177 lines
20 KiB
XML
177 lines
20 KiB
XML
<document id="05DD54EAB2B64B0684D19C6803AC7F8D" ID-DOI="10.11646/zootaxa.3990.2.2" ID-GBIF-Dataset="1740ec81-c55e-44ce-9a6d-cd62acd5d898" ID-ISSN="1175-5326" ID-Zenodo-Dep="242947" ID-ZooBank="BDF1BC11-4848-4858-99DC-0336A5FBEE2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267666231" checkinUser="plazi" docAuthor="Helwerda, Renate A." docDate="2015" docId="03FA878C951CFE44FF26E6D8027ECAE5" docLanguage="en" docName="zt03990p220.pdf" docOrigin="Zootaxa 3990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ringicula bella" docType="treatment" docVersion="9" lastPageNumber="209" masterDocId="FFC3FFF49510FE4AFFB1E574036DCA1D" masterDocTitle="Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines" masterLastPageNumber="220" masterPageNumber="197" pageNumber="209" updateTime="1698610896101" updateUser="plazi">
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<mods:titleInfo id="D360FBF58BE7F35A801784ED4D0482F3">
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<mods:title id="37BE86B6183A4C3218584EFFB0EED683">Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines</mods:title>
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</mods:titleInfo>
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<mods:name id="868091D3A21332AF7A473554E7C5F03E" type="personal">
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<mods:roleTerm id="5025BDB96362CBB879D4735D564DD23E">Author</mods:roleTerm>
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<mods:namePart id="C43DFE8D95DBC90DFD4DE6BE67480F8E">Helwerda, Renate A.</mods:namePart>
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<mods:typeOfResource id="8F4327E7DC6E69CAE02E539AF7BDF9CD">text</mods:typeOfResource>
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<mods:titleInfo id="C757C2EBB6C6F8B508C1C14757114FB0">
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<mods:title id="0ED9FB642F131994AA9CB55F767B40AE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="70DC19BE248BE624ED2DF54DA485A441">
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<mods:date id="8235EAFFFB5A3095EC3D7B7FDF1009D3">2015</mods:date>
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<mods:detail id="08ACA3C4261AB57783B3ED64964FF5D4" type="volume">
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<mods:number id="F8802D34A65D1661F67AD6BEFA03BA96">3990</mods:number>
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</mods:detail>
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<mods:detail id="035C5FB7D6D38F32CCCF345A2DAE66EB" type="issue">
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<mods:number id="3887DE2C1A07FE5D3E8648D474453821">2</mods:number>
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<mods:classification id="9F2066F7610AE51345683797A3A4772E">journal article</mods:classification>
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<mods:identifier id="AF747A2F50B6E68371BB84DA3EBE9E18" type="DOI">10.11646/zootaxa.3990.2.2</mods:identifier>
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<treatment id="03FA878C951CFE44FF26E6D8027ECAE5" ID-DOI="http://doi.org/10.5281/zenodo.6116531" ID-GBIF-Taxon="119638254" ID-Zenodo-Dep="6116531" LSID="urn:lsid:plazi:treatment:03FA878C951CFE44FF26E6D8027ECAE5" httpUri="http://treatment.plazi.org/id/03FA878C951CFE44FF26E6D8027ECAE5" lastPageId="14" lastPageNumber="209" pageId="12" pageNumber="209">
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<subSubSection id="C3496511951CFE46FF26E6D80227C9FA" pageId="12" pageNumber="209" type="nomenclature">
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<paragraph id="8BEC369A951CFE46FF26E6D802A3C9DB" blockId="12.[151,462,940,999]" box="[151,462,940,966]" pageId="12" pageNumber="209">
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<heading id="D0A481F6951CFE46FF26E6D802A3C9DB" bold="true" box="[151,462,940,966]" fontSize="11" level="1" pageId="12" pageNumber="209" reason="1">
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<emphasis id="B927EA88951CFE46FF26E6D802A3C9DB" bold="true" box="[151,462,940,966]" pageId="12" pageNumber="209">
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<taxonomicName id="4C534D19951CFE46FF26E6D8023DC9DB" ID-CoL="4T3TY" box="[151,336,940,966]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" higherTaxonomySource="CoL" kingdom="Animalia" order="Ellobiida" pageId="12" pageNumber="209" phylum="Mollusca" rank="species" species="bella">
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<emphasis id="B927EA88951CFE46FF26E6D8023DC9DB" bold="true" box="[151,336,940,966]" italics="true" pageId="12" pageNumber="209">Ringicula bella</emphasis>
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</taxonomicName>
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nov. spec.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEC369A951CFE46FF26E6BA0227C9FA" blockId="12.[151,462,940,999]" box="[151,330,974,999]" pageId="12" pageNumber="209">
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(
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<figureCitation id="13682A1F951CFE46FF2FE6BA022EC9FA" box="[158,323,974,999]" captionStart="FIGURE 23 – 28. 23" captionStartId="13.[151,250,1781,1803]" captionTargetBox="[239,1346,193,1758]" captionTargetId="figure@13.[239,1347,193,1760]" captionTargetPageId="13" captionText="FIGURE 23 – 28. 23. Ringicula bella nov. spec .. Holotype RGM 961.797. Locality Anda 1. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view. 24. Ringicula bella nov. spec .. Paratype RGM 961.750. Locality Tiep 2. (a) apical view, (b) apertural view, composite. 25. Ringicula spec 1. RGM 961.817. Locality Tiep 2. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view. 26. Ringicula cf. spec 1. RGM 961.753. Locality Tiep 2. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view. 27. Ringicula circumscripta nov. spec .. Holotype RGM 961.754. Locality Roxas. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view. 28. Ringicula spec 2. RGM 961.751. Locality Tiep 2. (a) rear view, (b) oblique apertural view, (c) apertural view, (d) basal view, (e) apical view." httpUri="https://zenodo.org/record/242952/files/figure.png" pageId="12" pageNumber="209">Figures 23–24</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3496511951CFE46FF26E16202D3CEFE" pageId="12" pageNumber="209" type="materials_examined">
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<paragraph id="8BEC369A951CFE46FF26E1620233CE4E" blockId="12.[151,1437,1046,2007]" pageId="12" pageNumber="209">
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<emphasis id="B927EA88951CFE46FF26E1620226CE32" bold="true" box="[151,331,1046,1071]" pageId="12" pageNumber="209">
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<typeStatus id="54E88838951CFE46FF26E16203BFCE32" box="[151,210,1046,1071]" pageId="12" pageNumber="209">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54E88838951CFE46FEE8E16202ACCE32" box="[345,449,1046,1071]" pageId="12" pageNumber="209" type="holotype">Holotype</typeStatus>
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<collectionCode id="ED42AE5F951CFE46FE7EE1630162CE32" box="[463,527,1047,1071]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.797 (
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<typeStatus id="54E88838951CFE46FD3CE16301ABCE32" box="[653,710,1047,1071]" pageId="12" pageNumber="209">Type</typeStatus>
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locality).
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<typeStatus id="54E88838951CFE46FCF7E16300C5CE32" box="[838,936,1047,1071]" pageId="12" pageNumber="209" type="paratype">Paratype</typeStatus>
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<collectionCode id="ED42AE5F951CFE46FC07E163009BCE32" box="[950,1014,1047,1071]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.750 (
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<typeStatus id="54E88838951CFE46FBC5E16307C3CE32" box="[1140,1198,1047,1071]" pageId="12" pageNumber="209">Type</typeStatus>
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locality) and
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<collectionCode id="ED42AE5F951CFE46FAEDE16306F1CE32" box="[1372,1436,1047,1071]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.802 (Anda4).
