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<document id="7B7BBB8B30B1CD5D4947ED49FAABD7EE" ID-DOI="10.11646/zootaxa.5343.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8324250" ID-ZooBank="20BF9AB4-F86D-42EA-AEAA-7C03EBA44EB6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1694069790509" checkinUser="plazi" docAuthor="Yamasaki, Takeshi, Hashimoto, Yoshiaki, Endo, Tomoji, Hyodo, Fujio, Itioka, Takao, Mohamed, Maryati &amp; Meleng, Paulus" docDate="2023" docId="03FA87E7FFB8FFC9FF7D388AFB6F899F" docLanguage="en" docName="zootaxa.5343.1.3.pdf" docOrigin="Zootaxa 5343 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Utivarachna kinabaluensis Deeleman-Reinhold 2001" docType="treatment" docVersion="2" lastPageNumber="73" masterDocId="FFC3FF9FFFB7FFDBFFEA3E15FFEC8816" masterDocTitle="Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species" masterLastPageNumber="73" masterPageNumber="55" pageNumber="70" updateTime="1694071302809" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4149DF8599E3F49AADEA4E4AC9FD012C">Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species</mods:title>
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<taxonomicName id="4C534D72FFB8FFD4FF7D388AFCCB8EAF" ID-CoL="7F64F" ID-ENA="1956988" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[151,807,1695,1722]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="70" phylum="Arthropoda" rank="species" species="kinabaluensis">
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<emphasis id="B927EAE3FFB8FFD4FF7D388AFE328EAC" bold="true" box="[151,478,1695,1722]" italics="true" pageId="15" pageNumber="70">Utivarachna kinabaluensis</emphasis>
<bibRefCitation id="EFC24B00FFB8FFD4FE0F388AFCCB8EAF" author="Deeleman-Reinhold, C. L." box="[485,807,1695,1721]" pageId="15" pageNumber="70" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
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<figureCitation id="13682A74FFB8FFD4FF7D38D6FEF88ECB" box="[151,276,1731,1757]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStart-3="FIGURE 27" captionStartId-0="16.[151,248,1037,1061]" captionStartId-1="16.[151,250,1623,1647]" captionStartId-2="17.[151,246,1110,1134]" captionStartId-3="17.[151,250,1503,1527]" captionTargetBox-0="[168,1436,309,1013]" captionTargetBox-1="[151,1435,1121,1599]" captionTargetBox-2="[155,1436,381,1087]" captionTargetBox-3="[151,1435,1195,1480]" captionTargetId-0="figure-91@16.[151,1436,309,1013]" captionTargetId-1="figure-131@16.[151,1435,1121,1599]" captionTargetId-2="figure-97@17.[151,1436,381,1087]" captionTargetId-3="figure-137@17.[151,1435,1195,1480]" captionTargetPageId-0="16" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="17" captionText-0="FIGURE 24. Utivarachna kinabaluensis Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15219).A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." captionText-1="FIGURE 25. Utivarachna kinabaluensis, male, holotype (RNMH.ARA.15219). A right palp, retrolateral view; B same, ventral view; C same, prolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." captionText-2="FIGURE 26. Utivarachna kinabaluensis Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15244). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." captionText-3="FIGURE 27. Utivarachna kinabaluensis, female, paratype (RNMH.ARA.15244). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.8324318" figureDoi-1="http://doi.org/10.5281/zenodo.8324320" figureDoi-2="http://doi.org/10.5281/zenodo.8324322" figureDoi-3="http://doi.org/10.5281/zenodo.8324326" httpUri-0="https://zenodo.org/record/8324318/files/figure.png" httpUri-1="https://zenodo.org/record/8324320/files/figure.png" httpUri-2="https://zenodo.org/record/8324322/files/figure.png" httpUri-3="https://zenodo.org/record/8324326/files/figure.png" pageId="15" pageNumber="70">Figs 2427</figureCitation>
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<taxonomicName id="4C534D72FFB8FFD4FF7D391DFD1C8F36" ID-CoL="7F64F" authority="Deeleman-Reinhold, 2001: 381" authorityName="Deeleman-Reinhold" authorityPageNumber="381" authorityYear="2001" box="[151,752,1800,1824]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="70" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFB8FFD4FF7D391DFE458F36" box="[151,425,1800,1824]" italics="true" pageId="15" pageNumber="70">Utivarachna kinabaluensis</emphasis>
<bibRefCitation id="EFC24B00FFB8FFD4FE45391DFD1C8F36" author="Deeleman-Reinhold, C. L." box="[431,752,1800,1824]" pageId="15" pageNumber="70" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001: 381</bibRefCitation>
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, figs 593597.
