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<mods:title id="57ED3440171A0C41E675AC0C93B3F533">“ Larger ” Benthic Foraminifera Of The Cenomanian. A Review Of The Identity And The Stratigraphic And Palaeogeographic Distribution Of Non-Fusiform Planispiral (Or Near-Planispiral) Forms</mods:title>
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<mods:namePart id="3423D1995C8D5F6E56E11417077ADB2D">SIMMONS, MICHAEL</mods:namePart>
<mods:affiliation id="DC9F762A5C15AF81DD0301D6780D2642">Halliburton, 97 Milton Park, Abingdon, OX 14 4 RW, UK &amp; The Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK</mods:affiliation>
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<paragraph id="8BF336A0FFF6A23CFF11FE1EA719C6AB" blockId="47.[151,771,389,480]" box="[151,596,456,480]" pageId="47" pageNumber="86">
<taxonomicName id="4C4C4D23FFF6A23CFF11FE1EA719C6AB" ID-CoL="BDQXN" authority="(Henson, 1948)" baseAuthorityName="Henson" baseAuthorityYear="1948" box="[151,596,456,480]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFF11FE1EA719C6AB" bold="true" box="[151,596,456,480]" pageId="47" pageNumber="86">
<emphasis id="B938EAB2FFF6A23CFF11FE1EA4EFC6AB" bold="true" box="[151,418,456,480]" italics="true" pageId="47" pageNumber="86">Hemicyclammina whitei</emphasis>
(
<bibRefCitation id="EFDD4B51FFF6A23CFE36FE1EA701C6AB" author="Henson, F. R. S." box="[432,588,456,480]" pageId="47" pageNumber="86" refId="ref83829" refString="Henson, F. R. S., 1948. Larger Imperforate Foraminifera of South-Western Asia. British Museum (Natural History), London, pp. xi- 127." type="book" year="1948">Henson, 1948</bibRefCitation>
)
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C356652BFFF6A23CFF11FDDCA7D9C52F" pageId="47" pageNumber="86" type="reference_group">
<paragraph id="8BF336A0FFF6A23CFF11FDDCA750C569" blockId="47.[151,541,522,546]" box="[151,541,522,546]" pageId="47" pageNumber="86">Reference Illustration &amp; Description</paragraph>
<paragraph id="8BF336A0FFF6A23CFF11FD9AA7D9C52F" blockId="47.[151,772,588,2069]" box="[151,660,588,612]" pageId="47" pageNumber="86">
Simmons &amp;
<bibRefCitation id="EFDD4B51FFF6A23CFE98FD9AA48AC52F" author="Bidgood, M. D. &amp; Simmons, M. D." box="[286,455,588,612]" pageId="47" pageNumber="86" refId="ref74158" refString="Bidgood, M. D. &amp; Simmons, M. D., 2022. Cenomanian planktonic foraminifera, bioevents and biozonation - a brief review. Newsletters on Stratigraphy, published online. DOI: 10.1127 / nos / 2022 / 0717" type="book" year="2022">Bidgood (2022)</bibRefCitation>
p. 27-30, figs. 1-4.
</paragraph>
</subSubSection>
<subSubSection id="C356652BFFF6A23CFF11FDBBA11DC54A" pageId="47" pageNumber="86" type="description">
<paragraph id="8BF336A0FFF6A23CFF11FDBBA439C49B" blockId="47.[151,772,588,2069]" pageId="47" pageNumber="86">
The status of
<taxonomicName id="4C4C4D23FFF6A23CFEABFDB8A4DDC5CE" baseAuthorityName="Henson" baseAuthorityYear="1948" box="[301,400,621,645]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFEABFDB8A4DDC5CE" box="[301,400,621,645]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
with respect to its senior synonymy with
<taxonomicName id="4C4C4D23FFF6A23CFF71FD59A4E9C5EC" authority="Maync" authorityName="Maync" authorityYear="1953" box="[247,420,654,679]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="sigali">
<emphasis id="B938EAB2FFF6A23CFF71FD59A41FC5ED" box="[247,338,654,678]" italics="true" pageId="47" pageNumber="86">H. sigali</emphasis>
Maync
</taxonomicName>
and a number of other taxa have been discussed by Simmons &amp;
<bibRefCitation id="EFDD4B51FFF6A23CFD81FD79A7F7C58C" author="Bidgood, M. D. &amp; Simmons, M. D." box="[519,698,687,711]" pageId="47" pageNumber="86" refId="ref74158" refString="Bidgood, M. D. &amp; Simmons, M. D., 2022. Cenomanian planktonic foraminifera, bioevents and biozonation - a brief review. Newsletters on Stratigraphy, published online. DOI: 10.1127 / nos / 2022 / 0717" type="book" year="2022">Bidgood (2022)</bibRefCitation>
. They noted that when
<bibRefCitation id="EFDD4B51FFF6A23CFEDBFD07A747C5A2" author="Maync, W." box="[349,522,721,745]" pageId="47" pageNumber="86" pagination="148 - 150" refId="ref88941" refString="Maync, W., 1953 b. Hemicyclammina sigali n. gen., n. sp. from the Cenomanian of Algeria. Contributions from the Cushman Foundation for Foraminiferal Research, 4: 148 - 150." type="journal article" year="1953">Maync (1953b)</bibRefCitation>
introduced the genus
<taxonomicName id="4C4C4D23FFF6A23CFF11FD24A41CC441" authorityName="Maync" authorityYear="1953" box="[151,337,754,778]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFF6A23CFF11FD24A41CC441" box="[151,337,754,778]" italics="true" pageId="47" pageNumber="86">Hemicyclammina</emphasis>
</taxonomicName>
, he was seemingly unaware of the publication of
<bibRefCitation id="EFDD4B51FFF6A23CFE95FCC5A4FBC460" author="Henson, F. R. S." box="[275,438,787,811]" pageId="47" pageNumber="86" refId="ref83829" refString="Henson, F. R. S., 1948. Larger Imperforate Foraminifera of South-Western Asia. British Museum (Natural History), London, pp. xi- 127." type="book" year="1948">Henson (1948)</bibRefCitation>
that introduced “
<taxonomicName id="4C4C4D23FFF6A23CFDFBFCC5A595C407" authorityName="Henson" authorityYear="1948" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFDFBFCC5A595C407" italics="true" pageId="47" pageNumber="86">Cyclammina whitei</emphasis>
</taxonomicName>
”. Examination of
<typeStatus id="54F78802FFF6A23CFE20FCE3A499C406" box="[422,468,821,845]" pageId="47" pageNumber="86">type</typeStatus>
and associated material indicates that this species is identical to
<taxonomicName id="4C4C4D23FFF6A23CFDB9FC80A7D3C426" authorityName="Maync" authorityYear="1953" box="[575,670,853,877]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="sigali">
<emphasis id="B938EAB2FFF6A23CFDB9FC80A7D3C426" box="[575,670,853,877]" italics="true" pageId="47" pageNumber="86">H. sigali</emphasis>
</taxonomicName>
and thus should be regarded as the
<typeStatus id="54F78802FFF6A23CFE3EFCA1A4ABC4C4" box="[440,486,887,911]" pageId="47" pageNumber="86">type</typeStatus>
species of the genus. See the Species Key Chart (Appendix) for diagnostic and other characteristics.
</paragraph>
<paragraph id="8BF336A0FFF6A23CFF11FC0FA467C148" blockId="47.[151,772,588,2069]" pageId="47" pageNumber="86">
The alveolar nature of the wall but with solid, pointed septa which do not reach the previous whorl in equatorial section is characteristic. The solid (“semi-” or “hemi-”) septa serve to distinguish the genus from
<taxonomicName id="4C4C4D23FFF6A23CFDF3FBEBA64EC31F" baseAuthorityName="Loeblich &amp; Tappan" baseAuthorityYear="1949" box="[629,771,1085,1108]" class="Globothalamea" family="Cyclamminidae" genus="Buccicrenata" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFF6A23CFDF3FBEBA64EC31F" box="[629,771,1085,1108]" italics="true" pageId="47" pageNumber="86">Buccicrenata</emphasis>
</taxonomicName>
with alveolar septa which are continuous outgrowths of the alveolar chamber wall.
<taxonomicName id="4C4C4D23FFF6A23CFE5FFBA9A7E5C3DC" authorityName="Yabe &amp; Hanzawa" authorityYear="1926" box="[473,680,1151,1175]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFF6A23CFE5FFBA9A7E5C3DC" box="[473,680,1151,1175]" italics="true" pageId="47" pageNumber="86">Pseudocyclammina</emphasis>
</taxonomicName>
is also similar but in addition to alveolar septa also has multiple apertures compared to the single slit of
<taxonomicName id="4C4C4D23FFF6A23CFDC4FB17A7B1C392" authorityName="Maync" authorityYear="1953" box="[578,764,1217,1241]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFF6A23CFDC4FB17A7B1C392" box="[578,764,1217,1241]" italics="true" pageId="47" pageNumber="86">Hemicyclammina</emphasis>
</taxonomicName>
.