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</paragraph>
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<paragraph id="8BEC369A951CFE46FF76E12A01EACEA2" blockId="12.[151,1437,1046,2007]" pageId="12" pageNumber="209">
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<emphasis id="B927EA88951CFE46FF76E12A02E9CE6A" bold="true" box="[199,388,1118,1143]" pageId="12" pageNumber="209">Other material.</emphasis>
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Anda1 (18)
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<collectionCode id="ED42AE5F951CFE46FDABE12B0137CE6A" box="[538,602,1119,1143]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.798, Anda2 (22)
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<collectionCode id="ED42AE5F951CFE46FCEAE12B00F6CE6A" box="[859,923,1119,1143]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.799, Anda3 (14)
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<collectionCode id="ED42AE5F951CFE46FB2CE12B07B0CE6A" box="[1181,1245,1119,1143]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.800, Anda4 (50)
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<collectionCode id="ED42AE5F951CFE46FF7CE1F70260CE86" box="[205,269,1155,1179]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.801, AndaDeVos (2)
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<collectionCode id="ED42AE5F951CFE46FD85E1F70119CE86" box="[564,628,1155,1179]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.803, AndaClif1 (1)
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<collectionCode id="ED42AE5F951CFE46FC3BE1F700A7CE86" box="[906,970,1155,1179]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.804, Tiep2 (3)
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<collectionCode id="ED42AE5F951CFE46FB1AE1F70786CE86" box="[1195,1259,1155,1179]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.805, Tiep3 (5)
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<collectionCode id="ED42AE5F951CFE46FF0FE1D30393CEA2" box="[190,254,1191,1215]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.822, Tiep4 (8)
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<collectionCode id="ED42AE5F951CFE46FE6CE1D30170CEA2" box="[477,541,1191,1215]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="12" pageNumber="209">RGM</collectionCode>
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961.823.
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</paragraph>
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<paragraph id="8BEC369A951CFE46FF76E1BE02D3CEFE" blockId="12.[151,1437,1046,2007]" box="[199,446,1226,1251]" pageId="12" pageNumber="209">
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<emphasis id="B927EA88951CFE46FF76E1BE0208CEFE" bold="true" box="[199,357,1226,1251]" pageId="12" pageNumber="209">
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<typeStatus id="54E88838951CFE46FF76E1BE026FCEFE" box="[199,258,1226,1251]" pageId="12" pageNumber="209">Type</typeStatus>
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locality.
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</emphasis>
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Anda1.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3496511951CFE46FF76E19A010ACF1A" box="[199,615,1262,1287]" pageId="12" pageNumber="209" type="etymology">
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<paragraph id="8BEC369A951CFE46FF76E19A010ACF1A" blockId="12.[151,1437,1046,2007]" box="[199,615,1262,1287]" pageId="12" pageNumber="209">
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<emphasis id="B927EA88951CFE46FF76E19A02C9CF1A" bold="true" box="[199,420,1262,1287]" pageId="12" pageNumber="209">Derivatio nominis.</emphasis>
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Pretty, adjective.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3496511951CFE46FF76E06600A3CF6E" pageId="12" pageNumber="209" type="diagnosis">
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<paragraph id="8BEC369A951CFE46FF76E06600A3CF6E" blockId="12.[151,1437,1046,2007]" pageId="12" pageNumber="209">
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<emphasis id="B927EA88951CFE46FF76E066022CCF36" bold="true" box="[199,321,1298,1323]" pageId="12" pageNumber="209">Diagnosis.</emphasis>
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Shell moderately large; H
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<quantity id="4CAB9B7F951CFE46FDC0E06601B5CF37" box="[625,728,1298,1323]" metricMagnitude="-3" metricUnit="m" metricValue="5.71" pageId="12" pageNumber="209" unit="mm" value="5.71">5.71 mm</quantity>
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,
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<geoCoordinate id="EE67505D951CFE46FD55E067005ACF36" box="[740,823,1298,1323]" direction="west" orientation="longitude" pageId="12" pageNumber="209" precision="555" value="-3.85">
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W
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<quantity id="4CAB9B7F951CFE46FCB7E066005ACF36" box="[774,823,1298,1323]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" pageId="12" pageNumber="209" unit="mm" value="3.85">3.85</quantity>
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</geoCoordinate>
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mm; protoconch slightly elevated, 1.2 whorl, DP
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<quantity id="4CAB9B7F951CFE46FADAE06703ABCF53" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="12" pageNumber="209" unit="mm" value="0.33">0.33 mm</quantity>
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; nucleus moderately small, DN
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<quantity id="4CAB9B7F951CFE46FD9EE04301FACF53" box="[559,663,1335,1359]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="209" unit="mm" value="0.17">0.17 mm</quantity>
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; teleoconch with spiral grooves; outer lip thick and crenulated; inner lip callused and with two nodules; columella callused and with two folds.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3496511951CFE44FF76E00A027ECAE5" lastPageId="14" lastPageNumber="211" pageId="12" pageNumber="209" type="description">
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<paragraph id="8BEC369A951CFE46FF76E00A06EDCCC6" blockId="12.[151,1437,1046,2007]" pageId="12" pageNumber="209">
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||
<emphasis id="B927EA88951CFE46FF76E00A0234CF8A" bold="true" box="[199,345,1406,1431]" pageId="12" pageNumber="209">Description.</emphasis>
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Shell rather high-spired, moderately large for its genus. Largest specimen has a total of 6.3 whorls, including a protoconch of 1.2 whorl. Protoconch rather small, smooth, weakly inclined. Teleoconch sculpted with thin, wavering spiral grooves and very weak, prosocyrt growth lines. Interspaces between spiral grooves decrease in width towards the base, whereas spiral grooves themselves increase in width towards the base. Spiral grooves intercalate along the whorls, increasing in number to 20–24 on the body whorl. Whorls convex, but not globose. Suture obscured by a narrow subsutural ridge, below which is a relatively broad groove. Aperture covered by thick callus, delineated by a rounded edge on the inner lip. Two, sometimes three rather small nodules present on parietal lip, lowermost nodule with a lamella extending into the aperture. Columellar lip bears two pronounced folds, lowermost fold crenulated. Outer lip thick, crenulated, with two subtle nodules present on the upper half of the lip. Exterior of the outer lip bears weak striations. Anterior and posterior sinus well-developed.
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</paragraph>
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<paragraph id="8BEC369A951CFE44FF76E392027ECAE5" blockId="12.[151,1437,1046,2007]" lastBlockId="14.[151,1436,151,248]" lastPageId="14" lastPageNumber="211" pageId="12" pageNumber="209">
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<emphasis id="B927EA88951CFE46FF76E3920212CCE2" bold="true" box="[199,383,1766,1791]" pageId="12" pageNumber="209">Differentiation.</emphasis>
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<taxonomicName id="4C534D19951CFE46FE34E392018CCCE2" authority="Nomura, 1939" authorityName="Nomura" authorityYear="1939" box="[389,737,1766,1791]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" higherTaxonomySource="CoL" kingdom="Animalia" order="Ellobiida" pageId="12" pageNumber="209" phylum="Mollusca" rank="species" species="niinoi">
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<emphasis id="B927EA88951CFE46FE34E3920154CCE3" box="[389,569,1766,1790]" italics="true" pageId="12" pageNumber="209">Ringicula niinoi</emphasis>
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<bibRefCitation id="EFC24B6B951CFE46FDF1E393018CCCE2" author="Nomura" box="[576,737,1767,1791]" pageId="12" pageNumber="209" refString="Nomura, S. (1939) Notes on some Opisthobranchiata based on the collection of the Saito Ho-on Kai Museum, chiefly collected from Northeast Honshu, Japan. Japanese Journal of Geology and Geography, 16, 11 - 27." type="journal article" year="1939">Nomura, 1939</bibRefCitation>
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</taxonomicName>
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is similar in its size and sculpture, but its spire is less elevated, its parietal lip bears only one, larger, nodule and its outer lip is less thick.
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<taxonomicName id="4C534D19951CFE46FBB0E27F0605CD3E" authority="Gould, 1860" authorityName="Gould" authorityYear="1860" box="[1025,1384,1802,1827]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" higherTaxonomySource="CoL" kingdom="Animalia" order="Ellobiida" pageId="12" pageNumber="209" phylum="Mollusca" rank="species" species="doliaris">
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<emphasis id="B927EA88951CFE46FBB0E27F07A2CD3E" box="[1025,1231,1803,1827]" italics="true" pageId="12" pageNumber="209">Ringicula doliaris</emphasis>
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<bibRefCitation id="EFC24B6B951CFE46FB68E27E0605CD3E" author="Gould" box="[1241,1384,1802,1827]" pageId="12" pageNumber="209" refString="Gould, A. A. (1860) Ringicula doliaris. In: Gould, A. A. (1859 - 1861) Descriptions of shells collected in the North Pacific Exploring Expedition under Captains Ringgold and Rodgers. Proceedings of the Boston Society for Natural History, 7, 323 - 336." type="journal article" year="1860">Gould, 1860</bibRefCitation>
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</taxonomicName>
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is a variable, not well-described species. The specimens figured by
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<bibRefCitation id="EFC24B6B951CFE46FCD6E25A0769CD5A" author="Habe" box="[871,1028,1838,1863]" pageId="12" pageNumber="209" refString="Habe, T. (1950 a) Ringiculidae and Retusidae (1) in Japan. Illustrated Catalogue of Japanese Shells, 2, 7 - 16." type="journal article" year="1950" yearSuffix="a">Habe (1950a)</bibRefCitation>
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,
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<bibRefCitation id="EFC24B6B951CFE46FBA0E25A07FACD5A" author="Hori" box="[1041,1175,1838,1863]" pageId="12" pageNumber="209" refString="Hori, S. (2000) Family Acteonidae. In: Okutani, T. (ed.), Marine Mollusks in Japan. Tokai University Press, Tokyo, pp. 732 - 735." type="book chapter" year="2000">Hori (2000)</bibRefCitation>
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and
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<bibRefCitation id="EFC24B6B951CFE46FB61E25A06F1CD5A" box="[1232,1436,1838,1863]" pageId="12" pageNumber="209" refString="Higo, S., Callomon, P. & Goto, Y. (2001) Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan, Types. Elle Scientific Publications, Osaka, 1 - 208." type="book chapter">
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Higo
|
||
<emphasis id="B927EA88951CFE46FAA1E25B0624CD5B" box="[1296,1353,1838,1862]" italics="true" pageId="12" pageNumber="209">et al.</emphasis>
|
||
(2001)
|
||
</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFC24B6B951CFE46FF2FE2260205CD76" box="[158,360,1874,1899]" pageId="12" pageNumber="209" refString="Higo, S., Callomon, P. & Goto, Y. (2001) Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan, Types. Elle Scientific Publications, Osaka, 1 - 208." type="book chapter">
|
||
Higo
|
||
<emphasis id="B927EA88951CFE46FF6EE227027BCD76" box="[223,278,1875,1899]" italics="true" pageId="12" pageNumber="209">et al.</emphasis>
|
||
(2001)
|
||
</bibRefCitation>
|
||
figured the
|
||
<typeStatus id="54E88838951CFE46FE41E2260139CD76" box="[496,596,1874,1899]" pageId="12" pageNumber="209" type="holotype">holotype</typeStatus>
|
||
) are slightly smaller, with a slightly less elevated spire, their spiral grooves are further apart, they have only one nodule on their parietal lip, their callus extends further onto the base and their outer lip is less thickened and smooth. The specimens figured by Willan (2010) are similar in shape, but smaller (height only
|
||
<quantity id="4CAB9B7F951CFE46FE97E2CB02F1CDCA" box="[294,412,1983,2007]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="12" pageNumber="209" unit="mm" value="4.2" valueMax="4.4" valueMin="4.0">4–4.4 mm</quantity>
|
||
), their spiral grooves are much further apart and they have only one nodule on their parietal lip.
|
||
<taxonomicName id="4C534D19951EFE44FF72E5EC0154CAAD" authority="Habe, 1950" authorityName="Habe" authorityYear="1950" box="[195,569,151,176]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" higherTaxonomySource="CoL" kingdom="Animalia" order="Ellobiida" pageId="14" pageNumber="211" phylum="Mollusca" rank="species" species="teramachii">
|
||
<emphasis id="B927EA88951EFE44FF72E5EC02DDCAAD" box="[195,432,152,176]" italics="true" pageId="14" pageNumber="211">Ringicula teramachii</emphasis>
|
||
Habe, 1950
|
||
</taxonomicName>
|
||
is similar in its shape and sculpture, but slightly smaller (
|
||
<typeStatus id="54E88838951EFE44FB70E5E30648CAAD" box="[1217,1317,151,176]" pageId="14" pageNumber="211" type="holotype">holotype</typeStatus>
|
||
height
|
||
<quantity id="4CAB9B7F951EFE44FAC8E5EC03AACAC9" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="14" pageNumber="211" unit="mm" value="4.9">4.9 mm</quantity>
|
||
), its callus on the inner lip is thin and lacks a nodule and its outer lip is also much less thickened and not crenulated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |