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<document id="7951C8C84B51DB49455C03DD00AD16DF" ID-CLB-Dataset="20712" ID-DOI="10.11646/zootaxa.5099.1.1" ID-GBIF-Dataset="3a781367-6b7e-4a8e-820e-dc4fc7795c33" ID-ISSN="1175-5326" ID-Zenodo-Dep="6036751" ID-ZooBank="2B4782D7-D3FD-4987-BBA4-6569AA7C8B6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644487542310" checkinUser="plazi" docAuthor="Liu, Jing, Lu, Xiangyi, Zhang, Qianwen, Wu, Xiangyi, Yang, Dongdong &amp; Bian, Xun" docDate="2022" docId="1279C969FFDEFFF1688F4795FDEAFAF7" docLanguage="en" docName="zootaxa.5099.1.1.pdf" docOrigin="Zootaxa 5099 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Capnogryllacris nigromarginata subsp. rectispina Liu, Lu &amp; Bian 2022, ssp. nov." docType="treatment" docVersion="8" lastPageNumber="17" masterDocId="EE40B111FFD3FFE168184228FFFAF872" masterDocTitle="Contribution to the knowledge of Chinese Gryllacrididae (Orthoptera) V: Further study on the Chinese Capnogryllacris and comment on the phylogenetic relationships of the Gryllacrididae" masterLastPageNumber="45" masterPageNumber="1" pageNumber="14" updateTime="1699308969733" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D740F3BBAA9A7C6F0B0138832EEE1EC1">Contribution to the knowledge of Chinese Gryllacrididae (Orthoptera) V: Further study on the Chinese Capnogryllacris and comment on the phylogenetic relationships of the Gryllacrididae</mods:title>
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<mods:namePart id="E35C89F1E7073016E3556ED9318A43A5">Liu, Jing</mods:namePart>
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<mods:namePart id="6F7A149A52C65005569C821482CD4877">Lu, Xiangyi</mods:namePart>
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<mods:namePart id="D0BD6C2C005B42D92AFC449EDB7314D7">Yang, Dongdong</mods:namePart>
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<mods:namePart id="B718BE9D1918D9BA4C9F664B76CF9222">Bian, Xun</mods:namePart>
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<treatment id="1279C969FFDEFFF1688F4795FDEAFAF7" ID-DOI="http://doi.org/10.5281/zenodo.6309098" ID-GBIF-Taxon="193331443" ID-Zenodo-Dep="6309098" LSID="urn:lsid:plazi:treatment:1279C969FFDEFFF1688F4795FDEAFAF7" httpUri="http://treatment.plazi.org/id/1279C969FFDEFFF1688F4795FDEAFAF7" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
<subSubSection id="D2CA2BF4FFDEFFEC688F4795FEA1FD8B" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="9A6F787FFFDEFFEC688F4795FC2CFDA5" blockId="13.[151,982,1469,1567]" box="[151,982,1469,1496]" pageId="13" pageNumber="14">
<heading id="C127CF13FFDEFFEC688F4795FC2CFDA5" bold="true" box="[151,982,1469,1496]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="A8A4A46DFFDEFFEC688F4795FC2CFDA5" bold="true" box="[151,982,1469,1496]" pageId="13" pageNumber="14">
<taxonomicName id="5DD003FCFFDEFFEC688F4795FC9FFDA5" authority="Liu, Lu &amp; Bian" authorityName="Liu, Lu &amp; Bian" authorityYear="2022" box="[151,869,1469,1496]" class="Insecta" family="Gryllacrididae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="nigromarginata" status="subsp. nov." subSpecies="rectispina">
<emphasis id="A8A4A46DFFDEFFEC688F4795FD64FDA5" bold="true" box="[151,670,1469,1496]" italics="true" pageId="13" pageNumber="14">Capnogryllacris nigromarginata rectispina</emphasis>
Liu, Lu &amp; Bian
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<taxonomicNameLabel id="B3971916FFDEFFEC6B754795FC2CFDA5" box="[877,982,1469,1495]" pageId="13" pageNumber="14" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
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<heading id="C127CF13FFDEFFEC688F47CAFEA1FD8B" bold="true" box="[151,347,1506,1529]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="A8A4A46DFFDEFFEC688F47CAFEA1FD8B" bold="true" box="[151,347,1506,1529]" pageId="13" pageNumber="14">Rflāä烟ẽȃ</emphasis>
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<subSubSection id="D2CA2BF4FFDEFFF1688F442DFC4CF9FB" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14" type="description">