<taxonomicName id="4C4C4D23FFF6A23CFF11FB34A485C3B1" authority="Redmond" authorityName="Redmond" box="[151,456,1250,1274]" class="Globothalamea" family="Everticyclamminidae" genus="Everticyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFF6A23CFF11FB34A414C3B1" box="[151,345,1250,1274]" italics="true" pageId="47" pageNumber="86">Everticyclammina</emphasis>
Redmond
</taxonomicName>
(Late Jurassic Early Cretaceous) is very similar in almost every respect, but its aperture is areal rather than extending upwards from the base as in
<taxonomicName id="4C4C4D23FFF6A23CFE9BFA93A49AC216" authorityName="Maync" authorityYear="1953" box="[285,471,1349,1373]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFF6A23CFE9BFA93A49AC216" box="[285,471,1349,1373]" italics="true" pageId="47" pageNumber="86">Hemicyclammina</emphasis>
</taxonomicName>
and which is also rather large, resulting in shorter septa. The poorly known genus
<taxonomicName id="4C4C4D23FFF6A23CFF11FA51A494C2D4" authority="Hillebrandt" authorityName="Hillebrandt" authorityYear="1971" box="[151,473,1415,1439]" class="Globothalamea" family="Cyclamminidae" genus="Alveocyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFF6A23CFF11FA51A418C2D4" box="[151,341,1415,1439]" italics="true" pageId="47" pageNumber="86">Alveocyclammina</emphasis>
Hillebrandt
</taxonomicName>
from the lower Albian of
<collectingCountry id="F35B7630FFF6A23CFF11FA7FA585C28B" box="[151,200,1449,1472]" name="Peru" pageId="47" pageNumber="86">Peru</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFF6A23CFF5BFA7FA4EFC28A" author="Hillebrandt, A." box="[221,418,1449,1473]" pageId="47" pageNumber="86" pagination="11 - 16" refId="ref83912" refString="Hillebrandt, A. v., 1971. Spirocyclinidae (Foraminiferen) aus der Unterkreide von Peru. Mitteilungen der Bayerischen Staatssammlung fur Palaontologie und Historische Geologie, 11: 11 - 16." type="journal article" year="1971">Hillebrandt, 1971</bibRefCitation>
) is also similar possessing an alveolar wall, but its septa are also alveolar and appear to be very short.
</paragraph>
<paragraph id="8BF336A0FFF6A23CFF11F9DAA592C024" blockId="47.[151,772,588,2069]" pageId="47" pageNumber="86">
The following poorly known taxa are considered by Simmons &amp;
<bibRefCitation id="EFDD4B51FFF6A23CFEA7F9FBA481C10E" author="Bidgood, M. D. &amp; Simmons, M. D." box="[289,460,1581,1605]" pageId="47" pageNumber="86" refId="ref74158" refString="Bidgood, M. D. &amp; Simmons, M. D., 2022. Cenomanian planktonic foraminifera, bioevents and biozonation - a brief review. Newsletters on Stratigraphy, published online. DOI: 10.1127 / nos / 2022 / 0717" type="book" year="2022">Bidgood (2022)</bibRefCitation>
to be probable synonyms of
<taxonomicName id="4C4C4D23FFF6A23CFF11F999A5B2C12D" authorityName="Henson" authorityYear="1948" box="[151,255,1614,1638]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFF11F999A5B2C12D" box="[151,255,1614,1638]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
:
<taxonomicName id="4C4C4D23FFF6A23CFE95F998A7D4C12D" authority="Hamaoui" authorityName="Hamaoui" box="[275,665,1614,1638]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="evoluta">
<emphasis id="B938EAB2FFF6A23CFE95F998A767C12D" box="[275,554,1614,1638]" italics="true" pageId="47" pageNumber="86">Hemicyclammina evoluta</emphasis>
Hamaoui
</taxonomicName>
,
<taxonomicName id="4C4C4D23FFF6A23CFD2AF998A4D8C1CC" authority="El-Dakkak" authorityName="El-Dakkak" authorityYear="1974" class="Globothalamea" family="Charentiidae" genus="Ismailia" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="neumannae">
<emphasis id="B938EAB2FFF6A23CFD2AF998A45FC1CC" italics="true" pageId="47" pageNumber="86">Ismailia neumannae</emphasis>
El-Dakkak
</taxonomicName>
, and
<taxonomicName id="4C4C4D23FFF6A23CFE64F9B9A5BCC1E3" authority="El-Dakkak." authorityName="El-Dakkak." class="Globothalamea" family="Charentiidae" genus="Sinainella" higherTaxonomySource="CoL,GBIF" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="aegyptiaca">
<emphasis id="B938EAB2FFF6A23CFE64F9B9A79EC1CC" box="[482,723,1647,1671]" italics="true" pageId="47" pageNumber="86">Sinainella aegyptiaca</emphasis>
El-Dakkak.
</taxonomicName>
<taxonomicName id="4C4C4D23FFF6A23CFF7EF947A4C7C1E3" authorityName="El-Dakkak" authorityYear="1974" box="[248,394,1681,1704]" class="Globothalamea" family="Charentiidae" genus="Ismailia" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="neumannae">
<emphasis id="B938EAB2FFF6A23CFF7EF947A4C7C1E3" box="[248,394,1681,1704]" italics="true" pageId="47" pageNumber="86">I. neumannae</emphasis>
</taxonomicName>
continues to be a name used in the Egyptian literature for specimens that might be
<taxonomicName id="4C4C4D23FFF6A23CFD1CF964A7B1C182" authorityName="Henson" authorityYear="1948" box="[666,764,1713,1737]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFD1CF964A7B1C182" box="[666,764,1713,1737]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
, although with external views that are not diagnostic (e.g., Shahin &amp; El Baz, 2021). A specimen illustrated from the late Cenomanian of
<collectingCountry id="F35B7630FFF6A23CFEFBF8C3A4F0C066" box="[381,445,1813,1837]" name="Egypt" pageId="47" pageNumber="86">Egypt</collectingCountry>
as
<taxonomicName id="4C4C4D23FFF6A23CFE6CF8C3A59FC005" authority="Neumann" authorityName="Neumann" class="Globothalamea" family="Charentiidae" genus="Charentia" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="cuvillieri">
<emphasis id="B938EAB2FFF6A23CFE6CF8C3A78FC066" box="[490,706,1813,1837]" italics="true" pageId="47" pageNumber="86">Charentia cuvillieri</emphasis>
Neumann
</taxonomicName>
by El-Sheikh and Hewaidy (1998) looks close to
<taxonomicName id="4C4C4D23FFF6A23CFD6FF8E1A595C024" authorityName="Henson" authorityYear="1948" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFD6FF8E1A595C024" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF336A0FFF6A23CFF11F8AEA01BC636" blockId="47.[151,772,588,2069]" lastBlockId="47.[816,1437,158,513]" pageId="47" pageNumber="86">
<taxonomicName id="4C4C4D23FFF6A23CFF11F8AEA48FC0DB" ID-CoL="826NQ" authority="Basha" authorityName="Basha" authorityYear="1978" box="[151,450,1912,1936]" class="Globothalamea" family="Charentiidae" genus="Charentia" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="hasaensis">
<emphasis id="B938EAB2FFF6A23CFF11F8AEA438C0DB" box="[151,373,1912,1936]" italics="true" pageId="47" pageNumber="86">Charentia hasaensis</emphasis>
Basha
</taxonomicName>
,
<taxonomicName id="4C4C4D23FFF6A23CFE49F8AEA5F7C0FA" ID-CoL="826NT" authority="Basha" authorityName="Basha" authorityYear="1978" class="Globothalamea" family="Charentiidae" genus="Charentia" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="rummanensis">
<emphasis id="B938EAB2FFF6A23CFE49F8AEA79EC0DB" box="[463,723,1912,1936]" italics="true" pageId="47" pageNumber="86">Charentia rummanensis</emphasis>
Basha
</taxonomicName>
and
<taxonomicName id="4C4C4D23FFF6A23CFF70F84FA4B2C0FA" ID-CoL="82WCM" authority="Basha" authorityName="Basha" authorityYear="1978" box="[246,511,1945,1969]" class="Globothalamea" family="Lituolidae" genus="Lituola" kingdom="Chromista" order="Lituolida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="hasaensis">
<emphasis id="B938EAB2FFF6A23CFF70F84FA4F9C0FA" box="[246,436,1945,1969]" italics="true" pageId="47" pageNumber="86">Lituola hasaensis</emphasis>
Basha
</taxonomicName>
are poorly known species introduced from material from the late Cenomanian of
<collectingCountry id="F35B7630FFF6A23CFF3EF80DA5B2C0B8" box="[184,255,2011,2035]" name="Jordan" pageId="47" pageNumber="86">Jordan</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFF6A23CFE97F80DA4D7C0B8" author="Basha, S. H." box="[273,410,2011,2035]" pageId="47" pageNumber="86" pagination="67 - 91" refId="ref73406" refString="Basha, S. H., 1978. Foraminifera from the Ajlun Group of East Jordan. Journal Geological Society of Iraq, 11: 67 - 91." type="journal article" year="1978">Basha, 1978</bibRefCitation>
). From the limited illustrations they may be partly synonymous with
<taxonomicName id="4C4C4D23FFF6A23CFDB4F82BA7D8CF5F" ID-CoL="82BK8" authorityName="Henson" authorityYear="1948" box="[562,661,2044,2068]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFDB4F82BA7D8CF5F" box="[562,661,2044,2068]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
. The
<typeStatus id="54F78802FFF6A23CFD53F82BA64FCF5E" box="[725,770,2045,2069]" pageId="47" pageNumber="86">type</typeStatus>
material needs to be re-examined. The same author reports but does not illustrate
<taxonomicName id="4C4C4D23FFF6A23CFBE8FF16A1BCC79C" ID-CoL="826NM" box="[1134,1265,191,215]" class="Globothalamea" family="Charentiidae" genus="Charentia" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="cuvillieri">
<emphasis id="B938EAB2FFF6A23CFBE8FF16A1BCC79C" box="[1134,1265,191,215]" italics="true" pageId="47" pageNumber="86">C. cuvillieri</emphasis>
</taxonomicName>
from the same rocks as his new species and he also reports the occurrence of
<taxonomicName id="4C4C4D23FFF6A23CFC15FED4A6BCC652" ID-CoL="82QQ5" authorityName="Maync" authorityYear="1953" box="[915,1009,257,281]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="sigali">
<emphasis id="B938EAB2FFF6A23CFC15FED4A6BCC652" box="[915,1009,257,281]" italics="true" pageId="47" pageNumber="86">H. sigali</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D23FFF6A23CFBAFFED4A1C0C652" ID-CoL="82BK8" authorityName="Henson" authorityYear="1948" box="[1065,1165,257,281]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFBAFFED4A1C0C652" box="[1065,1165,257,281]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
, but without illustration. Likewise,
<taxonomicName id="4C4C4D23FFF6A23CFC2EFEF4A025C671" ID-CoL="826NP" authority="Kerdany &amp; Eissa" authorityName="Kerdany &amp; Eissa" authorityYear="1973" box="[936,1384,290,314]" class="Globothalamea" family="Charentiidae" genus="Charentia" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="granulosa">
<emphasis id="B938EAB2FFF6A23CFC2EFEF4A1C2C671" box="[936,1167,290,314]" italics="true" pageId="47" pageNumber="86">Charentia granulosa</emphasis>
Kerdany &amp; Eissa
</taxonomicName>
, described from the late Cenomanian of
<collectingCountry id="F35B7630FFF6A23CFB56FE92A05DC617" box="[1232,1296,324,348]" name="Egypt" pageId="47" pageNumber="86">Egypt</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFF6A23CFAA7FE95A6C0C637" author="Kerdany, M. T. &amp; Eissa, R. A. &amp; Labib, F." pageId="47" pageNumber="86" pagination="89 - 96" refId="ref86364" refString="Kerdany, M. T., Eissa, R. A. &amp; Labib, F., 1973. Quelques foraminiferes Cenomaniens de la partie ouest de la region du Golfe de Suez (Egypte). Revue de Micropaleontologie, 16 (2): 89 - 96." type="journal article" year="1973">Kerdany et al., 1973</bibRefCitation>
), may include
<taxonomicName id="4C4C4D23FFF6A23CFBABFEB3A1C2C637" ID-CoL="82BK8" authorityName="Henson" authorityYear="1948" box="[1069,1167,356,380]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFBABFEB3A1C2C637" box="[1069,1167,356,380]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
amongst its
<typeStatus id="54F78802FFF6A23CFA90FEB3A01FC636" box="[1302,1362,357,381]" pageId="47" pageNumber="86">types</typeStatus>
.
</paragraph>
<paragraph id="8BF336A0FFF6A23CFCB6FE53A11DC54A" blockId="47.[816,1437,158,513]" pageId="47" pageNumber="86">
<bibRefCitation id="EFDD4B51FFF6A23CFCB6FE53A6A2C6D6" author="Dimitrova, E." box="[816,1007,389,413]" pageId="47" pageNumber="86" pagination="79 - 82" refId="ref79696" refString="Dimitrova, E., 1995. Foraminifera from the Cenomanian of the &quot; Cerovska Reka &quot; Section, Strandza region, Bulgaria. Geologica Balcanica, 25 (5 - 6): 79 - 82." type="journal article" year="1995">Dimitrova (1995)</bibRefCitation>
described “
<taxonomicName id="4C4C4D23FFF6A23CFBF2FE53A0DCC6D6" box="[1140,1425,389,413]" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="B938EAB2FFF6A23CFBF2FE53A025C6D6" box="[1140,1384,389,413]" italics="true" pageId="47" pageNumber="86">Pseudonummoloculina</emphasis>
sp.
</taxonomicName>
” from the Cenomanian of
<collectingCountry id="F35B7630FFF6A23CFBC1FE71A1EFC6F4" box="[1095,1186,423,447]" name="Bulgaria" pageId="47" pageNumber="86">Bulgaria</collectingCountry>
and compared it to
<taxonomicName id="4C4C4D23FFF6A23CFA00FE7EA6EDC6AB" ID-CoL="83DPH" authorityName="Calvez" authorityYear="1988" class="Tubothalamea" family="Miliolidae" genus="Pseudonummoloculina" kingdom="Chromista" order="Miliolida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="aurigerica">
<emphasis id="B938EAB2FFF6A23CFA00FE7EA6EDC6AB" italics="true" pageId="47" pageNumber="86">P. aurigerica</emphasis>
</taxonomicName>
. However, her illustration appears to have closer affinity to
<taxonomicName id="4C4C4D23FFF6A23CFC6EFE3CA104C54A" ID-CoL="82BK8" authorityName="Henson" authorityYear="1948" box="[1000,1097,489,513]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFC6EFE3CA104C54A" box="[1000,1097,489,513]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C356652BFFF6A23CFCB6FDFDA19EC5CE" pageId="47" pageNumber="86" type="distribution">
<paragraph id="8BF336A0FFF6A23CFCB6FDFDA10FC508" blockId="47.[816,1090,555,579]" box="[816,1090,555,579]" pageId="47" pageNumber="86">Stratigraphic Distribution</paragraph>
<paragraph id="8BF336A0FFF6A23CFCB6FDBBA19EC5CE" blockId="47.[816,1437,621,2068]" box="[816,1235,621,645]" pageId="47" pageNumber="86">Early Albian intra-Late Cenomanian.</paragraph>
</subSubSection>
<subSubSection id="C356652BFFF6A222FCB6FD58A585C652" lastPageId="49" lastPageNumber="88" pageId="47" pageNumber="86" type="description">
<paragraph id="8BF336A0FFF6A23CFCB6FD58A11AC359" blockId="47.[816,1437,621,2068]" pageId="47" pageNumber="86">
Simmons &amp;
<bibRefCitation id="EFDD4B51FFF6A23CFC42FD58A139C5ED" author="Bidgood, M. D. &amp; Simmons, M. D." box="[964,1140,654,678]" pageId="47" pageNumber="86" refId="ref74158" refString="Bidgood, M. D. &amp; Simmons, M. D., 2022. Cenomanian planktonic foraminifera, bioevents and biozonation - a brief review. Newsletters on Stratigraphy, published online. DOI: 10.1127 / nos / 2022 / 0717" type="book" year="2022">Bidgood (2022)</bibRefCitation>
have reviewed the stratigraphic distribution of
<taxonomicName id="4C4C4D23FFF6A23CFBB5FD66A1D4C58C" authorityName="Henson" authorityYear="1948" box="[1075,1177,687,711]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFBB5FD66A1D4C58C" box="[1075,1177,687,711]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
and its common synonym
<taxonomicName id="4C4C4D23FFF6A23CFCE8FD04A69DC5A2" authorityName="Maync" authorityYear="1953" box="[878,976,721,745]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="sigali">
<emphasis id="B938EAB2FFF6A23CFCE8FD04A69DC5A2" box="[878,976,721,745]" italics="true" pageId="47" pageNumber="86">H. sigali</emphasis>
</taxonomicName>
.
<taxonomicName id="4C4C4D23FFF6A23CFC65FD04A101C5A2" authorityName="Henson" authorityYear="1948" box="[995,1100,721,745]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFC65FD04A101C5A2" box="[995,1100,721,745]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
was first described (
<bibRefCitation id="EFDD4B51FFF6A23CFAC5FD07A62AC441" author="Henson, F. R. S." pageId="47" pageNumber="86" refId="ref83829" refString="Henson, F. R. S., 1948. Larger Imperforate Foraminifera of South-Western Asia. British Museum (Natural History), London, pp. xi- 127." type="book" year="1948">Henson, 1948</bibRefCitation>
) from the Dukhan-3 well in
<collectingCountry id="F35B7630FFF6A23CFB3BFD24A1B5C441" box="[1213,1272,754,778]" name="Qatar" pageId="47" pageNumber="86">Qatar</collectingCountry>
at a depth of 3542-3543. Although said to be “probably early Cenomanian”, this depth equates to the Mauddud/Nahr Umr Formation boundary and is of latest Albian age (
<bibRefCitation id="EFDD4B51FFF6A23CFAD3FC83A6A1C4C5" author="Bromhead, A. D. &amp; van Buchem, F. S. P. &amp; Simmons, M. D. &amp; Davies, R." pageId="47" pageNumber="86" pagination="119 - 162" refId="ref75589" refString="Bromhead, A. D., van Buchem, F. S. P., Simmons, M. D. &amp; Davies, R. B, 2022. Sequence stratigraphy, paleogeography, and petroleum plays of the Cenomanian-Turonian succession of the Arabian Plate. Journal of Petroleum Geology, 45 (2): 119 - 162." type="journal article" year="2022">Bromhead et al., 2022</bibRefCitation>
).
