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<document id="87B9E97918B4C734FF25D724F39AE581" ID-CLB-Dataset="20929" ID-DOI="10.21248/contrib.entomol.69.2.239-289" ID-GBIF-Dataset="37a37fe7-21ad-45a4-b0d9-880a2b03aa6f" ID-ISSN="0005-805X" ID-Zenodo-Dep="4755240" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620848784293" checkinUser="carolina" docAuthor="Assing, Volker, Brachat, Volker &amp; Meybohm, Heinrich" docDate="2019" docId="C4053B5AFFDDC915FF708D2982E4752D" docLanguage="en" docName="ContribEntomol.69.2.239-289.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 69 (2)" docSource="http://dx.doi.org/10.21248/contrib.entomol.69.2.239-289" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Cretotyphlus ASSING 2019, gen. nov." docType="treatment" docUuid="FD86523B-3B04-43D4-BD1B-40BC18EAD15F" docUuidSource="ZooBank" docVersion="10" lastPageNumber="263" masterDocId="383C4322FFC5C90DFF898E638163771A" masterDocTitle="Monograph of the Staphylinidae of Crete (Greece). Part II. Descriptions of new species (Insecta: Coleoptera)" masterLastPageNumber="289" masterPageNumber="239" pageNumber="263" updateTime="1698939163438" updateUser="ExternalLinkService">
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<mods:title id="3B828BA902B446A8CBBB08674CAA3FF8">Monograph of the Staphylinidae of Crete (Greece). Part II. Descriptions of new species (Insecta: Coleoptera)</mods:title>
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<mods:namePart id="1403D8B18374953FF006D51EA81E1C7F">Assing, Volker</mods:namePart>
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<mods:namePart id="A14D8986CA2842B7EBE86A19B1DB472B">Brachat, Volker</mods:namePart>
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<treatment id="C4053B5AFFDDC915FF708D2982E4752D" ID-DOI="http://doi.org/10.5281/zenodo.5911500" ID-GBIF-Taxon="192700063" ID-Zenodo-Dep="5911500" LSID="urn:lsid:zoobank.org:act:FD86523B-3B04-43D4-BD1B-40BC18EAD15F" httpUri="http://treatment.plazi.org/id/C4053B5AFFDDC915FF708D2982E4752D" lastPageNumber="263" pageId="24" pageNumber="263">
<subSubSection id="04B6D9C7FFDDC915FF708D298357747F" box="[249,564,842,869]" pageId="24" pageNumber="263" type="nomenclature">
<paragraph id="4C138A4CFFDDC915FF708D298357747F" blockId="24.[249,564,842,869]" box="[249,564,842,869]" pageId="24" pageNumber="263">
<heading id="175B3D20FFDDC915FF708D298357747F" box="[249,564,842,869]" centered="true" fontSize="12" level="2" pageId="24" pageNumber="263" reason="2">
<taxonomicName id="8BACF1CFFFDDC915FF708D2980AE747F" ID-CoL="8XKML" authority="ASSING" authorityName="ASSING" authorityYear="2019" box="[249,461,842,869]" class="Insecta" family="Staphylinidae" genus="Cretotyphlus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="7ED8565EFFDDC915FF708D298018747F" box="[249,379,842,869]" italics="true" pageId="24" pageNumber="263">Cretotyphlus</emphasis>
ASSING
</taxonomicName>
<taxonomicNameLabel id="65EBEB25FFDDC915FE5A8D298357747F" box="[467,564,842,869]" pageId="24" pageNumber="263" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="04B6D9C7FFDDC915FF5D8DE380C874A2" pageId="24" pageNumber="263" type="type_taxon">
<paragraph id="4C138A4CFFDDC915FF5D8DE380C874A2" blockId="24.[212,816,895,952]" pageId="24" pageNumber="263">
<emphasis id="7ED8565EFFDDC915FF5D8DE38001748D" bold="true" box="[212,354,895,920]" pageId="24" pageNumber="263">
<typeStatus id="931734EEFFDDC915FF5D8DE380697482" box="[212,266,896,920]" pageId="24" pageNumber="263">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="8BACF1CFFFDDC915FEFA8DE383DB7482" ID-CoL="8XYVF" authority="ASSING" authorityName="Assing &amp; Brachat &amp; Meybohm" authorityYear="2019" box="[371,696,896,920]" class="Insecta" family="Staphylinidae" genus="Cretotyphlus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="7ED8565EFFDDC915FEFA8DE3833D7482" box="[371,606,896,920]" italics="true" pageId="24" pageNumber="263">Cretotyphlus hamatus</emphasis>
ASSING
</taxonomicName>
<taxonomicNameLabel id="65EBEB25FFDDC915FD4A8DE3824E748D" box="[707,813,896,919]" pageId="24" pageNumber="263" rank="species">spec. nov.</taxonomicNameLabel>
; present designation.
