treatments-xml/data/03/FA/87/03FA87BBFFD8FFB9D6B93C4B014F0458.xml

<document id="61BF9765B5843B9906613D9AFD3D1B6B" ID-CLB-Dataset="68519" ID-DOI="10.11646/zootaxa.5150.2.5" ID-GBIF-Dataset="e099b088-6bbc-42eb-a57b-84bdfe32f64e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6621301" ID-ZooBank="D15CA296-D186-43EA-90D3-4979596A46F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654625951008" checkinUser="plazi" docAuthor="Rojas-Buffet, Carolina, Bouzan, Rodrigo Salvador, Sierwald, Petra, Brescovit, Antonio Do- Mingos &amp; Simó, Miguel" docDate="2022" docId="03FA87BBFFD8FFB9D6B93C4B014F0458" docLanguage="en" docName="zootaxa.5150.2.5.pdf" docOrigin="Zootaxa 5150 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sandalodesmus joachimadisi Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó 2022, n. sp." docType="treatment" docVersion="5" lastPageNumber="266" masterDocId="FFC3FFC3FFDAFFBFD62E3D250009074D" masterDocTitle="Sandalodesmus joachimadisi n. sp., the first Chelodesmidae member from Uruguay with an updated key to species of the genus (Diplopoda: Polydesmida)" masterLastPageNumber="268" masterPageNumber="260" pageNumber="262" updateTime="1699339059308" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A7FB084C94F3E888E95854A4BCB85D8F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D7CE020645EC8DEC7B57A992F5B0B021">
<mods:title id="45D474DF93E0C0204CDAB59319E0E04C">Sandalodesmus joachimadisi n. sp., the first Chelodesmidae member from Uruguay with an updated key to species of the genus (Diplopoda: Polydesmida)</mods:title>
</mods:titleInfo>
<mods:name id="A4B190BDD984CC9055AA945B5EE02C92" type="personal">
<mods:role id="0CDE34A5589AEB0648D51E965525B851">
<mods:roleTerm id="B0023F5180CA8101F49BE1456C9E5436">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="051F71ECE9AA1EC1CD9C5AD41A1CD3FA">Rojas-Buffet, Carolina</mods:namePart>
<mods:affiliation id="E10DDF5D744568F88EF183E1C4F928F0">Sección Entomología, Facultad de Ciencias, Universidad de la República, Montevideo, Uruguay. &amp; Departamento de Ecología y Biología Evolutiva, Instituto de Investigaciones Biológicas Clemente Estable, Ministerio de Educación y Cultura, Montevideo, Uruguay.</mods:affiliation>
</mods:name>
<mods:name id="1D0D13B180ADD539AB19AEDE60AD201E" type="personal">
<mods:role id="237338806470FAAC23CD371D9A45F1FE">
<mods:roleTerm id="F8A37C6757023B8E412F4CF2CDF3C92E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1CF3ECF2D967B7858AE369EB8D1D0488">Bouzan, Rodrigo Salvador</mods:namePart>
<mods:nameIdentifier id="F21BC39036BAC9E358069D5359A94089" type="ORCID">0000-0002-5331-7031</mods:nameIdentifier>
<mods:affiliation id="2555DF661B59D2424518EE845F7122F1">Laboratório de Coleções Zoológicas, Instituto Butantan, São Paulo, Brazil. &amp; Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; rodrigobouzan @ outlook. com; https: // orcid. org / 0000 - 0002 - 5331 - 7031</mods:affiliation>
<mods:nameIdentifier id="FD4DC72868CEA926E13D87B361167B03" type="email">rodrigobouzan@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name id="517E17405CC2ED94EE82393DC19987D6" type="personal">
<mods:role id="5EAFB1EAD9954BBEA2F72AAD90801D9A">
<mods:roleTerm id="A29C601F1F61F83272CE3BEAC263F1C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="66FED35E8C83DE3A0DD8AE594C218D6D">Sierwald, Petra</mods:namePart>
<mods:nameIdentifier id="AA8721BD0CFA6B5565082ADCAB2D4497" type="ORCID">0000-0003-2592-1298</mods:nameIdentifier>
<mods:affiliation id="A486E6C9B82874EBBEDC1EF481A017D4">Insect Division, Field Museum of Natural History, Chicago, IL. USA. &amp; psierwald @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 2592 - 1298</mods:affiliation>
<mods:nameIdentifier id="36382D22BCD670B51BCAB875510395A9" type="email">psierwald@fieldmuseum.org</mods:nameIdentifier>
</mods:name>
<mods:name id="3B1025E323761B756E1A7C970B0DA7C1" type="personal">
<mods:role id="CF697C2B1F0DB4F272CB944623B9B792">
<mods:roleTerm id="5375C46F9D482B584B646D040C68BD40">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="265B2063868EEB7452D43AC784139AB2">Brescovit, Antonio Do- Mingos</mods:namePart>
<mods:nameIdentifier id="D3D4B199A72E8E6509CC3C8DF4A19C3E" type="ORCID">0000-0002-1511-5324</mods:nameIdentifier>
<mods:affiliation id="B4BEDE338AEB48BD611C62AA29ED1767">Laboratório de Coleções Zoológicas, Instituto Butantan, São Paulo, Brazil. &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
<mods:nameIdentifier id="D16DBC34AC81D22E1B161B030B23E776" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
</mods:name>
<mods:name id="2ABB31653AF47833770094B0A363F700" type="personal">
<mods:role id="6D1A76C55EB53165DC00C1C3B2E31C9A">
<mods:roleTerm id="7726B7FC5902D258C329BB2736025053">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B7A78600FF2F836B27BEFD7CBF1A6AA">Simó, Miguel</mods:namePart>
<mods:nameIdentifier id="7BD1405C9A1ED87CB50FB5EAAAFD8D1C" type="ORCID">0000-0002-9477-5880</mods:nameIdentifier>
<mods:affiliation id="19ADD76524EEE8895EBC6921DC8D7B87">Sección Entomología, Facultad de Ciencias, Universidad de la República, Montevideo, Uruguay. &amp; simo @ fcien. edu. uy; https: // orcid. org / 0000 - 0002 - 9477 - 5880</mods:affiliation>
<mods:nameIdentifier id="BF05F7CA20D33ADF13E24002CF63ACFC" type="email">simo@fcien.edu.uy</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="CFC7FB01F7128347003B65A7EAC079B2">text</mods:typeOfResource>
<mods:relatedItem id="AB1E7641ACDDEE48E1D6B3D3B15D9B3E" type="host">
<mods:titleInfo id="A1D7208FBFDF5B868F949DEA2F7BFB43">
<mods:title id="2D4AA49438841611ECD87AE1E368AC3D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="509F31C96690D8B0CE4361520C395314">
<mods:date id="C46DB9EDFCD0C7A82D9D36284EA19795">2022</mods:date>
<mods:detail id="CAE5C8C8490D0C9DED5163B4EC22291D" type="pubDate">
<mods:number id="512ACD9C1572EC7D5620EC085D0283EE">2022-06-07</mods:number>
</mods:detail>
<mods:detail id="50717EE55918829E6BE02EE7F9B7EB0A" type="volume">
<mods:number id="3810BA373D76C771307FC8C084B9F993">5150</mods:number>
</mods:detail>
<mods:detail id="87A7E35E26C62C83EE946568D1FEC600" type="issue">
<mods:number id="A6E94F125DC0E2AAAF373E8C7D6FD0AE">2</mods:number>
</mods:detail>
<mods:extent id="76CEB6DBB130D28B10EB43219E34DE95" unit="page">
<mods:start id="0E2B0D09E63E6AE7CFD09242A89E7039">260</mods:start>
<mods:end id="91F148C31677AB50717209343D04BC01">268</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1F96EBA22AEE3D40B0E0219575EBF2AD">journal article</mods:classification>
<mods:identifier id="4E6E13617C0100A579E241E3643E1B80" type="CLB-Dataset">68519</mods:identifier>
<mods:identifier id="E2EDE37C3D9B2C627C7DEFE1CC7D0559" type="DOI">10.11646/zootaxa.5150.2.5</mods:identifier>
<mods:identifier id="ED20F3E5C7FD0664A41C098BE417A0E4" type="GBIF-Dataset">e099b088-6bbc-42eb-a57b-84bdfe32f64e</mods:identifier>
<mods:identifier id="B1D33B18FD68423408094693C2639777" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6C329B35052F48373D0FC9AE74E91969" type="Zenodo-Dep">6621301</mods:identifier>
<mods:identifier id="2BD7D2F4551602FD4A6D76192AEFADEB" type="ZooBank">D15CA296-D186-43EA-90D3-4979596A46F0</mods:identifier>
</mods:mods>
<treatment id="03FA87BBFFD8FFB9D6B93C4B014F0458" ID-DOI="http://doi.org/10.5281/zenodo.6621293" ID-GBIF-Taxon="195827685" ID-Zenodo-Dep="6621293" LSID="urn:lsid:plazi:treatment:03FA87BBFFD8FFB9D6B93C4B014F0458" httpUri="http://treatment.plazi.org/id/03FA87BBFFD8FFB9D6B93C4B014F0458" lastPageId="6" lastPageNumber="266" pageId="2" pageNumber="262">
<subSubSection id="C3496526FFD8FFBDD6B93C4B037D06C5" box="[151,884,366,393]" pageId="2" pageNumber="262" type="nomenclature">
<paragraph id="8BEC36ADFFD8FFBDD6B93C4B037D06C5" blockId="2.[151,884,366,429]" box="[151,884,366,393]" pageId="2" pageNumber="262">
<heading id="D0A481C1FFD8FFBDD6B93C4B037D06C5" bold="true" box="[151,884,366,393]" fontSize="11" level="1" pageId="2" pageNumber="262" reason="1">
<emphasis id="B927EABFFFD8FFBDD6B93C4B037D06C5" bold="true" box="[151,884,366,393]" pageId="2" pageNumber="262">
<taxonomicName id="4C534D2EFFD8FFBDD6B93C4B032C06C5" ID-CoL="993J7" authority="Rojas-Buffet &amp; Bouzan" authorityName="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó" authorityYear="2022" box="[151,805,366,393]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="262" phylum="Arthropoda" rank="species" species="joachimadisi" status="sp. nov.">
<emphasis id="B927EABFFFD8FFBDD6B93C4B01FE06C4" bold="true" box="[151,503,366,393]" italics="true" pageId="2" pageNumber="262">Sandalodesmus joachimadisi</emphasis>
Rojas-Buffet &amp; Bouzan
</taxonomicName>
, 
<taxonomicNameLabel id="A21457C4FFD8FFBDD51C3C4B037D06C5" box="[818,884,366,392]" pageId="2" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3496526FFD8FFBDD6B93CB700F106E0" box="[151,248,402,429]" pageId="2" pageNumber="262" type="description">
<paragraph id="8BEC36ADFFD8FFBDD6B93CB700F106E0" blockId="2.[151,884,366,429]" box="[151,248,402,429]" pageId="2" pageNumber="262">
<figureCitation id="13682A28FFD8FFBDD6B93CB700F106E0" box="[151,248,402,429]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1314,1338]" captionStartId-1="3.[151,250,1824,1848]" captionStartId-2="4.[151,250,1347,1371]" captionStartId-3="5.[151,250,697,721]" captionStartId-4="5.[151,250,1944,1968]" captionTargetBox-0="[151,1436,776,1290]" captionTargetBox-1="[269,1317,181,1800]" captionTargetBox-2="[151,1436,181,1323]" captionTargetBox-3="[155,1429,187,670]" captionTargetBox-4="[151,1427,818,1920]" captionTargetId-0="figure-208@2.[151,1436,776,1290]" captionTargetId-1="figure-20@3.[269,1317,181,1800]" captionTargetId-2="figure-17@4.[151,1436,181,1323]" captionTargetId-3="figure-20@5.[151,1436,181,673]" captionTargetId-4="figure-75@5.[151,1436,818,1920]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="5" captionText-0="FIGURE 1. Sandalodesmus joachimadisi n. sp. from Valle del Lunarejo, Rivera, Uruguay. A. Live male. B. Two specimens found under cow dung, one of them inside a nest." captionText-1="FIGURE 2. Somatic characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. B. Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. C. Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. D. Non type ♂ (FCE-Myr 0556), left antenna. E. Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. F. Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. G. Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. H. Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. Abbreviations: L: lingual lobes; M: mentum; O: ozopore; P: projection; S: stipes. Scale bars: 1.0 mm (A–D, G–H), 0.5 mm (E–F)." captionText-2="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." captionText-3="FIGURE 4. Gonopod drawings of Sandalodesmus joachimadisi n. sp., holotype ♂ (FCE-Myr 0664), left gonopod. A. Mesal view. B. Lateral view. C. Ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm." captionText-4="FIGURE 5. Distribution map of Sandalodesmus species." figureDoi-0="http://doi.org/10.5281/zenodo.6621305" figureDoi-1="http://doi.org/10.5281/zenodo.6621307" figureDoi-2="http://doi.org/10.5281/zenodo.6621311" figureDoi-3="http://doi.org/10.5281/zenodo.6621313" figureDoi-4="http://doi.org/10.5281/zenodo.6621315" httpUri-0="https://zenodo.org/record/6621305/files/figure.png" httpUri-1="https://zenodo.org/record/6621307/files/figure.png" httpUri-2="https://zenodo.org/record/6621311/files/figure.png" httpUri-3="https://zenodo.org/record/6621313/files/figure.png" httpUri-4="https://zenodo.org/record/6621315/files/figure.png" pageId="2" pageNumber="262">Figs 1–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3496526FFD8FFBDD6B93CFF058805BD" pageId="2" pageNumber="262" type="diagnosis">
<paragraph id="8BEC36ADFFD8FFBDD6B93CFF023D05C9" blockId="2.[151,1437,474,753]" pageId="2" pageNumber="262">
<emphasis id="B927EABFFFD8FFBDD6B93CFF011B06B9" bold="true" box="[151,274,474,500]" pageId="2" pageNumber="262">Diagnosis.</emphasis>
Males differ from all other species of the genus by the following combination of characters: prefemoral process (PfP) small, Z-shaped, twisted internally and directed parallel to acropodital branch (
<figureCitation id="13682A28FFD8FFBDD2B23CDB04F10555" box="[1180,1272,510,536]" captionStart="FIGURE 3" captionStartId="4.[151,250,1347,1371]" captionTargetBox="[151,1436,181,1323]" captionTargetId="figure-17@4.[151,1436,181,1323]" captionTargetPageId="4" captionText="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="2" pageNumber="262">Figs 3A</figureCitation>
, 
<figureCitation id="13682A28FFD8FFBDD32D3CDB05200555" box="[1283,1321,510,536]" captionStart="FIGURE 4" captionStartId="5.[151,250,697,721]" captionTargetBox="[155,1429,187,670]" captionTargetId="figure-20@5.[151,1436,181,673]" captionTargetPageId="5" captionText="FIGURE 4. Gonopod drawings of Sandalodesmus joachimadisi n. sp., holotype ♂ (FCE-Myr 0664), left gonopod. A. Mesal view. B. Lateral view. C. Ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6621313" httpUri="https://zenodo.org/record/6621313/files/figure.png" pageId="2" pageNumber="262">4A</figureCitation>
), prostatic groove (Pg) sinuous and S-shaped along the acropodite, ending upon an elongated prostatic apex (
<figureCitation id="13682A28FFD8FFBDD2D33F0705550571" box="[1277,1372,546,572]" captionStart="FIGURE 3" captionStartId="4.[151,250,1347,1371]" captionTargetBox="[151,1436,181,1323]" captionTargetId="figure-17@4.[151,1436,181,1323]" captionTargetPageId="4" captionText="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="2" pageNumber="262">Figs 3A</figureCitation>
, 
<figureCitation id="13682A28FFD8FFBDD3443F0705990571" box="[1386,1424,546,572]" captionStart="FIGURE 4" captionStartId="5.[151,250,697,721]" captionTargetBox="[155,1429,187,670]" captionTargetId="figure-20@5.[151,1436,181,673]" captionTargetPageId="5" captionText="FIGURE 4. Gonopod drawings of Sandalodesmus joachimadisi n. sp., holotype ♂ (FCE-Myr 0664), left gonopod. A. Mesal view. B. Lateral view. C. Ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6621313" httpUri="https://zenodo.org/record/6621313/files/figure.png" pageId="2" pageNumber="262">4A</figureCitation>
), acropodite process (Ap) broad and round, elongated, hood-shaped (
<figureCitation id="13682A28FFD8FFBDD58E3F630428052D" box="[928,1057,582,608]" captionStart="FIGURE 3" captionStartId="4.[151,250,1347,1371]" captionTargetBox="[151,1436,181,1323]" captionTargetId="figure-17@4.[151,1436,181,1323]" captionTargetPageId="4" captionText="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="2" pageNumber="262">Figs 3A–C</figureCitation>
, 
<figureCitation id="13682A28FFD8FFBDD21E3F63047F052D" box="[1072,1142,582,608]" captionStart="FIGURE 4" captionStartId="5.[151,250,697,721]" captionTargetBox="[155,1429,187,670]" captionTargetId="figure-20@5.[151,1436,181,673]" captionTargetPageId="5" captionText="FIGURE 4. Gonopod drawings of Sandalodesmus joachimadisi n. sp., holotype ♂ (FCE-Myr 0664), left gonopod. A. Mesal view. B. Lateral view. C. Ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6621313" httpUri="https://zenodo.org/record/6621313/files/figure.png" pageId="2" pageNumber="262">4A–C</figureCitation>
), solenomere (S) apically curved, sickle-shaped (
<figureCitation id="13682A28FFD8FFBDD7B73F4F01FF05C9" box="[409,502,618,644]" captionStart="FIGURE 3" captionStartId="4.[151,250,1347,1371]" captionTargetBox="[151,1436,181,1323]" captionTargetId="figure-17@4.[151,1436,181,1323]" captionTargetPageId="4" captionText="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="2" pageNumber="262">Figs 3A</figureCitation>
, 
<figureCitation id="13682A28FFD8FFBDD42C3F4F022105C9" box="[514,552,618,644]" captionStart="FIGURE 4" captionStartId="5.[151,250,697,721]" captionTargetBox="[155,1429,187,670]" captionTargetId="figure-20@5.[151,1436,181,673]" captionTargetPageId="5" captionText="FIGURE 4. Gonopod drawings of Sandalodesmus joachimadisi n. sp., holotype ♂ (FCE-Myr 0664), left gonopod. A. Mesal view. B. Lateral view. C. Ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6621313" httpUri="https://zenodo.org/record/6621313/files/figure.png" pageId="2" pageNumber="262">4A</figureCitation>
).
