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<document id="92F858DEEF2410EB2BD9DD5F7A8AF2B2" ID-DOI="10.5281/zenodo.5276149" ID-ISSN="0253-116X" ID-Zenodo-Dep="5276149" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630010787000" checkinUser="carolina" docAuthor="Assing, V." docDate="2009" docId="03A987C5FFA1FFEBFF2C170DFE1BB6D2" docLanguage="en" docName="LinzerbiolBeitr.2009.41.1.397-426.pdf" docOrigin="Linzer biologische Beiträge 41 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Ochthephilum besucheti" docType="treatment" docVersion="3" lastPageNumber="408" masterDocId="FF90FFBDFFABFFE0FF9D1365FFCCB760" masterDocTitle="On the Western Palaearctic and Middle Asian species of Ochthephilum STEPHENS, with notes on Cryptobium koltzei EPPELSHEIM (Coleoptera: Staphylinidae: Paederinae: Cryptobiina)" masterLastPageNumber="426" masterPageNumber="397" pageNumber="407" updateTime="1699136399403" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="84E1DFF227052E24FD0078A6A4342B11">On the Western Palaearctic and Middle Asian species of Ochthephilum STEPHENS, with notes on Cryptobium koltzei EPPELSHEIM (Coleoptera: Staphylinidae: Paederinae: Cryptobiina)</mods:title>
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<mods:namePart id="96900EF8AC27BF2AE24D85C6C9018064">Assing, V.</mods:namePart>
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<mods:title id="BE2AFDFE54B0AC6C15E2DDD4877A6CBC">Linzer biologische Beiträge</mods:title>
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<mods:date id="15418AF6F1DECD784FC7B67813A43E19">2009</mods:date>
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<mods:number id="1B507C03381BB60A950426D523F55F42">2009-08-30</mods:number>
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<mods:number id="F2C95B482540E9A1C6520B70066A3DC6">41</mods:number>
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<mods:identifier id="6D7ABB7EBDC38EEE11383A394DF432CF" type="DOI">10.5281/zenodo.5276149</mods:identifier>
<mods:identifier id="BE133AE118DA0EC48C449B4C034FED0E" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="28922CD92D579686E2A0268932E8D382" type="Zenodo-Dep">5276149</mods:identifier>
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<treatment id="03A987C5FFA1FFEBFF2C170DFE1BB6D2" LSID="urn:lsid:plazi:treatment:03A987C5FFA1FFEBFF2C170DFE1BB6D2" httpUri="http://treatment.plazi.org/id/03A987C5FFA1FFEBFF2C170DFE1BB6D2" lastPageId="11" lastPageNumber="408" pageId="10" pageNumber="407">
<subSubSection id="C31A6558FFA1FFEAFF2C170DFCDCB31E" box="[177,784,1128,1150]" pageId="10" pageNumber="407" type="nomenclature">
<paragraph id="8BBF36D3FFA1FFEAFF2C170DFCDCB31E" blockId="10.[177,784,1128,1150]" box="[177,784,1128,1150]" pageId="10" pageNumber="407">
<heading id="D0F781BFFFA1FFEAFF2C170DFCDCB31E" box="[177,784,1128,1150]" fontSize="8" level="2" pageId="10" pageNumber="407" reason="2">
<taxonomicName id="4C004D50FFA1FFEAFF2C170DFDAEB31E" ID-CoL="48FR7" authority="(BORDONI 1980)" baseAuthorityName="Bordoni" baseAuthorityYear="1980" box="[177,610,1128,1150]" class="Insecta" family="Staphylinidae" genus="Ochthephilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B974EAC1FFA1FFEAFF2C170DFDAEB31E" bold="true" box="[177,610,1128,1150]" pageId="10" pageNumber="407">
<emphasis id="B974EAC1FFA1FFEAFF2C170DFE7CB31E" bold="true" box="[177,432,1128,1150]" italics="true" pageId="10" pageNumber="407">Ochthephilum besucheti</emphasis>
(
<bibRefCitation id="EF914B22FFA1FFEAFE23170DFDEEB31C" author="BORDONI A" box="[446,546,1128,1150]" pageId="10" pageNumber="407" pagination="75 - 82" refId="ref20260" refString="BORDONI A. (1980): Studi sui Paederinae. VI. Nuove specie del mediterraneo orientale. - Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale di Milano 121: 75 - 82." type="journal article" year="1980">BORDONI</bibRefCitation>
1980)
</emphasis>
</taxonomicName>
(Fig. 23,
