185 lines
19 KiB
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185 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.925.39691" ID-GBIF-Dataset="bd72aac6-9c05-4ca2-bfc8-dbf5dc462aa4" ID-PMC="PMC7160194" ID-Pensoft-Pub="1313-2970-925-1" ID-Pensoft-UUID="DCC1953545B9557EA73102AE6D25711F" ID-PubMed="32317852" ID-ZooBank="BB966609087849A1B13C138C2495E6B7" ModsDocID="1313-2970-925-1" checkinTime="1586365656080" checkinUser="pensoft" docAuthor="Maddison, Wayne P., Maddison, David R., Derkarabetian, Shahan & Hedin, Marshal" docDate="2020" docId="41DCF2E718C0593DA4B5E528627B081E" docLanguage="en" docName="ZooKeys 925: 1-54" docOrigin="ZooKeys 925" docSource="http://dx.doi.org/10.3897/zookeys.925.39691" docTitle="Attinella dorsata" docType="treatment" docVersion="4" id="DCC1953545B9557EA73102AE6D25711F" lastPageNumber="1" masterDocId="DCC1953545B9557EA73102AE6D25711F" masterDocTitle="Sitticine jumping spiders: phylogeny, classification, and chromosomes (Araneae, Salticidae, Sitticini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="1" updateTime="1668168695961" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Sitticine jumping spiders: phylogeny, classification, and chromosomes (Araneae, Salticidae, Sitticini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Maddison, Wayne P.</mods:namePart>
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<mods:affiliation>Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-4953-4575</mods:nameIdentifier>
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<mods:nameIdentifier type="email">wmaddisn@mail.ubc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Maddison, David R.</mods:namePart>
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<mods:affiliation>Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Derkarabetian, Shahan</mods:namePart>
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<mods:affiliation>Department of Biology, San Diego State University, San Diego, CA 92182, USA & Department of Organismic and Evolutionary Biology, Harvard University, Cambridge MA 02138, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-9163-9277</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hedin, Marshal</mods:namePart>
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<mods:affiliation>Department of Biology, San Diego State University, San Diego, CA 92182, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>925</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>54</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.925.39691</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.925.39691</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-925-1</mods:identifier>
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<mods:identifier type="ZooBank">BB966609087849A1B13C138C2495E6B7</mods:identifier>
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<treatment ID-GBIF-Taxon="163540045" LSID="urn:lsid:plazi:treatment:41DCF2E718C0593DA4B5E528627B081E" httpUri="http://treatment.plazi.org/id/41DCF2E718C0593DA4B5E528627B081E" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="41DCF2E7-18C0-593D-A4B5-E528627B081E" authority="(Banks, 1895)" baseAuthorityName="Banks" baseAuthorityYear="1895" family="Salticidae" genus="Attinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attinella dorsata" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dorsata">Attinella dorsata (Banks, 1895)</taxonomicName>
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<figureCitation captionStart="Figures 1–14" captionStartId="F1" captionText="Figures 1 - 14. Subtribe Aillutticina (1 - 4) and the Jollas - Tomis clade of the subtribe Sitticina (5 - 14) 1 - 4 Aillutticus nitens, Uruguay (- 34.877, - 56.023): 1 - 3 male 4 female 5, 6 Tomis palpalis male and female, Ecuador (- 0.1996, - 77.7023) 7, 8 Jollas species: 7 J. cupreus male, Ecuador (- 0.675, - 76.397) 8 Jollas sp. female, Ecuador (- 0.7223, - 77.6408) 9 J. leucoproctus, Uruguay (- 34.94, - 54.95) 10 J. flabellatus, Uruguay (- 34.426, - 55.195) 11 - 14 Attinella dorsata male (11 - 13) and female (14), Canada (48.870, - 123.379). Also included in the Jollas - Tomis clade is Sittisax (Figs 99 - 103). Additional members of the Jollas - Tomis clade can be seen in Figs 108 - 128." figureDoi="10.3897/zookeys.925.39691.figures1-14" httpUri="https://binary.pensoft.net/fig/397983" pageId="0" pageNumber="1">Figures 11-14</figureCitation>