</paragraph>
<paragraph id="8BEC36ADFFD8FFBDD6E93FAB058805BD" blockId="2.[151,1437,474,753]" pageId="2" pageNumber="262">
<emphasis id="B927EABFFFD8FFBDD6E93FAB011D05E5" bold="true" box="[199,276,654,680]" pageId="2" pageNumber="262">Name.</emphasis>
The specific name is a patronymic in honor of the German ecologist and entomologist Joachim Ulrich Adis (1950–2007). He visited the Collection of Myriapoda of Facultad de Ciencias, Universidad de la República, 
<collectingRegion id="4997F84FFFD8FFBDD6B93FF3012805BD" box="[151,289,726,752]" country="Uruguay" name="Montevideo" pageId="2" pageNumber="262">Montevideo</collectingRegion>
, 
<collectingCountry id="F344763DFFD8FFBDD7033FF2018605BC" box="[301,399,727,753]" name="Uruguay" pageId="2" pageNumber="262">Uruguay</collectingCountry>
in 2005 and suggested the need for the study of the underexplored Uruguayan 
<taxonomicName id="4C534D2EFFD8FFBDD32D3FF3057405BD" box="[1283,1405,726,752]" class="Diplopoda" kingdom="Animalia" pageId="2" pageNumber="262" phylum="Arthropoda" rank="class">Diplopoda</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF2C6625FFD8FFBDD6B9380702850210" ID-DOI="http://doi.org/10.5281/zenodo.6621305" ID-Zenodo-Dep="6621305" httpUri="https://zenodo.org/record/6621305/files/figure.png" pageId="2" pageNumber="262" startId="2.[151,250,1314,1338]" targetBox="[151,1436,776,1290]" targetPageId="2">
<paragraph id="8BEC36ADFFD8FFBDD6B9380702850210" blockId="2.[151,1437,1314,1374]" pageId="2" pageNumber="262">
<emphasis id="B927EABFFFD8FFBDD6B93807011C0277" bold="true" box="[151,277,1314,1338]" pageId="2" pageNumber="262">FIGURE 1.</emphasis>
<taxonomicName id="4C534D2EFFD8FFBDD7323807024C0277" authority="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó, 2022" authorityName="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó" authorityYear="2022" box="[284,581,1314,1338]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="262" phylum="Arthropoda" rank="species" species="joachimadisi" status="sp. nov.">
<emphasis id="B927EABFFFD8FFBDD7323807024C0277" box="[284,581,1314,1338]" italics="true" pageId="2" pageNumber="262">Sandalodesmus joachimadisi</emphasis>
</taxonomicName>
<emphasis id="B927EABFFFD8FFBDD4623807028E0277" bold="true" box="[588,647,1314,1338]" pageId="2" pageNumber="262">
<taxonomicNameLabel id="A21457C4FFD8FFBDD4623807028E0277" box="[588,647,1314,1338]" pageId="2" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Valle del Lunarejo, Rivera, Uruguay. 
<emphasis id="B927EABFFFD8FFBDD2643807046A0277" bold="true" box="[1098,1123,1314,1338]" pageId="2" pageNumber="262">A.</emphasis>
Live male. 
<emphasis id="B927EABFFFD8FFBDD2CE380704FE0277" bold="true" box="[1248,1271,1314,1338]" pageId="2" pageNumber="262">B.</emphasis>
Two specimens found under cow dung, one of them inside a nest.
</paragraph>
</caption>
<subSubSection id="C3496526FFD8FFBDD6B938A9018A00D3" pageId="2" pageNumber="262" type="materials_examined">
<paragraph id="8BEC36ADFFD8FFBDD6B938A9018802EB" blockId="2.[151,1437,1420,1950]" box="[151,385,1420,1446]" pageId="2" pageNumber="262">
<heading id="D0A481C1FFD8FFBDD6B938A9018802EB" bold="true" box="[151,385,1420,1446]" fontSize="11" level="1" pageId="2" pageNumber="262" reason="1">
<emphasis id="B927EABFFFD8FFBDD6B938A9018802EB" bold="true" box="[151,385,1420,1446]" pageId="2" pageNumber="262">Material examined.</emphasis>
</heading>
</paragraph>
<paragraph id="8BEC36ADFFD8FFBDD6E9389502DB02A3" blockId="2.[151,1437,1420,1950]" pageId="2" pageNumber="262">
<materialsCitation id="3B3B3CF0FFD8FFBDD6E9389502DB02A3" ID-GBIF-Occurrence="3804719301" collectingDate="2018-12-27" collectorName="C. Rojas-Buffet" country="Uruguay" latitude="-31.189917" location="Rivera" longLatPrecision="1" longitude="-55.90117" pageId="2" pageNumber="262" specimenCount="1" specimenCount-male="1" stateProvince="Rivera" typeStatus="holotype">
<emphasis id="B927EABFFFD8FFBDD6E9389501340287" bold="true" box="[199,317,1456,1482]" pageId="2" pageNumber="262">
<typeStatus id="54E8880FFFD8FFBDD6E9389501310287" box="[199,312,1456,1482]" pageId="2" pageNumber="262" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F344763DFFD8FFBDD762389501DD0287" box="[332,468,1456,1482]" name="Uruguay" pageId="2" pageNumber="262">URUGUAY</collectingCountry>
● 
<specimenCount id="9D55FD24FFD8FFBDD42F389402100286" box="[513,537,1457,1483]" pageId="2" pageNumber="262" type="male">♂</specimenCount>
; Valle del Lunarejo, 
<collectingRegion id="4997F84FFFD8FFBDD530389503650287" box="[798,876,1456,1482]" country="Uruguay" name="Rivera" pageId="2" pageNumber="262">Rivera</collectingRegion>
, under cow dung, 
<geoCoordinate id="EE67506AFFD8FFBDD27B389504E10287" box="[1109,1256,1456,1482]" degrees="31" direction="south" minutes="11" orientation="latitude" pageId="2" pageNumber="262" precision="1" seconds="23.7" value="-31.189917">31°11'23.7&quot;S</geoCoordinate>
<geoCoordinate id="EE67506AFFD8FFBDD2D83895059E0286" box="[1270,1431,1456,1483]" degrees="55" direction="west" minutes="54" orientation="longitude" pageId="2" pageNumber="262" precision="1" seconds="04.2" value="-55.90117">55°54'04.2&quot;W</geoCoordinate>
; 
<date id="FFED106DFFD8FFBDD6B938F1012A02A3" box="[151,291,1492,1518]" pageId="2" pageNumber="262" value="2018-12-27">
<collectingDate id="EFA9E985FFD8FFBDD6B938F1012A02A3" box="[151,291,1492,1518]" pageId="2" pageNumber="262" value="2018-12-27">27.XII.2018</collectingDate>
</date>
; 
<collectorName id="26A6537BFFD8FFBDD70138F101E902A3" box="[303,480,1492,1518]" pageId="2" pageNumber="262">C. Rojas-Buffet</collectorName>
leg.; FCE-Myr 0664.
</materialsCitation>
</paragraph>
<paragraph id="8BEC36ADFFD8FFBDD6E938DD042C015F" blockId="2.[151,1437,1420,1950]" box="[199,1061,1527,1555]" pageId="2" pageNumber="262">
<emphasis id="B927EABFFFD8FFBDD6E938DD0141015F" bold="true" box="[199,328,1528,1554]" pageId="2" pageNumber="262">
<typeStatus id="54E8880FFFD8FFBDD6E938DD014A015F" box="[199,323,1528,1554]" pageId="2" pageNumber="262" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="F344763DFFD8FFBDD76138DD01DE015F" box="[335,471,1528,1554]" name="Uruguay" pageId="2" pageNumber="262">URUGUAY</collectingCountry>
● 
<specimenCount id="9D55FD24FFD8FFBDD7D938DD023E015E" box="[503,567,1528,1555]" pageId="2" pageNumber="262" type="female">3 ♀♀</specimenCount>
; same data as for 
<typeStatus id="54E8880FFFD8FFBDD52F38DD036E015F" box="[769,871,1528,1554]" pageId="2" pageNumber="262" type="holotype">holotype</typeStatus>
; FCE-Myr 0664.