<paragraph id="9A6F787FFFDEFFEC688F442DFEE1FE6D" blockId="13.[151,982,1469,1567]" box="[151,283,1541,1567]" pageId="13" pageNumber="14">
<figureCitation id="02EB64FAFFDEFFEC688F442DFEE1FE6D" box="[151,283,1541,1567]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStartId-0="16.[151,250,1692,1716]" captionStartId-1="17.[151,250,851,875]" captionTargetBox-0="[152,1436,666,1666]" captionTargetBox-1="[152,1436,181,827]" captionTargetId-0="figure-199@16.[152,1436,666,1666]" captionTargetId-1="figure-19@17.[152,1436,181,827]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." captionText-1="FIGURE 19. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: AB. habitus in lateral view; CD. tegmina in dorsal view: C. left tegmen, D. right tegmen." figureDoi-0="http://doi.org/10.5281/zenodo.6036824" figureDoi-1="http://doi.org/10.5281/zenodo.6036828" httpUri-0="https://zenodo.org/record/6036824/files/figure.png" httpUri-1="https://zenodo.org/record/6036828/files/figure.png" pageId="13" pageNumber="14">Figs. 1819</figureCitation>
</paragraph>
<paragraph id="9A6F787FFFDEFFEC688F4465FC73FEDD" blockId="13.[151,1436,1613,2036]" pageId="13" pageNumber="14">
<emphasis id="A8A4A46DFFDEFFEC688F4465FED3FE15" bold="true" box="[151,297,1613,1639]" pageId="13" pageNumber="14">Description.</emphasis>
Male. head ovoid. Face with scattered dots. Fastigium verticis slightly wider than scape, fastigium frontis separated from fastigium verticis by a fine suture (
<figureCitation id="02EB64FAFFDEFFEC6B094459FC83FEF9" box="[785,889,1649,1675]" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="13" pageNumber="14">Fig. 18A</figureCitation>
). Ocelli distinct, lateral ocelli suited on the lateral areas of fastigium frontis, and slightly smaller than median ocellus.
</paragraph>
<paragraph id="9A6F787FFFDEFFEC68DF4491FAC3FE85" blockId="13.[151,1436,1613,2036]" pageId="13" pageNumber="14">
Anterior margin of pronotum slightly projected in the middle, posterior margin almost straight; lateral lobes longer than deep, posterior margin rounded and directing downwards, humeral sinus indicated (
<figureCitation id="02EB64FAFFDEFFEC6CDD44F5FAD1FE85" box="[1221,1323,1757,1783]" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="13" pageNumber="14">Fig. 18B</figureCitation>
).
</paragraph>
<paragraph id="9A6F787FFFDEFFEC68DF452AFE6EFFFA" blockId="13.[151,1436,1613,2036]" pageId="13" pageNumber="14">Fore coxae swollen with 1 spine. Fore and middle femora unarmed, tibiae with 4 pairs of spurs and 1 pair of apical spurs on ventral surfaces. Hind femora with 1315 internal and 910 external spines on ventral surface; tibiae with 6 pairs of spines and 1 pair of apical spurs on dorsal surface, ventral surface with 1 pair of subapical spurs and 2 pairs of apical spurs.</paragraph>
<paragraph id="9A6F787FFFDEFFEF68DF45B9FB3DF917" blockId="13.[151,1436,1613,2036]" lastBlockId="14.[151,1437,151,609]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Tegmina surpassing the apex of abdomen, reaching the apical area of hind tibiae. Tegmina (
<figureCitation id="02EB64FAFFDEFFEC6CA445B9FABCFFD9" box="[1212,1350,1937,1963]" captionStart="FIGURE 19" captionStartId="17.[151,250,851,875]" captionTargetBox="[152,1436,181,827]" captionTargetId="figure-19@17.[152,1436,181,827]" captionTargetPageId="17" captionText="FIGURE 19. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: AB. habitus in lateral view; CD. tegmina in dorsal view: C. left tegmen, D. right tegmen." figureDoi="http://doi.org/10.5281/zenodo.6036828" httpUri="https://zenodo.org/record/6036828/files/figure.png" pageId="13" pageNumber="14">Fig. 19CD</figureCitation>
): radius releases RS behind middle area of tegmen, both forked near tip. In left tegmen (