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<emphasis id="B927EAE3FFB8FFD4FF7D395BFE2C8F7E" bold="true" box="[151,448,1870,1896]" pageId="15" pageNumber="70">Type material examined.</emphasis>
<materialsCitation id="3B3B3CACFFB8FFD4FE2D395BFC088F9A" collectingDate="1980-07-21" collectingDateMax="1980-07-27" collectingDateMin="1980-07-21" collectionCode="RMNH, R" collectorName="N. Borneo &amp; Kinabalu National Park, HQ &amp; Coll. Deeleman &amp; Deeleman" elevation="1550" location="Kinabalu National Park" pageId="15" pageNumber="70" specimenCount="1" typeStatus="holotype">
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(
<collectionCode id="ED42AE34FFB8FFD4FDB5395BFD548F7E" box="[607,696,1870,1896]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="70" type="Museum">RMNH</collectionCode>
.ARA. 15219;
<figureCitation id="13682A74FFB8FFD4FCB0395BFC538F7E" box="[858,959,1870,1896]" captionStart="FIGURE 24" captionStartId="16.[151,248,1037,1061]" captionTargetBox="[168,1436,309,1013]" captionTargetId="figure-91@16.[151,1436,309,1013]" captionTargetPageId="16" captionText="FIGURE 24. Utivarachna kinabaluensis Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15219).A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324318" httpUri="https://zenodo.org/record/8324318/files/figure.png" pageId="15" pageNumber="70">Fig. 24E</figureCitation>
),
<collectorName id="26A65327FFB8FFD4FC38395AFBA68F7F" box="[978,1098,1871,1897]" pageId="15" pageNumber="70">N. Borneo</collectorName>
,
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</quantity>
,
<date id="FFED1031FFB8FFD4FECD3967FE0C8F9A" box="[295,480,1906,1932]" pageId="15" pageNumber="70" value="1980-07-21" valueMax="1980-07-27" valueMin="1980-07-21">
<collectingDate id="EFA9E9D9FFB8FFD4FECD3967FE0C8F9A" box="[295,480,1906,1932]" pageId="15" pageNumber="70" value="1980-07-21" valueMax="1980-07-27" valueMin="1980-07-21">2127.VII.1980</collectingDate>
</date>
.,
<collectorName id="26A65327FFB8FFD4FE1A3967FD488F9A" box="[496,676,1906,1932]" pageId="15" pageNumber="70">Coll. Deeleman</collectorName>
, leg. C.L. &amp; P.
<collectionCode id="ED42AE34FFB8FFD4FCA43966FC8F8F9B" box="[846,867,1907,1933]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="70">R</collectionCode>
.
<collectorName id="26A65327FFB8FFD4FC853967FC088F9A" box="[879,996,1906,1932]" pageId="15" pageNumber="70">Deeleman</collectorName>
</materialsCitation>
.
<materialsCitation id="3B3B3CACFFB8FFD4FC1A3967FE6E8FA6" collectingDate="1980-07-21" collectingDateMax="1980-07-27" collectingDateMin="1980-07-21" collectionCode="RMNH" collectorName="N. Borneo &amp; Kinabalu National Park, HQ &amp; Coll. Deeleman &amp; Deeleman" elevation="1550" location="Kinabalu National Park" pageId="15" pageNumber="70" specimenCount="1" typeStatus="paratype">
<emphasis id="B927EAE3FFB8FFD4FC1A3967FBB18F9A" bold="true" box="[1008,1117,1906,1932]" pageId="15" pageNumber="70">
<typeStatus id="54E88853FFB8FFD4FC1A3967FBB18F9A" box="[1008,1117,1906,1932]" pageId="15" pageNumber="70">Paratype</typeStatus>
</emphasis>
: 1 ♁ (
<collectionCode id="ED42AE34FFB8FFD4FB423967FAED8F9A" box="[1192,1281,1906,1932]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="70" type="Museum">RMNH</collectionCode>
.ARA. 15244;
<figureCitation id="13682A74FFB8FFD4FF7D3983FF108FA6" box="[151,252,1942,1968]" captionStart="FIGURE 26" captionStartId="17.[151,246,1110,1134]" captionTargetBox="[155,1436,381,1087]" captionTargetId="figure-97@17.[151,1436,381,1087]" captionTargetPageId="17" captionText="FIGURE 26. Utivarachna kinabaluensis Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15244). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324322" httpUri="https://zenodo.org/record/8324322/files/figure.png" pageId="15" pageNumber="70">Fig. 26E</figureCitation>
), same data
