399 lines
71 KiB
XML
399 lines
71 KiB
XML
<document id="E13EC7873FB8A440A3C2A1C8950CB5D3" ID-CLB-Dataset="20417" ID-DOI="10.11646/zootaxa.5099.1.7" ID-GBIF-Dataset="fb0be913-eb7b-47bd-955a-5879d4c96e10" ID-ISSN="1175-5326" ID-Zenodo-Dep="6037021" ID-ZooBank="DADC9B9F-A342-4522-9EE8-B5965EE87B8A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644488494094" checkinUser="plazi" docAuthor="Shear, William A., Nosler, Philip & Marek, Paul E." docDate="2022" docId="03F8257FFFCA914DFF57FF6E538BF868" docLanguage="en" docName="zootaxa.5099.1.7.pdf" docOrigin="Zootaxa 5099 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amplaria oedipus Shear, Nosler & Marek 2022, new species" docType="treatment" docVersion="8" lastPageNumber="143" masterDocId="FFC15D07FFC8914BFFC0FFF95021FFCA" masterDocTitle="The millipede family Striariidae Bollman, 1893. IV. Amplaria oedipus, n. sp., with a secondary sexual modification of males unique among millipedes (Diplopoda Chordeumatida, Striarioidea)" masterLastPageNumber="145" masterPageNumber="137" pageNumber="139" updateTime="1699309484000" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="F464517AD33CF3806D8EC6AB71577CE5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="380E956AC000C5B6CD78E3C97811B737">
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<mods:title id="2FC4F168EF352E0C2015CA7F71D2635F">The millipede family Striariidae Bollman, 1893. IV. Amplaria oedipus, n. sp., with a secondary sexual modification of males unique among millipedes (Diplopoda Chordeumatida, Striarioidea)</mods:title>
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</mods:titleInfo>
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<mods:name id="2A07E3E380450E6B44FF46B637736D4B" type="personal">
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<mods:role id="3FD622EDF23AB5182F6CF44EE685E8E7">
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<mods:roleTerm id="1C265003DE13B87C3F6A35D02C96B1C4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3F54B9282AD2DE8715EFB0E70D3869B8">Shear, William A.</mods:namePart>
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<mods:affiliation id="30CC197D2C676043F81D0A4652FA47CE">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943 USA, current address: 1950 Price Drive, Farmville VA 23901 USA.</mods:affiliation>
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</mods:name>
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<mods:name id="5769B4A76EBD85A040525811EFCD4808" type="personal">
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<mods:role id="FF59799B06796884CDF9E6EAED7D13C1">
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<mods:roleTerm id="6A7812AE214DA99ADF247B2C333269B6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A66CEA0A37D60490416D2D025131E404">Nosler, Philip</mods:namePart>
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<mods:affiliation id="4A0B32CBB379FC9451AB5B63B84BB9A6">8921 Wooded Hills Court, Damascus OR 97089. philnosler @ gmail. com</mods:affiliation>
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</mods:name>
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<mods:name id="29BEF87F12B34D5CD466F71D3EA8802D" type="personal">
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<mods:role id="88ED0FC196BB68044B2B947497E901DA">
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<mods:roleTerm id="4CB384274D8E06850D25470A72CCF347">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3A5DC8C7DBA9CB9B1390490C42C2B338">Marek, Paul E.</mods:namePart>
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<mods:affiliation id="5A7881F5C10EF8521CFFD3DC5837465B">Department of Entomology, Virginia Tech, Blacksburg VA 24061 USA.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="2F6B381CF55E7B9E847A5DB7CE697005">text</mods:typeOfResource>
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<mods:title id="02BB8618FC7A54DC8A46E783D975FEFD">Zootaxa</mods:title>
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<mods:part id="88DE568DFDB4F30DFBEC727AA7CA05BE">
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<mods:date id="77172D46C5185AAF48BDAA21A65001F5">2022</mods:date>
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<mods:number id="29CE10AB755A65FCF9136DCEDCCF762C">2022-02-10</mods:number>
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<mods:number id="AB01EF49302317881714B709584D9D56">5099</mods:number>
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<mods:identifier id="0DB74CA770830674E9946B43819E140A" type="DOI">10.11646/zootaxa.5099.1.7</mods:identifier>
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<treatment id="03F8257FFFCA914DFF57FF6E538BF868" ID-DOI="http://doi.org/10.5281/zenodo.6037938" ID-GBIF-Taxon="193856928" ID-Zenodo-Dep="6037938" LSID="urn:lsid:plazi:treatment:03F8257FFFCA914DFF57FF6E538BF868" httpUri="http://treatment.plazi.org/id/03F8257FFFCA914DFF57FF6E538BF868" lastPageId="6" lastPageNumber="143" pageId="2" pageNumber="139">
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<subSubSection id="C34BC7E2FFCA9149FF57FF6E5314FF7B" box="[151,821,151,178]" pageId="2" pageNumber="139" type="nomenclature">
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<paragraph id="8BEE9469FFCA9149FF57FF6E5314FF7B" blockId="2.[151,821,151,213]" box="[151,821,151,178]" pageId="2" pageNumber="139">
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<heading id="D0A62305FFCA9149FF57FF6E5314FF7B" bold="true" box="[151,821,151,178]" fontSize="11" level="1" pageId="2" pageNumber="139" reason="1">
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<emphasis id="B925487BFFCA9149FF57FF6E5314FF7B" bold="true" box="[151,821,151,178]" italics="true" pageId="2" pageNumber="139">
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<taxonomicName id="4C51EFEAFFCA9149FF57FF6E52B9FF7B" authority="Shear, Nosler & Marek" authorityName="Shear, Nosler & Marek" authorityYear="2022" box="[151,664,151,178]" class="Diplopoda" family="Striariidae" genus="Amplaria" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="oedipus" status="sp. nov.">Amplaria oedipus Shear, Nosler & Marek</taxonomicName>
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,
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<taxonomicNameLabel id="A216F500FFCA9149FD65FF6E5314FF7B" box="[677,821,151,177]" pageId="2" pageNumber="139" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34BC7E2FFCA9149FF57FF4255A0FE42" pageId="2" pageNumber="139" type="description">
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<paragraph id="8BEE9469FFCA9149FF57FF42512CFF1F" blockId="2.[151,821,151,213]" box="[151,269,187,213]" pageId="2" pageNumber="139">
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<figureCitation id="136A88ECFFCA9149FF57FF425128FF1F" box="[151,265,187,213]" captionStart-0="FIGURES 1 and 2" captionStart-1="FIGURES 3–7" captionStart-2="FIGURES 8–11" captionStart-3="FIGURES 12, 13" captionStartId-0="3.[151,264,1943,1967]" captionStartId-1="4.[151,264,1860,1884]" captionStartId-2="5.[151,264,1781,1805]" captionStartId-3="6.[151,264,1282,1306]" captionTargetBox-0="[234,1354,181,1920]" captionTargetBox-1="[208,1379,181,1837]" captionTargetBox-2="[158,1430,309,1757]" captionTargetBox-3="[159,1418,196,1242]" captionTargetId-0="figure-19@3.[234,1354,181,1920]" captionTargetId-1="figure-22@4.[208,1379,181,1837]" captionTargetId-2="figure-75@5.[157,1430,309,1757]" captionTargetId-3="figure-22@6.[151,1436,181,1258]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 1 and 2. Amplaria oedipus n. sp., male paratype. Fig. 1. Head and anterior legs, ventral view; lab, labrum; lh, labral hook; ms, mandibular stipes; pf1, prefemur of first leg; pf2, prefemur of second leg; t1, tarsus of first leg. Fig. 2. Rings 26–29 and telson; numerals, ring numbers; c (numerals), crest numbers of right side of ring 26; tel, telson." captionText-1="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." captionText-2="FIGURES 8–11. Amplaria oedipus, n. sp., male paratype. Fig. 8. Gonopods, posterior view; aac, anterior angiocoxite; cx, gonopod coxa; fc, flagellocoxite; lcc, lateral lobe of colpocoxite; pac, posterior angiocoxite; pcc, posterior lobe of colpocoxite. The lateral lobe of the colpocoxite has collapsed due to drying for SEM. Fig. 9. Tip of left anterior angiocoxite, mesal view. Fig. 10. Tips of right anterior and posterior angiocoxites, posterior view; aac, anterior angiocoxite; fc, flagellocoxite; pac, posterior angiocoxite. Fig. 11. Left leg 9, anterior view; cp, coxal process; cx9, coxa of leg 9." captionText-3="FIGURES 12, 13. Amplaria oedipus, n. sp., male holotype. Fig. 12. Right gonopod, lateral view; aac, anterior angiocoxite; pac, posterior angiocoxite; lcc, lateral lobe of colpocoxite. Fig. 13. Right gonopod, mesal view; aac, anterior angiocoxite; fc, flagellocoxite; pcc, posterior lobe of colpocoxite; pac, posterior angiocoxite. Without scale." figureDoi-0="http://doi.org/10.5281/zenodo.6037025" figureDoi-1="http://doi.org/10.5281/zenodo.6037027" figureDoi-2="http://doi.org/10.5281/zenodo.6037035" figureDoi-3="http://doi.org/10.5281/zenodo.6037037" httpUri-0="https://zenodo.org/record/6037025/files/figure.png" httpUri-1="https://zenodo.org/record/6037027/files/figure.png" httpUri-2="https://zenodo.org/record/6037035/files/figure.png" httpUri-3="https://zenodo.org/record/6037037/files/figure.png" pageId="2" pageNumber="139">Figs 1–13</figureCitation>
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.
