD496517F1F9B1" box="[1404,1417,1545,1569]" gridcol="15" gridrow="2" pageId="7" pageNumber="84">1</td>
</tr>
<tr id="D9EC34B8FFE202C8FF72495D17F1F9D9" box="[163,1417,1585,1609]" gridrow="3" pageId="7" pageNumber="84" rowspan-11="1" rowspan-15="1" rowspan-7="1">
<th id="9A3D5DC4FFE202C8FF72495D1339F9D9" box="[163,321,1585,1609]" gridcol="0" gridrow="3" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF72495D1369F9D9" box="[163,273,1585,1609]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="sayanus">
<emphasis id="55A8EAE8FFE2FD30FF72495D1369F9D9" box="[163,273,1585,1609]" italics="true" pageId="7" pageNumber="84">A. sayanus</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FE58495D13D7F9D9" box="[393,431,1585,1609]" gridcol="1" gridrow="3" pageId="7" pageNumber="84">14</td>
<td id="9A3D5DC4FFE202C8FE0E495D107DF9D9" box="[479,517,1585,1609]" gridcol="2" gridrow="3" pageId="7" pageNumber="84">116</td>
<td id="9A3D5DC4FFE202C8FD89495D101DF9D9" box="[600,613,1585,1609]" gridcol="3" gridrow="3" pageId="7" pageNumber="84">2</td>
<td id="9A3D5DC4FFE202C8FD59495D10D9F9D9" box="[648,673,1585,1609]" gridcol="4" gridrow="3" pageId="7" pageNumber="84">35</td>
<td id="9A3D5DC4FFE202C8FD15495D1092F9D9" box="[708,746,1585,1609]" gridcol="5" gridrow="3" pageId="7" pageNumber="84">78</td>
<td id="9A3D5DC4FFE202C8FCDA495D115CF9D9" box="[779,804,1585,1609]" gridcol="6" gridrow="3" pageId="7" pageNumber="84">14</td>
<td id="9A3D5DC4FFE202C8FC7A495D11BCF9D9" box="[939,964,1585,1609]" gridcol="8" gridrow="3" pageId="7" pageNumber="84">15</td>
<td id="9A3D5DC4FFE202C8FC3B495D1668F9D9" box="[1002,1040,1585,1609]" gridcol="9" gridrow="3" pageId="7" pageNumber="84">113</td>
<td id="9A3D5DC4FFE202C8FBE1495D1645F9D9" box="[1072,1085,1585,1609]" gridcol="10" gridrow="3" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FB64495D16B6F9D9" box="[1205,1230,1585,1609]" gridcol="12" gridrow="3" pageId="7" pageNumber="84">3</td>
<td id="9A3D5DC4FFE202C8FB3B495D1768F9D9" box="[1258,1296,1585,1609]" gridcol="13" gridrow="3" pageId="7" pageNumber="84">62</td>
<td id="9A3D5DC4FFE202C8FAE7495D1737F9D9" box="[1334,1359,1585,1609]" gridcol="14" gridrow="3" pageId="7" pageNumber="84">65</td>
</tr>
<tr id="D9EC34B8FFE202C8FF72493517F1F9E1" box="[163,1417,1625,1649]" gridrow="4" pageId="7" pageNumber="84" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-3="1" rowspan-7="1">
<th id="9A3D5DC4FFE202C8FF7249351339F9E1" box="[163,321,1625,1649]" gridcol="0" gridrow="4" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF7249361339F9E1" authorityName="Muller &amp; Simons" authorityYear="2024" box="[163,321,1625,1649]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="retrodorsalis">
<emphasis id="55A8EAE8FFE2FD30FF7249361339F9E1" box="[163,321,1625,1649]" italics="true" pageId="7" pageNumber="84">A. retrodorsalis</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FE58493513D7F9E1" box="[393,431,1625,1649]" gridcol="1" gridrow="4" pageId="7" pageNumber="84">76</td>
<td id="9A3D5DC4FFE202C8FE0E4935107DF9E1" box="[479,517,1625,1649]" gridcol="2" gridrow="4" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FD59493510D9F9E1" box="[648,673,1625,1649]" gridcol="4" gridrow="4" pageId="7" pageNumber="84">5</td>
<td id="9A3D5DC4FFE202C8FD1549351092F9E1" box="[708,746,1625,1649]" gridcol="5" gridrow="4" pageId="7" pageNumber="84">59</td>
<td id="9A3D5DC4FFE202C8FCDA4935115CF9E1" box="[779,804,1625,1649]" gridcol="6" gridrow="4" pageId="7" pageNumber="84">11</td>
<td id="9A3D5DC4FFE202C8FC7A493511BCF9E1" box="[939,964,1625,1649]" gridcol="8" gridrow="4" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FC3B49351668F9E1" box="[1002,1040,1625,1649]" gridcol="9" gridrow="4" pageId="7" pageNumber="84">72</td>
<td id="9A3D5DC4FFE202C8FB64493516B6F9E1" box="[1205,1230,1625,1649]" gridcol="12" gridrow="4" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FB3B49351768F9E1" box="[1258,1296,1625,1649]" gridcol="13" gridrow="4" pageId="7" pageNumber="84">48</td>
<td id="9A3D5DC4FFE202C8FAE749351737F9E1" box="[1334,1359,1625,1649]" gridcol="14" gridrow="4" pageId="7" pageNumber="84">27</td>
