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<document id="ABDCED072697807CCC12C4851178FD17" ID-CLB-Dataset="8552" ID-DOI="10.5252/geodiversitas2020v42a23" ID-GBIF-Dataset="80fb3ed1-03d8-4294-8a2a-755927f26150" ID-ISSN="1638-9395" ID-Zenodo-Dep="4447671" ID-ZooBank="urn:lsid:zoobank.org:pub:54A81360-9A19-456B-AF72-E4868A4835BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610990408013" checkinUser="felipe" docAuthor="Akkemik, Ünal, Mantzouka, Dimitra &amp; Yildirim, Demet Kiran" docDate="2020" docId="03A987CEFFF8020883ACFC05FABDFC91" docLanguage="en" docName="Geodiversitas.42.23.427-441.pdf" docOrigin="Geodiversitas 42 (23)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="432" masterDocId="FF90FFB6FFFF020F832AFFB1FFA8FFA1" masterDocTitle="The first report of Lesbosoxylon Süss &amp; Velitzelos from the early-middle Miocene of eastern Anatolia" masterLastPageNumber="441" masterPageNumber="427" pageNumber="432" updateTime="1698924636178" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="97392E0206E1A6B0F0434745A1E4DF14">The first report of Lesbosoxylon Süss &amp; Velitzelos from the early-middle Miocene of eastern Anatolia</mods:title>
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<mods:namePart id="4B73371FFDA9D8FABE4705DB226334EF">Akkemik, Ünal</mods:namePart>
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<mods:namePart id="ACCBDE2DE3E733BB246779F717BF933E">Mantzouka, Dimitra</mods:namePart>
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<mods:namePart id="F9D5F615975FC817B97F570C5E799D1B">Yildirim, Demet Kiran</mods:namePart>
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<mods:title id="9670203E9F7CC8F5C05C1E0A1632C5C9">Geodiversitas</mods:title>
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<mods:date id="746CE4F990BBFF12ED97C109EF075A03">2020</mods:date>
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<mods:number id="D18F7F92312F0ACB401DBE86C922BBA9">2020-10-01</mods:number>
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<mods:number id="F366ED5AF084AACF1E23CD24995D3BD7">42</mods:number>
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<mods:classification id="D43B41E472BC80F81A03493EAE5798C2">journal article</mods:classification>
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<mods:identifier id="1C6F93F0E7FA5F933214232FBD71FA62" type="DOI">10.5252/geodiversitas2020v42a23</mods:identifier>
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<treatment id="03A987CEFFF8020883ACFC05FABDFC91" ID-DOI="http://doi.org/10.5281/zenodo.4488612" ID-GBIF-Taxon="177202486" ID-Zenodo-Dep="4488612" LSID="urn:lsid:plazi:treatment:03A987CEFFF8020883ACFC05FABDFC91" httpUri="http://treatment.plazi.org/id/03A987CEFFF8020883ACFC05FABDFC91" lastPageNumber="432" pageId="7" pageNumber="432">
<subSubSection id="C31A6553FFF8020883ACFC05FE52FC4E" pageId="7" pageNumber="432" type="nomenclature">
<paragraph id="8BBF36D8FFF8020883ACFC05FE52FC4E" blockId="7.[134,774,948,1007]" pageId="7" pageNumber="432">
<taxonomicName id="4C004D5BFFF8020883ACFC05FD01FC6E" authority="Akkemik &amp; Mantzouka" authorityName="Akkemik &amp; Mantzouka" authorityYear="2020" box="[134,681,948,975]" class="Pinopsida" family="Protopinaceae" genus="Lesbosoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="432" phylum="Tracheophyta" rank="species" species="kemaliyensis" status="sp. nov.">