<figureCitation id="02EB64FAFFDEFFEC6C0D459DFB86FFBD" box="[1045,1148,1973,1999]" captionStart="FIGURE 19" captionStartId="17.[151,250,851,875]" captionTargetBox="[152,1436,181,827]" captionTargetId="figure-19@17.[152,1436,181,827]" captionTargetPageId="17" captionText="FIGURE 19. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: AB. habitus in lateral view; CD. tegmina in dorsal view: C. left tegmen, D. right tegmen." figureDoi="http://doi.org/10.5281/zenodo.6036828" httpUri="https://zenodo.org/record/6036828/files/figure.png" pageId="13" pageNumber="14">Fig. 19C</figureCitation>
), media leaning on radius before basal quarter area, then releases MP, after short distance MA further divided two branches; cubitus anterior at base single branched, after divides into two branches behind basal quarter, the anterior branch makes a curvature and receives a short connection branch from MA and then remains single branched MP+CuA1 till tip (
<figureCitation id="02EB64FAFFDDFFEF6D124293FA88F8A7" box="[1290,1394,187,213]" captionStart="FIGURE 19" captionStartId="17.[151,250,851,875]" captionTargetBox="[152,1436,181,827]" captionTargetId="figure-19@17.[152,1436,181,827]" captionTargetPageId="17" captionText="FIGURE 19. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: AB. habitus in lateral view; CD. tegmina in dorsal view: C. left tegmen, D. right tegmen." figureDoi="http://doi.org/10.5281/zenodo.6036828" httpUri="https://zenodo.org/record/6036828/files/figure.png" pageId="14" pageNumber="15">Fig. 19D</figureCitation>
). In right tegmen, media and CuA1 with common stem, and fuse a short distance with radius after then M+CuA1 divided from R; MA arises from M+CuA1 before middle area of tegmen and connects with RS and then immediately divided; MP+CuA1 divides into MP and CuA1; CuA2 arises from base and does not divided further. Both tegmina: cubitus posterior undivided, free throughout; with 4 anal veins, the last two with common stem.
</paragraph>
<paragraph id="9A6F787FFFDDFFEF68DF4347FEBDFA13" blockId="14.[151,1437,151,609]" pageId="14" pageNumber="15">
Second and third abdominal tergites without stridulatory pegs (
<figureCitation id="02EB64FAFFDDFFEF6B974347FC0DF9FB" box="[911,1015,367,393]" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="14" pageNumber="15">Fig. 18D</figureCitation>
). Eighth abdominal tergite prolonged and apical area curved (
<figureCitation id="02EB64FAFFDDFFEF69BF43BBFDF7F9DF" box="[423,525,403,429]" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="14" pageNumber="15">Fig. 18E</figureCitation>
). Ninth abdominal tergite narrowing to apex with deep furrow in the middle, on both sides with a pair of oval swellings that terminate at apical margin into a compressed spine that in situ crossed each other before middle area (
<figureCitation id="02EB64FAFFDDFFEF6A1A43F3FD96F987" box="[514,620,475,501]" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="14" pageNumber="15">Fig. 18G</figureCitation>
). Cerci slender, its apices subacute. Subgenital plate longer than wide, lateral margins slightly terminating, posterior margin projected with 1 small triangular concavity in the middle (
<figureCitation id="02EB64FAFFDDFFEF6D7743D7FF31FA4F" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="14" pageNumber="15">Fig. 18H</figureCitation>
). Styli cylindrical, apical half curved outward, apices obtuse; inserted on apico-lateral margin of subgenital plate (
<figureCitation id="02EB64FAFFDDFFEF68C5406FFEC0FA13" box="[221,314,583,609]" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="14" pageNumber="15">Fig. 18I</figureCitation>
).
</paragraph>
<caption id="CEAF28F7FFDDFFEF688F456FFE0CFFD5" ID-DOI="http://doi.org/10.5281/zenodo.6036810" ID-Zenodo-Dep="6036810" httpUri="https://zenodo.org/record/6036810/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1863,1887]" targetBox="[151,1436,630,1837]" targetPageId="14">
<paragraph id="9A6F787FFFDDFFEF688F456FFE0CFFD5" blockId="14.[151,1437,1863,1960]" pageId="14" pageNumber="15">
<emphasis id="A8A4A46DFFDDFFEF688F456FFC3AFF2D" bold="true" box="[151,960,1863,1888]" pageId="14" pageNumber="15">
FIGURE 15.
<taxonomicName id="5DD003FCFFDDFFEF693C456FFD03FF2D" authority="Liu &amp; Lu &amp; Zhang &amp; Wu &amp; Yang &amp; Bian, 2022" authorityName="Liu &amp; Lu &amp; Zhang &amp; Wu &amp; Yang &amp; Bian" authorityYear="2022" box="[292,761,1863,1887]" class="Insecta" family="Gryllacrididae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="nigromarginata" status="subsp. nov." subSpecies="hainanensis">
<emphasis id="A8A4A46DFFDDFFEF693C456FFD03FF2D" bold="true" box="[292,761,1863,1887]" italics="true" pageId="14" pageNumber="15">Capnogryllacris nigromarginata hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3971916FFDDFFEF6AE64560FCA9FF12" box="[766,851,1864,1888]" pageId="14" pageNumber="15" rank="subSpecies">ssp. nov</taxonomicNameLabel>
.. Male: A.
</emphasis>
head in frontal view;
<emphasis id="A8A4A46DFFDDFFEF6C814560FB35FF2D" bold="true" box="[1177,1231,1863,1888]" pageId="14" pageNumber="15">BC.</emphasis>
head and pronotum:
<emphasis id="A8A4A46DFFDDFFEF688F4544FF54FFF6" bold="true" box="[151,174,1900,1924]" pageId="14" pageNumber="15">B.</emphasis>
dorsal view,
<emphasis id="A8A4A46DFFDDFFEF692E4543FEB4FFF1" bold="true" box="[310,334,1899,1923]" pageId="14" pageNumber="15">C.</emphasis>
lateral view;
<emphasis id="A8A4A46DFFDDFFEF69C24544FE08FFF6" bold="true" box="[474,498,1900,1924]" pageId="14" pageNumber="15">D.</emphasis>
second and third abdominal tergites in lateral view;
<emphasis id="A8A4A46DFFDDFFEF6C104544FBBAFFF6" bold="true" box="[1032,1088,1900,1924]" pageId="14" pageNumber="15">EH.</emphasis>
apex of abdomen:
<emphasis id="A8A4A46DFFDDFFEF6D1B4544FAE0FFF6" bold="true" box="[1283,1306,1900,1924]" pageId="14" pageNumber="15">E.</emphasis>
lateral view,
<emphasis id="A8A4A46DFFDDFFEF688F45B8FF50FFDA" bold="true" box="[151,170,1936,1960]" pageId="14" pageNumber="15">F.</emphasis>
dorsal view,
<emphasis id="A8A4A46DFFDDFFEF692A45A7FE96FFD5" bold="true" box="[306,364,1935,1959]" pageId="14" pageNumber="15">GH.</emphasis>
ventral view.
</paragraph>
</caption>
<caption id="CEAF28F7FFDCFFEE688F4132FDA5FB24" ID-DOI="http://doi.org/10.5281/zenodo.6036816" ID-Zenodo-Dep="6036816" httpUri="https://zenodo.org/record/6036816/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,794,818]" targetBox="[151,1428,181,769]" targetPageId="15">
<paragraph id="9A6F787FFFDCFFEE688F4132FDA5FB24" blockId="15.[151,1437,793,854]" pageId="15" pageNumber="16">
<emphasis id="A8A4A46DFFDCFFEE688F4132FC09FB40" bold="true" box="[151,1011,793,818]" pageId="15" pageNumber="16">
FIGURE 16.
<taxonomicName id="5DD003FCFFDCFFEE69314131FCFEFB43" authority="Liu &amp; Lu &amp; Zhang &amp; Wu &amp; Yang &amp; Bian, 2022" authorityName="Liu &amp; Lu &amp; Zhang &amp; Wu &amp; Yang &amp; Bian" authorityYear="2022" box="[297,772,793,818]" class="Insecta" family="Gryllacrididae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="nigromarginata" status="subsp. nov." subSpecies="hainanensis">
<emphasis id="A8A4A46DFFDCFFEE69314131FCFEFB43" bold="true" box="[297,772,793,818]" italics="true" pageId="15" pageNumber="16">Capnogryllacris nigromarginata hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3971916FFDCFFEE6B144132FC9EFB40" box="[780,868,794,818]" pageId="15" pageNumber="16" rank="subSpecies">ssp. nov</taxonomicNameLabel>
.. Male: AB.