</tr>
<tr id="D9EC34B8FFE202C8FF7249EE17F1F90A" box="[163,1417,1666,1690]" gridrow="5" pageId="7" pageNumber="84" rowspan-10="1" rowspan-7="1">
<th id="9A3D5DC4FFE202C8FF7249EE1339F90A" box="[163,321,1666,1690]" gridcol="0" gridrow="5" pageId="7" pageNumber="84">
<taxonomicName id="A0DC4D79FFE2FD30FF7249EE1355F909" box="[163,301,1666,1689]" family="Aphredoderidae" genus="Aphredoderus" kingdom="Animalia" order="Percopsiformes" pageId="7" pageNumber="84" phylum="Chordata" rank="species" species="mesotrema">
<emphasis id="55A8EAE8FFE2FD30FF7249EE1355F909" box="[163,301,1666,1689]" italics="true" pageId="7" pageNumber="84">A. mesotrema</emphasis>
</taxonomicName>
</th>
<td id="9A3D5DC4FFE202C8FE5849EE13D7F90A" box="[393,431,1666,1690]" gridcol="1" gridrow="5" pageId="7" pageNumber="84">169</td>
<td id="9A3D5DC4FFE202C8FE0E49EE107DF90A" box="[479,517,1666,1690]" gridcol="2" gridrow="5" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FD8949EE101DF90A" box="[600,613,1666,1690]" gridcol="3" gridrow="5" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FD5949EE10D9F90A" box="[648,673,1666,1690]" gridcol="4" gridrow="5" pageId="7" pageNumber="84">20</td>
<td id="9A3D5DC4FFE202C8FD1549EE1092F90A" box="[708,746,1666,1690]" gridcol="5" gridrow="5" pageId="7" pageNumber="84">112</td>
<td id="9A3D5DC4FFE202C8FCDA49EE115CF90A" box="[779,804,1666,1690]" gridcol="6" gridrow="5" pageId="7" pageNumber="84">23</td>
<td id="9A3D5DC4FFE202C8FC7A49EE11BCF90A" box="[939,964,1666,1690]" gridcol="8" gridrow="5" pageId="7" pageNumber="84">47</td>
<td id="9A3D5DC4FFE202C8FC3B49EE1668F90A" box="[1002,1040,1666,1690]" gridcol="9" gridrow="5" pageId="7" pageNumber="84">169</td>
<td id="9A3D5DC4FFE202C8FB5649EE16ECF90A" box="[1159,1172,1666,1690]" gridcol="11" gridrow="5" pageId="7" pageNumber="84">1</td>
<td id="9A3D5DC4FFE202C8FB6449EE16B6F90A" box="[1205,1230,1666,1690]" gridcol="12" gridrow="5" pageId="7" pageNumber="84">4</td>
<td id="9A3D5DC4FFE202C8FB3B49EE1768F90A" box="[1258,1296,1666,1690]" gridcol="13" gridrow="5" pageId="7" pageNumber="84">111</td>
<td id="9A3D5DC4FFE202C8FAE749EE1737F90A" box="[1334,1359,1666,1690]" gridcol="14" gridrow="5" pageId="7" pageNumber="84">46</td>
<td id="9A3D5DC4FFE202C8FAAD49EE17F1F90A" box="[1404,1417,1666,1690]" gridcol="15" gridrow="5" pageId="7" pageNumber="84">1</td>
</tr>
</table>
</paragraph>
<subSubSection id="2FC66571FFE2FD30FF16486F13CDF840" pageId="7" pageNumber="84" type="discussion">
<paragraph id="676336FAFFE2FD30FF16486F13CDF840" blockId="7.[151,1436,1794,2001]" pageId="7" pageNumber="84">
<emphasis id="55A8EAE8FFE2FD30FF16486F1345F88D" bold="true" box="[199,317,1795,1821]" pageId="7" pageNumber="84">Remarks:</emphasis>
We examined a photograph and radiograph of
<typeStatus id="B8678858FFE2FD30FC90486F11E0F88D" box="[833,920,1795,1821]" pageId="7" pageNumber="84" type="syntype">syntype</typeStatus>
ANSP 12690. The specimen is bent, deformed anterior to the dorsal fin, and the abdomen of the fish has been removed. Gilliams noted a yellow underside of the specimen and made no mention of a black lateral stripe on the caudal peduncle. We refer mainly to Gilliams’ pigment description of the specimen, except for our count of 3 anal spines rather than 2 using a radiograph. Specimens from the type locality are presumed extirpated. We included
<specimenCount id="71DAFD73FFE2FD30FD2C48FF11EDF83C" box="[765,917,1938,1965]" count="34" pageId="7" pageNumber="84" type="generic">34 specimens</specimenCount>
from the
<collectingRegion id="A518F818FFE2FD30FBD048FE1614F83C" box="[1025,1132,1938,1964]" country="United States of America" name="Delaware" pageId="7" pageNumber="84">Delaware</collectingRegion>
River in
<collectingRegion id="A518F818FFE2FD30FB0548FF172DF83D" box="[1236,1365,1939,1965]" country="United States of America" name="New Jersey" pageId="7" pageNumber="84">New Jersey</collectingRegion>
as the closest extant population.
</paragraph>
</subSubSection>
</treatment>
</document>
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