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<figureCitation captionStart="Figures 89–103" captionStartId="F8" captionText="Figures 89 - 103. Sitticines of Canada: the Jollas - Tomis clade, represented by the genera Attinella and Sittisax 89 - 93 Attinella concolor: 89 palp (Florida, Gainesville) 90, 91 ventral view of epigyne, dorsal view of cleared vulva (Florida, Gainesville) 92 male (Texas, 30.10, - 97.25) 93 female (Texas, 30.10, - 97.25) 94 - 98 Attinella dorsata: 94 palp (California, San Diego County) 95, 96 ventral view of epigyne, dorsal view of cleared vulva (British Columbia, Nanaimo) 97 male (California, Siskiyou County) 98 female (British Columbia, 48.870, - 123.379) 99 - 103 Sittisax ranieri: 99 palp (Northwest Territories, Tuktoyaktuk) 100, 101 ventral view of epigyne, dorsal view of cleared vulva (Nunavut, Baffin Island) 102 male (Saskatchewan, 55.27, - 105.19) 103 female (Ontario, Old Woman Bay)." figureDoi="10.3897/zookeys.925.39691.figures89-103" httpUri="https://binary.pensoft.net/fig/397990" pageId="0" pageNumber="1">, 94-98</figureCitation>
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<figureCitation captionStart="Figures 105–107" captionStartId="F10" captionText="Figures 105 - 107. Epigynes of Attinella dorsata and Tomis welchi 105 holotype of Attus dorsatus Banks, 1895, epigyne, ventral view 106, 107 holotype of Sitticus welchi Gertsch & Mulaik, 1936 106 epigyne, ventral view 107 cleared vulva, dorsal view." figureDoi="10.3897/zookeys.925.39691.figures105-107" httpUri="https://binary.pensoft.net/fig/397992" pageId="0" pageNumber="1">, 105</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Banks" authorityYear="1895" class="Insecta" family="Salticidae" genus="Attus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attus dorsatus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dorsatus">Attus dorsatus</taxonomicName>
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Banks, 1895 (holotype female in MCZ from California: Los Angeles, examined)
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Gertsch & Mulaik" authorityYear="1936" class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitticus absolutus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="absolutus">Sitticus absolutus</taxonomicName>
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Gertsch & Mulaik, 1936, synonymy restored
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitticus callidus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="callidus">Sitticus callidus</taxonomicName>
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Gertsch & Mulaik, 1936, synonymy restored
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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While females of this small Southwestern desert-dwelling species are indistinctly unmarked, males tend to be reddish with a narrow central longitudinal stripe (Figs
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<figureCitation captionStart="Figures 1–14" captionStartId="F1" captionText="Figures 1 - 14. Subtribe Aillutticina (1 - 4) and the Jollas - Tomis clade of the subtribe Sitticina (5 - 14) 1 - 4 Aillutticus nitens, Uruguay (- 34.877, - 56.023): 1 - 3 male 4 female 5, 6 Tomis palpalis male and female, Ecuador (- 0.1996, - 77.7023) 7, 8 Jollas species: 7 J. cupreus male, Ecuador (- 0.675, - 76.397) 8 Jollas sp. female, Ecuador (- 0.7223, - 77.6408) 9 J. leucoproctus, Uruguay (- 34.94, - 54.95) 10 J. flabellatus, Uruguay (- 34.426, - 55.195) 11 - 14 Attinella dorsata male (11 - 13) and female (14), Canada (48.870, - 123.379). Also included in the Jollas - Tomis clade is Sittisax (Figs 99 - 103). Additional members of the Jollas - Tomis clade can be seen in Figs 108 - 128." figureDoi="10.3897/zookeys.925.39691.figures1-14" httpUri="https://binary.pensoft.net/fig/397983" pageId="0" pageNumber="1">11-14</figureCitation>
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).