<figureCitation id="133B2A56FFA1FFEAFD58170DFCCBB31E" box="[709,775,1128,1150]" captionStart="Map 2" captionStartId="5.[177,222,1537,1558]" captionTargetBox="[177,1084,1003,1513]" captionTargetId="figure-64@5.[177,1084,1003,1513]" captionTargetPageId="5" captionText="Map 2: Distributions of Ochthephilum besuchti (BORDONI) (open circles), O. permutatum nov.sp. (filled circles), O. egregium (REITTER) (open squares), and O. hamatum ASSING (filled squares), based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5276153" httpUri="https://zenodo.org/record/5276153/files/figure.png" pageId="10" pageNumber="407">Map 2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A6558FFA1FFEAFF2C17EBFBF1B3D9" pageId="10" pageNumber="407" type="reference_group">
<paragraph id="8BBF36D3FFA1FFEAFF2C17EBFDA5B3C2" blockId="10.[177,1086,1166,1454]" box="[177,617,1166,1186]" pageId="10" pageNumber="407">
<taxonomicName id="4C004D50FFA1FFEAFF2C17EBFDD2B3C2" ID-CoL="6BL54" authority=": HOZMAN (1985)" authorityName="HOZMAN" authorityYear="1985" box="[177,542,1166,1186]" class="Insecta" family="Staphylinidae" genus="Cryptobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="egregium">
<emphasis id="B974EAC1FFA1FFEAFF2C17EBFEB5B3C2" box="[177,377,1166,1186]" italics="true" pageId="10" pageNumber="407">Cryptobium egregium</emphasis>
:
<bibRefCitation id="EF914B22FFA1FFEAFE1817EBFDD2B3C2" author="HOZMAN P" box="[389,542,1166,1186]" pageId="10" pageNumber="407" pagination="311 - 322" refId="ref20514" refString="HOZMAN P. (1985): Beitrag zur Kenntnis westpalaarktischer Arten der Gattung Cryptobium MANNHEIM [sic] (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 92: 311 - 322." type="journal article" year="1985">HOZMAN (1985)</bibRefCitation>
</taxonomicName>
; partim.
</paragraph>
<paragraph id="8BBF36D3FFA1FFEAFF2C17C0FBF1B3D9" blockId="10.[177,1086,1166,1454]" box="[177,1085,1187,1209]" pageId="10" pageNumber="407">
<materialsCitation id="3B683C8EFFA1FFEAFF2C17C0FBF1B3D9" box="[177,1085,1187,1209]" country="Turkey" location="Samsun-Bafra" pageId="10" pageNumber="407" specimenCount="1" stateProvince="Samsun" typeStatus="holotype">
T y p e m a t e r i a l e x a m i n e d
<typeStatus id="54BB8871FFA1FFEAFDF517C0FD79B3D9" box="[616,693,1189,1209]" pageId="10" pageNumber="407" type="holotype">Holotype</typeStatus>
3: &quot;
<collectingCountry id="F3177643FFA1FFEAFD6017C0FC8BB3D9" box="[765,839,1189,1209]" name="Turkey" pageId="10" pageNumber="407">Turquie</collectingCountry>
<collectingRegion id="49C4F831FFA1FFEAFCCA17C0FC6DB3D9" box="[855,929,1189,1209]" country="Turkey" name="Samsun" pageId="10" pageNumber="407">Samsun</collectingRegion>
,
<location id="8EDF6008FFA1FFEAFC2817C0FBF5B3D9" LSID="urn:lsid:plazi:treatment:03A987C5FFA1FFEBFF2C170DFE1BB6D2:8EDF6008FFA1FFEAFC2817C0FBF5B3D9" box="[949,1081,1189,1209]" country="Turkey" name="Samsun-Bafra" pageId="10" pageNumber="407" stateProvince="Samsun">Samsun-Bafra</location>
,
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A6558FFA1FFEBFF2C17D9FE1BB6D2" lastPageId="11" lastPageNumber="408" pageId="10" pageNumber="407" type="description">
<paragraph id="8BBF36D3FFA1FFEAFF2C17D9FBF0B3B0" blockId="10.[177,1086,1166,1454]" box="[177,1084,1212,1232]" pageId="10" pageNumber="407">
19.V.67, Cl. Besuchet /
<typeStatus id="54BB8871FFA1FFEAFE3317D9FDC1B3B0" box="[430,525,1212,1232]" pageId="10" pageNumber="407" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C004D50FFA1FFEAFD8617D9FD21B3B0" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[539,749,1212,1232]" class="Insecta" family="Staphylinidae" genus="Cryptobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="besucheti" status="sp. nov.">Cryptobium besucheti</taxonomicName>
<taxonomicNameLabel id="A24757BAFFA1FFEAFD6617D9FCFAB3B0" box="[763,822,1212,1232]" pageId="10" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
, Det. A. Bordoni 1977 /
</paragraph>
<paragraph id="8BBF36D3FFA1FFEAFF2C17B7FCDFB386" blockId="10.[177,1086,1166,1454]" box="[177,787,1234,1254]" pageId="10" pageNumber="407">
<taxonomicName id="4C004D50FFA1FFEAFF2C17B7FE22B386" authority="(Bordoni)" baseAuthorityName="Bordoni" baseAuthorityYear="1980" box="[177,494,1234,1254]" class="Insecta" family="Staphylinidae" genus="Ochthephilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="besucheti">Ochthephilum besucheti (Bordoni)</taxonomicName>
det. V. Assing 2008&quot; (MHNG).
</paragraph>
<paragraph id="8BBF36D3FFA1FFEAFF2C1797FEB1B2CE" blockId="10.[177,1086,1166,1454]" pageId="10" pageNumber="407">
A d d i t i o n a l m a t e r i a l e x a m i n e d
<collectingCountry id="F3177643FFA1FFEAFD261797FD39B266" box="[699,757,1266,1286]" name="Turkey" pageId="10" pageNumber="407">Turkey</collectingCountry>
: O r d u
<quantity id="4CF89B36FFA1FFEAFCF41797FC1FB266" box="[873,979,1265,1286]" metricMagnitude="4" metricUnit="m" metricValue="1.315" pageId="10" pageNumber="407" unit="km" value="13.15">13, 15 km</quantity>
S
<collectingRegion id="49C4F831FFA1FFEAFC691797FBEFB266" box="[1012,1059,1266,1286]" country="Turkey" name="Ordu" pageId="10" pageNumber="407">Ordu</collectingRegion>
, S Kabaduz,
<geoCoordinate id="EE345014FFA1FFEAFEA2166FFE45B27E" box="[319,393,1290,1310]" degrees="40" direction="north" minutes="49" orientation="latitude" pageId="10" pageNumber="407" precision="925" value="40.816666">40°49'N</geoCoordinate>
,
<geoCoordinate id="EE345014FFA1FFEAFE07166FFE2DB27E" box="[410,481,1290,1310]" degrees="37" direction="east" minutes="54" orientation="longitude" pageId="10" pageNumber="407" precision="925" value="37.9">37°54'E</geoCoordinate>
,
<quantity id="4CF89B36FFA1FFEAFE6F166FFDF8B27E" box="[498,564,1290,1310]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="10" pageNumber="407" unit="m" value="990.0">990 m</quantity>
, grassy road margin,
<date id="FFBE1013FFA1FFEAFC92166FFCB7B27E" box="[783,891,1290,1310]" pageId="10" pageNumber="407" value="2006-07-30">30.VII.2006</date>
, leg Assing (cAss). E r z u r u m:13, ca.
<quantity id="4CF89B36FFA1FFEAFE271647FE3FB256" box="[442,499,1314,1334]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="407" unit="km" value="35.0">35 km</quantity>
NW Tortum, Mescit Dağları, ca.
<geoCoordinate id="EE345014FFA1FFEAFC801647FCAAB256" box="[797,870,1314,1334]" degrees="40" direction="north" minutes="30" orientation="latitude" pageId="10" pageNumber="407" precision="925" value="40.5">40°30'N</geoCoordinate>
,
<geoCoordinate id="EE345014FFA1FFEAFCED1647FC7BB256" box="[880,951,1314,1334]" degrees="41" direction="east" minutes="25" orientation="longitude" pageId="10" pageNumber="407" precision="925" value="41.416668">41°25'E</geoCoordinate>
,
<quantity id="4CF89B36FFA1FFEAFC5C1647FBF5B256" box="[961,1081,1314,1334]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="10" pageNumber="407" unit="m" value="1850.0" valueMax="2000.0" valueMin="1700.0">1700-2000 m</quantity>
,
<date id="FFBE1013FFA1FFEAFF43165FFE8FB22E" box="[222,323,1338,1358]" pageId="10" pageNumber="407" value="1998-06-17">17.VI.1998</date>
, leg. Solodovnikov (cAss). E r z i n c a n: 233, Tercan, Euphrat,
<quantity id="4CF89B36FFA1FFEAFC76165FFBF5B22E" box="[1003,1081,1338,1358]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="407" unit="m" value="1400.0">1400 m</quantity>
,
<date id="FFBE1013FFA1FFEAFF431637FEF5B206" box="[222,313,1362,1382]" pageId="10" pageNumber="407" value="1986-06-06">6.VI.1986</date>
, leg. Besuchet, Löbl &amp; Burckhardt (MHNG, cAss).
<collectingCountry id="F3177643FFA1FFEAFC9A1637FC9DB206" box="[775,849,1362,1382]" name="Georgia" pageId="10" pageNumber="407">Georgia</collectingCountry>
: 13,
<collectingRegion id="49C4F831FFA1FFEAFC1E1637FC77B206" box="[899,955,1362,1382]" country="Georgia" name="Tbilisi" pageId="10" pageNumber="407">Tbilisi</collectingRegion>
env., Mzcheta [
<geoCoordinate id="EE345014FFA1FFEAFF79160FFEFCB21E" box="[228,304,1386,1406]" degrees="41" direction="north" minutes="51" orientation="latitude" pageId="10" pageNumber="407" precision="925" value="41.85">41°51'N</geoCoordinate>
,
<geoCoordinate id="EE345014FFA1FFEAFEA7160FFE4FB21E" box="[314,387,1386,1406]" degrees="44" direction="east" minutes="43" orientation="longitude" pageId="10" pageNumber="407" precision="925" value="44.716667">44°43'E</geoCoordinate>
],
<date id="FFBE1013FFA1FFEAFE0E160FFE34B21E" box="[403,504,1386,1406]" pageId="10" pageNumber="407" value="1988-06-26">26.VI.1988</date>
, leg. Wrase &amp; Schülke (cSch).
<collectingCountry id="F3177643FFA1FFEAFC88160FFCF7B21E" box="[789,827,1386,1406]" name="Iran" pageId="10" pageNumber="407">Iran</collectingCountry>
: 13 [&quot;
<taxonomicName id="4C004D50FFA1FFEAFCEB160FFEBCB2F6" authority="Reitter, Hozman" authorityName="Reitter, Hozman" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Cryptobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="egregium">Cryptobium egregium Reitter, Hozman</taxonomicName>
det. 1983&quot;],
<collectingRegion id="49C4F831FFA1FFEAFE7816E7FD9FB2F6" box="[485,595,1410,1430]" country="Iran" name="Mazandaran" pageId="10" pageNumber="407">Mazanderan</collectingRegion>
, Baladeh,
<geoCoordinate id="EE345014FFA1FFEAFD2D16E7FD36B2F6" box="[688,762,1410,1430]" degrees="36" direction="north" minutes="13" orientation="latitude" pageId="10" pageNumber="407" precision="925" value="36.216667">36°13'N</geoCoordinate>
,
<geoCoordinate id="EE345014FFA1FFEAFC9916E7FC80B2F6" box="[772,844,1410,1430]" degrees="51" direction="east" minutes="49" orientation="longitude" pageId="10" pageNumber="407" precision="925" value="51.816666">51°49'E</geoCoordinate>
,
<quantity id="4CF89B36FFA1FFEAFCCB16E7FC50B2F6" box="[854,924,1410,1430]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="407" unit="m" value="2200.0">2200 m</quantity>
,
<date id="FFBE1013FFA1FFEAFC3B16E7FBDEB2F6" box="[934,1042,1410,1430]" pageId="10" pageNumber="407" value="1974-07-12">12.VII.1974</date>
, leg. Senglet (MHNG).
</paragraph>
<paragraph id="8BBF36D3FFA1FFEBFF2C16DBFC43B64F" blockId="10.[177,1085,1470,1584]" lastBlockId="11.[177,1085,220,303]" lastPageId="11" lastPageNumber="408" pageId="10" pageNumber="407">
C o m m e n t: The original description of
<taxonomicName id="4C004D50FFA1FFEAFD1516DBFCC5B2B4" baseAuthorityName="Bordoni" baseAuthorityYear="1980" box="[648,777,1470,1492]" class="Insecta" family="Staphylinidae" genus="Ochthephilum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B974EAC1FFA1FFEAFD1516DBFCC5B2B4" box="[648,777,1470,1492]" italics="true" pageId="10" pageNumber="407">O. besucheti</emphasis>
</taxonomicName>
is based on a male
<typeStatus id="54BB8871FFA1FFEAFC7916DBFBF1B2B4" box="[996,1085,1470,1492]" pageId="10" pageNumber="407" type="holotype">holotype</typeStatus>
from &quot;
<collectingRegion id="49C4F831FFA1FFEAFF6E16B9FE8BB292" box="[243,327,1500,1522]" country="Turkey" name="Samsun" pageId="10" pageNumber="407">Samsun</collectingRegion>
, Bafra&quot; and
<specimenCount id="9D06FD5AFFA1FFEAFE5016B8FDF3B292" box="[461,575,1500,1522]" pageId="10" pageNumber="407" type="female">two female</specimenCount>
<typeStatus id="54BB8871FFA1FFEAFDD416B8FD66B292" box="[585,682,1501,1522]" pageId="10" pageNumber="407" type="paratype">paratypes</typeStatus>
from
<collectingRegion id="49C4F831FFA1FFEAFD7116B8FC88B292" box="[748,836,1501,1522]" country="Turkey" name="Erzurum" pageId="10" pageNumber="407">Erzurum</collectingRegion>
and
<collectingRegion id="49C4F831FFA1FFEAFCE616B9FC00B292" box="[891,972,1500,1522]" country="Turkey" name="Istanbul" pageId="10" pageNumber="407">Istanbul</collectingRegion>
(
<bibRefCitation id="EF914B22FFA1FFEAFC4016B9FF24B171" author="BORDONI A" pageId="10" pageNumber="407" pagination="75 - 82" refId="ref20260" refString="BORDONI A. (1980): Studi sui Paederinae. VI. Nuove specie del mediterraneo orientale. - Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale di Milano 121: 75 - 82." type="journal article" year="1980">BORDONI 1980</bibRefCitation>
). The
<typeStatus id="54BB8871FFA1FFEAFEB6169EFE48B171" box="[299,388,1531,1553]" pageId="10" pageNumber="407" type="holotype">holotype</typeStatus>
was examined and its aedeagus figured by
<bibRefCitation id="EF914B22FFA1FFEAFCD01699FC34B171" author="HOZMAN P" box="[845,1016,1531,1553]" pageId="10" pageNumber="407" pagination="311 - 322" refId="ref20514" refString="HOZMAN P. (1985): Beitrag zur Kenntnis westpalaarktischer Arten der Gattung Cryptobium MANNHEIM [sic] (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 92: 311 - 322." type="journal article" year="1985">HOZMAN (1985)</bibRefCitation>
. Since the species can be reliably distinguished from similar congeners only based on the male primary sexual characters, the identity of the
<specimenCount id="9D06FD5AFFA0FFEBFD1E13B9FCD9B791" box="[643,789,220,241]" pageId="11" pageNumber="408" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
, particularly that from
<collectingRegion id="49C4F831FFA0FFEBFB9E13B9FF18B671" country="Turkey" name="Istanbul" pageId="11" pageNumber="408">Istanbul</collectingRegion>
, is doubtful. For illustrations of the internal structures of the aedeagus see Fig. 23. Almost all the examined specimens have a uniformly blackish forebody.
</paragraph>
<paragraph id="8BBF36D3FFA0FFEBFF2C1224FE1BB6D2" blockId="11.[177,1085,320,434]" pageId="11" pageNumber="408">
D i s t r i b u t i o n: The species is currently known from several localities in northeastern
<collectingCountry id="F3177643FFA0FFEBFF62123AFE87B615" box="[255,331,351,373]" name="Turkey" pageId="11" pageNumber="408">Turkey</collectingCountry>
, one locality in
<collectingCountry id="F3177643FFA0FFEBFE73123AFD8EB615" box="[494,578,351,373]" name="Georgia" pageId="11" pageNumber="408">Georgia</collectingCountry>
, and one locality in northern
<collectingCountry id="F3177643FFA0FFEBFCF0123AFC54B614" box="[877,920,351,372]" name="Iran" pageId="11" pageNumber="408">Iran</collectingCountry>
, suggesting that it is a Caspian element (
<figureCitation id="133B2A56FFA0FFEBFE3A1218FE20B6F3" box="[423,492,381,403]" captionStart="Map 2" captionStartId="5.[177,222,1537,1558]" captionTargetBox="[177,1084,1003,1513]" captionTargetId="figure-64@5.[177,1084,1003,1513]" captionTargetPageId="5" captionText="Map 2: Distributions of Ochthephilum besuchti (BORDONI) (open circles), O. permutatum nov.sp. (filled circles), O. egregium (REITTER) (open squares), and O. hamatum ASSING (filled squares), based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5276153" httpUri="https://zenodo.org/record/5276153/files/figure.png" pageId="11" pageNumber="408">Map 2</figureCitation>
). The specimens from
<collectingCountry id="F3177643FFA0FFEBFD4B1218FCEBB6F3" box="[726,807,381,403]" name="Georgia" pageId="11" pageNumber="408">Georgia</collectingCountry>
and
<collectingCountry id="F3177643FFA0FFEBFCC7121BFC4FB6F3" box="[858,899,382,403]" name="Iran" pageId="11" pageNumber="408">Iran</collectingCountry>
represent the first records from these countries.
</paragraph>
</subSubSection>
</treatment>
</document>
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