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFB8FFCBFF2D39AFFC32890B" lastPageId="16" lastPageNumber="71" pageId="15" pageNumber="70" type="diagnosis">
<paragraph id="8BEC36F1FFB8FFCBFF2D39AFFC32890B" blockId="15.[151,1436,1870,2041]" lastBlockId="16.[151,1437,151,285]" lastPageId="16" lastPageNumber="71" pageId="15" pageNumber="70">
<emphasis id="B927EAE3FFB8FFD4FF2D39AFFEAE8FC2" bold="true" box="[199,322,1978,2004]" pageId="15" pageNumber="70">Diagnosis.</emphasis>
<taxonomicName id="4C534D72FFB8FFD4FEA139AFFD908FC2" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[331,636,1978,2004]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="70" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFB8FFD4FEA139AFFD908FC2" box="[331,636,1978,2004]" italics="true" pageId="15" pageNumber="70">Utivarachna kinabaluensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C534D72FFB8FFD4FCF339AEFB008FC2" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[793,1260,1978,2004]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="70" phylum="Arthropoda" rank="species" species="bucculenta">
<emphasis id="B927EAE3FFB8FFD4FCF339AEFC558FC2" box="[793,953,1978,2004]" italics="true" pageId="15" pageNumber="70">U. bucculenta</emphasis>
<bibRefCitation id="EFC24B00FFB8FFD4FC2839AFFB008FC2" author="Deeleman-Reinhold, C. L." box="[962,1260,1978,2004]" pageId="15" pageNumber="70" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C534D72FFB8FFD4FAC239AEFE628FEE" authority="Li, Zhang &amp; Yu, 2022" authorityName="Li, Zhang &amp; Yu" authorityYear="2022" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="70" phylum="Arthropoda" rank="species" species="fanjing">
<emphasis id="B927EAE3FFB8FFD4FAC239AEFA708FC2" box="[1320,1436,1978,2004]" italics="true" pageId="15" pageNumber="70">U. fanjing</emphasis>
<bibRefCitation id="EFC24B00FFB8FFD4FF7D39CBFE628FEE" author="Li, H. &amp; He, Y. &amp; Zhang, J. &amp; Yu, H." box="[151,398,2014,2041]" pageId="15" pageNumber="70" pagination="587 - 594" refId="ref9077" refString="Li, H., He, Y., Zhang, J. &amp; Yu, H. (2022) A new species of Utivarachna Kishida, 1940 from Fanjing Mountain Nature Reserve, Guizhou, China (Araneae: Trachelidae). Zootaxa, 5094 (4), 587 - 594. https: // doi. org / 10.11646 / zootaxa. 5094.4.4" type="journal article" year="2022">
Li, Zhang &amp; Yu,
<quantity id="4CAB9B14FFB8FFD4FEBC39CBFE628FEE" box="[342,398,2014,2040]" metricMagnitude="1" metricUnit="m" metricValue="5.135879999999999" pageId="15" pageNumber="70" unit="in" value="2022.0">2022</quantity>
</bibRefCitation>
</taxonomicName>
in having an apical RTA claw in males, and in copulatory organs. Regarding males, with the palp in retrolateral view, the RTA claw extends ventrally with a curved tip in
<taxonomicName id="4C534D72FFA7FFCBFBEA3E82FB5188A7" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[1024,1213,151,177]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFA7FFCBFBEA3E82FB5188A7" box="[1024,1213,151,177]" italics="true" pageId="16" pageNumber="71">U. kinabaluensis</emphasis>
</taxonomicName>
, whereas it extends anteriorly and is S-curved in
<taxonomicName id="4C534D72FFA7FFCBFE083EAEFD6C88C3" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[482,640,187,213]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="bucculenta">
<emphasis id="B927EAE3FFA7FFCBFE083EAEFD6C88C3" box="[482,640,187,213]" italics="true" pageId="16" pageNumber="71">U. bucculenta</emphasis>
</taxonomicName>
and extends dorsally without an apical curve in
<taxonomicName id="4C534D72FFA7FFCBFB4C3EAEFAF488C3" authorityName="Li, Zhang &amp; Yu" authorityYear="2022" box="[1190,1304,187,213]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="fanjing">
<emphasis id="B927EAE3FFA7FFCBFB4C3EAEFB5688C2" box="[1190,1210,187,212]" italics="true" pageId="16" pageNumber="71">U</emphasis>
.