</paragraph>
<paragraph id="8BEC36ADFFD8FFBDD6E93B39018A00D3" blockId="2.[151,1437,1420,1950]" pageId="2" pageNumber="262">
<emphasis id="B927EABFFFD8FFBDD6E93B39023B017B" bold="true" box="[199,562,1564,1590]" pageId="2" pageNumber="262">Additional material examined.</emphasis>
All from 
<collectingCountry id="F344763DFFD8FFBDD4B33B39032C017B" box="[669,805,1564,1590]" name="Uruguay" pageId="2" pageNumber="262">URUGUAY</collectingCountry>
● 
<specimenCount id="9D55FD24FFD8FFBDD56C3B390367017A" box="[834,878,1564,1591]" pageId="2" pageNumber="262" type="male">1 ♂</specimenCount>
; 
<collectingRegion id="4997F84FFFD8FFBDD5563B390487017A" box="[888,1166,1564,1591]" country="Uruguay" name="Tacuarembo" pageId="2" pageNumber="262">Tacuarembó Department</collectingRegion>
, 
<collectingRegion id="4997F84FFFD8FFBDD2B63B39052B017B" box="[1176,1314,1564,1590]" country="Uruguay" name="Tacuarembo" pageId="2" pageNumber="262">Tacuarembó</collectingRegion>
city, urban zone, 
<geoCoordinate id="EE67506AFFD8FFBDD6F23B6501790117" box="[220,368,1600,1626]" degrees="31" direction="south" minutes="44" orientation="latitude" pageId="2" pageNumber="262" precision="1" seconds="18.9" value="-31.738585">31°44'18.9&quot;S</geoCoordinate>
<geoCoordinate id="EE67506AFFD8FFBDD7573B6502120116" box="[377,539,1600,1627]" degrees="55" direction="west" minutes="58" orientation="longitude" pageId="2" pageNumber="262" precision="1" seconds="46.7" value="-55.97964">55°58'46.7&quot;W</geoCoordinate>
; 
<date id="FFED106DFFD8FFBDD4063B6502A20117" box="[552,683,1600,1626]" pageId="2" pageNumber="262" value="2013-11-14">14.XI.2013</date>
; A. Laborda leg.; FCE-Myr 0556 ● 
<specimenCount id="9D55FD24FFD8FFBDD2623B6504870116" box="[1100,1166,1600,1627]" pageId="2" pageNumber="262" type="female">2 ♀♀</specimenCount>
; same locality, walking at night, 
<geoCoordinate id="EE67506AFFD8FFBDD6D13B4101AB0133" box="[255,418,1636,1662]" degrees="31" direction="south" minutes="44" orientation="latitude" pageId="2" pageNumber="262" precision="1" seconds="19.63" value="-31.738787">31°44'19.63&quot;S</geoCoordinate>
<geoCoordinate id="EE67506AFFD8FFBDD7853B4102520132" box="[427,603,1636,1663]" degrees="55" direction="west" minutes="58" orientation="longitude" pageId="2" pageNumber="262" precision="1" seconds="46.60" value="-55.97961">55°58'46.60&quot;W</geoCoordinate>
; 
<date id="FFED106DFFD8FFBDD4473B4102FC0133" box="[617,757,1636,1662]" pageId="2" pageNumber="262" value="2018-12-27">27.XII.2018</date>
; D. Hagopián leg.; FCE-Myr 0665 ● 
<specimenCount id="9D55FD24FFD8FFBDD2823B4104F90132" box="[1196,1264,1636,1663]" pageId="2" pageNumber="262" type="male">4 ♂♂</specimenCount>
, 
<specimenCount id="9D55FD24FFD8FFBDD2D03B4105480132" box="[1278,1345,1636,1663]" pageId="2" pageNumber="262" type="female">3 ♀♀</specimenCount>
; 
<collectingRegion id="4997F84FFFD8FFBDD37C3B41011701EE" country="Uruguay" name="Rivera" pageId="2" pageNumber="262">Rivera Department</collectingRegion>
, Sierra de la Aurora, under stones, 
<geoCoordinate id="EE67506AFFD8FFBDD49A3BAD034101EF" box="[692,840,1672,1698]" degrees="31" direction="south" minutes="03" orientation="latitude" pageId="2" pageNumber="262" precision="1" seconds="18.8" value="-31.055222">31°03'18.8&quot;S</geoCoordinate>
<geoCoordinate id="EE67506AFFD8FFBDD57F3BAD03FB01EE" box="[849,1010,1672,1699]" degrees="55" direction="west" minutes="42" orientation="longitude" pageId="2" pageNumber="262" precision="1" seconds="58.5" value="-55.71625">55°42'58.5&quot;W</geoCoordinate>
; 
<date id="FFED106DFFD8FFBDD22E3BAD046701EF" box="[1024,1134,1672,1698]" pageId="2" pageNumber="262" value="1961-01-10">10.I.1961</date>
; C. Carbonell and L.C. de Zolessi leg.; FCE-Myr 0354 ● 
<specimenCount id="9D55FD24FFD8FFBDD7D93B89022D018A" box="[503,548,1708,1735]" pageId="2" pageNumber="262" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D55FD24FFD8FFBDD41F3B890257018A" box="[561,606,1708,1735]" pageId="2" pageNumber="262" type="female">1 ♀</specimenCount>
; Valle del Lunarejo, riparian forest, 
<geoCoordinate id="EE67506AFFD8FFBDD5D43B89047F018B" box="[1018,1142,1708,1734]" degrees="31" direction="south" minutes="11" orientation="latitude" pageId="2" pageNumber="262" precision="15" seconds="27" value="-31.190832">31°11'27''S</geoCoordinate>
<geoCoordinate id="EE67506AFFD8FFBDD2513B890503018A" box="[1151,1290,1708,1735]" degrees="55" direction="west" minutes="56" orientation="longitude" pageId="2" pageNumber="262" precision="15" seconds="20" value="-55.93889">55°56'20''W</geoCoordinate>
; 
<date id="FFED106DFFD8FFBDD3383B890591018B" box="[1302,1432,1708,1734]" pageId="2" pageNumber="262" value="2019-03-13">13.III.2019</date>
; C. Rojas-Buffet leg.; FCE-Myr 0768 ● 
<specimenCount id="9D55FD24FFD8FFBDD4763BF5029301A6" box="[600,666,1744,1771]" pageId="2" pageNumber="262" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="9D55FD24FFD8FFBDD4883BF502EE01A6" box="[678,743,1744,1771]" pageId="2" pageNumber="262" type="female">3 ♀♀</specimenCount>
; same locality, under cow dung, camping zone, 
<geoCoordinate id="EE67506AFFD8FFBDD3243BF5059501A7" box="[1290,1436,1744,1770]" degrees="31" direction="south" minutes="11" orientation="latitude" pageId="2" pageNumber="262" precision="1" seconds="18.8" value="-31.188555">31°11'18.8''S</geoCoordinate>
<geoCoordinate id="EE67506AFFD8FFBDD6B93BD1013E0042" box="[151,311,1780,1807]" degrees="55" direction="west" minutes="54" orientation="longitude" pageId="2" pageNumber="262" precision="1" seconds="19.1" value="-55.905308">55°54'19.1''W</geoCoordinate>
; 
<date id="FFED106DFFD8FFBDD76B3BD101CE0043" box="[325,455,1780,1806]" pageId="2" pageNumber="262" value="2019-03-13">13.III.2019</date>
; C. Rojas-Buffet and M. Simó leg.; FCE-Myr 0769 ● 
<specimenCount id="9D55FD24FFD8FFBDD26E3BD104790042" box="[1088,1136,1780,1807]" pageId="2" pageNumber="262" type="male">1 ♂</specimenCount>
; same locality, under cow dung, 
<geoCoordinate id="EE67506AFFD8FFBDD6F03A3D017A007E" box="[222,371,1816,1843]" degrees="31" direction="south" minutes="10" orientation="latitude" pageId="2" pageNumber="262" precision="1" seconds="21.5" value="-31.172638">31°10'21.5&quot;S</geoCoordinate>
<geoCoordinate id="EE67506AFFD8FFBDD7553A3D0215007E" box="[379,540,1816,1843]" degrees="55" direction="west" minutes="54" orientation="longitude" pageId="2" pageNumber="262" precision="1" seconds="49.4" value="-55.913723">55°54'49.4&quot;W</geoCoordinate>
; 
<date id="FFED106DFFD8FFBDD4063A3D02A2007F" box="[552,683,1816,1842]" pageId="2" pageNumber="262" value="2019-11-16">16.XI.2019</date>
; E. Lorier leg.; FCE-Myr 0770 ● 
<specimenCount id="9D55FD24FFD8FFBDD2033A3D0479007E" box="[1069,1136,1816,1843]" pageId="2" pageNumber="262" type="male">2 ♂♂</specimenCount>
; same locality, two meters from the stream, 
<geoCoordinate id="EE67506AFFD8FFBDD7703A1901FA001B" box="[350,499,1852,1878]" degrees="31" direction="south" minutes="10" orientation="latitude" pageId="2" pageNumber="262" precision="1" seconds="23.0" value="-31.173056">31°10'23.0&quot;S</geoCoordinate>
<geoCoordinate id="EE67506AFFD8FFBDD7D23A190294001A" box="[508,669,1852,1879]" degrees="55" direction="west" minutes="54" orientation="longitude" pageId="2" pageNumber="262" precision="1" seconds="06.7" value="-55.901863">55°54'06.7&quot;W</geoCoordinate>
; 
<date id="FFED106DFFD8FFBDD4853A190327001B" box="[683,814,1852,1878]" pageId="2" pageNumber="262" value="2018-11-26">26.XI.2018</date>
; M.J. Albo leg.; FCE-Myr 0771 ● 
<specimenCount id="9D55FD24FFD8FFBDD2EB3A19051F001A" box="[1221,1302,1852,1879]" pageId="2" pageNumber="262" type="male">13 ♂♂</specimenCount>
, 
<specimenCount id="9D55FD24FFD8FFBDD30B3A19055B001A" box="[1317,1362,1852,1879]" pageId="2" pageNumber="262" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="4997F84FFFD8FFBDD34D3A1901170036" country="Uruguay" name="Salto" pageId="2" pageNumber="262">Salto Department</collectingRegion>
, Olivera, grassland, collected in pitfall trap, 
<geoCoordinate id="EE67506AFFD8FFBDD50E3A4503BC0036" box="[800,949,1888,1915]" degrees="31" direction="south" minutes="05" orientation="latitude" pageId="2" pageNumber="262" precision="1" seconds="22.0" value="-31.089445">31°05'22.0&quot;S</geoCoordinate>
<geoCoordinate id="EE67506AFFD8FFBDD5903A4504690036" box="[958,1120,1888,1915]" degrees="56" direction="west" minutes="46" orientation="longitude" pageId="2" pageNumber="262" precision="1" seconds="57.0" value="-56.7825">56°46'57.0&quot;W</geoCoordinate>
; 
<date id="FFED106DFFD8FFBDD2433A4504F30037" box="[1133,1274,1888,1914]" pageId="2" pageNumber="262" value="2019-12-10">10.XII.2019</date>
; G. Pompozzi leg.; FCE-Myr 0767.