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</paragraph>
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<paragraph id="8BEE9469FFCA9149FF57FEFD52ADFEAE" blockId="2.[151,1437,259,2037]" pageId="2" pageNumber="139">
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<emphasis id="B925487BFFCA9149FF57FEFD50FEFED7" box="[151,223,260,285]" italics="true" pageId="2" pageNumber="139">Types:</emphasis>
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<materialsCitation id="3B399E34FFCA9149FF28FEFA52A9FEAE" ID-GBIF-Occurrence="3695140301" collectingDate="2019-09-25" collectingDateMax="2021-09-18" collectingDateMin="2019-09-25" county="Lane Co." location="Cape Mountain" municipality="Male" pageId="2" pageNumber="139" specimenCount="3" stateProvince="Oregon" typeStatus="holotype">
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<collectingMunicipality id="6B8A0E13FFCA9149FF28FEFA5103FED7" box="[232,290,259,285]" pageId="2" pageNumber="139">Male</collectingMunicipality>
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<typeStatus id="54EA2ACBFFCA9149FEEAFEFA51ADFED7" box="[298,396,259,285]" pageId="2" pageNumber="139" type="holotype">holotype</typeStatus>
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and female
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<typeStatus id="54EA2ACBFFCA9149FDDAFEFD525BFED4" box="[538,634,260,286]" pageId="2" pageNumber="139" type="paratype">paratype</typeStatus>
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from
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<location id="8E8EC2B2FFCA9149FD02FEFA5355FED7" LSID="urn:lsid:plazi:treatment:03F8257FFFCA914DFF57FF6E538BF868:8E8EC2B2FFCA9149FD02FEFA5355FED7" box="[706,884,259,285]" county="Lane Co." municipality="Male" name="Cape Mountain" pageId="2" pageNumber="139" stateProvince="Oregon">Cape Mountain</location>
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, near
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<location id="8E8EC2B2FFCA9149FC79FEFA543FFED7" LSID="urn:lsid:plazi:treatment:03F8257FFFCA914DFF57FF6E538BF868:8E8EC2B2FFCA9149FC79FEFA543FFED7" box="[953,1054,259,285]" county="Lane Co." municipality="Male" name="Florence" pageId="2" pageNumber="139" stateProvince="Oregon">Florence</location>
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, 44.0947°, -124.0756°,
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<collectingCounty id="628FECE5FFCA9149FAEDFEFD55B9FED7" box="[1325,1432,259,286]" pageId="2" pageNumber="139">Lane Co.</collectingCounty>
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,
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<collectingRegion id="49955A8BFFCA9149FF57FEDE50CFFE8B" box="[151,238,295,321]" country="United States of America" name="Oregon" pageId="2" pageNumber="139">Oregon</collectingRegion>
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, collected
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<date id="FFEFB2A9FFCA9149FEA7FEDE5262FE8B" box="[359,579,295,321]" pageId="2" pageNumber="139" value="2021-09-18">
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<collectingDate id="EFAB4B41FFCA9149FEA7FEDE5262FE8B" box="[359,579,295,321]" pageId="2" pageNumber="139" value="2021-09-18">18 September 2021</collectingDate>
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</date>
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by Phil Nosler; male
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<typeStatus id="54EA2ACBFFCA9149FCFDFEDE53BCFE8B" box="[829,925,295,321]" pageId="2" pageNumber="139" type="paratype">paratype</typeStatus>
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from same locality but 44.0954°, -124.9766, collected
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<date id="FFEFB2A9FFCA9149FEC3FEB351FCFEAF" box="[259,477,330,357]" pageId="2" pageNumber="139" value="2019-09-25">
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<collectingDate id="EFAB4B41FFCA9149FEC3FEB351FCFEAF" box="[259,477,330,357]" pageId="2" pageNumber="139" value="2019-09-25">25 September 2019</collectingDate>
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</date>
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by Phil Nosler
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BEE9469FFCA9149FF07FE9655A0FE42" blockId="2.[151,1437,259,2037]" box="[199,1409,366,393]" pageId="2" pageNumber="139">
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<materialsCitation id="3B399E34FFCA9149FF07FE96555CFE42" ID-GBIF-Occurrence="3695140302" box="[199,1405,366,393]" location="Academy of Sciences" municipality="All" pageId="2" pageNumber="139" specimenCount="1" stateProvince="California" typeStatus="paratype">
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Parts of male
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<typeStatus id="54EA2ACBFFCA9149FEA2FE9651E3FE43" box="[354,450,367,393]" pageId="2" pageNumber="139" type="paratype">paratype</typeStatus>
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on SEM stub WS36-15.