</paragraph>
</subSubSection>
<subSubSection id="04B6D9C7FFDDC915FF5D8D83826B7302" pageId="24" pageNumber="263" type="etymology">
<paragraph id="4C138A4CFFDDC915FF5D8D83826B7302" blockId="24.[212,816,992,1048]" pageId="24" pageNumber="263">
<emphasis id="7ED8565EFFDDC915FF5D8D83802874E2" bold="true" box="[212,331,992,1016]" pageId="24" pageNumber="263">Etymology</emphasis>
: The name (gender: masculine) is composed of
<collectingRegion id="8E6844AEFFDDC915FF668A6380497302" box="[239,298,1024,1048]" country="Greece" name="Kriti" pageId="24" pageNumber="263">Creta</collectingRegion>
and typhlus (from Greek typhlos: the blind).
</paragraph>
</subSubSection>
<subSubSection id="04B6D9C7FFDDC915FF5D8A5C84F176AD" pageId="24" pageNumber="263" type="description">
<paragraph id="4C138A4CFFDDC915FF5D8A5C80C372C2" blockId="24.[212,817,1087,1816]" pageId="24" pageNumber="263">
<emphasis id="7ED8565EFFDDC915FF5D8A5C8035734D" bold="true" box="[212,342,1087,1111]" pageId="24" pageNumber="263">Description</emphasis>
: Species of intermediate to relatively “large” size. Antenna 11-jointed and moderately incrassate, apical antennomeres without sensorial appendages, the apical three antennomeres larger, longer, forming a club, these antennomeres with dense long pubescence. Mandibles apically bifid. Gular sutures close, but not contiguous anteriorly (
<figureCitation id="D49796C9FFDDC915FE468B63837C7202" box="[463,543,1280,1304]" captionStart="Figs 7184" captionStartId="10.[212,252,2003,2025]" captionTargetBox="[217,1454,255,1986]" captionTargetId="figure-41@10.[212,1459,255,1994]" captionTargetPageId="10" captionText="Figs 7184: Kenotyphlus virgatus (7173), Cretotyphlus hamatus (7480), and C. idanus (8184): male sternite VIII (71, 77); aedeagus in lateral and in ventral, or in latero-ventral view (7273, 7879, 8384); habitus (74, 81); forebody in dorsal view (75); forebodyin ventral view (76); femalegenital segment (80);male meso-and metathorax and abdomen in ventralview (82). Scale bars: 74, 8182: 0.5 mm; 7173, 7580, 8384: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755254" httpUri="https://zenodo.org/record/4755254/files/figure.png" pageId="24" pageNumber="263">Figs 76</figureCitation>
,
<figureCitation id="D49796C9FFDDC915FDA78B6383287202" box="[558,587,1280,1304]" captionStart="Figs 8599" captionStartId="11.[174,214,2005,2027]" captionTargetBox="[186,1413,263,1983]" captionTargetId="figure-49@11.[174,1421,255,1994]" captionTargetPageId="11" captionText="Figs 8599: Cretotyphlus idanus (85), C. chanianus (8691), Pseudobium creticum (9296), and Gabrius candicus (9799): male sternite VIII (85, 88, 94); habitus (86, 92, 97); forebody in ventral view (87); aedeagus in lateral and in ventral view (8990, 9596, 9899); female genital segments (91); forebody (93). Scale bars: 9293, 97: 1.0 mm; 86: 0.5 mm; 9899: 0.2 mm; 85, 8791, 9496: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755258" httpUri="https://zenodo.org/record/4755258/files/figure.png" pageId="24" pageNumber="263">87</figureCitation>
). Labrum tridentate, i.e., with a pronounced median tooth and a smaller tooth on either side. Maxillary palpi with palpomeres II and III enlarged and somewhat flattened, III larger than II. Abdominal sternites without transverse sulcus at base (