</paragraph>
</subSubSection>
<caption id="DF2C6625FFD9FFBCD6B93A05046B00A0" ID-DOI="http://doi.org/10.5281/zenodo.6621307" ID-Zenodo-Dep="6621307" httpUri="https://zenodo.org/record/6621307/files/figure.png" pageId="3" pageNumber="263" startId="3.[151,250,1824,1848]" targetBox="[269,1317,181,1800]" targetPageId="3">
<paragraph id="8BEC36ADFFD9FFBCD6B93A05046B00A0" blockId="3.[151,1437,1824,2029]" pageId="3" pageNumber="263">
<emphasis id="B927EABFFFD9FFBCD6B93A0501060075" bold="true" box="[151,271,1824,1848]" pageId="3" pageNumber="263">FIGURE 2</emphasis>
. Somatic characters of 
<taxonomicName id="4C534D2EFFD9FFBCD42C3A0503220075" authority="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó, 2022" authorityName="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó" authorityYear="2022" box="[514,811,1824,1848]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="263" phylum="Arthropoda" rank="species" species="joachimadisi" status="sp. nov.">
<emphasis id="B927EABFFFD9FFBCD42C3A0503220075" box="[514,811,1824,1848]" italics="true" pageId="3" pageNumber="263">Sandalodesmus joachimadisi</emphasis>
</taxonomicName>
<emphasis id="B927EABFFFD9FFBCD51D3A0403870075" bold="true" box="[819,910,1824,1849]" pageId="3" pageNumber="263">
<taxonomicNameLabel id="A21457C4FFD9FFBCD51D3A0403640074" box="[819,877,1825,1849]" pageId="3" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
A.
</emphasis>
Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. 
<emphasis id="B927EABFFFD9FFBCD7363A6001260010" bold="true" box="[280,303,1861,1885]" pageId="3" pageNumber="263">B.</emphasis>
Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. 
<emphasis id="B927EABFFFD9FFBCD59D3A6103C20011" bold="true" box="[947,971,1860,1884]" pageId="3" pageNumber="263">C.</emphasis>
Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. 
<emphasis id="B927EABFFFD9FFBCD7763A4C016300CC" bold="true" box="[344,362,1897,1921]" pageId="3" pageNumber="263">D</emphasis>
. Non type ♂ (FCE-Myr 0556), left antenna. 
<emphasis id="B927EABFFFD9FFBCD51D3A4C034300CC" bold="true" box="[819,842,1897,1921]" pageId="3" pageNumber="263">E.</emphasis>
Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. 
<emphasis id="B927EABFFFD9FFBCD3A73A4C059500CC" bold="true" box="[1417,1436,1897,1921]" pageId="3" pageNumber="263">F.</emphasis>
Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. 
<emphasis id="B927EABFFFD9FFBCD4E23AA902EF00E9" bold="true" box="[716,742,1932,1956]" pageId="3" pageNumber="263">G.</emphasis>
Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. 
<emphasis id="B927EABFFFD9FFBCD7383A9401390084" bold="true" box="[278,304,1969,1993]" pageId="3" pageNumber="263">H.</emphasis>
Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. 
<emphasis id="B927EABFFFD9FFBCD20C3A9504D50084" bold="true" box="[1058,1244,1968,1993]" pageId="3" pageNumber="263">Abbreviations: L</emphasis>
: lingual lobes; 
<emphasis id="B927EABFFFD9FFBCD3533A94059C0084" bold="true" box="[1405,1429,1969,1993]" pageId="3" pageNumber="263">M</emphasis>
: mentum; 
<emphasis id="B927EABFFFD9FFBCD6D63AF1010500A1" bold="true" box="[248,268,2004,2028]" pageId="3" pageNumber="263">O</emphasis>
: ozopore; 
<emphasis id="B927EABFFFD9FFBCD7563AF0018E00A0" bold="true" box="[376,391,2005,2029]" pageId="3" pageNumber="263">P</emphasis>
: projection; 
<emphasis id="B927EABFFFD9FFBCD4273AF1021E00A1" bold="true" box="[521,535,2004,2028]" pageId="3" pageNumber="263">S</emphasis>
: stipes. 
<emphasis id="B927EABFFFD9FFBCD4443AF102E800A0" bold="true" box="[618,737,2004,2029]" pageId="3" pageNumber="263">Scale bars:</emphasis>
1.0 mm (
<emphasis id="B927EABFFFD9FFBCD56A3AF1037C00A0" bold="true" box="[836,885,2004,2029]" pageId="3" pageNumber="263">A–D</emphasis>
, 
<emphasis id="B927EABFFFD9FFBCD5AC3AF103BF00A0" bold="true" box="[898,950,2004,2029]" pageId="3" pageNumber="263">G–H</emphasis>
), 0.5 mm (
<emphasis id="B927EABFFFD9FFBCD2093AF0045D00A0" bold="true" box="[1063,1108,2005,2029]" pageId="3" pageNumber="263">E–F</emphasis>
).
</paragraph>
</caption>
<caption id="DF2C6625FFDEFFBBD6B9386604B802A6" ID-DOI="http://doi.org/10.5281/zenodo.6621311" ID-Zenodo-Dep="6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="4" pageNumber="264" startId="4.[151,250,1347,1371]" targetBox="[151,1436,181,1323]" targetPageId="4">
<paragraph id="8BEC36ADFFDEFFBBD6B9386604B802A6" blockId="4.[151,1437,1347,1515]" pageId="4" pageNumber="264">
<emphasis id="B927EABFFFDEFFBBD6B93866011C0216" bold="true" box="[151,277,1347,1371]" pageId="4" pageNumber="264">FIGURE 3.</emphasis>
Sexual characters of 
<taxonomicName id="4C534D2EFFDEFFBBD7D63866032B0216" authority="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó, 2022" authorityName="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó" authorityYear="2022" box="[504,802,1347,1371]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="264" phylum="Arthropoda" rank="species" species="joachimadisi" status="sp. nov.">
<emphasis id="B927EABFFFDEFFBBD7D63866032B0216" box="[504,802,1347,1371]" italics="true" pageId="4" pageNumber="264">Sandalodesmus joachimadisi</emphasis>
</taxonomicName>
<emphasis id="B927EABFFFDEFFBBD5043866038E0216" bold="true" box="[810,903,1347,1371]" pageId="4" pageNumber="264">
<taxonomicNameLabel id="A21457C4FFDEFFBBD5043866036F0217" box="[810,870,1347,1370]" pageId="4" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
A.
</emphasis>
Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. 
<emphasis id="B927EABFFFDEFFBBD6FD384200E20232" bold="true" box="[211,235,1383,1407]" pageId="4" pageNumber="264">B.</emphasis>
Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. 
<emphasis id="B927EABFFFDEFFBBD519384203460232" bold="true" box="[823,847,1383,1407]" pageId="4" pageNumber="264">C.</emphasis>
Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. 
<emphasis id="B927EABFFFDEFFBBD6B938AE00B902EE" bold="true" box="[151,176,1419,1443]" pageId="4" pageNumber="264">D.</emphasis>
Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. 
<emphasis id="B927EABFFFDEFFBBD4F238AE03A202EE" bold="true" box="[732,939,1419,1443]" pageId="4" pageNumber="264">Abbreviations: Ap:</emphasis>
acropodital process; 
<emphasis id="B927EABFFFDEFFBBD2AC38AE04A302EE" bold="true" box="[1154,1194,1419,1443]" pageId="4" pageNumber="264">Ca:</emphasis>
cannula; 
<emphasis id="B927EABFFFDEFFBBD32238AE053A02EE" bold="true" box="[1292,1331,1419,1443]" pageId="4" pageNumber="264">Cx:</emphasis>
coxa; 
<emphasis id="B927EABFFFDEFFBBD35838AE059502EE" bold="true" box="[1398,1436,1419,1443]" pageId="4" pageNumber="264">Ev:</emphasis>
external valve; 
<emphasis id="B927EABFFFDEFFBBD71C388A0158028A" bold="true" box="[306,337,1455,1479]" pageId="4" pageNumber="264">Iv:</emphasis>
internal valve; 
<emphasis id="B927EABFFFDEFFBBD7C0388A0211028A" bold="true" box="[494,536,1455,1479]" pageId="4" pageNumber="264">Op:</emphasis>
operculum; 
<emphasis id="B927EABFFFDEFFBBD4B9388A02BE028A" bold="true" box="[663,695,1455,1479]" pageId="4" pageNumber="264">Pf:</emphasis>
prefemur; 
<emphasis id="B927EABFFFDEFFBBD508388A035E028A" bold="true" box="[806,855,1455,1479]" pageId="4" pageNumber="264">PfP:</emphasis>
prefemoral process; 
<emphasis id="B927EABFFFDEFFBBD204388A0447028A" bold="true" box="[1066,1102,1455,1479]" pageId="4" pageNumber="264">Pg:</emphasis>
prostatic groove; 
<emphasis id="B927EABFFFDEFFBBD32A388A0512028A" bold="true" box="[1284,1307,1455,1479]" pageId="4" pageNumber="264">S:</emphasis>
solenomere; 
<emphasis id="B927EABFFFDEFFBBD6B938F600C202A6" bold="true" box="[151,203,1491,1515]" pageId="4" pageNumber="264">SpP:</emphasis>
spiniform process. 
<emphasis id="B927EABFFFDEFFBBD7BD38F6020302A6" bold="true" box="[403,522,1491,1515]" pageId="4" pageNumber="264">Scale bars:</emphasis>
0.5 mm (
<emphasis id="B927EABFFFDEFFBBD44338F6029402A6" bold="true" box="[621,669,1491,1515]" pageId="4" pageNumber="264">A–C</emphasis>
), 0.2 mm (
<emphasis id="B927EABFFFDEFFBBD52138F6032802A6" bold="true" box="[783,801,1491,1515]" pageId="4" pageNumber="264">D</emphasis>
). Photographed by: Damián Hagopián.
</paragraph>
</caption>
<subSubSection id="C3496526FFDEFFB9D6E93B3C02C70570" lastPageId="6" lastPageNumber="266" pageId="4" pageNumber="264" type="description">
<paragraph id="8BEC36ADFFDEFFBBD6E93B3C039200D6" blockId="4.[151,1437,1561,2020]" pageId="4" pageNumber="264">
<emphasis id="B927EABFFFDEFFBBD6E93B3C0194017E" bold="true" box="[199,413,1561,1587]" pageId="4" pageNumber="264">Description. Male</emphasis>
(
<typeStatus id="54E8880FFFDEFFBBD7823B3C021B017E" box="[428,530,1561,1587]" pageId="4" pageNumber="264" type="holotype">holotype</typeStatus>
, FCE-Myr 0664). Coloration in life: head reddish. Antennae reddish brown with distal end of antennomers yellowish, antennomere seven completely yellow (
<figureCitation id="13682A28FFDEFFBBD5DD3B180444011A" box="[1011,1101,1597,1623]" captionStart="FIGURE 2" captionStartId="3.[151,250,1824,1848]" captionTargetBox="[269,1317,181,1800]" captionTargetId="figure-20@3.[269,1317,181,1800]" captionTargetPageId="3" captionText="FIGURE 2. Somatic characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. B. Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. C. Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. D. Non type ♂ (FCE-Myr 0556), left antenna. E. Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. F. Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. G. Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. H. Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. Abbreviations: L: lingual lobes; M: mentum; O: ozopore; P: projection; S: stipes. Scale bars: 1.0 mm (A–D, G–H), 0.5 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6621307" httpUri="https://zenodo.org/record/6621307/files/figure.png" pageId="4" pageNumber="264">Fig. 2D</figureCitation>
). Body reddish, with paranota tip yellowish and metazonites whitish at posterior border (
<figureCitation id="13682A28FFDEFFBBD51F3B4403730136" box="[817,890,1633,1659]" captionStart="FIGURE 1" captionStartId="2.[151,250,1314,1338]" captionTargetBox="[151,1436,776,1290]" captionTargetId="figure-208@2.[151,1436,776,1290]" captionTargetPageId="2" captionText="FIGURE 1. Sandalodesmus joachimadisi n. sp. from Valle del Lunarejo, Rivera, Uruguay. A. Live male. B. Two specimens found under cow dung, one of them inside a nest." figureDoi="http://doi.org/10.5281/zenodo.6621305" httpUri="https://zenodo.org/record/6621305/files/figure.png" pageId="4" pageNumber="264">Figs 1</figureCitation>
, 
<figureCitation id="13682A28FFDEFFBBD5A43B4403D80136" box="[906,977,1633,1659]" captionStart="FIGURE 2" captionStartId="3.[151,250,1824,1848]" captionTargetBox="[269,1317,181,1800]" captionTargetId="figure-20@3.[269,1317,181,1800]" captionTargetPageId="3" captionText="FIGURE 2. Somatic characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. B. Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. C. Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. D. Non type ♂ (FCE-Myr 0556), left antenna. E. Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. F. Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. G. Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. H. Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. Abbreviations: L: lingual lobes; M: mentum; O: ozopore; P: projection; S: stipes. Scale bars: 1.0 mm (A–D, G–H), 0.5 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6621307" httpUri="https://zenodo.org/record/6621307/files/figure.png" pageId="4" pageNumber="264">2A–C</figureCitation>
). Gnathochilarium (
<figureCitation id="13682A28FFDEFFBBD29B3B4405240131" box="[1205,1325,1633,1660]" captionStart="FIGURE 2" captionStartId="3.[151,250,1824,1848]" captionTargetBox="[269,1317,181,1800]" captionTargetId="figure-20@3.[269,1317,181,1800]" captionTargetPageId="3" captionText="FIGURE 2. Somatic characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. B. Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. C. Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. D. Non type ♂ (FCE-Myr 0556), left antenna. E. Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. F. Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. G. Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. H. Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. Abbreviations: L: lingual lobes; M: mentum; O: ozopore; P: projection; S: stipes. Scale bars: 1.0 mm (A–D, G–H), 0.5 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6621307" httpUri="https://zenodo.org/record/6621307/files/figure.png" pageId="4" pageNumber="264">Fig. 2E, F</figureCitation>
): 
<emphasis id="B927EABFFFDEFFBBD36A3B4705940136" box="[1348,1437,1634,1659]" italics="true" pageId="4" pageNumber="264">mentum</emphasis>
(
<emphasis id="B927EABFFFDEFFBBD68F3BA000CD01D2" bold="true" box="[161,196,1669,1695]" pageId="4" pageNumber="264">M)</emphasis>
with smooth edges and small thick setae at central region; apical edge of 
<emphasis id="B927EABFFFDEFFBBD22A3BA3044D01D2" box="[1028,1092,1670,1695]" italics="true" pageId="4" pageNumber="264">stipes</emphasis>
(
<emphasis id="B927EABFFFDEFFBBD27A3BA0046401D2" bold="true" box="[1108,1133,1669,1695]" pageId="4" pageNumber="264">S)</emphasis>
and 
<emphasis id="B927EABFFFDEFFBBD28A3BA0053F01D2" box="[1188,1334,1669,1695]" italics="true" pageId="4" pageNumber="264">lingual lobes</emphasis>
(
<emphasis id="B927EABFFFDEFFBBD3693BA0056A01D2" bold="true" box="[1351,1379,1669,1695]" pageId="4" pageNumber="264">L)</emphasis>
with long and thin setae. Body rings: cuticle smooth; paranota with standard polydesmidan pore formula (5, 7, 9, 10, 12, 13, 15–19); paranota sub-triangular (
<figureCitation id="13682A28FFDEFFBBD41D3BE802A601AA" box="[563,687,1741,1767]" captionStart="FIGURE 2" captionStartId="3.[151,250,1824,1848]" captionTargetBox="[269,1317,181,1800]" captionTargetId="figure-20@3.[269,1317,181,1800]" captionTargetPageId="3" captionText="FIGURE 2. Somatic characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. B. Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. C. Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. D. Non type ♂ (FCE-Myr 0556), left antenna. E. Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. F. Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. G. Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. H. Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. Abbreviations: L: lingual lobes; M: mentum; O: ozopore; P: projection; S: stipes. Scale bars: 1.0 mm (A–D, G–H), 0.5 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6621307" httpUri="https://zenodo.org/record/6621307/files/figure.png" pageId="4" pageNumber="264">Fig. 2A–C</figureCitation>
); ozopores (
<emphasis id="B927EABFFFDEFFBBD5163BE8034701AA" bold="true" box="[824,846,1741,1767]" pageId="4" pageNumber="264">O</emphasis>
) situated posteriorly (
<figureCitation id="13682A28FFDEFFBBD26B3BE804B701AA" box="[1093,1214,1741,1767]" captionStart="FIGURE 2" captionStartId="3.[151,250,1824,1848]" captionTargetBox="[269,1317,181,1800]" captionTargetId="figure-20@3.[269,1317,181,1800]" captionTargetPageId="3" captionText="FIGURE 2. Somatic characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. B. Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. C. Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. D. Non type ♂ (FCE-Myr 0556), left antenna. E. Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. F. Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. G. Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. H. Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. Abbreviations: L: lingual lobes; M: mentum; O: ozopore; P: projection; S: stipes. Scale bars: 1.0 mm (A–D, G–H), 0.5 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6621307" httpUri="https://zenodo.org/record/6621307/files/figure.png" pageId="4" pageNumber="264">Fig. 2B, C</figureCitation>
). Stigma oval. Sternite 4 with a single pair of fused projections (
<emphasis id="B927EABFFFDEFFBBD4A03BD402A00046" bold="true" box="[654,681,1777,1803]" pageId="4" pageNumber="264">P)</emphasis>
(
<figureCitation id="13682A28FFDEFFBBD4963BD403180046" box="[696,785,1777,1803]" captionStart="FIGURE 2" captionStartId="3.[151,250,1824,1848]" captionTargetBox="[269,1317,181,1800]" captionTargetId="figure-20@3.[269,1317,181,1800]" captionTargetPageId="3" captionText="FIGURE 2. Somatic characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. B. Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. C. Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. D. Non type ♂ (FCE-Myr 0556), left antenna. E. Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. F. Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. G. Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. H. Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. Abbreviations: L: lingual lobes; M: mentum; O: ozopore; P: projection; S: stipes. Scale bars: 1.0 mm (A–D, G–H), 0.5 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6621307" httpUri="https://zenodo.org/record/6621307/files/figure.png" pageId="4" pageNumber="264">Fig. 2G</figureCitation>
). Other sternites without modifications (
<figureCitation id="13682A28FFDEFFBBD2FC3BD405250046" box="[1234,1324,1777,1803]" captionStart="FIGURE 2" captionStartId="3.[151,250,1824,1848]" captionTargetBox="[269,1317,181,1800]" captionTargetId="figure-20@3.[269,1317,181,1800]" captionTargetPageId="3" captionText="FIGURE 2. Somatic characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. B. Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. C. Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. D. Non type ♂ (FCE-Myr 0556), left antenna. E. Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. F. Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. G. Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. H. Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. Abbreviations: L: lingual lobes; M: mentum; O: ozopore; P: projection; S: stipes. Scale bars: 1.0 mm (A–D, G–H), 0.5 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6621307" httpUri="https://zenodo.org/record/6621307/files/figure.png" pageId="4" pageNumber="264">Fig. 2H</figureCitation>
). Anterior sternites covered with short and thick setae, posterior ones sparsely covered with thin setae. Leg pair 2 coxae each with a rectangular-shaped genital papilla. Walking legs reddish white-brown and without modifications (
<figureCitation id="13682A28FFDEFFBBD3693A1C0582001E" box="[1351,1419,1849,1875]" captionStart="FIGURE 1" captionStartId="2.[151,250,1314,1338]" captionTargetBox="[151,1436,776,1290]" captionTargetId="figure-208@2.[151,1436,776,1290]" captionTargetPageId="2" captionText="FIGURE 1. Sandalodesmus joachimadisi n. sp. from Valle del Lunarejo, Rivera, Uruguay. A. Live male. B. Two specimens found under cow dung, one of them inside a nest." figureDoi="http://doi.org/10.5281/zenodo.6621305" httpUri="https://zenodo.org/record/6621305/files/figure.png" pageId="4" pageNumber="264">Fig. 1</figureCitation>
). Gonopod aperture on seventh body ring oval, posterior edge straight and without folds, but slightly elevated in anal view. Telson anteriorly reddish, posterior border yellowish (
<figureCitation id="13682A28FFDEFFBBD5183AA4038700D6" box="[822,910,1921,1947]" captionStart="FIGURE 2" captionStartId="3.[151,250,1824,1848]" captionTargetBox="[269,1317,181,1800]" captionTargetId="figure-20@3.[269,1317,181,1800]" captionTargetPageId="3" captionText="FIGURE 2. Somatic characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), anterior body rings, dorsal view. B. Holotype ♂ (FCE-Myr 0664), middle body rings, dorsal view. C. Holotype ♂ (FCE-Myr 0664), posterior body rings, dorsal view. D. Non type ♂ (FCE-Myr 0556), left antenna. E. Non type ♂ (FCE-Myr 0556), ventral gnathochilarium. F. Non type ♂ (FCE-Myr 0556), dorsal gnathochilarium. G. Non type ♂ (FCE-Myr 0767), sternite 4 with a single pair of fused projections. H. Non type ♂ (FCE-Myr 0767), posterior sternites without modifications. Abbreviations: L: lingual lobes; M: mentum; O: ozopore; P: projection; S: stipes. Scale bars: 1.0 mm (A–D, G–H), 0.5 mm (E–F)." figureDoi="http://doi.org/10.5281/zenodo.6621307" httpUri="https://zenodo.org/record/6621307/files/figure.png" pageId="4" pageNumber="264">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8BEC36ADFFDEFFB9D6E93A80028E0650" blockId="4.[151,1437,1561,2020]" lastBlockId="6.[151,1437,151,789]" lastPageId="6" lastPageNumber="266" pageId="4" pageNumber="264">
Gonopods (
<figureCitation id="13682A28FFDEFFBBD7623A80019F00F2" box="[332,406,1957,1983]" captionStart="FIGURE 3" captionStartId="4.[151,250,1347,1371]" captionTargetBox="[151,1436,181,1323]" captionTargetId="figure-17@4.[151,1436,181,1323]" captionTargetPageId="4" captionText="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="4" pageNumber="264">Figs 3</figureCitation>
, 
<figureCitation id="13682A28FFDEFFBBD7883A8001BE00F2" box="[422,439,1957,1983]" captionStart="FIGURE 4" captionStartId="5.[151,250,697,721]" captionTargetBox="[155,1429,187,670]" captionTargetId="figure-20@5.[151,1436,181,673]" captionTargetPageId="5" captionText="FIGURE 4. Gonopod drawings of Sandalodesmus joachimadisi n. sp., holotype ♂ (FCE-Myr 0664), left gonopod. A. Mesal view. B. Lateral view. C. Ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6621313" httpUri="https://zenodo.org/record/6621313/files/figure.png" pageId="4" pageNumber="264">4</figureCitation>
): coxae (
<emphasis id="B927EABFFFDEFFBBD40A3A80024F00F2" bold="true" box="[548,582,1957,1983]" pageId="4" pageNumber="264">Cx</emphasis>
) almost same length as telopodite, prominent in lateral view (
<figureCitation id="13682A28FFDEFFBBD3393A80059100F2" box="[1303,1432,1957,1983]" captionStart="FIGURE 3" captionStartId="4.[151,250,1347,1371]" captionTargetBox="[151,1436,181,1323]" captionTargetId="figure-17@4.[151,1436,181,1323]" captionTargetPageId="4" captionText="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="4" pageNumber="264">Figs 3A–C</figureCitation>
, 
<figureCitation id="13682A28FFDEFFBBD6B93AEC00AE00AE" box="[151,167,1993,2019]" captionStart="FIGURE 4" captionStartId="5.[151,250,697,721]" captionTargetBox="[155,1429,187,670]" captionTargetId="figure-20@5.[151,1436,181,673]" captionTargetPageId="5" captionText="FIGURE 4. Gonopod drawings of Sandalodesmus joachimadisi n. sp., holotype ♂ (FCE-Myr 0664), left gonopod. A. Mesal view. B. Lateral view. C. Ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6621313" httpUri="https://zenodo.org/record/6621313/files/figure.png" pageId="4" pageNumber="264">4</figureCitation>
), with two anterior macrosetae (
<figureCitation id="13682A28FFDEFFBBD43B3AEC027B00AE" box="[533,626,1993,2019]" captionStart="FIGURE 3" captionStartId="4.[151,250,1347,1371]" captionTargetBox="[151,1436,181,1323]" captionTargetId="figure-17@4.[151,1436,181,1323]" captionTargetPageId="4" captionText="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="4" pageNumber="264">Figs 3B</figureCitation>
, 
<figureCitation id="13682A28FFDEFFBBD4503AEC02A900AE" box="[638,672,1993,2019]" captionStart="FIGURE 4" captionStartId="5.[151,250,697,721]" captionTargetBox="[155,1429,187,670]" captionTargetId="figure-20@5.[151,1436,181,673]" captionTargetPageId="5" captionText="FIGURE 4. Gonopod drawings of Sandalodesmus joachimadisi n. sp., holotype ♂ (FCE-Myr 0664), left gonopod. A. Mesal view. B. Lateral view. C. Ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6621313" httpUri="https://zenodo.org/record/6621313/files/figure.png" pageId="4" pageNumber="264">4B</figureCitation>
) and with a single, large and slender spiniform process (
<emphasis id="B927EABFFFDEFFBBD30F3AEC055800AE" bold="true" box="[1313,1361,1993,2019]" pageId="4" pageNumber="264">SpP</emphasis>
) (Figs Total length (
<typeStatus id="54E8880FFFDCFFB9D7003DB2019D07FC" box="[302,404,151,177]" pageId="6" pageNumber="266" type="holotype">holotype</typeStatus>
, FCE-Myr 0664) (fragmented specimen): 39.8. Width: 4.47. Antennomere lengths (1 to 7): 0.42; 0.94; 0.93; 0.95; 0.97; 1.00; 0.24. Podomeres lengths of leg 10 (1 to 7): 0.49; 0.80; 1.53; 0.53; 0.63; 1.00; 0.30. Gonopod aperture: 1.0 long, 1.9 wide. Telson: 1.01 long. Gonopod: 1.69 long, 1.83 wide. Gonocoxae: 0.83 long, 1.09 wide. Telopodite: 1.57 long, 0.72 wide.
</paragraph>
<caption id="DF2C6625FFDFFFBAD6B93F9C04A70454" ID-DOI="http://doi.org/10.5281/zenodo.6621313" ID-Zenodo-Dep="6621313" httpUri="https://zenodo.org/record/6621313/files/figure.png" pageId="5" pageNumber="265" startId="5.[151,250,697,721]" targetBox="[155,1429,187,670]" targetPageId="5">
<paragraph id="8BEC36ADFFDFFFBAD6B93F9C04A70454" blockId="5.[151,1437,697,794]" pageId="5" pageNumber="265">
<emphasis id="B927EABFFFDFFFBAD6B93F9C0107059C" bold="true" box="[151,270,697,721]" pageId="5" pageNumber="265">FIGURE 4</emphasis>
. Gonopod drawings of 
<taxonomicName id="4C534D2EFFDFFFBAD42C3F9C0323059C" authority="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó, 2022" authorityName="Rojas-Buffet &amp; Bouzan &amp; Sierwald &amp; Brescovit &amp; Simó" authorityYear="2022" box="[514,810,697,721]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="265" phylum="Arthropoda" rank="species" species="joachimadisi" status="sp. nov.">
<emphasis id="B927EABFFFDFFFBAD42C3F9C0323059C" box="[514,810,697,721]" italics="true" pageId="5" pageNumber="265">Sandalodesmus joachimadisi</emphasis>
</taxonomicName>
<emphasis id="B927EABFFFDFFFBAD51F3F9F0365059F" bold="true" box="[817,876,698,722]" pageId="5" pageNumber="265">
<taxonomicNameLabel id="A21457C4FFDFFFBAD51F3F9F0365059F" box="[817,876,698,722]" pageId="5" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype ♂ (FCE-Myr 0664), left gonopod. 
<emphasis id="B927EABFFFDFFFBAD3103F9C055E059C" bold="true" box="[1342,1367,697,721]" pageId="5" pageNumber="265">A.</emphasis>
Mesal view. 
<emphasis id="B927EABFFFDFFFBAD6FB3FFB00E505BB" bold="true" box="[213,236,734,758]" pageId="5" pageNumber="265">B.</emphasis>
Lateral view. 
<emphasis id="B927EABFFFDFFFBAD7AF3FF8019305B8" bold="true" box="[385,410,733,757]" pageId="5" pageNumber="265">C.</emphasis>
Ventral view. 
<emphasis id="B927EABFFFDFFFBAD41E3FF8030805B8" bold="true" box="[560,769,733,757]" pageId="5" pageNumber="265">Abbreviations: Ap:</emphasis>
acropodital process; 
<emphasis id="B927EABFFFDFFFBAD5F03FF8040C05B8" bold="true" box="[990,1029,733,757]" pageId="5" pageNumber="265">Ca:</emphasis>
cannula; 
<emphasis id="B927EABFFFDFFFBAD2443FF8049B05B8" bold="true" box="[1130,1170,733,757]" pageId="5" pageNumber="265">Cx:</emphasis>
coxa; 
<emphasis id="B927EABFFFDFFFBAD2F63FF804F005B8" bold="true" box="[1240,1273,733,757]" pageId="5" pageNumber="265">Pf:</emphasis>
prefemur; 
<emphasis id="B927EABFFFDFFFBAD3423FF8059405B8" bold="true" box="[1388,1437,733,757]" pageId="5" pageNumber="265">PfP:</emphasis>
prefemoral process; 
<emphasis id="B927EABFFFDFFFBAD7483E2701820457" bold="true" box="[358,395,770,794]" pageId="5" pageNumber="265">Pg:</emphasis>
prostatic groove; 
<emphasis id="B927EABFFFDFFFBAD46C3E2402500454" bold="true" box="[578,601,769,793]" pageId="5" pageNumber="265">S:</emphasis>
solenomere; 
<emphasis id="B927EABFFFDFFFBAD4CF3E24031C0454" bold="true" box="[737,789,769,793]" pageId="5" pageNumber="265">SpP:</emphasis>
spiniform process. 
<emphasis id="B927EABFFFDFFFBAD5F33E2404450457" bold="true" box="[989,1100,769,794]" pageId="5" pageNumber="265">Scale bars</emphasis>
: 0.5 mm.
</paragraph>
</caption>
<caption id="DF2C6625FFDFFFBAD6B93ABD02D500FD" ID-DOI="http://doi.org/10.5281/zenodo.6621315" ID-Zenodo-Dep="6621315" box="[151,732,1944,1968]" httpUri="https://zenodo.org/record/6621315/files/figure.png" pageId="5" pageNumber="265" startId="5.[151,250,1944,1968]" targetBox="[151,1427,818,1920]" targetPageId="5">
<paragraph id="8BEC36ADFFDFFFBAD6B93ABD02D500FD" blockId="5.[151,732,1944,1968]" box="[151,732,1944,1968]" pageId="5" pageNumber="265">
<emphasis id="B927EABFFFDFFFBAD6B93ABD011A00FD" bold="true" box="[151,275,1944,1968]" pageId="5" pageNumber="265">FIGURE 5.</emphasis>
Distribution map of 
<taxonomicName id="4C534D2EFFDFFFBAD7C63ABD02D500FD" box="[488,732,1944,1968]" class="Diplopoda" family="Chelodesmidae" genus="Sandalodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="265" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B927EABFFFDFFFBAD7C63ABD028F00FD" box="[488,646,1944,1968]" italics="true" pageId="5" pageNumber="265">Sandalodesmus</emphasis>
species.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BEC36ADFFDCFFB9D6E93C0202C70570" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="266">
<emphasis id="B927EABFFFDCFFB9D6E93C020117060C" bold="true" box="[199,286,295,321]" pageId="6" pageNumber="266">Female</emphasis>
(
<typeStatus id="54E8880FFFDCFFB9D7013C02019D060C" box="[303,404,295,321]" pageId="6" pageNumber="266" type="paratype">paratype</typeStatus>
, FCE-Myr 0664). Coloration and body features as in male except that the coxae on body ring 3 with a small process. Posterior margin of vulvae aperture smooth, without projections. Vulvae (
<figureCitation id="13682A28FFDCFFB9D3283C6E05540628" box="[1286,1373,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1347,1371]" captionTargetBox="[151,1436,181,1323]" captionTargetId="figure-17@4.[151,1436,181,1323]" captionTargetPageId="4" captionText="FIGURE 3. Sexual characters of Sandalodesmus joachimadisi n. sp. A. Holotype ♂ (FCE-Myr 0664), left gonopod, mesal view. B. Holotype ♂ (FCE-Myr 0664), left gonopod, lateral view. C. Holotype ♂ (FCE-Myr 0664), left gonopod, ventral view. D. Paratype ♀ (FCE-Myr 0664), left vulva, ventral view. Abbreviations: Ap: acropodital process; Ca: cannula; Cx: coxa; Ev: external valve; Iv: internal valve; Op: operculum; Pf: prefemur; PfP: prefemoral process; Pg: prostatic groove; S: solenomere; SpP: spiniform process. Scale bars: 0.5 mm (A–C), 0.2 mm (D). Photographed by: Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.6621311" httpUri="https://zenodo.org/record/6621311/files/figure.png" pageId="6" pageNumber="266">Fig. 3D</figureCitation>
) oval shaped in ventral view and elliptical shaped in lateral view. External and internal valves unequal in size, external valve (
<emphasis id="B927EABFFFDCFFB9D6CB3CB6010F06E0" bold="true" box="[229,262,403,429]" pageId="6" pageNumber="266">Ev</emphasis>
) longer than internal valve (
<emphasis id="B927EABFFFDCFFB9D4663CB6026806E0" bold="true" box="[584,609,403,429]" pageId="6" pageNumber="266">Iv</emphasis>
), in ventral/distal view. Internal edges of both valves irregular in the medial portion. Both valves with scattered setae in comparison with operculum (
<emphasis id="B927EABFFFDCFFB9D5D73C930417069D" bold="true" box="[1017,1054,438,464]" pageId="6" pageNumber="266">Op</emphasis>
). Operculum about 1/3 of the size of valves, typical for chelodesmids. Total length: 38.04. Width total: 4.95. Telson 1.3 long. Posterior margin of the vulvae aperture: 1.57 wide. Vulvae: 0.79 long, 0.55 wide. External valve: 0.72 long, 0.21 wide. Internal valve: 0.68 long, 0.31 wide. Operculum: 0.12 long, 0.31 wide.
</paragraph>
</subSubSection>
<subSubSection id="C3496526FFDCFFB9D6E93F63029E052C" box="[199,663,582,609]" pageId="6" pageNumber="266" type="distribution">
<paragraph id="8BEC36ADFFDCFFB9D6E93F63029E052C" blockId="6.[151,1437,151,789]" box="[199,663,582,609]" pageId="6" pageNumber="266">
<emphasis id="B927EABFFFDCFFB9D6E93F630168052D" bold="true" box="[199,353,582,608]" pageId="6" pageNumber="266">Distribution.</emphasis>
Northern 
<collectingCountry id="F344763DFFDCFFB9D7FA3F62023F052C" box="[468,566,583,609]" name="Uruguay" pageId="6" pageNumber="266">Uruguay</collectingCountry>
(
<figureCitation id="13682A28FFDCFFB9D46B3F620280052C" box="[581,649,582,609]" captionStart="FIGURE 5" captionStartId="5.[151,250,1944,1968]" captionTargetBox="[151,1427,818,1920]" captionTargetId="figure-75@5.[151,1436,818,1920]" captionTargetPageId="5" captionText="FIGURE 5. Distribution map of Sandalodesmus species." figureDoi="http://doi.org/10.5281/zenodo.6621315" httpUri="https://zenodo.org/record/6621315/files/figure.png" pageId="6" pageNumber="266">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3496526FFDCFFB9D6E93F4E014F0458" pageId="6" pageNumber="266" type="biology_ecology">
<paragraph id="8BEC36ADFFDCFFB9D6E93F4E014F0458" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="266">
<emphasis id="B927EABFFFDCFFB9D6E93F4E018E05C9" bold="true" box="[199,391,618,645]" pageId="6" pageNumber="266">Natural history.</emphasis>
The species is usually associated with urban environments, such as built-up areas or camping areas. It can also be found in natural areas, such as riparian forest or grasslands. Specimens were found under stones, under decaying tree trunks, under cow dung, or walking on grass. Some specimens were found together in nests (
<figureCitation id="13682A28FFDCFFB9D6CE3FF2013F05BC" box="[224,310,727,753]" captionStart="FIGURE 1" captionStartId="2.[151,250,1314,1338]" captionTargetBox="[151,1436,776,1290]" captionTargetId="figure-208@2.[151,1436,776,1290]" captionTargetPageId="2" captionText="FIGURE 1. Sandalodesmus joachimadisi n. sp. from Valle del Lunarejo, Rivera, Uruguay. A. Live male. B. Two specimens found under cow dung, one of them inside a nest." figureDoi="http://doi.org/10.5281/zenodo.6621305" httpUri="https://zenodo.org/record/6621305/files/figure.png" pageId="6" pageNumber="266">Fig. 1B</figureCitation>
) that may have been built by the millipedes. Frequently more than one individual was found at the collection sites.
</paragraph>
</subSubSection>
</treatment>
</document>