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<normalizedToken originalValue="Prószyński">Proszynski</normalizedToken>
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(2017a) rejected
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<bibRefCitation author="Richman, DB" journalOrPublisher="Peckhamia" pageId="0" pageNumber="1" pagination="1 - 125" refId="B59" refString="Richman, DB, 1979. Jumping spiders of the United States and Canada: Changes in the key and list (1). Peckhamia 1: 1 - 125" title="Jumping spiders of the United States and Canada: Changes in the key and list (1)." volume="1" year="1979">
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<normalizedToken originalValue="Richman’s">Richman's</normalizedToken>
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(1979)
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</bibRefCitation>
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synonymy of
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<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1895" family="Salticidae" genus="Attinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attinella dorsata" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dorsata">
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<emphasis italics="true" pageId="0" pageNumber="1">Attinella dorsata</emphasis>
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</taxonomicName>
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(Banks, 1895) with
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<taxonomicName authorityName="Gertsch & Mulaik" authorityYear="1936" class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitticus absolutus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="absolutus">
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<emphasis italics="true" pageId="0" pageNumber="1">Sitticus absolutus</emphasis>
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</taxonomicName>
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, saying that
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<taxonomicName class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="dorsata" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dorsata">
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<emphasis italics="true" pageId="0" pageNumber="1">dorsata</emphasis>
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</taxonomicName>
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is unidentifiable. That statement is false, given that the type specimen is in the MCZ and in good condition. The specimen (examined) has a relatively wide carapace with single thin longitudinal pale line dorsally, long fourth leg, no retromarginal cheliceral tooth, and epigyne (Fig.
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<figureCitation captionStart="Figures 105–107" captionStartId="F10" captionText="Figures 105 - 107. Epigynes of Attinella dorsata and Tomis welchi 105 holotype of Attus dorsatus Banks, 1895, epigyne, ventral view 106, 107 holotype of Sitticus welchi Gertsch & Mulaik, 1936 106 epigyne, ventral view 107 cleared vulva, dorsal view." figureDoi="10.3897/zookeys.925.39691.figures105-107" httpUri="https://binary.pensoft.net/fig/397992" pageId="0" pageNumber="1">105</figureCitation>
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) with a single anterior opening that leads posteriorly through a single duct that splits before the spermathecae, which are visible as two small medial pear-shapes flanked by slightly larger chambers. In these respects, it clearly falls within our current concept of
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<taxonomicName authorityName="Gertsch & Mulaik" authorityYear="1936" class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitticus absolutus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="absolutus">
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<emphasis italics="true" pageId="0" pageNumber="1">Sitticus absolutus</emphasis>
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</taxonomicName>
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as a common, widespread, and relatively uniform species from Texas to California north to Canada (see illustrations by
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<bibRefCitation author="Gertsch, WJ" journalOrPublisher="American Museum Novitates" pageId="0" pageNumber="1" pagination="1 - 21" refId="B16" refString="Gertsch, WJ, Mulaik, S, 1936. Diagnoses of new southern Spiders. American Museum Novitates 851: 1 - 21" title="Diagnoses of new southern Spiders." volume="851" year="1936">Gertsch and Mulaik 1936</bibRefCitation>
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,
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<normalizedToken originalValue="Prószyński">Proszynski</normalizedToken>
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1973). Even if future work were to show that the Californian populations (type locality of
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<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1895" class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="dorsatus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dorsatus">
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<emphasis italics="true" pageId="0" pageNumber="1">dorsatus</emphasis>
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</taxonomicName>
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) and Texan populations (type locality of
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<taxonomicName authorityName="Gertsch & Mulaik" authorityYear="1936" class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="absolutus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="absolutus">
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<emphasis italics="true" pageId="0" pageNumber="1">absolutus</emphasis>
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</taxonomicName>
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) represent distinct species, they are extremely closely related, certainly congeneric.
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<taxonomicName authorityName="Banks" authorityYear="1895" class="Insecta" family="Miridae" genus="Attus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attus dorsatus" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dorsatus">
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<emphasis italics="true" pageId="0" pageNumber="1">Attus dorsatus</emphasis>
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</taxonomicName>
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is a member of these Californian populations, and for this reason the synonymy of
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<taxonomicName authorityName="Proszynski" authorityYear="2017" class="Arachnida" family="Salticidae" genus="Sittiab" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sittiab" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Sittiab</emphasis>
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</taxonomicName>
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(type species
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<taxonomicName authorityName="Gertsch & Mulaik" authorityYear="1936" class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitticus absolutus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="absolutus">
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<emphasis italics="true" pageId="0" pageNumber="1">Sitticus absolutus</emphasis>
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</taxonomicName>
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) with
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<taxonomicName authorityName="Banks" authorityYear="1905" family="Salticidae" genus="Attinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attinella" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Attinella</emphasis>
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</taxonomicName>
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(type species
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<taxonomicName authorityName="Banks" authorityYear="1895" class="Insecta" family="Miridae" genus="Attus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attus dorsatus" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dorsatus">
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<emphasis italics="true" pageId="0" pageNumber="1">Attus dorsatus</emphasis>
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</taxonomicName>
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) is assured.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">Canada: British Columbia: Summerland (1 male, CNC), Galiano Island (2 males, 3 females, UBC-SEM), Nanaimo (1 female). U.S.A.: CALIFORNIA: Humboldt Co., Orleans (1 male, UBC-SEM), Siskyou Co., Beaver Creek and Klamath River (1 male, UBC-SEM), San Diego Co., Johnson Canyon (1 male 1 female, UBC-SEM), El Dorado Co., Camino (1 female, UBC-SEM), Inyo Co., Gilbert Summit (1 female, UBC-SEM); Utah: Millard Co., Sevier Lake (1 male, UBC-SEM); Colorado: Morgan Co., Jackson Lake (1 male, UBC-SEM), Jefferson Co., Golden (2 females, UBC-SEM); Texas: Jim Hogg Co., Guerra (1 female, UBC-SEM), Pecos Co., Fort Stockton (1 female, UBC-SEM).</paragraph>
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<caption doi="10.3897/zookeys.925.39691.figures105-107" httpUri="https://binary.pensoft.net/fig/397992" pageId="0" pageNumber="1" start="Figures 105–107" startId="F10">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figures 105-107.</emphasis>
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Epigynes of
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<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1895" family="Salticidae" genus="Attinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attinella dorsata" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dorsata">
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<emphasis italics="true" pageId="0" pageNumber="1">Attinella dorsata</emphasis>
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</taxonomicName>
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and
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<taxonomicName family="Salticidae" genus="Tomis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomis welchi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="welchi">
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<emphasis italics="true" pageId="0" pageNumber="1">Tomis welchi</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="1">105</emphasis>
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holotype of
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<taxonomicName authorityName="Banks" authorityYear="1895" class="Insecta" family="Miridae" genus="Attus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Attus dorsatus" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dorsatus">
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<emphasis italics="true" pageId="0" pageNumber="1">Attus dorsatus</emphasis>
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</taxonomicName>
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Banks, 1895, epigyne, ventral view
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<emphasis bold="true" pageId="0" pageNumber="1">106, 107</emphasis>
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holotype of
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<taxonomicName authorityName="Gertsch & Mulaik" authorityYear="1936" class="Arachnida" family="Salticidae" genus="Sitticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitticus welchi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="welchi">
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<emphasis italics="true" pageId="0" pageNumber="1">Sitticus welchi</emphasis>
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</taxonomicName>
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Gertsch & Mulaik, 1936
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<emphasis bold="true" pageId="0" pageNumber="1">106</emphasis>
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epigyne, ventral view
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<emphasis bold="true" pageId="0" pageNumber="1">107</emphasis>
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cleared vulva, dorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |