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<document id="AF47554048DA9E75CC40B8EC97FCDFB5" ID-CLB-Dataset="29785" ID-DOI="10.11646/zootaxa.4441.3.1" ID-GBIF-Dataset="80ac1017-65a3-434c-8436-e8701f7761df" ID-ISSN="1175-5326" ID-Zenodo-Dep="1301610" ID-ZooBank="16AB230A-AFCD-484B-ABAD-7CEEDD1532B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530166255974" checkinUser="plazi" docAuthor="Hu, Shao-Ji, Cotton, Adam M., Condamine, Fabien L., Duan, Kuang, Wang, Rong-Jiang, Hsu, Yu-Feng, Zhang, Xin &amp; Cao, Jun" docDate="2018" docId="290C2D1AFFDE6D730D95FF79F2304C57" docLanguage="en" docName="zootaxa.4441.3.1.pdf" docOrigin="Zootaxa 4441 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Graphium (Pazala) mandarinus subsp. fangana" docType="treatment" docVersion="7" lastPageNumber="434" masterDocId="D5355562FFFF6D520D02FFEEF6544F07" masterDocTitle="Revision of Pazala Moore, 1888: The Graphium (Pazala) mandarinus (Oberthür, 1879) Group, with Treatments of Known Taxa and Descriptions of New Species and New Subspecies (Lepidoptera: Papilionidae)" masterLastPageNumber="446" masterPageNumber="401" pageNumber="434" updateTime="1698551520226" updateUser="ExternalLinkService">
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<mods:title id="3030483ECD01308B6D2067D7A1DAA7FE">Revision of Pazala Moore, 1888: The Graphium (Pazala) mandarinus (Oberthür, 1879) Group, with Treatments of Known Taxa and Descriptions of New Species and New Subspecies (Lepidoptera: Papilionidae)</mods:title>
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<mods:namePart id="47C91CE9AD89267F8D63A22B0823086A">Hu, Shao-Ji</mods:namePart>
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<mods:namePart id="82A1A679770C291134516F5236CC4349">Condamine, Fabien L.</mods:namePart>
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<mods:namePart id="9055ACFB2B7992B0E8AC6B3DA4174C2E">Duan, Kuang</mods:namePart>
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<treatment id="290C2D1AFFDE6D730D95FF79F2304C57" ID-DOI="http://doi.org/10.5281/zenodo.5963646" ID-GBIF-Taxon="150784679" ID-Zenodo-Dep="5963646" LSID="urn:lsid:plazi:treatment:290C2D1AFFDE6D730D95FF79F2304C57" httpUri="http://treatment.plazi.org/id/290C2D1AFFDE6D730D95FF79F2304C57" lastPageNumber="434" pageId="33" pageNumber="434">
<subSubSection id="E9BFCF87FFDE6D730D95FF79F72B4FD4" pageId="33" pageNumber="434" type="nomenclature">
<paragraph id="A11A9C0CFFDE6D730D95FF79F5304FB5" blockId="33.[151,868,151,211]" box="[151,868,151,178]" pageId="33" pageNumber="434">
<heading id="FA522B60FFDE6D730D95FF79F5304FB5" bold="true" box="[151,868,151,178]" fontSize="11" level="1" pageId="33" pageNumber="434" reason="1">
<taxonomicName id="66A5E78FFFDE6D730D95FF79F5304FB5" authority="(K. Okano, 1986)" baseAuthorityName="K. Okano" baseAuthorityYear="1986" box="[151,868,151,178]" class="Insecta" family="Papilionidae" genus="Graphium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="mandarinus" subGenus="Pazala" subSpecies="fangana">
<emphasis id="93D1401EFFDE6D730D95FF79F5304FB5" bold="true" box="[151,868,151,178]" pageId="33" pageNumber="434">
<emphasis id="93D1401EFFDE6D730D95FF79F7414FB6" bold="true" box="[151,277,151,177]" italics="true" pageId="33" pageNumber="434">Graphium</emphasis>
(
<emphasis id="93D1401EFFDE6D730C27FF79F7234FB6" bold="true" box="[293,375,151,177]" italics="true" pageId="33" pageNumber="434">Pazala</emphasis>
)
<emphasis id="93D1401EFFDE6D730C8AFF79F4D34FB6" bold="true" box="[392,647,151,177]" italics="true" pageId="33" pageNumber="434">mandarinus fangana</emphasis>
(
<bibRefCitation id="C534E1FDFFDE6D730F95FF76F5084FB5" box="[663,860,152,178]" pageId="33" pageNumber="434" refId="ref22721" refString="Okano, K. (1986) Two new subspecies of butterflies from Thailand. Tokurana, 11, 1 - 8." type="journal article">K. Okano, 1986</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A11A9C0CFFDE6D730D95FF54F72B4FD4" blockId="33.[151,868,151,211]" box="[151,383,186,211]" pageId="33" pageNumber="434">
(
<figureCitation id="399E8089FFDE6D730D9CFF54F75A4FD4" box="[158,270,186,211]" captionStart="FIGURE 25" captionStartId="30.[151,250,1583,1605]" captionTargetBox="[151,1436,714,1561]" captionTargetId="figure@30.[151,1436,714,1561]" captionTargetPageId="30" captionText="FIGURE 25. Three known subspecies of G. (P.) mandarinus (Oberthür, 1879) from Sichuan and Indo-China; upperside in the first row, underside in the second row; scale bar = 10 mm. A: ssp. kimurai Murayama, 1982, ♂, Doi Inthanon, Chiang Mai, N. Thailand; B: ssp. kimurai Murayama, 1982, ♂, Mt. Ashaemyinanaukmyin, W. Shan State, Myanmar; C: ditto, ♀; D: ssp. fangana (K. Okano, 1986), ♂, Doi Pu Meun, Chiang Mai, N. Thailand; E: ditto, ♀; F: ssp. mandarinus, ♂, Baoxing, W. Sichuan. China; G: ssp. mandarinus, ♀, Daba Shan, W. Sichuan. China." httpUri="https://zenodo.org/record/1301660/files/figure.png" pageId="33" pageNumber="434">Figure 25</figureCitation>
, D and E)
</paragraph>
</subSubSection>
<subSubSection id="E9BFCF87FFDE6D730D95FEECF7474D36" pageId="33" pageNumber="434" type="reference_group">
<paragraph id="A11A9C0CFFDE6D730D95FEECF4E24E51" blockId="33.[151,1435,258,342]" pageId="33" pageNumber="434">
<taxonomicName id="66A5E78FFFDE6D730D95FEECF41D4E1F" authority="K. Okano, 1986" authorityName="K. Okano" authorityYear="1986" box="[151,585,258,280]" class="Insecta" family="Papilionidae" genus="Pazala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="glycerion" subSpecies="fangana">
<emphasis id="93D1401EFFDE6D730D95FEECF7C94E1F" box="[151,413,258,280]" italics="true" pageId="33" pageNumber="434">Pazala glycerion fangana</emphasis>
<bibRefCitation id="C534E1FDFFDE6D730CA6FEEDF41D4E1F" box="[420,585,258,280]" pageId="33" pageNumber="434" refId="ref22721" refString="Okano, K. (1986) Two new subspecies of butterflies from Thailand. Tokurana, 11, 1 - 8." type="journal article">K. Okano, 1986</bibRefCitation>
</taxonomicName>
; Tokurana, 11 (3): 1, f. 3; TL: “near Fang, alt.
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, (about
<geoCoordinate id="C491FACBFFDE6D7309CEFEECF37E4E1F" box="[1228,1322,258,280]" direction="north" orientation="latitude" pageId="33" pageNumber="434" precision="925" value="20.333334">20°20 N</geoCoordinate>
,
<geoCoordinate id="C491FACBFFDE6D730837FEECF3DB4E1F" box="[1333,1423,258,280]" direction="east" orientation="longitude" pageId="33" pageNumber="434" precision="925" value="99.833336">99°50 E</geoCoordinate>
), Chieng Mai Prov., North
<collectingCountry id="D9B2DC9CFFDE6D730CD7FECFF4654E30" box="[469,561,289,311]" name="Thailand" pageId="33" pageNumber="434">Thailand</collectingCountry>
” [The co-ordinates stated in the original description are inaccurate, the correct coordinates are approximately
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,
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].
</paragraph>
<paragraph id="A11A9C0CFFDE6D730D95FE6CF7474D36" blockId="33.[151,1437,386,849]" pageId="33" pageNumber="434">
<emphasis id="93D1401EFFDE6D730D95FE6CF7F04E9C" bold="true" box="[151,420,386,411]" pageId="33" pageNumber="434">Diagnostic characters:</emphasis>
Very similar to the preceding subspecies in appearance, forewing length: male 26.031.0 mm (mean = 29.2 ±
<quantity id="665D31E9FFDE6D730C80FE49F7884EB9" box="[386,476,423,447]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="434" unit="mm" value="1.2">1.2 mm</quantity>
,
<emphasis id="93D1401EFFDE6D730CEAFE49F7A24EB9" box="[488,502,423,446]" italics="true" pageId="33" pageNumber="434">n</emphasis>
= 29), female 32.0 mm (only one female). Forewing, apex slightly blunt, also not produced, termen straight; vein M3 and the base of vein Cu1 tinged with black scales in both sexes. Hindwing, submarginal black short lines better developed than the preceding subspecies, which may reach space sc+r
<quantity id="665D31E9FFDE6D730834FDEEF30E4D0D" box="[1334,1370,497,526]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="33" pageNumber="434" unit="in" value="1.0">1 in</quantity>
some specimens.
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<paragraph id="A11A9C0CFFDE6D730DC5FDD2F4674DC7" blockId="33.[151,1437,386,849]" pageId="33" pageNumber="434">
<emphasis id="93D1401EFFDE6D730DC5FDD2F7364D52" bold="true" box="[199,354,572,597]" pageId="33" pageNumber="434">Distribution:</emphasis>
Only known from a few mountains in northern
<collectingCountry id="D9B2DC9CFFDE6D730E8DFDD2F5A74D52" box="[911,1011,572,597]" name="Thailand" pageId="33" pageNumber="434">Thailand</collectingCountry>
near Mae Ai and Fang districts. The restricted range of this taxon effectively inside the potential range of the previous subspecies needs further study. It is notable that
<taxonomicName id="66A5E78FFFDE6D730C3BFD6BF7954D9A" box="[313,449,644,669]" class="Insecta" family="Papilionidae" genus="Pazala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="glycerion" subSpecies="kimurai">
ssp.
<emphasis id="93D1401EFFDE6D730C68FD6AF7954D9A" box="[362,449,644,669]" italics="true" pageId="33" pageNumber="434">kimurai</emphasis>
</taxonomicName>
is absent from the mountains where
<taxonomicName id="66A5E78FFFDE6D730E58FD6BF5E24D9A" authorityName="K. Okano" authorityYear="1986" box="[858,950,645,669]" class="Insecta" family="Papilionidae" genus="Pazala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="glycerion" subSpecies="fangana">
<emphasis id="93D1401EFFDE6D730E58FD6BF5E24D9A" box="[858,950,645,669]" italics="true" pageId="33" pageNumber="434">fangana</emphasis>
</taxonomicName>
occurs, and the taxa are genetically distinct as well as separable on wing pattern.
</paragraph>
<paragraph id="A11A9C0CFFDE6D730DC5FD22F2D64C2A" blockId="33.[151,1437,386,849]" pageId="33" pageNumber="434">
<emphasis id="93D1401EFFDE6D730DC5FD22F7194DE2" bold="true" box="[199,333,716,741]" pageId="33" pageNumber="434">Phenology:</emphasis>
This subspecies is univoltine, flying only in February and March, unlike
<taxonomicName id="66A5E78FFFDE6D73099FFD23F37C4DE2" box="[1181,1320,716,741]" class="Insecta" family="Papilionidae" genus="Pazala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="glycerion" subSpecies="kimurai">
ssp.
<emphasis id="93D1401EFFDE6D7309D3FD22F37C4DE2" box="[1233,1320,716,741]" italics="true" pageId="33" pageNumber="434">kimurai</emphasis>
</taxonomicName>
. Recently deceased Prasobsuk Sukkit (pers. comm.) confirmed that he and local collectors have looked for
<taxonomicName id="66A5E78FFFDE6D730809FD1FF3C84C0F" authorityName="K. Okano" authorityYear="1986" box="[1291,1436,752,776]" class="Insecta" family="Papilionidae" genus="Pazala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="glycerion" subSpecies="fangana">
ssp.
<emphasis id="93D1401EFFDE6D730842FD1EF3C84C0F" box="[1344,1436,752,776]" italics="true" pageId="33" pageNumber="434">fangana</emphasis>
</taxonomicName>
throughout the year over many years, and never found it other than in February to March.
</paragraph>
</subSubSection>
<subSubSection id="E9BFCF87FFDE6D730DC5FCD6F2304C57" box="[199,1124,824,849]" pageId="33" pageNumber="434" type="biology_ecology">
<paragraph id="A11A9C0CFFDE6D730DC5FCD6F2304C57" blockId="33.[151,1437,386,849]" box="[199,1124,824,849]" pageId="33" pageNumber="434">
<emphasis id="93D1401EFFDE6D730DC5FCD6F7194C56" bold="true" box="[199,333,824,849]" pageId="33" pageNumber="434">Host plant:</emphasis>
Unknown, presumably a plant species belonging to family
<taxonomicName id="66A5E78FFFDE6D730EEBFCD7F2344C57" box="[1001,1120,825,848]" family="Lauraceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Laurales" pageId="33" pageNumber="434" phylum="Magnoliopsida" rank="family">Lauraceae</taxonomicName>
.
</paragraph>
</subSubSection>
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