<emphasis id="B927EAE3FFA7FFCBFB233EAEFAF488C3" box="[1225,1304,187,213]" italics="true" pageId="16" pageNumber="71">fanjing</emphasis>
</taxonomicName>
. Regarding females, the bursae are large and elongate oval, tapering terminally in
<taxonomicName id="4C534D72FFA7FFCBFC4F3ECAFB8D88EF" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[933,1121,223,249]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFA7FFCBFC4F3ECAFB8D88EF" box="[933,1121,223,249]" italics="true" pageId="16" pageNumber="71">U. kinabaluensis</emphasis>
</taxonomicName>
, whereas they are small and short tube-like in
<taxonomicName id="4C534D72FFA7FFCBFEB73F16FE17890B" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[349,507,259,285]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="bucculenta">
<emphasis id="B927EAE3FFA7FFCBFEB73F16FE17890B" box="[349,507,259,285]" italics="true" pageId="16" pageNumber="71">U. bucculenta</emphasis>
</taxonomicName>
, and large and long tube-like in
<taxonomicName id="4C534D72FFA7FFCBFC8C3F16FC3B890B" authorityName="Li, Zhang &amp; Yu" authorityYear="2022" box="[870,983,259,285]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="fanjing">
<emphasis id="B927EAE3FFA7FFCBFC8C3F16FC3B890B" box="[870,983,259,285]" italics="true" pageId="16" pageNumber="71">U. fanjing</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF2C6679FFA7FFCBFF7D3A18FA7D8C5E" ID-DOI="http://doi.org/10.5281/zenodo.8324318" ID-Zenodo-Dep="8324318" httpUri="https://zenodo.org/record/8324318/files/figure.png" pageId="16" pageNumber="71" startId="16.[151,248,1037,1061]" targetBox="[168,1436,309,1013]" targetPageId="16" targetType="figure">
<paragraph id="8BEC36F1FFA7FFCBFF7D3A18FA7D8C5E" blockId="16.[151,1436,1037,1097]" pageId="16" pageNumber="71">
<emphasis id="B927EAE3FFA7FFCBFF7D3A18FEF08C33" bold="true" box="[151,284,1037,1061]" pageId="16" pageNumber="71">FIGURE 24.</emphasis>
<taxonomicName id="4C534D72FFA7FFCBFECA3A18FCDA8C33" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[288,822,1037,1061]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFA7FFCBFECA3A18FDC78C33" box="[288,555,1037,1061]" italics="true" pageId="16" pageNumber="71">Utivarachna kinabaluensis</emphasis>
<bibRefCitation id="EFC24B00FFA7FFCBFDDA3A18FCDA8C33" author="Deeleman-Reinhold, C. L." box="[560,822,1037,1061]" pageId="16" pageNumber="71" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
, male, holotype (RNMH.ARA.15219).
<emphasis id="B927EAE3FFA7FFCBFB503A18FB208C33" bold="true" box="[1210,1228,1037,1061]" pageId="16" pageNumber="71">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFA7FFCBFF7D3A24FF448C5F" bold="true" box="[151,168,1073,1097]" pageId="16" pageNumber="71">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFA7FFCBFE9C3A24FE648C5F" bold="true" box="[374,392,1073,1097]" pageId="16" pageNumber="71">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFA7FFCBFCDF3A24FCAB8C5F" bold="true" box="[821,839,1073,1097]" pageId="16" pageNumber="71">D</emphasis>
leg I, left, prolateral view;
<emphasis id="B927EAE3FFA7FFCBFBBC3A24FB8B8C5F" bold="true" box="[1110,1127,1073,1097]" pageId="16" pageNumber="71">E</emphasis>
label. Scale bars AD: 1 mm.
</paragraph>
</caption>
<caption id="DF2C6679FFA7FFCBFF7D3842FA9F8E85" ID-DOI="http://doi.org/10.5281/zenodo.8324320" ID-Zenodo-Dep="8324320" httpUri="https://zenodo.org/record/8324320/files/figure.png" pageId="16" pageNumber="71" startId="16.[151,250,1623,1647]" targetBox="[151,1435,1121,1599]" targetPageId="16" targetType="figure">
<paragraph id="8BEC36F1FFA7FFCBFF7D3842FA9F8E85" blockId="16.[151,1436,1623,1683]" pageId="16" pageNumber="71">
<emphasis id="B927EAE3FFA7FFCBFF7D3842FEF38E79" bold="true" box="[151,287,1623,1647]" pageId="16" pageNumber="71">FIGURE 25.</emphasis>
<taxonomicName id="4C534D72FFA7FFCBFECF3842FDD98E79" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[293,565,1623,1647]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFA7FFCBFECF3842FDD98E79" box="[293,565,1623,1647]" italics="true" pageId="16" pageNumber="71">Utivarachna kinabaluensis</emphasis>
</taxonomicName>
, male, holotype (RNMH.ARA.15219).
<emphasis id="B927EAE3FFA7FFCBFC223842FC368E79" bold="true" box="[968,986,1623,1647]" pageId="16" pageNumber="71">A</emphasis>
right palp, retrolateral view;
<emphasis id="B927EAE3FFA7FFCBFB153842FAFC8E79" bold="true" box="[1279,1296,1623,1647]" pageId="16" pageNumber="71">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFA7FFCBFF3C386EFF048E85" bold="true" box="[214,232,1659,1683]" pageId="16" pageNumber="71">C</emphasis>
same, prolateral view;
<emphasis id="B927EAE3FFA7FFCBFE3F386EFE0B8E85" bold="true" box="[469,487,1659,1683]" pageId="16" pageNumber="71">D</emphasis>
same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum.
</paragraph>
</caption>
<subSubSection id="C349657AFFA7FFC9FF2D38CAFC6788EF" lastPageId="18" lastPageNumber="73" pageId="16" pageNumber="71" type="description">
<paragraph id="8BEC36F1FFA7FFCBFF2D38CAFCF98FE0" blockId="16.[151,1437,1759,2038]" pageId="16" pageNumber="71">
<emphasis id="B927EAE3FFA7FFCBFF2D38CAFE9A8EEF" bold="true" box="[199,374,1759,1785]" pageId="16" pageNumber="71">Measurements</emphasis>
(
<typeStatus id="54E88853FFA7FFCBFE6F38CAFE058EEF" box="[389,489,1759,1785]" pageId="16" pageNumber="71">holotype</typeStatus>
♁/
<typeStatus id="54E88853FFA7FFCBFDEC38F5FD818EED" box="[518,621,1760,1787]" pageId="16" pageNumber="71">paratype</typeStatus>
<specimenCount id="9D55FD78FFA7FFCBFD9F38CAFD658EEF" box="[629,649,1759,1785]" pageId="16" pageNumber="71" type="female"></specimenCount>
). Carapace length 3.15/2.90; cephalic width 0.91/1.72; thoracic width 2.35/2.13. Diameter of eyes: AME 0.18/0.17; ALE 0.20/0.18; PME 0.15/0.13; PLE 0.18/0.13. Interdistances between eyes: AMEAME 0.05/0.06; AMEALE 0.13/0.15; ALEALE 0.67/0.67; PMEPME 0.22/0.25; PME PLE 0.33/0.33; PLEPLE 0.90/1.12; ALEPLE 0.16/0.17; AMEPME 0.07/0.06. MOA: length 0.35/0.30; anterior width 0.38/0.40; posterior width 0.51/0.52. Clypeus height 0.42/0.41. Abdomen length 2.85/3.03; width 2.05/2.06. Legs: I 8.62 (2.46, 0.83, 2.20, 2.06, 1.07)/7.30 (2.16, 0.75, 1.77, 1.65, 0.97); II 7.90 (2.33, 0.87, 1.90, 1.80, 1.00)/6.41 (1.87, 0.75, 1.50, 1.47, 0.82); III 5.66 (1.70, 0.65, 1.22, 1.46, 0.63)/4.94 (1.45, 0.67, 1.00, 1.25, 0.57); IV 7.97 (2.30, 0.75, 1.85, 2.17, 0.90)/6.65 (1.86, 0.65, 1.52, 1.87, 0.75).
</paragraph>
<paragraph id="8BEC36F1FFA6FFCAFF2D3E82FEA38973" blockId="17.[151,1436,151,357]" pageId="17" pageNumber="72">
<emphasis id="B927EAE3FFA6FFCAFF2D3E82FEE888A7" bold="true" box="[199,260,151,177]" pageId="17" pageNumber="72">Male</emphasis>
(
<figureCitation id="13682A74FFA6FFCAFEFD3E82FE4A88A7" box="[279,422,151,177]" captionStart="FIGURE 24" captionStartId="16.[151,248,1037,1061]" captionTargetBox="[168,1436,309,1013]" captionTargetId="figure-91@16.[151,1436,309,1013]" captionTargetPageId="16" captionText="FIGURE 24. Utivarachna kinabaluensis Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15219).A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324318" httpUri="https://zenodo.org/record/8324318/files/figure.png" pageId="17" pageNumber="72">Fig. 24AD</figureCitation>
). Carapace entirely granulated, dorsally oval; AER almost straight in dorsal view; PER recurved in dorsal view; lateral margins of carapace strongly constricted between cephalic and thoracic parts; lateral margins of thoracic part weakly undulating. Chelicerae sparsely granulated, with five prolateral and five retrolateral teeth. Clypeus with small wedge-shaped projection between chelicerae. Pedicel short.Abdomen oval; entire dorsum and epigastric area sclerotized. Legs granulated; legs I and II elongated, with small cusps located ventrally from patella to tarsus.
</paragraph>
<caption id="DF2C6679FFA6FFCAFF7D3A43FA948C84" ID-DOI="http://doi.org/10.5281/zenodo.8324322" ID-Zenodo-Dep="8324322" httpUri="https://zenodo.org/record/8324322/files/figure.png" pageId="17" pageNumber="72" startId="17.[151,246,1110,1134]" targetBox="[155,1436,381,1087]" targetPageId="17" targetType="figure">
<paragraph id="8BEC36F1FFA6FFCAFF7D3A43FA948C84" blockId="17.[151,1436,1110,1171]" pageId="17" pageNumber="72">
<emphasis id="B927EAE3FFA6FFCAFF7D3A43FEF68C78" bold="true" box="[151,282,1110,1134]" pageId="17" pageNumber="72">FIGURE 26.</emphasis>
<taxonomicName id="4C534D72FFA6FFCAFEF53A42FCC08C79" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[287,812,1111,1135]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="72" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFA6FFCAFEF53A42FDC98C79" box="[287,549,1111,1135]" italics="true" pageId="17" pageNumber="72">Utivarachna kinabaluensis</emphasis>
<bibRefCitation id="EFC24B00FFA6FFCAFDC03A42FCC08C79" author="Deeleman-Reinhold, C. L." box="[554,812,1111,1135]" pageId="17" pageNumber="72" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
, female, paratype (RNMH.ARA.15244).
<emphasis id="B927EAE3FFA6FFCAFB573A43FB238C78" bold="true" box="[1213,1231,1110,1134]" pageId="17" pageNumber="72">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFA6FFCAFF7D3A6EFF4B8C85" bold="true" box="[151,167,1147,1171]" pageId="17" pageNumber="72">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFA6FFCAFE9B3A6FFE6F8C84" bold="true" box="[369,387,1146,1170]" pageId="17" pageNumber="72">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFA6FFCAFCCD3A6EFCD58C85" bold="true" box="[807,825,1147,1171]" pageId="17" pageNumber="72">D</emphasis>
leg I, left, prolateral view;
<emphasis id="B927EAE3FFA6FFCAFBA83A6EFBBE8C85" bold="true" box="[1090,1106,1147,1171]" pageId="17" pageNumber="72">E</emphasis>
label. Scale bars AD: 1 mm.
</paragraph>
</caption>
<caption id="DF2C6679FFA6FFCAFF7D3BCAFB088E0A" ID-DOI="http://doi.org/10.5281/zenodo.8324326" ID-Zenodo-Dep="8324326" httpUri="https://zenodo.org/record/8324326/files/figure.png" pageId="17" pageNumber="72" startId="17.[151,250,1503,1527]" targetBox="[151,1435,1195,1480]" targetPageId="17" targetType="figure">
<paragraph id="8BEC36F1FFA6FFCAFF7D3BCAFB088E0A" blockId="17.[151,1436,1503,1564]" pageId="17" pageNumber="72">
<emphasis id="B927EAE3FFA6FFCAFF7D3BCAFEF68DEE" bold="true" box="[151,282,1503,1528]" pageId="17" pageNumber="72">FIGURE 27</emphasis>
.
<taxonomicName id="4C534D72FFA6FFCAFECD3BF5FDD58DEE" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[295,569,1504,1528]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="72" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFA6FFCAFECD3BF5FDD58DEE" box="[295,569,1504,1528]" italics="true" pageId="17" pageNumber="72">Utivarachna kinabaluensis</emphasis>
</taxonomicName>
, female, paratype (RNMH.ARA.15244).
<emphasis id="B927EAE3FFA6FFCAFC093BCAFC198DE1" bold="true" box="[995,1013,1503,1527]" pageId="17" pageNumber="72">A</emphasis>
epigyne, ventral view;
<emphasis id="B927EAE3FFA6FFCAFB0D3BF5FB148DEE" bold="true" box="[1255,1272,1504,1528]" pageId="17" pageNumber="72">B</emphasis>
internal organs, ventral view;
<emphasis id="B927EAE3FFA6FFCAFEC83816FED88E0D" bold="true" box="[290,308,1539,1563]" pageId="17" pageNumber="72">C</emphasis>
same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca.
</paragraph>
</caption>
<paragraph id="8BEC36F1FFA6FFCAFF2D387BFC1A8EE2" blockId="17.[151,1437,1646,2032]" pageId="17" pageNumber="72">
Palp (
<figureCitation id="13682A74FFA6FFCAFEE0387BFE7B8E9F" box="[266,407,1646,1673]" captionStart="FIGURE 25" captionStartId="16.[151,250,1623,1647]" captionTargetBox="[151,1435,1121,1599]" captionTargetId="figure-131@16.[151,1435,1121,1599]" captionTargetPageId="16" captionText="FIGURE 25. Utivarachna kinabaluensis, male, holotype (RNMH.ARA.15219). A right palp, retrolateral view; B same, ventral view; C same, prolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." figureDoi="http://doi.org/10.5281/zenodo.8324320" httpUri="https://zenodo.org/record/8324320/files/figure.png" pageId="17" pageNumber="72">Fig. 25AD</figureCitation>
). Cymbium longer than wide, pear-shaped, with convex retrolateral margin. Bulb inverted pear-shaped; prolateral half weakly sclerotized with subtegulum. Embolus coils stored between cymbium and bulb. Sperm duct U-curved in posterior part of bulb, running retrolaterally to dorsal side with weak curve. RTA strongly extended, apically developed in semi-transparent plate with long apical claw.
</paragraph>
<paragraph id="8BEC36F1FFA6FFCAFF2D38EBFB7B8F76" blockId="17.[151,1437,1646,2032]" pageId="17" pageNumber="72">
Coloration and setation (
<figureCitation id="13682A74FFA6FFCAFE3538EBFD808F0E" box="[479,620,1790,1816]" captionStart="FIGURE 24" captionStartId="16.[151,248,1037,1061]" captionTargetBox="[168,1436,309,1013]" captionTargetId="figure-91@16.[151,1436,309,1013]" captionTargetPageId="16" captionText="FIGURE 24. Utivarachna kinabaluensis Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15219).A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324318" httpUri="https://zenodo.org/record/8324318/files/figure.png" pageId="17" pageNumber="72">Fig. 24AD</figureCitation>
). Carapace dark brown, covered with fine setae. Chelicerae dark brown, covered with fine setae; prolateral margin with long setae. Abdomen covered with fine setae; sclerotized dorsum dark brown, and epigastric area brown. Legs brown to light brown, covered with fine setae.
</paragraph>
<paragraph id="8BEC36F1FFA6FFCAFF2D397FFE308FBE" blockId="17.[151,1437,1646,2032]" pageId="17" pageNumber="72">
<emphasis id="B927EAE3FFA6FFCAFF2D397FFEF28F92" bold="true" box="[199,286,1898,1924]" pageId="17" pageNumber="72">Female</emphasis>
(
<figureCitation id="13682A74FFA6FFCAFEC5397FFE508F92" box="[303,444,1898,1924]" captionStart="FIGURE 26" captionStartId="17.[151,246,1110,1134]" captionTargetBox="[155,1436,381,1087]" captionTargetId="figure-97@17.[151,1436,381,1087]" captionTargetPageId="17" captionText="FIGURE 26. Utivarachna kinabaluensis Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15244). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324322" httpUri="https://zenodo.org/record/8324322/files/figure.png" pageId="17" pageNumber="72">Fig. 26AD</figureCitation>
). Somatic characteristics almost same as in male. Legs less granulated, without distinct ventral cusps in legs I and II.
</paragraph>
<paragraph id="8BEC36F1FFA6FFC9FF2D39A7FE0288C3" blockId="17.[151,1437,1646,2032]" lastBlockId="18.[151,1436,151,393]" lastPageId="18" lastPageNumber="73" pageId="17" pageNumber="72">
Copulatory organs (
<figureCitation id="13682A74FFA6FFCAFE4439A7FDD08FDA" box="[430,572,1970,1996]" captionStart="FIGURE 27" captionStartId="17.[151,250,1503,1527]" captionTargetBox="[151,1435,1195,1480]" captionTargetId="figure-137@17.[151,1435,1195,1480]" captionTargetPageId="17" captionText="FIGURE 27. Utivarachna kinabaluensis, female, paratype (RNMH.ARA.15244). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8324326" httpUri="https://zenodo.org/record/8324326/files/figure.png" pageId="17" pageNumber="72">Fig. 27AC</figureCitation>
). Copulatory atrium almost as long as wide. Copulatory openings located at anterior corner of atrium; copulatory ducts extending towards anterior, with strong curvature and then helical coiling. Bursae inverted pear-shaped, extending posteriorly, tapering. Spermathecae almost oval. Fertilization ducts straight, extending posteriorly.
</paragraph>
<paragraph id="8BEC36F1FFA5FFC9FF2D3ECAFC6788EF" blockId="18.[151,1436,151,393]" box="[199,907,223,249]" pageId="18" pageNumber="73">
Coloration and setation (
<figureCitation id="13682A74FFA5FFC9FE363ECAFD8488EF" box="[476,616,223,249]" captionStart="FIGURE 26" captionStartId="17.[151,246,1110,1134]" captionTargetBox="[155,1436,381,1087]" captionTargetId="figure-97@17.[151,1436,381,1087]" captionTargetPageId="17" captionText="FIGURE 26. Utivarachna kinabaluensis Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15244). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars AD: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8324322" httpUri="https://zenodo.org/record/8324322/files/figure.png" pageId="18" pageNumber="73">Fig. 26AD</figureCitation>
). Almost same as in male.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFA5FFC9FF2D3F17FCA7890B" box="[199,843,258,285]" pageId="18" pageNumber="73" type="distribution">
<paragraph id="8BEC36F1FFA5FFC9FF2D3F17FCA7890B" blockId="18.[151,1436,151,393]" box="[199,843,258,285]" pageId="18" pageNumber="73">
<emphasis id="B927EAE3FFA5FFC9FF2D3F17FE8D890A" bold="true" box="[199,353,258,284]" pageId="18" pageNumber="73">Distribution.</emphasis>
<collectingRegion id="4997F813FFA5FFC9FE8D3F16FE47890B" box="[359,427,259,285]" country="Malaysia" name="Sabah" pageId="18" pageNumber="73">Sabah</collectingRegion>
(known only from the
<typeStatus id="54E88853FFA5FFC9FD453F16FD33890B" box="[687,735,259,285]" pageId="18" pageNumber="73">type</typeStatus>
locality).
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFA5FFC9FF2D3F32FB6F899F" pageId="18" pageNumber="73" type="discussion">
<paragraph id="8BEC36F1FFA5FFC9FF2D3F32FB6F899F" blockId="18.[151,1436,151,393]" pageId="18" pageNumber="73">
<emphasis id="B927EAE3FFA5FFC9FF2D3F32FED78957" bold="true" box="[199,315,295,321]" pageId="18" pageNumber="73">Remarks.</emphasis>
Although
<taxonomicName id="4C534D72FFA5FFC9FE5D3F32FD9B8957" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[439,631,295,321]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFA5FFC9FE5D3F32FE278956" box="[439,459,295,320]" italics="true" pageId="18" pageNumber="73">U</emphasis>
.
<emphasis id="B927EAE3FFA5FFC9FE373F32FD9B8957" box="[477,631,295,321]" italics="true" pageId="18" pageNumber="73">kinabaluensis</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="4C534D72FFA5FFC9FCC43F32FC248957" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[814,968,295,321]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B927EAE3FFA5FFC9FCC43F32FC248957" box="[814,968,295,321]" italics="true" pageId="18" pageNumber="73">kinabaluensis</emphasis>
</taxonomicName>
-group, this species is very similar to
<taxonomicName id="4C534D72FFA5FFC9FA6B3F32FEE98973" authorityName="Dankittipakul, Tavano &amp; Singtripop" authorityYear="2011" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="galyaniae">
<emphasis id="B927EAE3FFA5FFC9FA6B3F32FEE98973" italics="true" pageId="18" pageNumber="73">U. galyaniae</emphasis>
</taxonomicName>
, which belongs to the
<taxonomicName id="4C534D72FFA5FFC9FDEF3F5EFD678973" authorityName="Kishida" authorityYear="1940" box="[517,651,331,357]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="fukasawana">
<emphasis id="B927EAE3FFA5FFC9FDEF3F5EFD678973" box="[517,651,331,357]" italics="true" pageId="18" pageNumber="73">fukasawana</emphasis>
</taxonomicName>
-group. The possibility of a synonymous relationship between the two species should be considered in a future study based on the
<typeStatus id="54E88853FFA5FFC9FCD93F7AFC8F899F" box="[819,867,367,393]" pageId="18" pageNumber="73">type</typeStatus>
material of
<taxonomicName id="4C534D72FFA5FFC9FC013F7AFB90899F" authorityName="Dankittipakul, Tavano &amp; Singtripop" authorityYear="2011" box="[1003,1148,367,393]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="73" phylum="Arthropoda" rank="species" species="galyaniae">
<emphasis id="B927EAE3FFA5FFC9FC013F7AFB90899F" box="[1003,1148,367,393]" italics="true" pageId="18" pageNumber="73">U. galyaniae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>