<emphasis id="B974EACAFFF8020883ACFC05FE35FC6E" bold="true" box="[134,413,948,975]" italics="true" pageId="7" pageNumber="432">Lesbosoxylon kemaliyensis</emphasis>
Akkemik &amp; Mantzouka
</taxonomicName>
,
<taxonomicNameLabel id="A24757B1FFF802088199FC04FCAEFC6E" box="[691,774,949,975]" pageId="7" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="133B2A5DFFF8020882B2FC65FE59FC4F" box="[408,497,980,1007]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="6.[230,241,1863,1880]" captionStartId-1="8.[132,143,1962,1979]" captionStartId-2="10.[132,143,1962,1979]" captionStartId-3="12.[132,143,1940,1957]" captionTargetBox-0="[428,1160,175,1838]" captionTargetBox-1="[179,1411,215,1928]" captionTargetBox-2="[181,1407,215,1906]" captionTargetBox-3="[188,1400,216,1915]" captionTargetId-0="figure-87@6.[428,1160,175,1838]" captionTargetId-1="figure-26@8.[747,1318,703,1273]" captionTargetId-2="figure-26@10.[745,1313,683,1253]" captionTargetId-3="figure-139@12.[739,1306,725,1295]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="10" captionTargetPageId-3="12" captionText-0="FIG. 3. — The petrified wood used in description (specimen KEM01, 7 × 8 × 19 cm, Istanbul University-Cerrahpasa, Faculty of Forestry)." captionText-1="FIG. 4. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, B, transversal section with gradual transition from earlywood to latewood (arrow); C, traumatic resin canals in transversal section (arrow); D, uniseriate rays in tangential section (arrow); E, fusiform ray with horizontal resin canal, and resinous cells around the ray (arrow). Scale bars: A-C, E, 200 µm; D, 100 µm." captionText-2="FIG. 5. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, Biseriate ray (black arrow) and tracheids with resin material (white arrow); B, a tall fusiform ray (arrow); C, D, big rays with idioblast-like cells (asterisk); E, heterocelluler ray cells in a ray; F, G, H, I, axial parenchyma cells with smooth end walls (white arrows), and slightly nodular end walls (black arrow). Scale bars: A, B, E, 100 µm; C, D, F-I, 20 µm." captionText-3="FIG. 6. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, radial section with axial resin canal (arrow); B-D, 1-3 seriate of tracheidal pitting, and crassulae formation common (white arrow); E, pinoid type cross-field pitting, 1-2 pits per cross-field on one row, and smooth end wall of ray cell; F, six pits on two rows in a cross-field (arrow); G, pit border on ray tracheids (arrow); H, one-two seriate of smooth-walled ray tracheids (arrow). Scale bars: A, 200 µm; B, 100 µm; C-G, 20 µm; H, 30 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4447679" figureDoi-1="http://doi.org/10.5281/zenodo.4447681" figureDoi-2="http://doi.org/10.5281/zenodo.4447683" figureDoi-3="http://doi.org/10.5281/zenodo.4447685" httpUri-0="https://zenodo.org/record/4447679/files/figure.png" httpUri-1="https://zenodo.org/record/4447681/files/figure.png" httpUri-2="https://zenodo.org/record/4447683/files/figure.png" httpUri-3="https://zenodo.org/record/4447685/files/figure.png" pageId="7" pageNumber="432">Figs 3-6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31A6553FFF8020883DCFBA6FD3DFB89" box="[246,661,1047,1064]" pageId="7" pageNumber="432" type="description">