<figureCitation id="D49796C9FFDDC915FF528BC3804B72A2" box="[219,296,1440,1464]" captionStart="Figs 7184" captionStartId="10.[212,252,2003,2025]" captionTargetBox="[217,1454,255,1986]" captionTargetId="figure-41@10.[212,1459,255,1994]" captionTargetPageId="10" captionText="Figs 7184: Kenotyphlus virgatus (7173), Cretotyphlus hamatus (7480), and C. idanus (8184): male sternite VIII (71, 77); aedeagus in lateral and in ventral, or in latero-ventral view (7273, 7879, 8384); habitus (74, 81); forebody in dorsal view (75); forebodyin ventral view (76); femalegenital segment (80);male meso-and metathorax and abdomen in ventralview (82). Scale bars: 74, 8182: 0.5 mm; 7173, 7580, 8384: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755254" httpUri="https://zenodo.org/record/4755254/files/figure.png" pageId="24" pageNumber="263">Fig. 82</figureCitation>
). Procoxal cavities without suture anteriorly. Tarsi three-jointed.
</paragraph>
<paragraph id="4C138A4CFFDDC915FF5D8B8283AA71AD" blockId="24.[212,817,1087,1816]" pageId="24" pageNumber="263">
♂: posterior sternites (VVIII or VIVIII) each with extensive median impression, these impressions gradually becoming more pronounced towards sternite VIII (
<figureCitation id="D49796C9FFDDC915FF528823804B7142" box="[219,296,1600,1624]" captionStart="Figs 7184" captionStartId="10.[212,252,2003,2025]" captionTargetBox="[217,1454,255,1986]" captionTargetId="figure-41@10.[212,1459,255,1994]" captionTargetPageId="10" captionText="Figs 7184: Kenotyphlus virgatus (7173), Cretotyphlus hamatus (7480), and C. idanus (8184): male sternite VIII (71, 77); aedeagus in lateral and in ventral, or in latero-ventral view (7273, 7879, 8384); habitus (74, 81); forebody in dorsal view (75); forebodyin ventral view (76); femalegenital segment (80);male meso-and metathorax and abdomen in ventralview (82). Scale bars: 74, 8182: 0.5 mm; 7173, 7580, 8384: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755254" httpUri="https://zenodo.org/record/4755254/files/figure.png" pageId="24" pageNumber="263">Fig. 82</figureCitation>
); sternite VIII (
<figureCitation id="D49796C9FFDDC915FE5288238349714D" box="[475,554,1600,1624]" captionStart="Figs 7184" captionStartId="10.[212,252,2003,2025]" captionTargetBox="[217,1454,255,1986]" captionTargetId="figure-41@10.[212,1459,255,1994]" captionTargetPageId="10" captionText="Figs 7184: Kenotyphlus virgatus (7173), Cretotyphlus hamatus (7480), and C. idanus (8184): male sternite VIII (71, 77); aedeagus in lateral and in ventral, or in latero-ventral view (7273, 7879, 8384); habitus (74, 81); forebody in dorsal view (75); forebodyin ventral view (76); femalegenital segment (80);male meso-and metathorax and abdomen in ventralview (82). Scale bars: 74, 8182: 0.5 mm; 7173, 7580, 8384: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755254" httpUri="https://zenodo.org/record/4755254/files/figure.png" pageId="24" pageNumber="263">Figs 77</figureCitation>
,
<figureCitation id="D49796C9FFDDC915FDB3882383E37142" box="[570,640,1600,1624]" captionStart="Figs 8599" captionStartId="11.[174,214,2005,2027]" captionTargetBox="[186,1413,263,1983]" captionTargetId="figure-49@11.[174,1421,255,1994]" captionTargetPageId="11" captionText="Figs 8599: Cretotyphlus idanus (85), C. chanianus (8691), Pseudobium creticum (9296), and Gabrius candicus (9799): male sternite VIII (85, 88, 94); habitus (86, 92, 97); forebody in ventral view (87); aedeagus in lateral and in ventral view (8990, 9596, 9899); female genital segments (91); forebody (93). Scale bars: 9293, 97: 1.0 mm; 86: 0.5 mm; 9899: 0.2 mm; 85, 8791, 9496: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755258" httpUri="https://zenodo.org/record/4755258/files/figure.png" pageId="24" pageNumber="263">85, 88</figureCitation>
) symmetric (or nearly so) and with modified pubescence; aedeagus with long, slender, and apically acute ventral process, with slender dorsal portion, and without parameres.
</paragraph>
<paragraph id="4C138A4CFFDDC915FF5D88A2834B7002" blockId="24.[212,817,1087,1816]" pageId="24" pageNumber="263">
♀: sternite VIII angularly produced posteriorly; genital segment with more or less distinct pair of sclerotized internal structures (
<figureCitation id="D49796C9FFDDC915FE2C896380967002" box="[421,501,1792,1816]" captionStart="Figs 7184" captionStartId="10.[212,252,2003,2025]" captionTargetBox="[217,1454,255,1986]" captionTargetId="figure-41@10.[212,1459,255,1994]" captionTargetPageId="10" captionText="Figs 7184: Kenotyphlus virgatus (7173), Cretotyphlus hamatus (7480), and C. idanus (8184): male sternite VIII (71, 77); aedeagus in lateral and in ventral, or in latero-ventral view (7273, 7879, 8384); habitus (74, 81); forebody in dorsal view (75); forebodyin ventral view (76); femalegenital segment (80);male meso-and metathorax and abdomen in ventralview (82). Scale bars: 74, 8182: 0.5 mm; 7173, 7580, 8384: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755254" httpUri="https://zenodo.org/record/4755254/files/figure.png" pageId="24" pageNumber="263">Figs 80</figureCitation>
,
<figureCitation id="D49796C9FFDDC915FE77896383787002" box="[510,539,1792,1816]" captionStart="Figs 8599" captionStartId="11.[174,214,2005,2027]" captionTargetBox="[186,1413,263,1983]" captionTargetId="figure-49@11.[174,1421,255,1994]" captionTargetPageId="11" captionText="Figs 8599: Cretotyphlus idanus (85), C. chanianus (8691), Pseudobium creticum (9296), and Gabrius candicus (9799): male sternite VIII (85, 88, 94); habitus (86, 92, 97); forebody in ventral view (87); aedeagus in lateral and in ventral view (8990, 9596, 9899); female genital segments (91); forebody (93). Scale bars: 9293, 97: 1.0 mm; 86: 0.5 mm; 9899: 0.2 mm; 85, 8791, 9496: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755258" httpUri="https://zenodo.org/record/4755258/files/figure.png" pageId="24" pageNumber="263">91</figureCitation>
).
</paragraph>
<paragraph id="4C138A4CFFDDC915FF5D895C84F176AD" blockId="24.[212,817,1855,2104]" lastBlockId="24.[854,1459,256,440]" pageId="24" pageNumber="263">
<emphasis id="7ED8565EFFDDC915FF5D895C80C17042" bold="true" box="[212,418,1855,1880]" pageId="24" pageNumber="263">Comparative notes</emphasis>
: Based on the morphogical characters indicated above,
<taxonomicName id="8BACF1CFFFDDC915FE018903836F7062" authorityName="ASSING" authorityYear="2019" box="[392,524,1888,1912]" class="Insecta" family="Staphylinidae" genus="Cretotyphlus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="7ED8565EFFDDC915FE018903836F7062" box="[392,524,1888,1912]" italics="true" pageId="24" pageNumber="263">Cretotyphlus</emphasis>
</taxonomicName>
is assigned to the
<taxonomicName id="8BACF1CFFFDDC915FD58890380757082" authorityName="Coiffait" authorityYear="1963" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="tribe" tribe="Metrotyphlini">Metrotyphlini</taxonomicName>
. Using the key to West Palaearctic
<taxonomicName id="8BACF1CFFFDDC915FD0689E382537082" authorityName="Fauvel" authorityYear="1874" box="[655,816,1920,1944]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Leptotyphlinae">Leptotyphlinae</taxonomicName>
in
<bibRefCitation id="283DF7BDFFDDC915FF7989C380C770A2" author="COIffAIT, H." box="[240,420,1952,1976]" pageId="24" pageNumber="263" pagination="1 - 651" refId="ref47598" refString="COIffAIT, H. 1972: Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites, sous-familles: Xantholininae et Leptotyphlinae. - Supplement a la Nouvelle Revue d'Entomologie 2 (2): 1 - 651." type="journal article" year="1972">COIffAIT (1972)</bibRefCitation>
,
<taxonomicName id="8BACF1CFFFDDC915FE2789C3835170A2" authorityName="ASSING" authorityYear="2019" box="[430,562,1952,1976]" class="Insecta" family="Staphylinidae" genus="Cretotyphlus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="7ED8565EFFDDC915FE2789C3835170A2" box="[430,562,1952,1976]" italics="true" pageId="24" pageNumber="263">Cretotyphlus</emphasis>
</taxonomicName>
would key out together with
<taxonomicName id="8BACF1CFFFDDC915FE9A89A380F470C2" authorityName="Coiffait" authorityYear="1955" box="[275,407,1984,2008]" class="Insecta" family="Staphylinidae" genus="Gynotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="7ED8565EFFDDC915FE9A89A380F470C2" box="[275,407,1984,2008]" italics="true" pageId="24" pageNumber="263">Gynotyphlus</emphasis>
</taxonomicName>
COIffAIT, 1955, a genus currently including two species of doubtful status from Corfu and the north of mainland
<collectingCountry id="34BBCADCFFDDC915FE42866383747F02" box="[459,535,2048,2072]" name="Greece" pageId="24" pageNumber="263">Greece</collectingCountry>
, as well as a common and widespread polymorphic and parthenogenetic species,
<taxonomicName id="8BACF1CFFFDDC915FCDF8F6385C37602" authority="(DODERO, 1900)" baseAuthorityName="DODERO" baseAuthorityYear="1900" box="[854,1184,256,280]" class="Insecta" family="Staphylinidae" genus="Gynotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="species" species="perpusillus">
<emphasis id="7ED8565EFFDDC915FCDF8F6382857602" box="[854,998,256,280]" italics="true" pageId="24" pageNumber="263">G. perpusillus</emphasis>
(DODERO, 1900)
</taxonomicName>
. The new genus is distinguished from
<taxonomicName id="8BACF1CFFFDDC915FC618F43850F7622" authorityName="Coiffait" authorityYear="1955" box="[1000,1132,288,312]" class="Insecta" family="Staphylinidae" genus="Gynotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="7ED8565EFFDDC915FC618F43850F7622" box="[1000,1132,288,312]" italics="true" pageId="24" pageNumber="263">Gynotyphlus</emphasis>
</taxonomicName>
by larger body size, longer, less compact, and less incrassate antennae with less transverse antennomeres, longer mandibles, the ventral aspect of the head (
<taxonomicName id="8BACF1CFFFDDC915FC178FE385417682" authorityName="Coiffait" authorityYear="1955" box="[926,1058,384,408]" class="Insecta" family="Staphylinidae" genus="Gynotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="7ED8565EFFDDC915FC178FE385417682" box="[926,1058,384,408]" italics="true" pageId="24" pageNumber="263">Gynotyphlus</emphasis>
</taxonomicName>
: gular sutures distinctly contiguous anteriorly), and the absence of sulci on the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="04B6D9C7FFDDC915FCDF8FBC82E4752D" pageId="24" pageNumber="263" type="distribution">
<paragraph id="4C138A4CFFDDC915FCDF8FBC82E4752D" blockId="24.[854,1459,479,567]" pageId="24" pageNumber="263">
<emphasis id="7ED8565EFFDDC915FCDF8FBC82BC76ED" bold="true" box="[854,991,479,503]" pageId="24" pageNumber="263">Distribution</emphasis>
: The genus currently includes three endemic species in
<collectingRegion id="8E6844AEFFDDC915FC428C6385647502" box="[971,1031,512,536]" country="Greece" name="Kriti" pageId="24" pageNumber="263">Crete</collectingRegion>
, all of them apparently very local and rare.
</paragraph>
</subSubSection>
</treatment>
</document>