<bibRefCitation id="EFDD4B51FFF6A23CFB87FCA1A1EFC4C5" author="Henson, F. R. S." box="[1025,1186,886,910]" pageId="47" pageNumber="86" refId="ref83829" refString="Henson, F. R. S., 1948. Larger Imperforate Foraminifera of South-Western Asia. British Museum (Natural History), London, pp. xi- 127." type="book" year="1948">Henson (1948)</bibRefCitation>
also recorded the species from the Nahr Umr Formation at Rumaila-
<quantity id="4CB49B45FFF6A23CFAB1FC41A02FC4E4" box="[1335,1378,919,943]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="47" pageNumber="86" unit="in" value="1.0">1 in</quantity>
<collectingCountry id="F35B7630FFF6A23CFAEDFC4EA0D4C4E4" box="[1387,1433,920,943]" name="Iraq" pageId="47" pageNumber="86">Iraq</collectingCountry>
, suggestive of an Albian age (
<bibRefCitation id="EFDD4B51FFF6A23CFBFAFC6EA015C49B" author="Aqrawi, A. A. M. &amp; Mahdi, T. A. &amp; Sherwani, G. H. &amp; Horbury, A. D." box="[1148,1368,952,976]" pageId="47" pageNumber="86" refId="ref71971" refString="Aqrawi, A. A. M., Mahdi, T. A., Sherwani, G. H. &amp; Horbury, A. D., 2010. Characterisation of the Mid-Cretaceous Mishrif Reservoir of the Southern Mesopotamian Basin, Iraq. American Association of Petroleum Geologists, Search and Discovery Article # 50264." type="book" year="2010">Aqrawi et al., 2010</bibRefCitation>
). The
<typeStatus id="54F78802FFF6A23CFCB6FC0CA613C4B9" box="[816,862,986,1010]" pageId="47" pageNumber="86">type</typeStatus>
material of
<taxonomicName id="4C4C4D23FFF6A23CFC65FC0CA10DC4BA" authorityName="Maync" authorityYear="1953" box="[995,1088,985,1009]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="sigali">
<emphasis id="B938EAB2FFF6A23CFC65FC0CA10DC4BA" box="[995,1088,985,1009]" italics="true" pageId="47" pageNumber="86">H. sigali</emphasis>
</taxonomicName>
is from the middle Cenomanian of
<collectingCountry id="F35B7630FFF6A23CFCCBFC2CA6D0C359" box="[845,925,1018,1042]" name="Algeria" pageId="47" pageNumber="86">Algeria</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFF6A23CFC2BFC2DA10AC359" author="Maync, W." box="[941,1095,1018,1043]" pageId="47" pageNumber="86" pagination="148 - 150" refId="ref88941" refString="Maync, W., 1953 b. Hemicyclammina sigali n. gen., n. sp. from the Cenomanian of Algeria. Contributions from the Cushman Foundation for Foraminiferal Research, 4: 148 - 150." type="journal article" year="1953">Maync, 1953b</bibRefCitation>
).
</paragraph>
<paragraph id="8BF336A0FFF6A23CFCB6FBCDA621C2D4" blockId="47.[816,1437,621,2068]" pageId="47" pageNumber="86">
Records commonly show a general composited Albian-Cenomanian age range. See, for example:
<bibRefCitation id="EFDD4B51FFF6A23CFA8AFBEAA635C33E" author="Ansary, S. E. &amp; Andrawis, S. F. &amp; Fahmy, S." pageId="47" pageNumber="86" pagination="1 - 17" refId="ref71816" refString="Ansary, S. E., Andrawis, S. F. &amp; Fahmy, S., 1962. Biostratigraphic studies of the Upper Cretaceous sections in the G. P. C. wells in the Eastern Desert and Sinai. Bulletin of the 3 rd Arab Petroleum Congress, Beirut, pp. 1 - 17." type="proceedings paper" year="1962">Ansary et al. (1962)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFF6A23CFC0CFB8BA153C33E" author="Wynd, J. G." box="[906,1054,1117,1141]" pageId="47" pageNumber="86" refId="ref102381" refString="Wynd, J. G., 1965. Biofacies of the Iranian Oil Consortium Agreement Area. IOOC Report 1082 (Unpublished)." type="book" year="1965">Wynd (1965)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFF6A23CFBB7FB8BA180C33E" author="Sampo, M." box="[1073,1229,1117,1141]" pageId="47" pageNumber="86" refId="ref96442" refString="Sampo, M., 1969. Microfacies and Microfossils of Zagros Area, Southwestern Iran. E. J. Brill. 102 pp." type="book" year="1969">Sampò (1969)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFF6A23CFB66FB8BA0DBC33E" author="Kalantari, A." box="[1248,1430,1117,1141]" pageId="47" pageNumber="86" refId="ref85776" refString="Kalantari, A., 1992. Lithostratigraphy and Microfacies of Zagros Orogenic Area S. W. Iran. National Iranian Oil Company, Geological Laboratories, Publication 12, Tehran." type="book" year="1992">Kalantari (1992)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFF6A23CFCB6FBA9A69EC3DC" author="Shirazi, M. P. N." box="[816,979,1151,1175]" pageId="47" pageNumber="86" pagination="709 - 714" refId="ref98703" refString="Shirazi, M. P. N., 2009. Albian-Cenomanian Zonation (Foraminifers and Calcareous Algae) in the Northern Fars, Iran. American Journal of Applied Sciences, 6 (4): 709 - 714." type="journal article" year="2009">Shirazi (2009)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFF6A23CFC6CFBA9A197C3DC" author="Shirazi, M. P. N. &amp; Bahrami, M. &amp; Rezaee, B." box="[1002,1242,1151,1175]" pageId="47" pageNumber="86" pagination="777 - 783" refId="ref98744" refString="Shirazi, M. P. N., Bahrami, M., Rezaee, B. &amp; Gha-ramani, S., 2011. Microbiostratigraphy of Kazhdumi formation in the Northwestern Shiraz (Southwest Iran) on the basis of foraminifera and calcareous algae. Acta Geologica Sinica-English Edition, 85 (4): 777 - 783." type="journal article" year="2011">Shirazi et al. (2011)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFF6A23CFB77FBA9A630C3FC" author="Omidvar, M. &amp; Mehrabi, H. &amp; Sajjadi, F." pageId="47" pageNumber="86" pagination="158 - 177" refId="ref92031" refString="Omidvar, M., Mehrabi, H. &amp; Sajjadi, F., 2014 a. Depositional environment and biostratigraphy of the upper Sarvak Formation in Ahwaz Oilfield (Well No. 63). Sedimentary Facies, 7 (2): 158 - 177." type="journal article" year="2014">Omidvar et al. (2014a</bibRefCitation>
, b) from the Iranian Zagros;
<bibRefCitation id="EFDD4B51FFF6A23CFB55FB76A0D5C3F3" author="Saint-Marc, P." box="[1235,1432,1184,1208]" pageId="47" pageNumber="86" pagination="85 - 94" refId="ref95980" refString="Saint-Marc, P., 1970. Sur quelques foraminiferes Cenomaniens et Turoniens du Liban. Revue de Micropaleontologie, 13 (2): 85 - 94." type="journal article" year="1970">Saint-Marc (1970</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FFF6A23CFCB6FB17A63FC392" author="Saint-Marc, P." box="[816,882,1217,1241]" pageId="47" pageNumber="86" pagination="1 - 342" refId="ref96009" refString="Saint-Marc, P., 1974 a. Etude stratigraphique et micropaleontologique de l'Albien, du Cenomanien et du Turonien du Liban. Notes et Memoires sur le Moyen- Orient, XIII, Museum National d'Histoire Naturelle, Paris, pp. 1 - 342." type="book chapter" year="1974">
1974
<bibRefCitation id="EFDD4B51FFF6A23CFCE0FB14A63FC392" author="Arnaud, A. &amp; Berthout &amp; Brunt, L. &amp; Cherchi, A. &amp; Chiocchini, M. &amp; De Castro, P." box="[870,882,1218,1241]" pageId="47" pageNumber="86" pagination="383 - 393" refId="ref72126" refString="Arnaud, A., Berthout, Brunt, L., Cherchi, A., Chiocchini, M., De Castro, P. &amp; 9 others, 1981. Tableau de repartition stratigraphique des grands foraminiferes caracteristiques du Cretace Moyen de la region Mediterraneenne. Cretaceous Research, 2: 383 - 393." type="journal article" year="1981">a</bibRefCitation>
</bibRefCitation>
, 1981) from
<collectingCountry id="F35B7630FFF6A23CFB96FB17A13DC392" box="[1040,1136,1217,1241]" name="Lebanon" pageId="47" pageNumber="86">Lebanon</collectingCountry>
;
<bibRefCitation id="EFDD4B51FFF6A23CFB07FB17A0DAC392" author="Simmons, M. D. &amp; Hart, M. B." box="[1153,1431,1217,1241]" pageId="47" pageNumber="86" pagination="176 - 207" refId="ref98973" refString="Simmons, M. D. &amp; Hart, M. B., 1987. The biostratigraphy and microfacies of the Early to mid-Cretaceous carbonates of Wadi Mi'aidin, Central Oman Mountains. In: Hart, M. B. (ed), Micropalaeontology of Carbonate Environments, John Wiley &amp; Sons: 176 - 207." type="book chapter" year="1987">Simmons &amp; Hart (1987)</bibRefCitation>
,
<bibRefCitation id="EFDD4B51FFF6A23CFCB6FB34A147C3B1" author="Forbes, G. A. &amp; Jansen, H. S. M. &amp; Schreurs, J." box="[816,1034,1250,1274]" pageId="47" pageNumber="86" refId="ref81251" refString="Forbes, G. A., Jansen, H. S. M. &amp; Schreurs, J., 2010. Lexicon of Oman subsurface stratigraphy: reference guide to the stratigraphy of Oman's hydrocarbon basins. GeoArabia Special Publication, 5, Gulf Petro-Link, Bahrain." type="book" year="2010">Forbes et al. (2010)</bibRefCitation>
from
<collectingCountry id="F35B7630FFF6A23CFBD6FB34A1D9C3B1" box="[1104,1172,1250,1274]" name="Oman" pageId="47" pageNumber="86">Oman</collectingCountry>
; and general summaries by
<bibRefCitation id="EFDD4B51FFF6A23CFCD3FAD5A16CC250" author="Saint-Marc, P." box="[853,1057,1283,1307]" pageId="47" pageNumber="86" pagination="317 - 325" refId="ref96089" refString="Saint-Marc, P., 1977. Repartition stratigraphique des grands Foraminiferes benthiques de l'Aptien, de l'Albien, du Cenomanien et du Turonien dans les regions mediterraneennes. Revista Espanola de Micropaleontologia, IX (3): 317 - 325." type="journal article" year="1977">Saint-Marc (1977)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFF6A23CFBB6FAD5A02AC250" author="Sartorio, D. &amp; Venturini, S." box="[1072,1383,1283,1307]" pageId="47" pageNumber="86" refId="ref96576" refString="Sartorio, D., &amp; Venturini, S., 1988. Southern Tethys Biofacies. Agip S. p. A., 235 pp." type="book" year="1988">Sartorio &amp; Venturini (1988)</bibRefCitation>
, and
<bibRefCitation id="EFDD4B51FFF6A23CFCB6FAF2A162C277" author="Schroeder, R. &amp; Van Buchem, F. S. P. &amp; Cherchi, A. &amp; Baghbani, D. &amp; Vincent, B. &amp; Immenhauser, A. &amp; Granier, B." box="[816,1071,1316,1340]" pageId="47" pageNumber="86" pagination="359" refId="ref98334" refString="Schroeder, R., Van Buchem, F. S. P., Cherchi, A., Baghbani, D., Vincent, B., Immenhauser, A. &amp; Granier, B., 2010. Revised orbitolinid biostratigraphic zonation for the Barremian - Aptian of the eastern Arabian Turonian succession in Gabal Nezzazat, southwestern Sinai, Egypt. Revue de Paleobiologie, 26 (2): 359." type="journal article" year="2010">Schroeder et al. (2010)</bibRefCitation>
. Statements that the species does not range above the Albian (e.g.,
<bibRefCitation id="EFDD4B51FFF6A23CFB10FA93A0D8C216" author="Afghah, M. &amp; Dookh, R." box="[1174,1429,1349,1373]" pageId="47" pageNumber="86" pagination="5 - 19" refId="ref70352" refString="Afghah, M. &amp; Dookh, R., 2014. Microbiostratigraphy of the Sarvak Formation in the east and north-east of Shiraz (Kuh-e-Siah and Kuh-e-Pichakan), SW of Iranian Journal of Science I. A. U. (JSIAU), 24: 5 - 19." type="journal article" year="2014">Afghah &amp; Dookh, 2014</bibRefCitation>
) should be discounted at both a local and inter-regional level.
</paragraph>
<paragraph id="8BF336A0FFF6A23CFCB6FA7FA6F2C066" blockId="47.[816,1437,621,2068]" pageId="47" pageNumber="86">
<bibRefCitation id="EFDD4B51FFF6A23CFCB6FA7FA6BEC28A" author="Hart, M. B. &amp; Callapez, P. M. &amp; Fisher, J. K. &amp; Hannant, K. &amp; Monteiro, J. F. &amp; Price, G. D. &amp; Watkinson, M. P." box="[816,1011,1449,1473]" pageId="47" pageNumber="86" pagination="311 - 326" refId="ref83592" refString="Hart, M. B., Callapez, P. M., Fisher, J. K., Hannant, K., Monteiro, J. F., Price, G. D. &amp; Watkinson, M. P., 2005. Micropalaeontology and Stratigraphy of the Cenomanian / Turonian boundary in the Lusitanian Basin, Portugal. Journal of Iberian Geology, 31 (2): 311 - 326." type="journal article" year="2005">Hart et al. (2005)</bibRefCitation>
recorded
<taxonomicName id="4C4C4D23FFF6A23CFBE3FA7CA189C28A" authorityName="Maync" authorityYear="1953" box="[1125,1220,1449,1473]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="sigali">
<emphasis id="B938EAB2FFF6A23CFBE3FA7CA189C28A" box="[1125,1220,1449,1473]" italics="true" pageId="47" pageNumber="86">H. sigali</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C4C4D23FFF6A23CFB77FA7CA01BC28A" authorityName="Henson" authorityYear="1948" box="[1265,1366,1449,1473]" class="Globothalamea" family="Cyclamminidae" genus="Cyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFF6A23CFB77FA7CA01BC28A" box="[1265,1366,1449,1473]" italics="true" pageId="47" pageNumber="86">H. whitei</emphasis>
</taxonomicName>
) from strata in
<collectingCountry id="F35B7630FFF6A23CFC14FA1CA6A1C2A9" box="[914,1004,1482,1506]" name="Portugal" pageId="47" pageNumber="86">Portugal</collectingCountry>
confidently assigned to the
<taxonomicName id="4C4C4D23FFF6A23CFAA0FA1DA0D1C2A9" authorityName="Tethyan" baseAuthorityName="Spath" baseAuthorityYear="1926" box="[1318,1436,1483,1506]" class="Cephalopoda" family="Acanthoceratidae" genus="Calycoceras" kingdom="Animalia" pageId="47" pageNumber="86" phylum="Mollusca" rank="species" species="guerangeri">
<emphasis id="B938EAB2FFF6A23CFAA0FA1DA0D1C2A9" box="[1318,1436,1483,1506]" italics="true" pageId="47" pageNumber="86">guerangeri</emphasis>
</taxonomicName>
and
<emphasis id="B938EAB2FFF6A23CFCD8FA3DA6ADC148" box="[862,992,1515,1539]" italics="true" pageId="47" pageNumber="86">geslinianum</emphasis>
ammonite zones of the late Cenomanian, following earlier records by
<bibRefCitation id="EFDD4B51FFF6A23CFBFEF9DAA06EC16F" author="Berthou, P-Y." box="[1144,1315,1548,1572]" pageId="47" pageNumber="86" pagination="1 - 168" refId="ref74017" refString="Berthou, P-Y., 1973. Le Cenomanien de l'Estremadure Portugaise. Servicos Geologicos de Portugal, 23: 1 - 168." type="journal article" year="1973">Berthou (1973)</bibRefCitation>
;
<bibRefCitation id="EFDD4B51FFF6A23CFAB1F9DAA635C10E" author="Lauverjat, J." pageId="47" pageNumber="86" pagination="109 - 114" refId="ref87514" refString="Lauverjat, J., 1976. Le Cenomanien de la Vallee du Mondego (Portugal). Limite avec le Turonien. Evolution Ouest-Est, implications paleogeographiques. Geologie Mediterraneenne, 1: 109 - 114." type="journal article" year="1976">Lauverjat (1976)</bibRefCitation>
and Crosaz-Galletti (1979).
<bibRefCitation id="EFDD4B51FFF6A23CFB48F9FBA0D0C10E" author="Saint-Marc, P." box="[1230,1437,1581,1605]" pageId="47" pageNumber="86" pagination="103 - 131" refId="ref96200" refString="Saint-Marc, P., 1981. Lebanon. In: Reyment, R. A. &amp; Bengtson, P. (eds.) Aspects of Mid-Cretaceous Regional Geology. IGCP Projects 58, Academic Press, pp. 103 - 131." type="book chapter" year="1981">Saint-Marc (1981)</bibRefCitation>
reported the species from latest Cenomanian strata in
<collectingCountry id="F35B7630FFF6A23CFCB6F9B9A6C0C1CC" box="[816,909,1647,1671]" name="Lebanon" pageId="47" pageNumber="86">Lebanon</collectingCountry>
with the ammonites
<taxonomicName id="4C4C4D23FFF6A23CFB18F9B9A16DC1E3" authority="(Blackenhorn)" baseAuthorityName="Blackenhorn" class="Cephalopoda" family="Acanthoceratidae" genus="Eucalycoceras" kingdom="Animalia" pageId="47" pageNumber="86" phylum="Mollusca" rank="species" species="palaestinense">
<emphasis id="B938EAB2FFF6A23CFB18F9B9A636C1E3" italics="true" pageId="47" pageNumber="86">Eucalycoceras palaestinense</emphasis>
(Blackenhorn)
</taxonomicName>
and
<taxonomicName id="4C4C4D23FFF6A23CFBDCF946A632C182" authority="(Spath)" baseAuthorityName="Spath" baseAuthorityYear="1931" class="Cephalopoda" family="Acanthoceratidae" genus="Protacanthoceras" kingdom="Animalia" pageId="47" pageNumber="86" phylum="Mollusca" rank="species" species="angolaense">
<emphasis id="B938EAB2FFF6A23CFBDCF946A0D0C1E3" box="[1114,1437,1680,1704]" italics="true" pageId="47" pageNumber="86">Protacanthoceras angolaense</emphasis>
(Spath)
</taxonomicName>
and planktonic foraminifera
<taxonomicName id="4C4C4D23FFF6A23CFB38F967A6A7C1A1" baseAuthorityName="Trujillo" baseAuthorityYear="1960" class="Globothalamea" family="Hedbergellidae" genus="Helvetoglobotruncana" kingdom="Chromista" order="Rotaliida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="praehelvetica">
<emphasis id="B938EAB2FFF6A23CFB38F967A6A7C1A1" italics="true" pageId="47" pageNumber="86">Helvetoglobotruncana praehelvetica</emphasis>
</taxonomicName>
(Trujillo) and
<taxonomicName id="4C4C4D23FFF6A23CFB1AF904A01DC1A1" box="[1180,1360,1746,1770]" class="Globothalamea" family="Hedbergellidae" genus="Whiteinella" kingdom="Chromista" order="Rotaliida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="undetermined">
<emphasis id="B938EAB2FFF6A23CFB1AF904A05AC1A1" box="[1180,1303,1746,1770]" italics="true" pageId="47" pageNumber="86">Whiteinella</emphasis>
spp.
</taxonomicName>
These records confer an upper range age limit of intra-late Cenomanian.
</paragraph>
<paragraph id="8BF336A0FFF6A23CFCB6F8E0A076CF5F" blockId="47.[816,1437,621,2068]" pageId="47" pageNumber="86">
Pre-Albian records of the species are believed to be of a potentially ancestral form (“
<taxonomicName id="4C4C4D23FFF6A23CFBEEF881A06FC024" authorityName="Maync" authorityYear="1953" box="[1128,1314,1879,1903]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFF6A23CFBEEF881A06FC024" box="[1128,1314,1879,1903]" italics="true" pageId="47" pageNumber="86">Hemicyclammina</emphasis>
</taxonomicName>
? sp.”) and include those of
<bibRefCitation id="EFDD4B51FFF6A23CFC60F8AEA19DC0DB" author="Hosseini, S. A. &amp; Conrad, M. A. &amp; Clavel, B. &amp; Carras, N." box="[998,1232,1912,1936]" pageId="47" pageNumber="86" pagination="257 - 288" refId="ref84131" refString="Hosseini, S. A., Conrad, M. A., Clavel, B. &amp; Carras, N., 2016. Berriasian-Aptian shallow water carbonates in the Zagros fold-thrust belt, SW Iran: Integrated Srisotope dating and biostratigraphy. Cretaceous Research, 57: 257 - 288." type="journal article" year="2016">Hosseini et al. (2016)</bibRefCitation>
who illustrate material under the name
<taxonomicName id="4C4C4D23FFF6A23CFBA7F84CA132C0FA" authorityName="Maync" authorityYear="1953" box="[1057,1151,1945,1969]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="47" pageNumber="86" phylum="Foraminifera" rank="species" species="sigali">
<emphasis id="B938EAB2FFF6A23CFBA7F84CA132C0FA" box="[1057,1151,1945,1969]" italics="true" pageId="47" pageNumber="86">H. sigali</emphasis>
</taxonomicName>
from the Barremian Gadvan Formation of the Iranian Zagros. However, the specimens are very small (
<quantity id="4CB49B45FFF6A23CFBA7F80DA1F8C0B8" box="[1057,1205,2011,2035]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="47" pageNumber="86" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2 0.3 mm</quantity>
in diameter), and the presence of an alveolar wall is not demonstrated.
</paragraph>
<caption id="DF336628FFE9A223FEF9FCE5A1F8C402" ID-DOI="http://doi.org/10.5281/zenodo.10834255" ID-Zenodo-Dep="10834255" box="[383,1205,819,841]" httpUri="https://zenodo.org/record/10834255/files/figure.png" pageId="48" pageNumber="87" startId="48.[383,416,819,840]" targetBox="[218,1338,168,798]" targetPageId="48" targetType="figure">
<paragraph id="8BF336A0FFE9A223FEF9FCE5A1F8C402" blockId="48.[383,1205,819,841]" box="[383,1205,819,841]" pageId="48" pageNumber="87">
<emphasis id="B938EAB2FFE9A223FEF9FCE5A488C403" bold="true" box="[383,453,819,840]" pageId="48" pageNumber="87">Fig. 34</emphasis>
Cenomanian paleogeographic distribution of
<taxonomicName id="4C4C4D23FFE9A223FCF8FCE5A1E3C402" authorityName="Schlagintweit &amp; Yazdi-Moghadam" authorityYear="2023" box="[894,1198,819,841]" class="Globothalamea" family="Cyclamminidae" genus="Pseudocyclammina" kingdom="Chromista" order="Loftusiida" pageId="48" pageNumber="87" phylum="Foraminifera" rank="species" species="sarvakensis">Pseudocyclammina sarvakensis</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF336628FFE9A223FF48F92EA19BC002" ID-DOI="http://doi.org/10.5281/zenodo.10834257" ID-Zenodo-Dep="10834257" httpUri="https://zenodo.org/record/10834257/files/figure.png" pageId="48" pageNumber="87" startId="48.[206,239,1784,1805]" targetBox="[218,1372,915,1763]" targetPageId="48" targetType="figure">
<paragraph id="8BF336A0FFE9A223FF48F92EA19BC002" blockId="48.[206,1371,1784,1865]" pageId="48" pageNumber="87">
<emphasis id="B938EAB2FFE9A223FF48F92EA45EC046" bold="true" box="[206,275,1784,1805]" pageId="48" pageNumber="87">Fig. 35</emphasis>
Representative illustrations of
<taxonomicName id="4C4C4D23FFE9A223FDB9F92EA665C045" baseAuthorityName="Henson" baseAuthorityYear="1948" box="[575,808,1784,1806]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="48" pageNumber="87" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFE9A223FDB9F92EA665C045" box="[575,808,1784,1806]" italics="true" pageId="48" pageNumber="87">Hemicyclammina whitei</emphasis>
</taxonomicName>
:
<emphasis id="B938EAB2FFE9A223FCB3F92EA60CC046" bold="true" box="[821,833,1784,1805]" pageId="48" pageNumber="87">a</emphasis>
Axial view,
<bibRefCitation id="EFDD4B51FFE9A223FC3AF92EA1DCC045" author="Whittaker, J. E. &amp; Jones, R. W. &amp; Banner, F. T." box="[956,1169,1784,1806]" pageId="48" pageNumber="87" refId="ref102083" refString="Whittaker, J. E., Jones, R. W. &amp; Banner, F. T., 1998. Key Mesozoic benthic foraminifera of the Middle East. Natural History Museum, London, 237 pp." type="book" year="1998">Whittaker et al. (1998</bibRefCitation>
, pl. 12, fig. 1a, Qatar - see also Simmons &amp;
<bibRefCitation id="EFDD4B51FFE9A223FE21F8C0A778C067" author="Bidgood, M. D. &amp; Simmons, M. D." box="[423,565,1814,1836]" pageId="48" pageNumber="87" refId="ref74158" refString="Bidgood, M. D. &amp; Simmons, M. D., 2022. Cenomanian planktonic foraminifera, bioevents and biozonation - a brief review. Newsletters on Stratigraphy, published online. DOI: 10.1127 / nos / 2022 / 0717" type="book" year="2022">Bidgood, 2022</bibRefCitation>
, fig. 1a);
<emphasis id="B938EAB2FFE9A223FD15F8C0A7EDC060" bold="true" box="[659,672,1814,1835]" pageId="48" pageNumber="87">b</emphasis>
Axial section,
<bibRefCitation id="EFDD4B51FFE9A223FCB4F8C0A14BC067" author="Whittaker, J. E. &amp; Jones, R. W. &amp; Banner, F. T." box="[818,1030,1814,1836]" pageId="48" pageNumber="87" refId="ref102083" refString="Whittaker, J. E., Jones, R. W. &amp; Banner, F. T., 1998. Key Mesozoic benthic foraminifera of the Middle East. Natural History Museum, London, 237 pp." type="book" year="1998">Whittaker et al. (1998</bibRefCitation>
, pl. 59, fig. 8, U.A.E. - see also Simmons &amp;
<bibRefCitation id="EFDD4B51FFE9A223FEC1F8E5A495C002" author="Bidgood, M. D. &amp; Simmons, M. D." box="[327,472,1843,1865]" pageId="48" pageNumber="87" refId="ref74158" refString="Bidgood, M. D. &amp; Simmons, M. D., 2022. Cenomanian planktonic foraminifera, bioevents and biozonation - a brief review. Newsletters on Stratigraphy, published online. DOI: 10.1127 / nos / 2022 / 0717" type="book" year="2022">Bidgood, 2022</bibRefCitation>
, fig. 3c);
<emphasis id="B938EAB2FFE9A223FDB2F8E2A772C002" bold="true" box="[564,575,1844,1865]" pageId="48" pageNumber="87">c</emphasis>
Equatorial section, Simmons &amp;
<bibRefCitation id="EFDD4B51FFE9A223FCFEF8E5A147C002" author="Bidgood, M. D. &amp; Simmons, M. D." box="[888,1034,1843,1865]" pageId="48" pageNumber="87" refId="ref74158" refString="Bidgood, M. D. &amp; Simmons, M. D., 2022. Cenomanian planktonic foraminifera, bioevents and biozonation - a brief review. Newsletters on Stratigraphy, published online. DOI: 10.1127 / nos / 2022 / 0717" type="book" year="2022">Bidgood (2022</bibRefCitation>
, fig. 3a, Abu Dhabi).
</paragraph>
</caption>
<paragraph id="8BF336A0FFE8A222FF11FF48A585C652" blockId="49.[151,771,158,281]" pageId="49" pageNumber="88">
A similar, slight larger (
<quantity id="4CB49B45FFE8A222FE25FF48A4B5C7FE" box="[419,504,158,182]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="49" pageNumber="88" unit="mm" value="0.3">0.3 mm</quantity>
diameter) specimen has been illustrated by
<bibRefCitation id="EFDD4B51FFE8A222FEF2FF69A7DAC79C" author="Ozkan, R. &amp; Altiner, D." box="[372,663,191,215]" pageId="49" pageNumber="88" pagination="153 - 178" refId="ref92491" refString="Ozkan, R. &amp; Altiner, D., 2019. The Cretaceous Mardin Group carbonates in southeast Turkey: lithostratigraphy, foraminiferal biostratigraphy, microfacies and sequence stratigraphic evolution. Cretaceous Research, 98: 153 - 178." type="journal article" year="2019">Özkan and Altiner (2019)</bibRefCitation>
from the early Aptian of south-east
<collectingCountry id="F35B7630FFE8A222FE3AFF36A744C7B3" box="[444,521,224,248]" name="Turkey" pageId="49" pageNumber="88">Turkey</collectingCountry>
as “
<taxonomicName id="4C4C4D23FFE8A222FDBBFF36A7BAC7B3" authorityName="Maync" authorityYear="1953" box="[573,759,224,248]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="49" pageNumber="88" phylum="Foraminifera" rank="genus">
<emphasis id="B938EAB2FFE8A222FDBBFF36A7BAC7B3" box="[573,759,224,248]" italics="true" pageId="49" pageNumber="88">Hemicyclammina</emphasis>
</taxonomicName>
? sp.”.
</paragraph>
</subSubSection>
<subSubSection id="C356652BFFE8A222FF11FE95A766C4E4" pageId="49" pageNumber="88" type="distribution">
<paragraph id="8BF336A0FFE8A222FF11FE95A713C610" blockId="49.[151,606,323,347]" box="[151,606,323,347]" pageId="49" pageNumber="88">Cenomanian Paleogeographic Distribution</paragraph>
<paragraph id="8BF336A0FFE8A222FF11FE53A766C4E4" blockId="49.[151,772,389,943]" pageId="49" pageNumber="88">
Broadly across Neotethys, the Atlantic and the Caribbean.
<taxonomicName id="4C4C4D23FFE8A222FF11FE7EA449C6F4" baseAuthorityName="Henson" baseAuthorityYear="1948" box="[151,260,423,447]" class="Globothalamea" family="Cyclamminidae" genus="Hemicyclammina" kingdom="Chromista" order="Loftusiida" pageId="49" pageNumber="88" phylum="Foraminifera" rank="species" species="whitei">
<emphasis id="B938EAB2FFE8A222FF11FE7EA449C6F4" box="[151,260,423,447]" italics="true" pageId="49" pageNumber="88">H. whitei</emphasis>
</taxonomicName>
has a broad palaeogeographic distribution across Neotethys and seemed to thrive in marly sediments deposited on middle outer shelves. Confirmed records in addition to those previously mentioned above include from
<collectingCountry id="F35B7630FFE8A222FF55FDFDA459C508" box="[211,276,555,579]" name="Brazil" pageId="49" pageNumber="88">Brazil</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE8A222FEA2FDFDA707C508" author="Berthou, P-Y. &amp; Bengtson, P." box="[292,586,555,580]" pageId="49" pageNumber="88" pagination="1 - 88" refId="ref73875" refString="Berthou, P-Y. &amp; Bengtson, P., 1988. Stratigraphic Correlation by Microfacies of the Cenomanian - Coniacian of the Sergipe Basin, Brazil. Fossils and Strata no. 21, pp. 1 - 88. Oslo Universitetsforlaget." type="journal article" year="1988">Berthou &amp; Bengtson, 1988</bibRefCitation>
);
<collectingCountry id="F35B7630FFE8A222FDE2FDFDA7F8C508" box="[612,693,555,579]" name="Mexico" pageId="49" pageNumber="88">Mexico</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE8A222FD43FDFDA475C52F" author="Omana, L. &amp; Lopez-Doncel, R. &amp; Torres, J. R. &amp; Alencaster, G. &amp; Lopez-Caballero, I." pageId="49" pageNumber="88" pagination="691 - 725" refId="ref91622" refString="Omana, L., Lopez-Doncel, R., Torres, J. R., Alencaster, G. &amp; Lopez-Caballero, I. 2019. Mid - late Cenomanian larger benthic foraminifers from the El Abra Formation W Valles-San Luis Potosi Platform, central - eastern Mexico: Taxonomy, biostratigraphy and paleoenvironmental implications. Boletin de la Sociedad Geologica Mexicana, 71 (3): 691 - 725." type="journal article" year="2019">Omaña et al., 2019</bibRefCitation>
);
<collectingCountry id="F35B7630FFE8A222FED4FD9BA4FEC52F" box="[338,435,589,612]" name="Morocco" pageId="49" pageNumber="88">Morocco</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE8A222FE45FD9AA7D6C52F" author="Andreu, B. &amp; Bilotte, M. &amp; Ettachfini, E. M. &amp; Fessard, N." box="[451,667,588,612]" pageId="49" pageNumber="88" pagination="521 - 540" refId="ref71680" refString="Andreu, B., Bilotte, M., Ettachfini, E. M. &amp; Grambast- Fessard, N., 1996. Microfauna (foraminifera, ostracods) and microflora (algae, charaphyta) of the upper Albian? Cenomanian-Turonian in the Essaouira Basin (western High Atlas, Morocco): biostratigraphy and palaeoecology. Geologie de l'Afrique et de l'Atlantique Sud, Bulletin des Centres de Recherches Exploration-Production d'Elf-Aquitaine, 16: 521 - 540." type="journal article" year="1996">Andreu et al., 1996</bibRefCitation>
);
<collectingCountry id="F35B7630FFE8A222FD30FD9AA64FC52F" box="[694,770,588,612]" name="Turkey" pageId="49" pageNumber="88">Turkey</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE8A222FF18FDBBA4EFC5CE" author="Bignot, G. &amp; Poisson, A." box="[158,418,621,645]" pageId="49" pageNumber="88" pagination="71 - 77" refId="ref74202" refString="Bignot, G. &amp; Poisson, A., 1974. Le Cenomanien du flanc Oriental du Katran Dag (= Sam Dag) pres d'Anatalya (Turquie). Bulletin of the Mining Research and Exploration Institute of Turkey, 82: 71 - 77." type="journal article" year="1974">Bignot &amp; Poisson, 1974</bibRefCitation>
);
<collectingCountry id="F35B7630FFE8A222FE3FFDBBA74DC5CE" box="[441,512,621,645]" name="Jordan" pageId="49" pageNumber="88">Jordan</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE8A222FD88FDBBA580C5ED" author="Weidich, K. F. &amp; Al-Harithi, T." pageId="49" pageNumber="88" pagination="587 - 619" refId="ref101853" refString="Weidich, K. F. &amp; Al-Harithi, T., 1990. Agglutinated foraminifera from the Albian and Cenomanian of Jordan. In: Palaeoecology, Biostratigraphy, Paleoceanography and Taxonomy of Agglutinated Foraminifera (C. Hemleben et al., eds.). NATO ASI Series, Kluwer Academic Publishers: 587 - 619" type="book chapter" year="1990">Weidich &amp; Al-Harithi, 1990</bibRefCitation>
);
<collectingCountry id="F35B7630FFE8A222FF6AFD58A432C5ED" box="[236,383,654,678]" name="Saudi Arabia" pageId="49" pageNumber="88">Saudi Arabia</collectingCountry>
(Dr. Wyn Hughes, pers. comm., 2022); Abu Dhabi (
<bibRefCitation id="EFDD4B51FFE8A222FEF5FD79A745C58C" author="Banner, F. T." box="[371,520,687,711]" pageId="49" pageNumber="88" pagination="242 - 290" refId="ref73119" refString="Banner, F. T., 1970. A synopsis of the Spirocyclinidae. Revista Espanola Micropaleontologia, 11 (3): 242 - 290." type="journal article" year="1970">Banner, 1970</bibRefCitation>
), and
<collectingCountry id="F35B7630FFE8A222FDD4FD79A7E4C58C" box="[594,681,687,711]" name="Somalia" pageId="49" pageNumber="88">Somalia</collectingCountry>
(
<bibRefCitation id="EFDD4B51FFE8A222FD3AFD79A580C5A2" author="Luger, P." pageId="49" pageNumber="88" refId="ref88061" refString="Luger, P., 2018. Micropalaeontology (Foraminiferida, Ostracoda), biostratigraphy and facies development of the Cretaceous of Northern Somalia - including a contribution concerning the geodynamic development of eastern Gondwana during the Cretaceous to basal Paleocene. Unpublished PhD thesis, Documenta Naturae Abhandlungen, 2 Vols." type="book" year="2018">Luger, 2018</bibRefCitation>
). Unillustrated records are known from numerous intermediate locations (Pyrenees
<bibRefCitation id="EFDD4B51FFE8A222FD9FFD24A786C441" author="Peybernes, B." box="[537,715,754,778]" pageId="49" pageNumber="88" pagination="491 - 499" refId="ref93070" refString="Peybernes, B., 1984. Foraminiferes benthiques nouveaux de l'Albien superieur du Massif du Turbon (Pyrenees espagnole). Benthos ' 83; 2 nd lnternational Symposium Benthic Foraminifera (Pau): 491 - 499." type="journal article" year="1984">Peybernès, 1984</bibRefCitation>
;
<collectingCountry id="F35B7630FFE8A222FD5BFD24A584C460" name="Tunisia" pageId="49" pageNumber="88">Tunisia</collectingCountry>
<bibRefCitation id="EFDD4B51FFE8A222FF77FCC5A747C460" author="Robaszynski, F. &amp; Zagrarni, M. F. &amp; Caron, M. &amp; Amedro, F." box="[241,522,787,811]" pageId="49" pageNumber="88" pagination="1 - 15" refId="ref95223" refString="Robaszynski, F., Zagrarni, M. F., Caron, M. &amp; Amedro, F., 2010. The global bio-events at the Cenomanian-Turonian transition in the reduced Bahloul Formation of Bou Ghanem (central Tunisia). Cretaceous Research, 31: 1 - 15." type="journal article" year="2010">Robaszynski et al., 2010</bibRefCitation>
;
<collectingCountry id="F35B7630FFE8A222FD98FCC5A711C460" box="[542,604,787,811]" name="Libya" pageId="49" pageNumber="88">Libya</collectingCountry>
<bibRefCitation id="EFDD4B51FFE8A222FD05FCC5A409C407" author="Megerisi, M. &amp; Mamgain, V. D." pageId="49" pageNumber="88" refId="ref89167" refString="Megerisi, M. &amp; Mamgain, V. D., 1980. The Upper Cretaceous Tertiary Formations of Northern Libya, a synthesis. Department of Geological Research &amp; Mining, Industrial Research Centre Bulletin, Tripoli, 12, 85 p." type="book" year="1980">Megerisi &amp; Mamgain, 1980</bibRefCitation>
;
<collectingCountry id="F35B7630FFE8A222FEDEFCE2A4C4C407" box="[344,393,820,844]" name="Italy" pageId="49" pageNumber="88">Italy</collectingCountry>
<bibRefCitation id="EFDD4B51FFE8A222FE2BFCE2A7C4C407" author="Simone, L. &amp; Bravi, S. &amp; Carannante, G. &amp; Masucci, I. &amp; Pomoni-Papaioannou, F." box="[429,649,820,844]" pageId="49" pageNumber="88" pagination="6 - 23" refId="ref99377" refString="Simone, L., Bravi, S., Carannante, G., Masucci, I. &amp; Pomoni-Papaioannou, F., 2012. Arid versus wet climatic evidence in the &quot; middle Cretaceous &quot; calcareous successions of the Southern Apennines (Italy). Cretaceous Research, 36: 6 - 23." type="journal article" year="2012">Simone et al., 2012</bibRefCitation>
;
<collectingCountry id="F35B7630FFE8A222FD1AFCE2A7A7C407" box="[668,746,820,844]" name="Croatia" pageId="49" pageNumber="88">Croatia</collectingCountry>
<bibRefCitation id="EFDD4B51FFE8A222FF11FC83A436C426" author="Husinec, A. &amp; Velic, I. &amp; Fucek, L. &amp; Vlahovic, I. &amp; Maticec, D. &amp; Ostric, N. &amp; Korbar, T." box="[151,379,853,877]" pageId="49" pageNumber="88" pagination="155 - 171" refId="ref84655" refString="Husinec, A., Velic, I., Fucek, L., Vlahovic, I., Maticec, D., Ostric, N. &amp; Korbar, T., 2000. Mid Cretaceous orbitolinid (Foraminiferida) record from the islands of Cres and Losinj (Croatia) and its regional stratigraphic correlation. Cretaceous Research, 21: 155 - 171." type="journal article" year="2000">Husinec et al., 2000</bibRefCitation>
;
<collectingCountry id="F35B7630FFE8A222FE09FC83A49EC426" box="[399,467,853,877]" name="Serbia" pageId="49" pageNumber="88">Serbia</collectingCountry>
<bibRefCitation id="EFDD4B51FFE8A222FE7EFC83A447C4C5" author="Radoicic, R. &amp; Schlagintweit, F." pageId="49" pageNumber="88" pagination="39 - 51" refId="ref93827" refString="Radoicic, R. &amp; Schlagintweit, F., 2007. Neomeris mokragorensis sp. nov. (Calcareous alga, Dasycladales) from the Cretaceous of Serbia, Montenegro and the Northern Calcareous Alps, (Gosau Group, Austria). Annales Geologiques de la Peninsule Balkanique, 68: 39 - 51." type="journal article" year="2007">Radoičić and Schlagintweit, 2007</bibRefCitation>
;
<collectingCountry id="F35B7630FFE8A222FE9CFCA0A425C4C5" box="[282,360,886,910]" name="Kuwait" pageId="49" pageNumber="88">Kuwait</collectingCountry>
<bibRefCitation id="EFDD4B51FFE8A222FE0EFCA0A72FC4C5" author="Youssef, A. H. &amp; Al-Sahlan, G. &amp; Kadar, A. P. &amp; Karam, K. A. &amp; Packer, S. R. &amp; Starkie, S. &amp; Stead, D." box="[392,610,886,910]" pageId="49" pageNumber="88" pagination="1 - 26" refId="ref102924" refString="Youssef, A. H., Al-Sahlan, G., Kadar, A. P., Karam, K. A., Packer, S. R., Starkie, S. &amp; Stead, D., 2019. Sequence stratigraphic framework of the Wara and Ahmadi Formations, onshore Kuwait. Stratigraphy, 16 (1): 1 - 26." type="journal article" year="2019">Youssef et al., 2019</bibRefCitation>
), and possibly Tibet (
<bibRefCitation id="EFDD4B51FFE8A222FF5BFC41A756C4E4" author="BouDagher-Fadel, M. K. &amp; Hu, X. &amp; Price, G. D. &amp; Sun, G. &amp; Wang, J-G &amp; An, W." box="[221,539,919,943]" pageId="49" pageNumber="88" pagination="188 - 207" refId="ref75121" refString="BouDagher-Fadel, M. K., Hu, X., Price, G. D., Sun, G., Wang, J-G &amp; An, W., 2017. Foraminiferal biostratigraphy and palaeoenvironmental analysis of the Mid-Cretaceous limestones in the southern Tibetan Plateau Journal of Foraminiferal Research, 47 (2): 188 - 207." type="journal article" year="2017">BouDagher-Fadel et al., 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>