<paragraph id="8BBF36D8FFF8020883DCFBA6FD3DFB89" blockId="7.[246,661,1047,1064]" box="[246,661,1047,1064]" pageId="7" pageNumber="432">Plant Fossil Names Registry Number: PFN001780</paragraph>
</subSubSection>
<subSubSection id="C31A6553FFF8020883AEFBE8FCA0FB2A" pageId="7" pageNumber="432" type="materials_examined">
<paragraph id="8BBF36D8FFF8020883AEFBE8FCA0FB2A" blockId="7.[132,776,1112,1163]" pageId="7" pageNumber="432">
<materialsCitation id="3B683C85FFF8020883AEFBE8FCA0FB2A" ID-GBIF-Occurrence="3028143301" collectionCode="KEM" county="University-Cerrahpasa" location="Department of Forest Botany" municipality="Faculty of Forestry" pageId="7" pageNumber="432" specimenCode="KEM01" specimenCount="1" stateProvince="Istanbul" typeStatus="holotype">
<typeStatus id="54BB887AFFF8020883AEFBE8FF4DFBCE" box="[132,229,1113,1136]" pageId="7" pageNumber="432" type="holotype">HOLOTYPE</typeStatus>
. — Specimen
<specimenCode id="DBA69EA3FFF802088249FBE9FE18FBD1" box="[355,432,1112,1136]" collectionCode="KEM" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15230" lsid="urn:lsid:biocol.org:col:15230" name="Kemerovo University" pageId="7" pageNumber="432" type="Herbarium">KEM01</specimenCode>
; repository:
<collectingRegion id="49C4F83AFFF80208813DFBE9FDF7FBD1" box="[535,607,1112,1136]" country="Turkey" name="Istanbul" pageId="7" pageNumber="432">Istanbul</collectingRegion>
<collectingCounty id="62DE4E54FFF80208814BFBE9FF05FB2B" pageId="7" pageNumber="432">University-Cerrahpasa</collectingCounty>
,
<collectingMunicipality id="6BDBACA2FFF80208839EFBC2FEF2FB2A" box="[180,346,1139,1163]" pageId="7" pageNumber="432">Faculty of Forestry</collectingMunicipality>
,
<location id="8EDF6003FFF80208824AFBC2FDCEFB2A" LSID="urn:lsid:plazi:treatment:03A987CEFFF8020883ACFC05FABDFC91:8EDF6003FFF80208824AFBC2FDCEFB2A" box="[352,614,1139,1163]" county="University-Cerrahpasa" municipality="Faculty of Forestry" name="Department of Forest Botany" pageId="7" pageNumber="432" stateProvince="Istanbul">Department of Forest Botany</location>
(designated here).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A6553FFF8020883AEFB12FDCFFB74" pageId="7" pageNumber="432" type="etymology">
<paragraph id="8BBF36D8FFF8020883AEFB12FDCFFB74" blockId="7.[132,775,1186,1237]" pageId="7" pageNumber="432">
ETYMOLOGY. — The epithet “
<taxonomicName id="4C004D5BFFF802088287FB13FDB7FB1B" authorityName="Akkemik &amp; Mantzouka" authorityYear="2020" box="[429,543,1186,1210]" class="Pinopsida" family="Protopinaceae" genus="Lesbosoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="432" phylum="Tracheophyta" rank="species" species="kemaliyensis">
<emphasis id="B974EACAFFF802088287FB13FDB7FB1B" box="[429,543,1186,1210]" italics="true" pageId="7" pageNumber="432">kemaliyensis</emphasis>
</taxonomicName>
” comes from the name of the main city “Kemaliye” nearby the fossil site.
</paragraph>
</subSubSection>
<subSubSection id="C31A6553FFF8020883AEFB5CFD32FAC5" pageId="7" pageNumber="432" type="materials_examined">
<paragraph id="8BBF36D8FFF8020883AEFB5CFE56FAA5" blockId="7.[132,510,1260,1284]" box="[132,510,1260,1284]" pageId="7" pageNumber="432">
<collectionCode id="ED11AE1DFFF8020883AEFB5CFEBFFAA2" box="[132,279,1261,1284]" pageId="7" pageNumber="432">TYPE HORIZON</collectionCode>
. — Divriği Formation.
</paragraph>
<paragraph id="8BBF36D8FFF8020883AEFAADFE14FA95" blockId="7.[132,444,1308,1332]" box="[132,444,1308,1332]" pageId="7" pageNumber="432">AGE. — Early-middle Miocene.</paragraph>
<paragraph id="8BBF36D8FFF8020883AEFAFDFD32FAC5" blockId="7.[132,666,1356,1380]" box="[132,666,1356,1380]" pageId="7" pageNumber="432">
<collectionCode id="ED11AE1DFFF8020883AEFAFDFEBFFAC3" box="[132,279,1356,1379]" pageId="7" pageNumber="432">TYPE LOCALITY</collectionCode>
. — Kemaliye (Eğin),
<collectingRegion id="49C4F83AFFF8020882C5FAFDFDE0FAC5" box="[495,584,1356,1380]" country="Turkey" name="Erzincan" pageId="7" pageNumber="432">Erzincan</collectingRegion>
,
<collectingCountry id="F3177648FFF80208817BFAFDFD3EFAC5" box="[593,662,1356,1380]" name="Turkey" pageId="7" pageNumber="432">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A6553FFF8020883AEFACDFD55F93D" pageId="7" pageNumber="432" type="diagnosis">
<paragraph id="8BBF36D8FFF8020883AEFACDFD55F93D" blockId="7.[131,777,1403,1692]" pageId="7" pageNumber="432">DIAGNOSIS. — Coniferous wood with resin canals. Growth ring boundaries distinct, transition from earlywood to latewood mostly gradual, axial and radial resin canals with thin-walled epithelial cells, latewood tracheids thin to thick walled, bordered pits on radial walls of tracheids 1-2(-3) seriate, crassulae common, rays heterocellular, uniseriate, partly biseriate, uniseriate rays up to 27 cells high, fusiform rays up to 30 seriate high. Axial parenchyma occasionally present. Ray tracheids 2-3 rows, cell walls of ray tracheids smooth. Cross-field pitting pinoid, 1-2(-6) pits per cross-field, horizontal and end walls of ray parenchyma cells smooth and slightly nodular, horizontal walls occasionally pitted, indentures not observed.</paragraph>
</subSubSection>
<subSubSection id="C31A6553FFF8020883AEF963FABDFC91" pageId="7" pageNumber="432" type="description">
<paragraph id="8BBF36D8FFF8020883AEF963FEBDF94B" blockId="7.[130,776,1746,2028]" box="[132,277,1746,1772]" pageId="7" pageNumber="432">DESCRIPTION</paragraph>
<paragraph id="8BBF36D8FFF8020883AEF943FABDFC91" blockId="7.[130,776,1746,2028]" lastBlockId="7.[810,1457,215,816]" pageId="7" pageNumber="432">
The petrified wood sample has grey color; dimensions are 7 × 8 ×
<quantity id="4CF89B3DFFF8020883A8F8A3FF69F88D" box="[130,193,1810,1836]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="432" unit="cm" value="19.0">19 cm</quantity>
(
<figureCitation id="133B2A5DFFF8020883E7F8A3FEABF88D" box="[205,259,1810,1836]" captionStart="FIG" captionStartId="6.[230,241,1863,1880]" captionTargetBox="[428,1160,175,1838]" captionTargetId="figure-87@6.[428,1160,175,1838]" captionTargetPageId="6" captionText="FIG. 3. — The petrified wood used in description (specimen KEM01, 7 × 8 × 19 cm, Istanbul University-Cerrahpasa, Faculty of Forestry)." figureDoi="http://doi.org/10.5281/zenodo.4447679" httpUri="https://zenodo.org/record/4447679/files/figure.png" pageId="7" pageNumber="432">Fig.3</figureCitation>
). In cross section,growth ring boundaries are distinct, transition from earlywood to latewood is mostly gradual (
<figureCitation id="133B2A5DFFF802088192F880FF3FF8CD" captionStart="FIG" captionStartId="8.[132,143,1962,1979]" captionTargetBox="[179,1411,215,1928]" captionTargetId="figure-26@8.[747,1318,703,1273]" captionTargetPageId="8" captionText="FIG. 4. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, B, transversal section with gradual transition from earlywood to latewood (arrow); C, traumatic resin canals in transversal section (arrow); D, uniseriate rays in tangential section (arrow); E, fusiform ray with horizontal resin canal, and resinous cells around the ray (arrow). Scale bars: A-C, E, 200 µm; D, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4447681" httpUri="https://zenodo.org/record/4447681/files/figure.png" pageId="7" pageNumber="432">Fig.4A, B</figureCitation>
). Measured diameters of tracheids and pits are given inTable 1. Axial resin canals with thin walled epithelial cells present mostly in latewood zone (
<figureCitation id="133B2A5DFFF8020882B2F820FDAFF80D" box="[408,519,1937,1964]" captionStart="FIG" captionStartId="8.[132,143,1962,1979]" captionTargetBox="[179,1411,215,1928]" captionTargetId="figure-26@8.[747,1318,703,1273]" captionTargetPageId="8" captionText="FIG. 4. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, B, transversal section with gradual transition from earlywood to latewood (arrow); C, traumatic resin canals in transversal section (arrow); D, uniseriate rays in tangential section (arrow); E, fusiform ray with horizontal resin canal, and resinous cells around the ray (arrow). Scale bars: A-C, E, 200 µm; D, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4447681" httpUri="https://zenodo.org/record/4447681/files/figure.png" pageId="7" pageNumber="432">Fig. 4A, B</figureCitation>
). Traumatic resin canals present (
<figureCitation id="133B2A5DFFF8020883F1F800FE80F86A" box="[219,296,1969,1995]" captionStart="FIG" captionStartId="8.[132,143,1962,1979]" captionTargetBox="[179,1411,215,1928]" captionTargetId="figure-26@8.[747,1318,703,1273]" captionTargetPageId="8" captionText="FIG. 4. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, B, transversal section with gradual transition from earlywood to latewood (arrow); C, traumatic resin canals in transversal section (arrow); D, uniseriate rays in tangential section (arrow); E, fusiform ray with horizontal resin canal, and resinous cells around the ray (arrow). Scale bars: A-C, E, 200 µm; D, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4447681" httpUri="https://zenodo.org/record/4447681/files/figure.png" pageId="7" pageNumber="432">Fig. 4C</figureCitation>
). In tangential section, rays uniseriate (
<figureCitation id="133B2A5DFFF802088180F800FD52F86D" box="[682,762,1969,1996]" captionStart="FIG" captionStartId="8.[132,143,1962,1979]" captionTargetBox="[179,1411,215,1928]" captionTargetId="figure-26@8.[747,1318,703,1273]" captionTargetPageId="8" captionText="FIG. 4. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, B, transversal section with gradual transition from earlywood to latewood (arrow); C, traumatic resin canals in transversal section (arrow); D, uniseriate rays in tangential section (arrow); E, fusiform ray with horizontal resin canal, and resinous cells around the ray (arrow). Scale bars: A-C, E, 200 µm; D, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4447681" httpUri="https://zenodo.org/record/4447681/files/figure.png" pageId="7" pageNumber="432">Fig.4D</figureCitation>
), and fusiform rays present (
<figureCitation id="133B2A5DFFF80208828BF860FE5AF84D" box="[417,498,2001,2028]" captionStart="FIG" captionStartId="8.[132,143,1962,1979]" captionTargetBox="[179,1411,215,1928]" captionTargetId="figure-26@8.[747,1318,703,1273]" captionTargetPageId="8" captionText="FIG. 4. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, B, transversal section with gradual transition from earlywood to latewood (arrow); C, traumatic resin canals in transversal section (arrow); D, uniseriate rays in tangential section (arrow); E, fusiform ray with horizontal resin canal, and resinous cells around the ray (arrow). Scale bars: A-C, E, 200 µm; D, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4447681" httpUri="https://zenodo.org/record/4447681/files/figure.png" pageId="7" pageNumber="432">Fig. 4E</figureCitation>
). Rays occasionally biseri-ate or partly biseriate (
<figureCitation id="133B2A5DFFF80208870EFF66FBD1FF50" box="[1060,1145,215,242]" captionStart="FIG" captionStartId="10.[132,143,1962,1979]" captionTargetBox="[181,1407,215,1906]" captionTargetId="figure-26@10.[745,1313,683,1253]" captionTargetPageId="10" captionText="FIG. 5. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, Biseriate ray (black arrow) and tracheids with resin material (white arrow); B, a tall fusiform ray (arrow); C, D, big rays with idioblast-like cells (asterisk); E, heterocelluler ray cells in a ray; F, G, H, I, axial parenchyma cells with smooth end walls (white arrows), and slightly nodular end walls (black arrow). Scale bars: A, B, E, 100 µm; C, D, F-I, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4447683" httpUri="https://zenodo.org/record/4447683/files/figure.png" pageId="7" pageNumber="432">Fig. 5A</figureCitation>
). Big rays with idioblast-like cells present (
<figureCitation id="133B2A5DFFF8020880ECFF46FBA8FEB3" box="[966,1024,247,274]" captionStart="FIG" captionStartId="10.[132,143,1962,1979]" captionTargetBox="[181,1407,215,1906]" captionTargetId="figure-26@10.[745,1313,683,1253]" captionTargetPageId="10" captionText="FIG. 5. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, Biseriate ray (black arrow) and tracheids with resin material (white arrow); B, a tall fusiform ray (arrow); C, D, big rays with idioblast-like cells (asterisk); E, heterocelluler ray cells in a ray; F, G, H, I, axial parenchyma cells with smooth end walls (white arrows), and slightly nodular end walls (black arrow). Scale bars: A, B, E, 100 µm; C, D, F-I, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4447683" httpUri="https://zenodo.org/record/4447683/files/figure.png" pageId="7" pageNumber="432">Fig 5</figureCitation>
B-D). Ray cells uniform in some rays, and heterocellular, wider or narrower (
<figureCitation id="133B2A5DFFF8020887F0FEA6FAB2FE93" box="[1242,1306,279,306]" captionStart="FIG" captionStartId="10.[132,143,1962,1979]" captionTargetBox="[181,1407,215,1906]" captionTargetId="figure-26@10.[745,1313,683,1253]" captionTargetPageId="10" captionText="FIG. 5. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, Biseriate ray (black arrow) and tracheids with resin material (white arrow); B, a tall fusiform ray (arrow); C, D, big rays with idioblast-like cells (asterisk); E, heterocelluler ray cells in a ray; F, G, H, I, axial parenchyma cells with smooth end walls (white arrows), and slightly nodular end walls (black arrow). Scale bars: A, B, E, 100 µm; C, D, F-I, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4447683" httpUri="https://zenodo.org/record/4447683/files/figure.png" pageId="7" pageNumber="432">Fig. 5</figureCitation>
D-E). Height of uniseriate rays up to 27 cells (
<figureCitation id="133B2A5DFFF8020887ACFE86FB70FEF0" box="[1158,1240,311,337]" captionStart="FIG" captionStartId="8.[132,143,1962,1979]" captionTargetBox="[179,1411,215,1928]" captionTargetId="figure-26@8.[747,1318,703,1273]" captionTargetPageId="8" captionText="FIG. 4. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, B, transversal section with gradual transition from earlywood to latewood (arrow); C, traumatic resin canals in transversal section (arrow); D, uniseriate rays in tangential section (arrow); E, fusiform ray with horizontal resin canal, and resinous cells around the ray (arrow). Scale bars: A-C, E, 200 µm; D, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4447681" httpUri="https://zenodo.org/record/4447681/files/figure.png" pageId="7" pageNumber="432">Fig. 4C</figureCitation>
), height of fusiform rays up to 30 cells and wider (
<figureCitation id="133B2A5DFFF802088741FEE6FB16FED0" box="[1131,1214,343,369]" captionStart="FIG" captionStartId="8.[132,143,1962,1979]" captionTargetBox="[179,1411,215,1928]" captionTargetId="figure-26@8.[747,1318,703,1273]" captionTargetPageId="8" captionText="FIG. 4. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, B, transversal section with gradual transition from earlywood to latewood (arrow); C, traumatic resin canals in transversal section (arrow); D, uniseriate rays in tangential section (arrow); E, fusiform ray with horizontal resin canal, and resinous cells around the ray (arrow). Scale bars: A-C, E, 200 µm; D, 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4447681" httpUri="https://zenodo.org/record/4447681/files/figure.png" pageId="7" pageNumber="432">Fig. 4D</figureCitation>
and
<figureCitation id="133B2A5DFFF8020887D8FEE6FAE0FED0" box="[1266,1352,343,370]" captionStart="FIG" captionStartId="10.[132,143,1962,1979]" captionTargetBox="[181,1407,215,1906]" captionTargetId="figure-26@10.[745,1313,683,1253]" captionTargetPageId="10" captionText="FIG. 5. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, Biseriate ray (black arrow) and tracheids with resin material (white arrow); B, a tall fusiform ray (arrow); C, D, big rays with idioblast-like cells (asterisk); E, heterocelluler ray cells in a ray; F, G, H, I, axial parenchyma cells with smooth end walls (white arrows), and slightly nodular end walls (black arrow). Scale bars: A, B, E, 100 µm; C, D, F-I, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4447683" httpUri="https://zenodo.org/record/4447683/files/figure.png" pageId="7" pageNumber="432">Fig. 5D</figureCitation>
). Average height of rays 4-15 cells. Radial resin canals with thin–walled epithelial cells. Axial parenchyma cells occasionally present, and generally smooth transverse end walls (
<figureCitation id="133B2A5DFFF80208863AFE07FAF8FE70" box="[1296,1360,438,465]" captionStart="FIG" captionStartId="10.[132,143,1962,1979]" captionTargetBox="[181,1407,215,1906]" captionTargetId="figure-26@10.[745,1313,683,1253]" captionTargetPageId="10" captionText="FIG. 5. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, Biseriate ray (black arrow) and tracheids with resin material (white arrow); B, a tall fusiform ray (arrow); C, D, big rays with idioblast-like cells (asterisk); E, heterocelluler ray cells in a ray; F, G, H, I, axial parenchyma cells with smooth end walls (white arrows), and slightly nodular end walls (black arrow). Scale bars: A, B, E, 100 µm; C, D, F-I, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4447683" httpUri="https://zenodo.org/record/4447683/files/figure.png" pageId="7" pageNumber="432">Fig. 5</figureCitation>
F-G and I), and slightly nodular (
<figureCitation id="133B2A5DFFF802088714FE67FB3FFE50" box="[1086,1175,470,497]" captionStart="FIG" captionStartId="10.[132,143,1962,1979]" captionTargetBox="[181,1407,215,1906]" captionTargetId="figure-26@10.[745,1313,683,1253]" captionTargetPageId="10" captionText="FIG. 5. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, Biseriate ray (black arrow) and tracheids with resin material (white arrow); B, a tall fusiform ray (arrow); C, D, big rays with idioblast-like cells (asterisk); E, heterocelluler ray cells in a ray; F, G, H, I, axial parenchyma cells with smooth end walls (white arrows), and slightly nodular end walls (black arrow). Scale bars: A, B, E, 100 µm; C, D, F-I, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4447683" httpUri="https://zenodo.org/record/4447683/files/figure.png" pageId="7" pageNumber="432">Fig. 5H</figureCitation>
). Axial resin canal visible in radial section (
<figureCitation id="133B2A5DFFF8020880CDFE47FB92FDB1" box="[999,1082,502,528]" captionStart="FIG" captionStartId="12.[132,143,1940,1957]" captionTargetBox="[188,1400,216,1915]" captionTargetId="figure-139@12.[739,1306,725,1295]" captionTargetPageId="12" captionText="FIG. 6. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, radial section with axial resin canal (arrow); B-D, 1-3 seriate of tracheidal pitting, and crassulae formation common (white arrow); E, pinoid type cross-field pitting, 1-2 pits per cross-field on one row, and smooth end wall of ray cell; F, six pits on two rows in a cross-field (arrow); G, pit border on ray tracheids (arrow); H, one-two seriate of smooth-walled ray tracheids (arrow). Scale bars: A, 200 µm; B, 100 µm; C-G, 20 µm; H, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.4447685" httpUri="https://zenodo.org/record/4447685/files/figure.png" pageId="7" pageNumber="432">Fig. 6A</figureCitation>
). Tracheid pitting in radial walls of earlywood 1-2 (-3) seriate opposite and crassulae formation common (
<figureCitation id="133B2A5DFFF8020880B0FD87FC7EFDF0" box="[922,982,566,593]" captionStart="FIG" captionStartId="10.[132,143,1962,1979]" captionTargetBox="[181,1407,215,1906]" captionTargetId="figure-26@10.[745,1313,683,1253]" captionTargetPageId="10" captionText="FIG. 5. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, Biseriate ray (black arrow) and tracheids with resin material (white arrow); B, a tall fusiform ray (arrow); C, D, big rays with idioblast-like cells (asterisk); E, heterocelluler ray cells in a ray; F, G, H, I, axial parenchyma cells with smooth end walls (white arrows), and slightly nodular end walls (black arrow). Scale bars: A, B, E, 100 µm; C, D, F-I, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4447683" httpUri="https://zenodo.org/record/4447683/files/figure.png" pageId="7" pageNumber="432">Fig. 5</figureCitation>
B-D). Cross-field pitting pinoid in 1(-2) rows of usually 1-2(-6) pits per cross-field (
<figureCitation id="133B2A5DFFF8020887EEFDE7FA9AFDD1" box="[1220,1330,598,624]" captionStart="FIG" captionStartId="12.[132,143,1940,1957]" captionTargetBox="[188,1400,216,1915]" captionTargetId="figure-139@12.[739,1306,725,1295]" captionTargetPageId="12" captionText="FIG. 6. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, radial section with axial resin canal (arrow); B-D, 1-3 seriate of tracheidal pitting, and crassulae formation common (white arrow); E, pinoid type cross-field pitting, 1-2 pits per cross-field on one row, and smooth end wall of ray cell; F, six pits on two rows in a cross-field (arrow); G, pit border on ray tracheids (arrow); H, one-two seriate of smooth-walled ray tracheids (arrow). Scale bars: A, 200 µm; B, 100 µm; C-G, 20 µm; H, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.4447685" httpUri="https://zenodo.org/record/4447685/files/figure.png" pageId="7" pageNumber="432">Fig. 6E, F</figureCitation>
). End walls of ray parenchyma cells smooth, no indentures observed. Rays heterocellular. Ray tracheids present, 1-3 seriate, smooth walled (
<figureCitation id="133B2A5DFFF8020880A9FD04FC72FD71" box="[899,986,693,720]" captionStart="FIG" captionStartId="12.[132,143,1940,1957]" captionTargetBox="[188,1400,216,1915]" captionTargetId="figure-139@12.[739,1306,725,1295]" captionTargetPageId="12" captionText="FIG. 6. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, radial section with axial resin canal (arrow); B-D, 1-3 seriate of tracheidal pitting, and crassulae formation common (white arrow); E, pinoid type cross-field pitting, 1-2 pits per cross-field on one row, and smooth end wall of ray cell; F, six pits on two rows in a cross-field (arrow); G, pit border on ray tracheids (arrow); H, one-two seriate of smooth-walled ray tracheids (arrow). Scale bars: A, 200 µm; B, 100 µm; C-G, 20 µm; H, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.4447685" httpUri="https://zenodo.org/record/4447685/files/figure.png" pageId="7" pageNumber="432">Fig. 6G</figureCitation>
). Ray cells smooth walled (
<figureCitation id="133B2A5DFFF80208862DFD04FAEEFD6E" box="[1287,1350,693,719]" captionStart="FIG" captionStartId="12.[132,143,1940,1957]" captionTargetBox="[188,1400,216,1915]" captionTargetId="figure-139@12.[739,1306,725,1295]" captionTargetPageId="12" captionText="FIG. 6. — Wood features of Lesbosoxylon kemaliyensis Akkemik &amp; Mantzouka, sp. nov.: A, radial section with axial resin canal (arrow); B-D, 1-3 seriate of tracheidal pitting, and crassulae formation common (white arrow); E, pinoid type cross-field pitting, 1-2 pits per cross-field on one row, and smooth end wall of ray cell; F, six pits on two rows in a cross-field (arrow); G, pit border on ray tracheids (arrow); H, one-two seriate of smooth-walled ray tracheids (arrow). Scale bars: A, 200 µm; B, 100 µm; C-G, 20 µm; H, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.4447685" httpUri="https://zenodo.org/record/4447685/files/figure.png" pageId="7" pageNumber="432">Fig. 6</figureCitation>
E-H) and occasionally pitted. These descriptive features are also provided in Appendix A based on “Criteria used according the IAWA for fossil conifers” by
<bibRefCitation id="EF914B29FFF802088702FCA4FAB8FC91" author="WHEELER E. A. &amp; BAAS P. &amp; GASSON P. E. &amp; EDS" box="[1064,1296,789,816]" pageId="7" pageNumber="432" pagination="1 - 70" refId="ref11967" refString="WHEELER E. A., BAAS P. &amp; GASSON P. E. (EDS) 2004. - IAWA list of microscopic features for softwood identification. IAWA Journal 25: 1 - 70. https: // doi. org / 10.1163 / 22941932 - 90000349" type="journal article" year="2004">
Wheeler
<emphasis id="B974EACAFFF8020887A0FCA7FB15FC8E" box="[1162,1213,789,815]" italics="true" pageId="7" pageNumber="432">et al.</emphasis>
(2004)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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