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<collectingMunicipality id="6B8A0E13FFCA9149FD1AFE9752DFFE42" box="[730,766,366,392]" pageId="2" pageNumber="139">All</collectingMunicipality>
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specimens deposited in
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<collectingRegion id="49955A8BFFCA9149FBCFFE9754A0FE42" box="[1039,1153,366,392]" country="United States of America" name="California" pageId="2" pageNumber="139">California</collectingRegion>
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<location id="8E8EC2B2FFCA9149FB46FE97555CFE42" LSID="urn:lsid:plazi:treatment:03F8257FFFCA914DFF57FF6E538BF868:8E8EC2B2FFCA9149FB46FE97555CFE42" box="[1158,1405,366,392]" municipality="All" name="Academy of Sciences" pageId="2" pageNumber="139" stateProvince="California">Academy of Sciences</location>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34BC7E2FFCA9149FF07FE6B53C7FE1A" pageId="2" pageNumber="139" type="diagnosis">
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<paragraph id="8BEE9469FFCA9149FF07FE6B53C7FE1A" blockId="2.[151,1437,259,2037]" pageId="2" pageNumber="139">
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<emphasis id="B925487BFFCA9149FF07FE6B5165FE66" bold="true" box="[199,324,402,428]" pageId="2" pageNumber="139">Diagnosis:</emphasis>
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This species cannot be confused with any other, due to the unique modification of the tarsi of male legpairs 5 and 6, which are enormously swollen and pyriform (
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<figureCitation id="136A88ECFFCA9149FC97FE4F53F9FE1A" box="[855,984,438,464]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Figs 3, 6, 7</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34BC7E2FFCA9149FF07FE235290FDF1" pageId="2" pageNumber="139" type="etymology">
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<paragraph id="8BEE9469FFCA9149FF07FE235290FDF1" blockId="2.[151,1437,259,2037]" pageId="2" pageNumber="139">
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<emphasis id="B925487BFFCA9149FF07FE235173FE3E" bold="true" box="[199,338,474,500]" pageId="2" pageNumber="139">Etymology:</emphasis>
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The species epithet is a noun in apposition, a Greek proper name. Oedipus is a well-known figure from Greek mythology and the subject of two tragedies by Sophocles. The name Oedipus means “swollen foot” and is therefore appropriate for this unusual species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34BC7E2FFCA9149FF07FDBD5458F9E9" pageId="2" pageNumber="139" type="description">
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<paragraph id="8BEE9469FFCA9149FF07FDBD5467FC93" blockId="2.[151,1437,259,2037]" pageId="2" pageNumber="139">
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<emphasis id="B925487BFFCA9149FF07FDBD517AFD94" bold="true" box="[199,347,580,606]" pageId="2" pageNumber="139">Description:</emphasis>
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<emphasis id="B925487BFFCA9149FEA4FDBC522EFD95" box="[356,527,581,607]" italics="true" pageId="2" pageNumber="139">
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Male
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<typeStatus id="54EA2ACBFFCA9149FE67FDBC522AFD95" box="[423,523,581,607]" pageId="2" pageNumber="139" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Body with 30 rings (including telson). Length, about
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, width
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||
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|
||
at midbody. Ocularium with 12 or 13 ommatidia. Labrum (
|
||
<emphasis id="B925487BFFCA9149FCD4FD905318FD49" bold="true" box="[788,825,617,643]" pageId="2" pageNumber="139">lab</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FC87FD9153A6FD49" box="[839,903,616,643]" captionStart="FIGURES 1 and 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[234,1354,181,1920]" captionTargetId="figure-19@3.[234,1354,181,1920]" captionTargetPageId="3" captionText="FIGURES 1 and 2. Amplaria oedipus n. sp., male paratype. Fig. 1. Head and anterior legs, ventral view; lab, labrum; lh, labral hook; ms, mandibular stipes; pf1, prefemur of first leg; pf2, prefemur of second leg; t1, tarsus of first leg. Fig. 2. Rings 26–29 and telson; numerals, ring numbers; c (numerals), crest numbers of right side of ring 26; tel, telson." figureDoi="http://doi.org/10.5281/zenodo.6037025" httpUri="https://zenodo.org/record/6037025/files/figure.png" pageId="2" pageNumber="139">Fig. 1</figureCitation>
|
||
) with acute spines (
|
||
<emphasis id="B925487BFFCA9149FBACFD9054A2FD49" bold="true" box="[1132,1155,617,643]" pageId="2" pageNumber="139">lh</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FB51FD9154F0FD49" box="[1169,1233,616,643]" captionStart="FIGURES 1 and 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[234,1354,181,1920]" captionTargetId="figure-19@3.[234,1354,181,1920]" captionTargetPageId="3" captionText="FIGURES 1 and 2. Amplaria oedipus n. sp., male paratype. Fig. 1. Head and anterior legs, ventral view; lab, labrum; lh, labral hook; ms, mandibular stipes; pf1, prefemur of first leg; pf2, prefemur of second leg; t1, tarsus of first leg. Fig. 2. Rings 26–29 and telson; numerals, ring numbers; c (numerals), crest numbers of right side of ring 26; tel, telson." figureDoi="http://doi.org/10.5281/zenodo.6037025" httpUri="https://zenodo.org/record/6037025/files/figure.png" pageId="2" pageNumber="139">Fig. 1</figureCitation>
|
||
) at distal corners. Mandibular stipes (
|
||
<emphasis id="B925487BFFCA9149FEB3FD7551B4FD6C" bold="true" box="[371,405,652,678]" pageId="2" pageNumber="139">ms</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FE64FD7551C5FD6C" box="[420,484,652,678]" captionStart="FIGURES 1 and 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[234,1354,181,1920]" captionTargetId="figure-19@3.[234,1354,181,1920]" captionTargetPageId="3" captionText="FIGURES 1 and 2. Amplaria oedipus n. sp., male paratype. Fig. 1. Head and anterior legs, ventral view; lab, labrum; lh, labral hook; ms, mandibular stipes; pf1, prefemur of first leg; pf2, prefemur of second leg; t1, tarsus of first leg. Fig. 2. Rings 26–29 and telson; numerals, ring numbers; c (numerals), crest numbers of right side of ring 26; tel, telson." figureDoi="http://doi.org/10.5281/zenodo.6037025" httpUri="https://zenodo.org/record/6037025/files/figure.png" pageId="2" pageNumber="139">Fig. 1</figureCitation>
|
||
) lacking projections, rimmed with rounded teeth. Collum white, with 10 low crests, sixth crest on each side vaguely noticeable. Metazonite ornamentation (Fig. 2) typical of the family, with 12 crests, prominent and equal, wider spacing between crests 2 and 3, and 4 and 5 on each side. Rings with two longitudinal bands of purplish brown pigment; crests 2 and 3 of each side within these pigmented bands thereby making crests appear with darker pigment. Segmental setae short, distally with a brush of small spinules. Telson (
|
||
<emphasis id="B925487BFFCA9149FADFFCE2551CFCFF" bold="true" box="[1311,1341,795,821]" pageId="2" pageNumber="139">tel</emphasis>
|
||
, Fig. 2) slightly elongated, 3-lobed. Cerotegument lacking in all specimens so far examined.
|
||
</paragraph>
|
||
<paragraph id="8BEE9469FFCA9149FF07FC9B5094FB28" blockId="2.[151,1437,259,2037]" pageId="2" pageNumber="139">
|
||
Legpair 1 white, reduced, smallest of all legs (
|
||
<figureCitation id="136A88ECFFCA9149FD29FC9B530FFCB7" box="[745,814,866,893]" captionStart="FIGURES 1 and 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[234,1354,181,1920]" captionTargetId="figure-19@3.[234,1354,181,1920]" captionTargetPageId="3" captionText="FIGURES 1 and 2. Amplaria oedipus n. sp., male paratype. Fig. 1. Head and anterior legs, ventral view; lab, labrum; lh, labral hook; ms, mandibular stipes; pf1, prefemur of first leg; pf2, prefemur of second leg; t1, tarsus of first leg. Fig. 2. Rings 26–29 and telson; numerals, ring numbers; c (numerals), crest numbers of right side of ring 26; tel, telson." figureDoi="http://doi.org/10.5281/zenodo.6037025" httpUri="https://zenodo.org/record/6037025/files/figure.png" pageId="2" pageNumber="139">Fig. 1</figureCitation>
|
||
), needle-like setae lacking, but femora with flattened specialized setae. Tarsi of legpair 1 (
|
||
<emphasis id="B925487BFFCA9149FDF1FC7F5268FC6A" bold="true" box="[561,585,902,928]" pageId="2" pageNumber="139">t1</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FD96FC7F52B7FC6A" box="[598,662,902,928]" captionStart="FIGURES 1 and 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[234,1354,181,1920]" captionTargetId="figure-19@3.[234,1354,181,1920]" captionTargetPageId="3" captionText="FIGURES 1 and 2. Amplaria oedipus n. sp., male paratype. Fig. 1. Head and anterior legs, ventral view; lab, labrum; lh, labral hook; ms, mandibular stipes; pf1, prefemur of first leg; pf2, prefemur of second leg; t1, tarsus of first leg. Fig. 2. Rings 26–29 and telson; numerals, ring numbers; c (numerals), crest numbers of right side of ring 26; tel, telson." figureDoi="http://doi.org/10.5281/zenodo.6037025" httpUri="https://zenodo.org/record/6037025/files/figure.png" pageId="2" pageNumber="139">Fig. 1</figureCitation>
|
||
) with comb setae. Remaining legpairs with purplish brown pigment. Legpair 2 (
|
||
<figureCitation id="136A88ECFFCA9149FED2FC535178FC0E" box="[274,345,938,964]" captionStart="FIGURES 1 and 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[234,1354,181,1920]" captionTargetId="figure-19@3.[234,1354,181,1920]" captionTargetPageId="3" captionText="FIGURES 1 and 2. Amplaria oedipus n. sp., male paratype. Fig. 1. Head and anterior legs, ventral view; lab, labrum; lh, labral hook; ms, mandibular stipes; pf1, prefemur of first leg; pf2, prefemur of second leg; t1, tarsus of first leg. Fig. 2. Rings 26–29 and telson; numerals, ring numbers; c (numerals), crest numbers of right side of ring 26; tel, telson." figureDoi="http://doi.org/10.5281/zenodo.6037025" httpUri="https://zenodo.org/record/6037025/files/figure.png" pageId="2" pageNumber="139">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FEA7FC535157FC0E" box="[359,374,938,964]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">4</figureCitation>
|
||
) with
|
||
<emphasis id="B925487BFFCA9149FE7FFC52524BFC0E" box="[447,618,938,964]" italics="true" pageId="2" pageNumber="139">vasa deferentia</emphasis>
|
||
(
|
||
<emphasis id="B925487BFFCA9149FDBAFC5352B9FC0E" bold="true" box="[634,664,938,964]" pageId="2" pageNumber="139">vd</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FD66FC5352C9FC0E" box="[678,744,938,964]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 4</figureCitation>
|
||
) opening from unmodified coxae (
|
||
<emphasis id="B925487BFFCA9149FBB0FC5354B8FC0E" bold="true" box="[1136,1177,938,964]" pageId="2" pageNumber="139">cx2</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FB67FC5354CAFC0E" box="[1191,1259,938,964]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 4</figureCitation>
|
||
); with rounded, bulging trochanters (
|
||
<emphasis id="B925487BFFCA9149FE42FC375187FC22" bold="true" box="[386,422,974,1000]" pageId="2" pageNumber="139">tr2</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FE75FC3751D9FC22" box="[437,504,974,1000]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 4</figureCitation>
|
||
) with short processes bearing modified setae (
|
||
<emphasis id="B925487BFFCA9149FBC6FC37540AFC22" bold="true" box="[1030,1067,974,1000]" pageId="2" pageNumber="139">trp</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FBFAFC37545EFC22" box="[1082,1151,974,1000]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 4</figureCitation>
|
||
); and with enlarged, flattened prefemora (
|
||
<emphasis id="B925487BFFCA9149FEA0FC0851A7FBC1" bold="true" box="[352,390,1009,1035]" pageId="2" pageNumber="139">pf2</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FE55FC0851FBFBC6" box="[405,474,1009,1036]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 4</figureCitation>
|
||
). Legpair 3 (
|
||
<figureCitation id="136A88ECFFCA9149FDABFC0852EEFBC6" box="[619,719,1009,1036]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Figs 3, 5</figureCitation>
|
||
) with short, curved coxal flasks (
|
||
<emphasis id="B925487BFFCA9149FB8BFC085440FBC1" bold="true" box="[1099,1121,1009,1035]" pageId="2" pageNumber="139">cf</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FBB0FC085493FBC6" box="[1136,1202,1009,1036]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 5</figureCitation>
|
||
) distally with curled setae; and with telopodites not enlarged but femora flattened, lower surface with evenly scattered small tubercles (
|
||
<figureCitation id="136A88ECFFCA9149FF60FBC050C2FB99" box="[160,227,1081,1107]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 3</figureCitation>
|
||
). Postcoxal bars of ring 3 long, distally expanded, meeting or crossing in midline. Legpairs 4–7 (
|
||
<figureCitation id="136A88ECFFCA9149FACEFBC0556EFB99" box="[1294,1359,1081,1107]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 3</figureCitation>
|
||
) much enlarged, with flattened femora; legpairs 5 and 6 with greatly swollen pyriform tarsi (
|
||
<emphasis id="B925487BFFCA9149FB99FBA454B6FBBC" bold="true" box="[1113,1175,1116,1143]" pageId="2" pageNumber="139">t5, t6</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FB65FBA45505FBBD" box="[1189,1316,1117,1143]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Figs 3, 6, 7</figureCitation>
|
||
) about
|
||
<quantity id="4CA9398CFFCA9149FAB9FBA550E2FB51" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="139" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long and
|
||
<quantity id="4CA9398CFFCA9149FEEEFB7851A2FB51" box="[302,387,1153,1179]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="139" unit="mm" value="0.4">0.4 mm</quantity>
|
||
in diameter, tapering proximally to articulation with tibiae and tapering distally at tarsungulum (
|
||
<figureCitation id="136A88ECFFCA9149FF60FB5D50C2FB75" box="[160,227,1188,1215]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 7</figureCitation>
|
||
), but nearly spherical in distal view (
|
||
<figureCitation id="136A88ECFFCA9149FDBFFB5D52E2FB75" box="[639,707,1188,1215]" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 6</figureCitation>
|
||
). Legpair 7 without coxal lobes covering bases of gonopods (
|
||
<figureCitation id="136A88ECFFCA9149FAAFFB5D5086FB28" captionStart="FIGURES 3–7" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[208,1379,181,1837]" captionTargetId="figure-22@4.[208,1379,181,1837]" captionTargetPageId="4" captionText="FIGURES 3–7. Amplaria oedipus, n. sp., male paratype. Fig. 3. Legpairs 3–7, ventral view; cx3, third leg coxa; pf(numeral), prefemora of legs 4-7; t(numeral), greatly inflated tarsi of legs 5, 6. Fig. 4. Bases of legpair 2, ventral view; cx2, coxa of second leg; pf2, prefemur of leg 2; tr2, trochanter of second leg; trp, trochanteral process; vd, opening of vas deferens. Fig. 5. Bases of legpair 3; cf, coxal flask of third leg; cx3, coxa of third leg; s3, sternum of third leg. Fig. 6. Greatly inflated tarsus of left fifth leg, distal view; t5, tarsus of fifth leg. Fig. 7. Greatly inflated tarsus of right sixth leg, lateral view; t6, tarsus of sixth leg." figureDoi="http://doi.org/10.5281/zenodo.6037027" httpUri="https://zenodo.org/record/6037027/files/figure.png" pageId="2" pageNumber="139">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEE9469FFCA9149FF07FB155459FA16" blockId="2.[151,1437,259,2037]" pageId="2" pageNumber="139">
|
||
Gonopods (
|
||