</emphasis>
habitus in lateral view;
<emphasis id="A8A4A46DFFDCFFEE6CE84132FADDFB40" bold="true" box="[1264,1319,794,818]" pageId="15" pageNumber="16">CD.</emphasis>
tegmina in dorsal view:
<emphasis id="A8A4A46DFFDCFFEE69024116FEC9FB24" bold="true" box="[282,307,830,854]" pageId="15" pageNumber="16">C.</emphasis>
left tegmen,
<emphasis id="A8A4A46DFFDCFFEE69A04116FE2BFB24" bold="true" box="[440,465,830,854]" pageId="15" pageNumber="16">D.</emphasis>
right tegmen.
</paragraph>
</caption>
<caption id="CEAF28F7FFDCFFEE688F45A7FCFBFF82" ID-DOI="http://doi.org/10.5281/zenodo.6036822" ID-Zenodo-Dep="6036822" httpUri="https://zenodo.org/record/6036822/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1935,1959]" targetBox="[152,1436,878,1912]" targetPageId="15">
<paragraph id="9A6F787FFFDCFFEE688F45A7FCFBFF82" blockId="15.[151,1437,1935,2032]" pageId="15" pageNumber="16">
<emphasis id="A8A4A46DFFDCFFEE688F45A7FC3AFFD5" bold="true" box="[151,960,1935,1960]" pageId="15" pageNumber="16">
FIGURE 17.
<taxonomicName id="5DD003FCFFDCFFEE693C45A7FD03FFD5" authority="Liu &amp; Lu &amp; Zhang &amp; Wu &amp; Yang &amp; Bian, 2022" authorityName="Liu &amp; Lu &amp; Zhang &amp; Wu &amp; Yang &amp; Bian" authorityYear="2022" box="[292,761,1935,1960]" class="Insecta" family="Gryllacrididae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="nigromarginata" status="subsp. nov." subSpecies="hainanensis">
<emphasis id="A8A4A46DFFDCFFEE693C45A7FD03FFD5" bold="true" box="[292,761,1935,1960]" italics="true" pageId="15" pageNumber="16">Capnogryllacris nigromarginata hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B3971916FFDCFFEE6AE645B8FCA9FFDA" box="[766,851,1936,1960]" pageId="15" pageNumber="16" rank="subSpecies">ssp. nov</taxonomicNameLabel>
.. Male: A.
</emphasis>
head in frontal view;
<emphasis id="A8A4A46DFFDCFFEE6C8145B8FB35FFD5" bold="true" box="[1177,1231,1935,1960]" pageId="15" pageNumber="16">BC.</emphasis>
head and pronotum:
<emphasis id="A8A4A46DFFDCFFEE688F459CFF54FFBE" bold="true" box="[151,174,1972,1996]" pageId="15" pageNumber="16">B.</emphasis>
lateral view,
<emphasis id="A8A4A46DFFDCFFEE692F459BFEAAFFB9" bold="true" box="[311,336,1971,1995]" pageId="15" pageNumber="16">C.</emphasis>
dorsal view;
<emphasis id="A8A4A46DFFDCFFEE69C2459BFDE9FFB9" bold="true" box="[474,531,1971,1995]" pageId="15" pageNumber="16">DG.</emphasis>
apex of abdomen:
<emphasis id="A8A4A46DFFDCFFEE6ACD459CFD14FFBE" bold="true" box="[725,750,1972,1996]" pageId="15" pageNumber="16">D.</emphasis>
lateral view,
<emphasis id="A8A4A46DFFDCFFEE6B6F459CFC74FFBE" bold="true" box="[887,910,1972,1996]" pageId="15" pageNumber="16">E.</emphasis>
apical view,
<emphasis id="A8A4A46DFFDCFFEE6C0C459CFBDDFFBE" bold="true" box="[1044,1063,1972,1996]" pageId="15" pageNumber="16">F.</emphasis>
apico-ventral view,
<emphasis id="A8A4A46DFFDCFFEE6CEF459BFAEBFFB9" bold="true" box="[1271,1297,1971,1995]" pageId="15" pageNumber="16">G.</emphasis>
ventral view;
<emphasis id="A8A4A46DFFDCFFEE688F45F0FF32FF82" bold="true" box="[151,200,2008,2032]" pageId="15" pageNumber="16">HI.</emphasis>
tegmina in dorsal view:
<emphasis id="A8A4A46DFFDCFFEE69DB45F0FE27FF82" bold="true" box="[451,477,2008,2032]" pageId="15" pageNumber="16">H.</emphasis>
left tegmen,
<emphasis id="A8A4A46DFFDCFFEE6A7B45F0FD89FF82" bold="true" box="[611,627,2008,2032]" pageId="15" pageNumber="16">I.</emphasis>
right tegmen.
</paragraph>
</caption>
<paragraph id="9A6F787FFFC3FFF168DF42BFFE58F8C3" blockId="16.[151,1437,151,645]" box="[199,418,151,177]" pageId="16" pageNumber="17">
<emphasis id="A8A4A46DFFC3FFF168DF42BFFEDFF8C3" bold="true" box="[199,293,151,177]" pageId="16" pageNumber="17">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="9A6F787FFFC3FFF168DF4292FD4FF917" blockId="16.[151,1437,151,645]" pageId="16" pageNumber="17">
<emphasis id="A8A4A46DFFC3FFF168DF4292FEB5F8A6" bold="true" box="[199,335,186,212]" pageId="16" pageNumber="17">Coloration.</emphasis>
Body brown. Ocelli pale. Genae bright mauve. Internal margins of antennal sockets black. Dorsal surface of head with 1 pair of black spots behind eyes (
<figureCitation id="02EB64FAFFC3FFF16B1B42F7FC96F88B" box="[771,876,223,249]" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="16" pageNumber="17">Fig. 18A</figureCitation>
). Margins of pronotum black which interrupted at the anterior margin of lateral lobes; anterior and posterior margins widened, but posterior spots extending the middle area (
<figureCitation id="02EB64FAFFC3FFF168CD430FFEC0F933" box="[213,314,295,321]" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="16" pageNumber="17">Fig. 18B</figureCitation>
). Dorsal surface of abdomen brownish (
<figureCitation id="02EB64FAFFC3FFF16AE0430FFCA7F933" box="[760,861,295,321]" captionStart="FIGURE 18" captionStartId="16.[151,250,1692,1716]" captionTargetBox="[152,1436,666,1666]" captionTargetId="figure-199@16.[152,1436,666,1666]" captionTargetPageId="16" captionText="FIGURE 18. Capnogryllacris nigromarginata rectispina ssp. nov.. Male: A. head in frontal view; BC. head and pronotum: B. dorsal view, C. lateral view; D. second and third abdominal tergites in lateral view; EI. apex of abdomen: E. lateral view, F. dorsal view, G. dorso-apical view, H. ventral and slightly apical view, I. ventral view." figureDoi="http://doi.org/10.5281/zenodo.6036824" httpUri="https://zenodo.org/record/6036824/files/figure.png" pageId="16" pageNumber="17">Fig. 18E</figureCitation>
). Apical areas of femora and basal areas of tibiae of all legs black. Spines of tibiae blackish or black.
</paragraph>
<paragraph id="9A6F787FFFC3FFF168DF4347FC4CF9FB" blockId="16.[151,1437,151,645]" box="[199,950,366,393]" pageId="16" pageNumber="17">
<emphasis id="A8A4A46DFFC3FFF168DF4347FE3FF9FA" bold="true" box="[199,453,366,393]" pageId="16" pageNumber="17">Measurements (mm).</emphasis>
Male BL 34.0, PL 10.0, TL 40.0, HFL 22.0.
</paragraph>
</subSubSection>
<subSubSection id="D2CA2BF4FFC3FFF168DF43BAFD60F9DC" box="[199,666,402,430]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="9A6F787FFFC3FFF168DF43BAFD60F9DC" blockId="16.[151,1437,151,645]" box="[199,666,402,430]" pageId="16" pageNumber="17">
<emphasis id="A8A4A46DFFC3FFF168DF43BAFE9BF9DE" bold="true" box="[199,353,402,428]" pageId="16" pageNumber="17">Distribution.</emphasis>
<collectingRegion id="5814B69DFFC3FFF1697F43BBFE3DF9DF" box="[359,455,403,429]" country="China" name="Guangxi" pageId="16" pageNumber="17">Guangxi</collectingRegion>
(Fangchenggang).
</paragraph>
</subSubSection>
<subSubSection id="D2CA2BF4FFC3FFF168DF439EFA79F9A3" box="[199,1411,438,466]" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="9A6F787FFFC3FFF168DF439EFA79F9A3" blockId="16.[151,1437,151,645]" box="[199,1411,438,466]" pageId="16" pageNumber="17">
<emphasis id="A8A4A46DFFC3FFF168DF439EFE4BF9A2" bold="true" box="[199,433,438,464]" pageId="16" pageNumber="17">Material examined.</emphasis>
<materialsCitation id="2AB87222FFC3FFF169AF439FFA79F9A3" ID-GBIF-Occurrence="3464448313" box="[439,1411,439,466]" collectingDate="2021-05-06" location="Fangchenggang" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="456BC6DDFFC3FFF169AF439FFDDEF9A3" box="[439,548,439,465]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
: male (XZ163),
<location id="9F0F2EA4FFC3FFF16AC4439FFC6AF9A3" LSID="urn:lsid:plazi:treatment:1279C969FFDEFFF1688F4795FDEAFAF7:9F0F2EA4FFC3FFF16AC4439FFC6AF9A3" box="[732,912,439,465]" name="Fangchenggang" pageId="16" pageNumber="17" stateProvince="Guangxi">Fangchenggang</location>
,
<collectingRegion id="5814B69DFFC3FFF16B83439FFC04F9A3" box="[923,1022,439,465]" country="China" name="Guangxi" pageId="16" pageNumber="17">Guangxi</collectingRegion>
,
<date id="EE6E5EBFFFC3FFF16C12439FFB61F9A3" box="[1034,1179,439,465]" pageId="16" pageNumber="17" value="2021-05-06">
<collectingDate id="FE2AA757FFC3FFF16C12439FFB61F9A3" box="[1034,1179,439,465]" pageId="16" pageNumber="17" value="2021-05-06">May 6, 2021</collectingDate>
</date>
, coll. by Yang Chen.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D2CA2BF4FFC3FFF168DF43F2FF04FA4F" pageId="16" pageNumber="17" type="discussion">
<paragraph id="9A6F787FFFC3FFF168DF43F2FF04FA4F" blockId="16.[151,1437,151,645]" pageId="16" pageNumber="17">
<emphasis id="A8A4A46DFFC3FFF168DF43F2FEB6F986" bold="true" box="[199,332,474,500]" pageId="16" pageNumber="17">Discussion.</emphasis>
The new species differs from
<taxonomicName id="5DD003FCFFC3FFF16ABD43F3FA8BF987" authority="nigromarginata (Karny, 1928)" authorityName="nigromarginata (Karny" authorityYear="1928" baseAuthorityName="Karny" baseAuthorityYear="1928" box="[677,1393,475,501]" class="Insecta" family="Gryllacrididae" genus="Capnogryllacris" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="nigromarginata" subSpecies="nigromarginata">
<emphasis id="A8A4A46DFFC3FFF16ABD43F3FB31F986" box="[677,1227,475,501]" italics="true" pageId="16" pageNumber="17">Capnogryllacris nigromarginata nigromarginata</emphasis>
(Karny, 1928)
</taxonomicName>
by: dorsal surface of head with 1 pair of small black spots, black spots of anterior and posterior margin of pronotum widened.
</paragraph>
</subSubSection>
<subSubSection id="D2CA2BF4FFC3FFF168DF406FFDEAFAF7" pageId="16" pageNumber="17" type="etymology">
<paragraph id="9A6F787FFFC3FFF168DF406FFDEAFAF7" blockId="16.[151,1437,151,645]" pageId="16" pageNumber="17">
<emphasis id="A8A4A46DFFC3FFF168DF406FFEB4FA13" bold="true" box="[199,334,583,609]" pageId="16" pageNumber="17">Etymology.</emphasis>
The new subspecies name is from Latin word “
<emphasis id="A8A4A46DFFC3FFF16B42406FFC7EFA12" box="[858,900,583,608]" italics="true" pageId="16" pageNumber="17">rect</emphasis>
” and “
<emphasis id="A8A4A46DFFC3FFF16BCA406FFBFBFA12" box="[978,1025,583,608]" italics="true" pageId="16" pageNumber="17">spin</emphasis>
” referring to the spines of male ninth abdominal tergite almost straight.
</paragraph>
</subSubSection>
</treatment>
</document>