<figureCitation id="136A88ECFFCA9149FE8BFB1551E1FACC" box="[331,448,1260,1286]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1781,1805]" captionTargetBox="[158,1430,309,1757]" captionTargetId="figure-75@5.[157,1430,309,1757]" captionTargetPageId="5" captionText="FIGURES 8–11. Amplaria oedipus, n. sp., male paratype. Fig. 8. Gonopods, posterior view; aac, anterior angiocoxite; cx, gonopod coxa; fc, flagellocoxite; lcc, lateral lobe of colpocoxite; pac, posterior angiocoxite; pcc, posterior lobe of colpocoxite. The lateral lobe of the colpocoxite has collapsed due to drying for SEM. Fig. 9. Tip of left anterior angiocoxite, mesal view. Fig. 10. Tips of right anterior and posterior angiocoxites, posterior view; aac, anterior angiocoxite; fc, flagellocoxite; pac, posterior angiocoxite. Fig. 11. Left leg 9, anterior view; cp, coxal process; cx9, coxa of leg 9." figureDoi="http://doi.org/10.5281/zenodo.6037035" httpUri="https://zenodo.org/record/6037035/files/figure.png" pageId="2" pageNumber="139">Figs 8–10</figureCitation>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FE0CFB155235FACC" box="[460,532,1260,1286]" captionStart="FIGURES 12, 13" captionStartId="6.[151,264,1282,1306]" captionTargetBox="[159,1418,196,1242]" captionTargetId="figure-22@6.[151,1436,181,1258]" captionTargetPageId="6" captionText="FIGURES 12, 13. Amplaria oedipus, n. sp., male holotype. Fig. 12. Right gonopod, lateral view; aac, anterior angiocoxite; pac, posterior angiocoxite; lcc, lateral lobe of colpocoxite. Fig. 13. Right gonopod, mesal view; aac, anterior angiocoxite; fc, flagellocoxite; pcc, posterior lobe of colpocoxite; pac, posterior angiocoxite. Without scale." figureDoi="http://doi.org/10.5281/zenodo.6037037" httpUri="https://zenodo.org/record/6037037/files/figure.png" pageId="2" pageNumber="139">12, 13</figureCitation>
|
||
) with structure typical of
|
||
<emphasis id="B925487BFFCA9149FCFFFB155390FACC" box="[831,945,1260,1286]" italics="true" pageId="2" pageNumber="139">
|
||
<taxonomicName id="4C51EFEAFFCA9149FCFFFB15538DFACC" authorityName="Chamberlin" authorityYear="1941" box="[831,940,1260,1286]" class="Diplopoda" family="Striariidae" genus="Amplaria" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="139" phylum="Arthropoda" rank="genus">Amplaria</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Anterior angiocoxites (
|
||
<emphasis id="B925487BFFCA9149FB7FFB1554C6FACC" bold="true" box="[1215,1255,1260,1286]" pageId="2" pageNumber="139">aac</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FB37FB15554DFACC" box="[1271,1388,1260,1286]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1781,1805]" captionTargetBox="[158,1430,309,1757]" captionTargetId="figure-75@5.[157,1430,309,1757]" captionTargetPageId="5" captionText="FIGURES 8–11. Amplaria oedipus, n. sp., male paratype. Fig. 8. Gonopods, posterior view; aac, anterior angiocoxite; cx, gonopod coxa; fc, flagellocoxite; lcc, lateral lobe of colpocoxite; pac, posterior angiocoxite; pcc, posterior lobe of colpocoxite. The lateral lobe of the colpocoxite has collapsed due to drying for SEM. Fig. 9. Tip of left anterior angiocoxite, mesal view. Fig. 10. Tips of right anterior and posterior angiocoxites, posterior view; aac, anterior angiocoxite; fc, flagellocoxite; pac, posterior angiocoxite. Fig. 11. Left leg 9, anterior view; cp, coxal process; cx9, coxa of leg 9." figureDoi="http://doi.org/10.5281/zenodo.6037035" httpUri="https://zenodo.org/record/6037035/files/figure.png" pageId="2" pageNumber="139">Figs 8, 10</figureCitation>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FAB9FB155092FAE3" captionStart="FIGURES 12, 13" captionStartId="6.[151,264,1282,1306]" captionTargetBox="[159,1418,196,1242]" captionTargetId="figure-22@6.[151,1436,181,1258]" captionTargetPageId="6" captionText="FIGURES 12, 13. Amplaria oedipus, n. sp., male holotype. Fig. 12. Right gonopod, lateral view; aac, anterior angiocoxite; pac, posterior angiocoxite; lcc, lateral lobe of colpocoxite. Fig. 13. Right gonopod, mesal view; aac, anterior angiocoxite; fc, flagellocoxite; pcc, posterior lobe of colpocoxite; pac, posterior angiocoxite. Without scale." figureDoi="http://doi.org/10.5281/zenodo.6037037" httpUri="https://zenodo.org/record/6037037/files/figure.png" pageId="2" pageNumber="139">12, 13</figureCitation>
|
||
) slightly curved, distally hook-like, with small, bifurcate posterior distal process (
|
||
<figureCitation id="136A88ECFFCA9149FB89FAF654AFFAE3" box="[1097,1166,1295,1321]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1781,1805]" captionTargetBox="[158,1430,309,1757]" captionTargetId="figure-75@5.[157,1430,309,1757]" captionTargetPageId="5" captionText="FIGURES 8–11. Amplaria oedipus, n. sp., male paratype. Fig. 8. Gonopods, posterior view; aac, anterior angiocoxite; cx, gonopod coxa; fc, flagellocoxite; lcc, lateral lobe of colpocoxite; pac, posterior angiocoxite; pcc, posterior lobe of colpocoxite. The lateral lobe of the colpocoxite has collapsed due to drying for SEM. Fig. 9. Tip of left anterior angiocoxite, mesal view. Fig. 10. Tips of right anterior and posterior angiocoxites, posterior view; aac, anterior angiocoxite; fc, flagellocoxite; pac, posterior angiocoxite. Fig. 11. Left leg 9, anterior view; cp, coxal process; cx9, coxa of leg 9." figureDoi="http://doi.org/10.5281/zenodo.6037035" httpUri="https://zenodo.org/record/6037035/files/figure.png" pageId="2" pageNumber="139">Fig. 9</figureCitation>
|
||
). Posterior angiocoxites (
|
||
<emphasis id="B925487BFFCA9149FF61FACA50EAFA87" bold="true" box="[161,203,1331,1357]" pageId="2" pageNumber="139">pac</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FF17FACA5169FA87" box="[215,328,1331,1357]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1781,1805]" captionTargetBox="[158,1430,309,1757]" captionTargetId="figure-75@5.[157,1430,309,1757]" captionTargetPageId="5" captionText="FIGURES 8–11. Amplaria oedipus, n. sp., male paratype. Fig. 8. Gonopods, posterior view; aac, anterior angiocoxite; cx, gonopod coxa; fc, flagellocoxite; lcc, lateral lobe of colpocoxite; pac, posterior angiocoxite; pcc, posterior lobe of colpocoxite. The lateral lobe of the colpocoxite has collapsed due to drying for SEM. Fig. 9. Tip of left anterior angiocoxite, mesal view. Fig. 10. Tips of right anterior and posterior angiocoxites, posterior view; aac, anterior angiocoxite; fc, flagellocoxite; pac, posterior angiocoxite. Fig. 11. Left leg 9, anterior view; cp, coxal process; cx9, coxa of leg 9." figureDoi="http://doi.org/10.5281/zenodo.6037035" httpUri="https://zenodo.org/record/6037035/files/figure.png" pageId="2" pageNumber="139">Figs 8–10</figureCitation>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FE92FACA51B6FA87" box="[338,407,1331,1357]" captionStart="FIGURES 12, 13" captionStartId="6.[151,264,1282,1306]" captionTargetBox="[159,1418,196,1242]" captionTargetId="figure-22@6.[151,1436,181,1258]" captionTargetPageId="6" captionText="FIGURES 12, 13. Amplaria oedipus, n. sp., male holotype. Fig. 12. Right gonopod, lateral view; aac, anterior angiocoxite; pac, posterior angiocoxite; lcc, lateral lobe of colpocoxite. Fig. 13. Right gonopod, mesal view; aac, anterior angiocoxite; fc, flagellocoxite; pcc, posterior lobe of colpocoxite; pac, posterior angiocoxite. Without scale." figureDoi="http://doi.org/10.5281/zenodo.6037037" httpUri="https://zenodo.org/record/6037037/files/figure.png" pageId="2" pageNumber="139">12, 13</figureCitation>
|
||
) about half the length of the anterior angiocoxites, long, projecting anterior branch sheaths at least 3 flagellocoxites (
|
||
<emphasis id="B925487BFFCA9149FE59FAAE518EFABB" bold="true" box="[409,431,1367,1393]" pageId="2" pageNumber="139">fc</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FE7CFAAE5223FABB" box="[444,514,1367,1393]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1781,1805]" captionTargetBox="[158,1430,309,1757]" captionTargetId="figure-75@5.[157,1430,309,1757]" captionTargetPageId="5" captionText="FIGURES 8–11. Amplaria oedipus, n. sp., male paratype. Fig. 8. Gonopods, posterior view; aac, anterior angiocoxite; cx, gonopod coxa; fc, flagellocoxite; lcc, lateral lobe of colpocoxite; pac, posterior angiocoxite; pcc, posterior lobe of colpocoxite. The lateral lobe of the colpocoxite has collapsed due to drying for SEM. Fig. 9. Tip of left anterior angiocoxite, mesal view. Fig. 10. Tips of right anterior and posterior angiocoxites, posterior view; aac, anterior angiocoxite; fc, flagellocoxite; pac, posterior angiocoxite. Fig. 11. Left leg 9, anterior view; cp, coxal process; cx9, coxa of leg 9." figureDoi="http://doi.org/10.5281/zenodo.6037035" httpUri="https://zenodo.org/record/6037035/files/figure.png" pageId="2" pageNumber="139">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FDCEFAAE520CFABB" box="[526,557,1367,1393]" captionStart="FIGURES 12, 13" captionStartId="6.[151,264,1282,1306]" captionTargetBox="[159,1418,196,1242]" captionTargetId="figure-22@6.[151,1436,181,1258]" captionTargetPageId="6" captionText="FIGURES 12, 13. Amplaria oedipus, n. sp., male holotype. Fig. 12. Right gonopod, lateral view; aac, anterior angiocoxite; pac, posterior angiocoxite; lcc, lateral lobe of colpocoxite. Fig. 13. Right gonopod, mesal view; aac, anterior angiocoxite; fc, flagellocoxite; pcc, posterior lobe of colpocoxite; pac, posterior angiocoxite. Without scale." figureDoi="http://doi.org/10.5281/zenodo.6037037" httpUri="https://zenodo.org/record/6037037/files/figure.png" pageId="2" pageNumber="139">13</figureCitation>
|
||
). Posterior lobes of colpocoxites (
|
||
<emphasis id="B925487BFFCA9149FC68FAAE53F1FABB" bold="true" box="[936,976,1367,1393]" pageId="2" pageNumber="139">pcc</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FC1EFAAE5402FABB" box="[990,1059,1367,1393]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1781,1805]" captionTargetBox="[158,1430,309,1757]" captionTargetId="figure-75@5.[157,1430,309,1757]" captionTargetPageId="5" captionText="FIGURES 8–11. Amplaria oedipus, n. sp., male paratype. Fig. 8. Gonopods, posterior view; aac, anterior angiocoxite; cx, gonopod coxa; fc, flagellocoxite; lcc, lateral lobe of colpocoxite; pac, posterior angiocoxite; pcc, posterior lobe of colpocoxite. The lateral lobe of the colpocoxite has collapsed due to drying for SEM. Fig. 9. Tip of left anterior angiocoxite, mesal view. Fig. 10. Tips of right anterior and posterior angiocoxites, posterior view; aac, anterior angiocoxite; fc, flagellocoxite; pac, posterior angiocoxite. Fig. 11. Left leg 9, anterior view; cp, coxal process; cx9, coxa of leg 9." figureDoi="http://doi.org/10.5281/zenodo.6037035" httpUri="https://zenodo.org/record/6037035/files/figure.png" pageId="2" pageNumber="139">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FBEFFAAE546AFABB" box="[1071,1099,1367,1393]" captionStart="FIGURES 12, 13" captionStartId="6.[151,264,1282,1306]" captionTargetBox="[159,1418,196,1242]" captionTargetId="figure-22@6.[151,1436,181,1258]" captionTargetPageId="6" captionText="FIGURES 12, 13. Amplaria oedipus, n. sp., male holotype. Fig. 12. Right gonopod, lateral view; aac, anterior angiocoxite; pac, posterior angiocoxite; lcc, lateral lobe of colpocoxite. Fig. 13. Right gonopod, mesal view; aac, anterior angiocoxite; fc, flagellocoxite; pcc, posterior lobe of colpocoxite; pac, posterior angiocoxite. Without scale." figureDoi="http://doi.org/10.5281/zenodo.6037037" httpUri="https://zenodo.org/record/6037037/files/figure.png" pageId="2" pageNumber="139">13</figureCitation>
|
||
) with finely spiculate surface. Lateral lobes of colpocoxite (
|
||
<emphasis id="B925487BFFCA9149FE29FA82522BFA5F" bold="true" box="[489,522,1403,1429]" pageId="2" pageNumber="139">lcc</emphasis>
|
||
) large. Ninth legpair (
|
||
<figureCitation id="136A88ECFFCA9149FCCAFA83537DFA5F" box="[778,860,1402,1429]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1781,1805]" captionTargetBox="[158,1430,309,1757]" captionTargetId="figure-75@5.[157,1430,309,1757]" captionTargetPageId="5" captionText="FIGURES 8–11. Amplaria oedipus, n. sp., male paratype. Fig. 8. Gonopods, posterior view; aac, anterior angiocoxite; cx, gonopod coxa; fc, flagellocoxite; lcc, lateral lobe of colpocoxite; pac, posterior angiocoxite; pcc, posterior lobe of colpocoxite. The lateral lobe of the colpocoxite has collapsed due to drying for SEM. Fig. 9. Tip of left anterior angiocoxite, mesal view. Fig. 10. Tips of right anterior and posterior angiocoxites, posterior view; aac, anterior angiocoxite; fc, flagellocoxite; pac, posterior angiocoxite. Fig. 11. Left leg 9, anterior view; cp, coxal process; cx9, coxa of leg 9." figureDoi="http://doi.org/10.5281/zenodo.6037035" httpUri="https://zenodo.org/record/6037035/files/figure.png" pageId="2" pageNumber="139">Fig. 11</figureCitation>
|
||
) with mesally hooked coxal process (
|
||
<emphasis id="B925487BFFCA9149FAD0FA82550DFA5F" bold="true" box="[1296,1324,1403,1429]" pageId="2" pageNumber="139">cp</emphasis>
|
||
,
|
||
<figureCitation id="136A88ECFFCA9149FAFCFA8355AFFA5F" box="[1340,1422,1402,1429]" captionStart="FIGURES 8–11" captionStartId="5.[151,264,1781,1805]" captionTargetBox="[158,1430,309,1757]" captionTargetId="figure-75@5.[157,1430,309,1757]" captionTargetPageId="5" captionText="FIGURES 8–11. Amplaria oedipus, n. sp., male paratype. Fig. 8. Gonopods, posterior view; aac, anterior angiocoxite; cx, gonopod coxa; fc, flagellocoxite; lcc, lateral lobe of colpocoxite; pac, posterior angiocoxite; pcc, posterior lobe of colpocoxite. The lateral lobe of the colpocoxite has collapsed due to drying for SEM. Fig. 9. Tip of left anterior angiocoxite, mesal view. Fig. 10. Tips of right anterior and posterior angiocoxites, posterior view; aac, anterior angiocoxite; fc, flagellocoxite; pac, posterior angiocoxite. Fig. 11. Left leg 9, anterior view; cp, coxal process; cx9, coxa of leg 9." figureDoi="http://doi.org/10.5281/zenodo.6037035" httpUri="https://zenodo.org/record/6037035/files/figure.png" pageId="2" pageNumber="139">Fig. 11</figureCitation>
|
||
), telopodite setose, flattened, mesally with recess receiving coxal process. Telopodites fitting into shallow depressions in ventrolateral margins of seventh diplotergite. Tenth leg coxae with coxal glands.
|
||
</paragraph>
|
||
<paragraph id="8BEE9469FFCA9149FF07FA1F5458F9E9" blockId="2.[151,1437,259,2037]" pageId="2" pageNumber="139">
|
||
<emphasis id="B925487BFFCA9149FF07FA1F51B1F9CA" box="[199,400,1510,1536]" italics="true" pageId="2" pageNumber="139">
|
||
Female
|
||
<typeStatus id="54EA2ACBFFCA9149FEE4FA1E51AAF9CA" box="[292,395,1511,1536]" pageId="2" pageNumber="139" type="paratype">paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Similar to male but with smaller collum, less projecting pygidium and lacking the greatly inflated tarsi on the 5th and 6th legpairs, and other male secondary sexual modifications.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34BC7E2FFCA914DFF07F9D4538BF868" lastPageId="6" lastPageNumber="143" pageId="2" pageNumber="139" type="distribution">
|
||
<paragraph id="8BEE9469FFCA9149FF07F9D453E4F98D" blockId="2.[151,1437,259,2037]" box="[199,965,1581,1608]" pageId="2" pageNumber="139">
|
||
<emphasis id="B925487BFFCA9149FF07F9D45142F98D" bold="true" box="[199,355,1581,1607]" pageId="2" pageNumber="139">Distribution:</emphasis>
|
||
Known only from Cape Mountain, Lane Co.,
|
||
<collectingRegion id="49955A8BFFCA9149FCA9F9D453E0F98D" box="[873,961,1581,1607]" country="United States of America" name="Oregon" pageId="2" pageNumber="139">Oregon</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEE9469FFCA9149FF07F9A85410F91C" blockId="2.[151,1437,259,2037]" pageId="2" pageNumber="139">
|
||
<emphasis id="B925487BFFCA9149FF07F9A85132F9A1" bold="true" box="[199,275,1617,1643]" pageId="2" pageNumber="139">Notes:</emphasis>
|
||
Collections were made by hand in early fall to late spring. Most searches were made along the main trail to the peak of Cape Mountain (accessed from the parking lot at Dry Lake Horse Camp). All specimens were collected from the vicinity of the peak. Litter and soil were examined, as well as the undersides of logs, rocks, and bark. Specimens were fixed in 90% isopropanol and preserved in 70-80% ethanol.
|
||
</paragraph>
|
||
<paragraph id="8BEE9469FFCA9149FF07F9195199F83F" blockId="2.[151,1437,259,2037]" pageId="2" pageNumber="139">
|
||
The habitat at this location is a Sitka Spruce (
|
||
<taxonomicName id="4C51EFEAFFCA9149FD1DF918547AF931" authority="(Bong.) Carriere" authorityName="Carriere" baseAuthorityName="Bong." box="[733,1115,1760,1787]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="139" phylum="Tracheophyta" rank="species" species="sitchensis">
|
||
<emphasis id="B925487BFFCA9149FD1DF91853B2F930" box="[733,915,1760,1786]" italics="true" pageId="2" pageNumber="139">Picea sitchensis</emphasis>
|
||
(Bong.) Carrière
|
||
</taxonomicName>
|
||
) forest. In
|
||
<collectingRegion id="49955A8BFFCA9149FB1DF9195515F930" box="[1245,1332,1760,1786]" country="United States of America" name="Oregon" pageId="2" pageNumber="139">Oregon</collectingRegion>
|
||
, this is a unique ecosystem found only in very close proximity to the Pacific coast, where the climate is mild and wet (Franklin and Dyrness 1988); Cape Mountain lies approximately
|
||
<quantity id="4CA9398CFFCA9149FCE1F8DE5378F88B" box="[801,857,1831,1858]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="139" unit="km" value="5.0">5 km</quantity>
|
||
from the coast. Sitka Spruce is the dominant conifer in this habitat, and is supplemented by Western Red Cedar (
|
||
<taxonomicName id="4C51EFEAFFCA9149FCFDF8B254B8F8AC" authority="Donn ex D. Don" authorityName="D. Don" box="[829,1177,1867,1894]" class="Pinopsida" family="Cupressaceae" genus="Thuja" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="139" phylum="Tracheophyta" rank="species" species="plicata">
|
||
<emphasis id="B925487BFFCA9149FCFDF8B253F5F8AF" box="[829,980,1867,1893]" italics="true" pageId="2" pageNumber="139">Thuja plicata</emphasis>
|
||
Donn ex D. Don
|
||
</taxonomicName>
|
||
) and Red Alder (
|
||
<taxonomicName id="4C51EFEAFFCA9149FA9CF8B25106F843" authority="Bong." authorityName="Bong." class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="139" phylum="Tracheophyta" rank="species" species="rubra">
|
||
<emphasis id="B925487BFFCA9149FA9CF8B250F6F843" italics="true" pageId="2" pageNumber="139">Alnus rubra</emphasis>
|
||
Bong.
|
||
</taxonomicName>
|
||
) in disturbed areas. The undergrowth consists of dense Salal (
|
||
<taxonomicName id="4C51EFEAFFCA9149FBC4F896550AF840" authority="Pursh" authorityName="Pursh" box="[1028,1323,1903,1930]" class="Magnoliopsida" family="Ericaceae" genus="Gaultheria" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="139" phylum="Tracheophyta" rank="species" species="shallon">
|
||
<emphasis id="B925487BFFCA9149FBC4F89654FDF843" box="[1028,1244,1903,1929]" italics="true" pageId="2" pageNumber="139">Gaultheria shallon</emphasis>
|
||
Pursh
|
||
</taxonomicName>
|
||
), Western Sword-fern (
|
||
<taxonomicName id="4C51EFEAFFCA9149FEE6F86B52F9F867" authority="(Kaulf.) C. Presl" authorityName="C. Presl" baseAuthorityName="Kaulf." box="[294,728,1938,1965]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="139" phylum="Tracheophyta" rank="species" species="munitum">
|
||
<emphasis id="B925487BFFCA9149FEE6F86B5239F866" box="[294,536,1938,1964]" italics="true" pageId="2" pageNumber="139">Polystichum munitum</emphasis>
|
||
(Kaulf.) C. Presl
|
||
</taxonomicName>
|
||
), Salmonberry (
|
||
<taxonomicName id="4C51EFEAFFCA9149FC4AF86B54B6F867" authority="Pursh" authorityName="Pursh" box="[906,1175,1938,1965]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="139" phylum="Tracheophyta" rank="species" species="spectabilis">
|
||
<emphasis id="B925487BFFCA9149FC4AF86B546FF866" box="[906,1102,1938,1964]" italics="true" pageId="2" pageNumber="139">Rubus spectabilis</emphasis>
|
||
Pursh
|
||
</taxonomicName>
|
||
), Red Elderberry (
|
||
<taxonomicName id="4C51EFEAFFCA9149FAA3F86A51E3F81B" authority="Linnaeus" authorityName="Linnaeus" class="Magnoliopsida" family="Adoxaceae" genus="Sambucus" kingdom="Plantae" order="Dipsacales" pageId="2" pageNumber="139" phylum="Tracheophyta" rank="species" species="racemosa">
|
||
<emphasis id="B925487BFFCA9149FAA3F86A516FF81A" italics="true" pageId="2" pageNumber="139">Sambucus racemosa</emphasis>
|
||
Linnaeus
|
||
</taxonomicName>
|
||
), Evergreen Huckleberry (
|
||
<taxonomicName id="4C51EFEAFFCA9149FD30F84E542CF81B" authority="Pursh" authorityName="Pursh" box="[752,1037,1974,2001]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="139" phylum="Tracheophyta" rank="species" species="ovatum">
|
||
<emphasis id="B925487BFFCA9149FD30F84E53E1F81A" box="[752,960,1975,2000]" italics="true" pageId="2" pageNumber="139">Vaccinium ovatum</emphasis>
|
||
Pursh
|
||
</taxonomicName>
|
||
), and Red Huckleberry (
|
||
<taxonomicName id="4C51EFEAFFCA9149FAE8F84E516FF83F" authority="Sm." authorityName="Sm." class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="139" phylum="Tracheophyta" rank="species" species="parvifolium">
|
||
<emphasis id="B925487BFFCA9149FAE8F84E513BF83E" italics="true" pageId="2" pageNumber="139">Vaccinium parvifolium</emphasis>
|
||
Sm.
|
||
</taxonomicName>
|
||
in Rees).
|
||
</paragraph>
|
||
<paragraph id="8BEE9469FFCD914EFF07FF6E52F4FED7" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="142">
|
||
<taxonomicName id="4C51EFEAFFCD914EFF07FF6E51A7FF7B" box="[199,390,151,177]" class="Diplopoda" family="Striariidae" genus="Amplaria" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="oedipu">
|
||
<emphasis id="B925487BFFCD914EFF07FF6E51A7FF7B" box="[199,390,151,177]" italics="true" pageId="5" pageNumber="142">Amplaria oedipu</emphasis>
|
||
</taxonomicName>
|
||
s was always found at or near the litter surface. The male
|
||
<typeStatus id="54EA2ACBFFCD914EFBE3FF6E54A2FF7B" box="[1059,1155,151,177]" pageId="5" pageNumber="142" type="paratype">paratype</typeStatus>
|
||
was found underneath a small, broken branch which also supported platydesmid and rhiscosomid millipedes. The male
|
||
<typeStatus id="54EA2ACBFFCD914EFB79FF42553AFF1F" box="[1209,1307,187,213]" pageId="5" pageNumber="142" type="holotype">holotype</typeStatus>
|
||
and female
|
||
<typeStatus id="54EA2ACBFFCD914EFF57FF2650D6FF33" box="[151,247,223,249]" pageId="5" pageNumber="142" type="paratype">paratype</typeStatus>
|
||
were found in leaf litter.
|
||
<taxonomicName id="4C51EFEAFFCD914EFDD8FF2652C0FF33" authorityName="Shear & Nosler & Marek" authorityYear="2022" box="[536,737,223,249]" class="Diplopoda" family="Striariidae" genus="Amplaria" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="142" phylum="Arthropoda" rank="species" species="oedipus">
|
||
<emphasis id="B925487BFFCD914EFDD8FF2652C0FF33" box="[536,737,223,249]" italics="true" pageId="5" pageNumber="142">Amplaria oedipus</emphasis>
|
||
</taxonomicName>
|
||
was found in places rich with other chordeumatidan families, including
|
||
<taxonomicName id="4C51EFEAFFCD914EFEC8FEFA515FFED7" authorityName="Verhoeff" authorityYear="1909" box="[264,382,259,285]" class="Diplopoda" family="Caseyidae" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="142" phylum="Arthropoda" rank="family">Caseyidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C51EFEAFFCD914EFE49FEFA522AFED7" authorityName="Loomis" authorityYear="1966" box="[393,523,259,285]" class="Diplopoda" family="Tingupidae" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="142" phylum="Arthropoda" rank="family">Tingupidae</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C51EFEAFFCD914EFD85FEFA52F3FED7" authorityName="Cook" authorityYear="1896" box="[581,722,259,285]" class="Diplopoda" family="Conotylidae" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="142" phylum="Arthropoda" rank="family">Conotylidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEE9469FFCE914DFF07FA69529AF900" blockId="6.[151,1437,1424,1955]" pageId="6" pageNumber="143">
|
||
When the first male was collected by PN and sent to WS for identification we suspected the strange tarsi might have been due to a parasite, such as a nematode or a fly larva. However, dissection of one of the tarsi revealed neither, just amorphous tissue. Also, why would a parasitic attack be limited to just those tarsi? Another consideration was that a random mutation may have occurred in the single male. The collection by PN of another male with the exact same greatly inflated tarsi on the 5th and 6th legpairs spoke against both the parasite and mutation hypotheses. We therefore think that the evidence at hand supports our conclusion that the modifications are normal for the mature males of this species. In our experience, such a tarsal condition is not known from any other species of the class
|
||
<taxonomicName id="4C51EFEAFFCE914DFF14F975516CF96C" authorityName="de Blainville in Gervais" authorityYear="1844" box="[212,333,1676,1702]" class="Diplopoda" kingdom="Animalia" pageId="6" pageNumber="143" phylum="Arthropoda" rank="class">Diplopoda</taxonomicName>
|
||
, though enlarged other podomeres (usually the prefemora) are not unusual. Aside from the enlarged tarsi,
|
||
<taxonomicName id="4C51EFEAFFCE914DFF15F948516AF900" authorityName="Shear & Nosler & Marek" authorityYear="2022" box="[213,331,1712,1739]" class="Diplopoda" family="Striariidae" genus="Amplaria" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="oedipus">
|
||
<emphasis id="B925487BFFCE914DFF15F948516AF900" box="[213,331,1712,1739]" italics="true" pageId="6" pageNumber="143">A. oedipus</emphasis>
|
||
</taxonomicName>
|
||
is a typical species of
|
||
<emphasis id="B925487BFFCE914DFD89F949529AF900" box="[585,699,1712,1738]" italics="true" pageId="6" pageNumber="143">
|
||
<taxonomicName id="4C51EFEAFFCE914DFD89F9495297F900" authorityName="Chamberlin" authorityYear="1941" box="[585,694,1712,1738]" class="Diplopoda" family="Striariidae" genus="Amplaria" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="143" phylum="Arthropoda" rank="genus">Amplaria</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEE9469FFCE914DFF07F92D54D6F8FD" blockId="6.[151,1437,1424,1955]" pageId="6" pageNumber="143">
|
||
The unique nature of this modification recalls the eastern North American species
|
||
<taxonomicName id="4C51EFEAFFCE914DFBB8F92D523FF8D9" authority="Shear, 2011" authorityName="Shear" authorityYear="2011" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="peponocranium">
|
||
<emphasis id="B925487BFFCE914DFBB8F92D50EEF8D8" italics="true" pageId="6" pageNumber="143">Pseudotremia peponocranium</emphasis>
|
||
<bibRefCitation id="EFC0E998FFCE914DFF18F901517FF8D8" author="Shear, W. A." box="[216,350,1784,1810]" pageId="6" pageNumber="143" pagination="1 - 38" refId="ref6014" refString="Shear, W. A. (2011) Cave millipedesof the United States. X. New species and records of the genus Pseudotremia Cope. 2. Species from Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae). Zootaxa, 3109, 1 - 38." type="journal article" year="2011">Shear, 2011</bibRefCitation>
|
||
(Cleidogonidae)
|
||
</taxonomicName>
|
||
which differs from all other members of its genus in having a large, swollen, quadrate projection on the front of the head in both males and females (
|
||
<bibRefCitation id="EFC0E998FFCE914DFC77F8E55442F8FD" author="Shear, W. A." box="[951,1123,1820,1847]" pageId="6" pageNumber="143" pagination="1 - 38" refId="ref6014" refString="Shear, W. A. (2011) Cave millipedesof the United States. X. New species and records of the genus Pseudotremia Cope. 2. Species from Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae). Zootaxa, 3109, 1 - 38." type="journal article" year="2011">Shear 2011: 15</bibRefCitation>
|
||
, figs 34, 35).
|
||
</paragraph>
|
||
<paragraph id="8BEE9469FFCE914DFF07F8B9538BF868" blockId="6.[151,1437,1424,1955]" pageId="6" pageNumber="143">
|
||
A single male and
|
||
<specimenCount id="9D575FE0FFCE914DFE61F8B85201F890" box="[417,544,1856,1883]" pageId="6" pageNumber="143" type="female">two female</specimenCount>
|
||
<taxonomicName id="4C51EFEAFFCE914DFDEAF8B952B5F890" authorityName="Chamberlin" authorityYear="1941" box="[554,660,1856,1882]" class="Diplopoda" family="Striariidae" genus="Amplaria" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="143" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B925487BFFCE914DFDEAF8B952B5F890" box="[554,660,1856,1882]" italics="true" pageId="6" pageNumber="143">Amplaria</emphasis>
|
||
</taxonomicName>
|
||
of another, undescribed species were collected at a lower elevation on Cape Mountain (
|
||
<geoCoordinate id="EE65F2AEFFCE914DFEB8F89D51D6F8B4" box="[376,503,1892,1918]" degrees="44.0910" direction="north" orientation="latitude" pageId="6" pageNumber="143" precision="5" value="44.091">44.0910°N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate id="EE65F2AEFFCE914DFDCAF89D5281F8B4" box="[522,672,1892,1918]" degrees="124.0737" direction="west" orientation="longitude" pageId="6" pageNumber="143" precision="5" value="-124.0737">124.0737°W</geoCoordinate>
|
||
). The gonopods of this species, to be described later, are similar but distinct from those of
|
||
<taxonomicName id="4C51EFEAFFCE914DFE4EF8705224F868" authorityName="Shear & Nosler & Marek" authorityYear="2022" box="[398,517,1928,1955]" class="Diplopoda" family="Striariidae" genus="Amplaria" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="oedipus">
|
||
<emphasis id="B925487BFFCE914DFE4EF8705224F868" box="[398,517,1928,1955]" italics="true" pageId="6" pageNumber="143">A. oedipus</emphasis>
|
||
</taxonomicName>
|
||
and the male lacks the enlarged tarsi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |