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<document id="E543BD1EC8CD1A7CBC2BAA5A82D780C6" ID-CLB-Dataset="45956" ID-DOI="10.11646/zootaxa.3790.2.4" ID-GBIF-Dataset="fb357b0e-637b-4303-9414-b6d2809daff8" ID-ISSN="1175-5326" ID-Zenodo-Dep="29899" ID-ZooBank="BDA1931C-FEDB-4142-8A63-2765593621A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460411323498" checkinUser="plazi" docAuthor="Jäger, Peter" docDate="2014" docId="03DA87EFFFA63975FF69FA6C85A51918" docLanguage="en" docName="zt03790p356.pdf" docOrigin="Zootaxa 3790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cebrennus rungsi Jager 2000" docType="treatment" docVersion="10" lastPageNumber="326" masterDocId="FFE3FF97FFA33972FFFEFFEE86631D40" masterDocTitle="Cebrennus Simon, 1880 (Araneae: Sparassidae): a revisionary up-date with the description of four new species and an updated identification key for all species" masterLastPageNumber="356" masterPageNumber="319" pageNumber="324" updateTime="1698300605956" updateUser="ExternalLinkService">
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<mods:title id="42D685449A855C39C0CDE0AACBA7BA1A">Cebrennus Simon, 1880 (Araneae: Sparassidae): a revisionary up-date with the description of four new species and an updated identification key for all species</mods:title>
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<treatment id="03DA87EFFFA63975FF69FA6C85A51918" ID-DOI="http://doi.org/10.5281/zenodo.6124551" ID-GBIF-Taxon="119423555" ID-Zenodo-Dep="6124551" LSID="urn:lsid:plazi:treatment:03DA87EFFFA63975FF69FA6C85A51918" httpUri="http://treatment.plazi.org/id/03DA87EFFFA63975FF69FA6C85A51918" lastPageId="7" lastPageNumber="326" pageId="5" pageNumber="324">
<subSubSection id="C3696572FFA63977FF69FA6C87C318FE" pageId="5" pageNumber="324" type="nomenclature">
<paragraph id="8BCC36F9FFA63977FF69FA6C847418DD" blockId="5.[151,535,1410,1470]" box="[151,535,1410,1437]" pageId="5" pageNumber="324">
<heading id="D0848195FFA63977FF69FA6C847418DD" bold="true" box="[151,535,1410,1437]" fontSize="11" level="1" pageId="5" pageNumber="324" reason="1">
<taxonomicName id="4C734D7AFFA63977FF69FA6C847418DD" ID-CoL="RYYG" authority="Jager, 2000" authorityName="Jager" authorityYear="2000" box="[151,535,1410,1437]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="324" phylum="Arthropoda" rank="species" species="rungsi">
<emphasis id="B907EAEBFFA63977FF69FA6C847418DD" bold="true" box="[151,535,1410,1437]" pageId="5" pageNumber="324">
<emphasis id="B907EAEBFFA63977FF69FA6C871318DC" bold="true" box="[151,368,1410,1436]" italics="true" pageId="5" pageNumber="324">Cebrennus rungsi</emphasis>
(
<bibRefCitation id="EFE24B08FFA63977FE81FA6D846C18DD" author="Jager" box="[383,527,1411,1437]" pageId="5" pageNumber="324" refString="Jager, P. (2000) The huntsman spider genus Cebrennus: four new species and a preliminary key to known species (Araneae: Sparassidae: Sparassinae). Revue Arachnologique, 13, 163 - 186." type="journal article" year="2000">Jäger, 2000</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCC36F9FFA63977FF69FA4B87C318FE" blockId="5.[151,535,1410,1470]" box="[151,416,1445,1470]" pageId="5" pageNumber="324">
<figureCitation id="13482A7CFFA63977FF69FA4B877518FE" box="[151,278,1445,1470]" captionStart="FIGURES 12 17. 12 13" captionStartId="6.[151,264,1915,1938]" captionTargetBox="[237,1386,200,1893]" captionTargetId="figure@6.[200,1387,193,1894]" captionTargetPageId="6" captionText="FIGURES 12 17. 12 13 Cebrennus rungsi Jäger, 2000, males from Anti-Atlas, Morocco; 14 Cebrennus spec. A, male from Beni Abbes Algeria; 15 17 Cebrennus castaneitarsis Simon, 1880, males (15 Algeria, 16 17 holotype male from Wahran, Algeria). Left male palp (12 16 ventral, 17 retrolateral). DE—distal part of embolus, K—embolic kink, PE—proximal part of embolus, RTA—retrolateral tibial apophysis." httpUri="https://zenodo.org/record/398876/files/figure.png" pageId="5" pageNumber="324">Figs 1213</figureCitation>
,
<figureCitation id="13482A7CFFA63977FEDCFA48870818FE" box="[290,363,1446,1470]" captionStart="FIGURES 88 100" captionStartId="23.[151,264,1906,1929]" captionTargetBox="[202,1368,204,1861]" captionTargetId="figure@23.[201,1369,193,1872]" captionTargetPageId="23" captionText="FIGURES 88 100. Cebrennus spp., habitus (photos in ethanol; 88, 92, 95, 97 dorsal, 89, 98 ventral, 91, 94, 100 frontal, 90, 93, 96, 99 lateral). 88 91 Cebrennus kochi (O. Pickard-Cambridge, 1872), female from Sede Boquer, Israel; 92 93 Cebrennus rungsi Jäger, 2000, male from Anti-Atlas, Morocco; 94 96 Cebrennus atlas spec. nov., holotype female from Atlas, Morocco; 97 100 Cebrennus flagellatus spec. nov., holotype male from Afghanistan." httpUri="https://zenodo.org/record/398886/files/figure.png" pageId="5" pageNumber="324">9293</figureCitation>
,
<figureCitation id="13482A7CFFA63977FE88FA4887C318FE" box="[374,416,1446,1470]" captionStart="FIGURE 173" captionStartId="35.[151,249,736,758]" captionTargetBox="[151,1436,225,688]" captionTargetId="figure@35.[151,1436,225,688]" captionTargetPageId="35" captionText="FIGURE 173. Distribution records of Cebrennus spp. (according to Jäger 2000 and references therein, this paper and Reimoser 1919). Note that the exact locality for C. laurae spec. nov. is unknown." httpUri="https://zenodo.org/record/398898/files/figure.png" pageId="5" pageNumber="324">173</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3696572FFA63977FF69FA0385AB1B43" box="[151,968,1517,1539]" pageId="5" pageNumber="324" type="reference_group">
<paragraph id="8BCC36F9FFA63977FF69FA0385AB1B43" blockId="5.[151,968,1517,1539]" box="[151,968,1517,1539]" pageId="5" pageNumber="324">
<treatmentCitationGroup id="AB6311D7FFA63977FF69FA0385AB1B43" box="[151,968,1517,1539]" pageId="5" pageNumber="324">
<taxonomicName id="4C734D7AFFA63977FF69FA0387991B43" ID-CoL="RYYG" authority="Jager, 2000: 172" authorityName="Jager" authorityPageNumber="172" authorityYear="2000" box="[151,506,1517,1539]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="324" phylum="Arthropoda" rank="species" species="rungsi">
<emphasis id="B907EAEBFFA63977FF69FA0387291B43" box="[151,330,1517,1539]" italics="true" pageId="5" pageNumber="324">Cebrennus rungsi</emphasis>
<treatmentCitation id="0AD210E8FFA63977FEAFFA0087991B43" author="Jager" box="[337,506,1517,1539]" page="172" pageId="5" pageNumber="324" year="2000">
<bibRefCitation id="EFE24B08FFA63977FEAFFA0087AA1B43" author="Jager" box="[337,457,1517,1539]" pageId="5" pageNumber="324" refString="Jager, P. (2000) The huntsman spider genus Cebrennus: four new species and a preliminary key to known species (Araneae: Sparassidae: Sparassinae). Revue Arachnologique, 13, 163 - 186." type="journal article" year="2000">Jäger, 2000</bibRefCitation>
: 172
</treatmentCitation>
</taxonomicName>
, figs 3441 (description of male and female).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3696572FFA63977FF69F9C183201A60" pageId="5" pageNumber="324" type="materials_examined">
<paragraph id="8BCC36F9FFA63977FF69F9C183201A60" blockId="5.[151,1436,1583,2005]" pageId="5" pageNumber="324">
<emphasis id="B907EAEBFFA63977FF69F9C185741B08" bold="true" box="[151,791,1583,1608]" pageId="5" pageNumber="324">
Material examined.
<collectingCountry id="F3647669FFA63977FE77F9C184451B08" box="[393,550,1583,1608]" name="Morocco" pageId="5" pageNumber="324">MOROCCO</collectingCountry>
:
<emphasis id="B907EAEBFFA63977FDCCF9C185741B08" bold="true" box="[562,791,1583,1608]" italics="true" pageId="5" pageNumber="324">Souss-Massa-Drâa:</emphasis>
</emphasis>
<specimenCount id="9D75FD70FFA63977FCE1F9DE85091B08" box="[799,874,1583,1608]" pageId="5" pageNumber="324" type="male">1 male</specimenCount>
(PJ 3391),
<collectingCountry id="F3647669FFA63977FC10F9C1825D1B08" box="[1006,1086,1583,1608]" name="Morocco" pageId="5" pageNumber="324">Agadir</collectingCountry>
, Hering leg.
<date id="FFCD1039FFA63977FB2EF9DE83FF1B08" box="[1232,1436,1583,1608]" pageId="5" pageNumber="324" value="1986-11-02">2 November 1986</date>
(
<collectionCode id="ED62AE3CFFA63977FF61F9BA86821B2C" box="[159,225,1620,1644]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="5" pageNumber="324">ZMB</collectionCode>
);
<specimenCount id="9D75FD70FFA63977FF06F9BA87241B2D" box="[248,327,1620,1645]" pageId="5" pageNumber="324" type="male">1 male</specimenCount>
(PJ 3396, SD 645), Anti-Atlas Range, Ighil, ca. 110 air km ESE of
<collectingCountry id="F3647669FFA63977FB91F9BA82DC1B2D" box="[1135,1215,1620,1645]" name="Morocco" pageId="5" pageNumber="324">Agadir</collectingCountry>
,
<geoCoordinate id="EE47503EFFA63977FB33F9BB83111B2C" box="[1229,1394,1620,1644]" direction="north" orientation="latitude" pageId="5" pageNumber="324" precision="1" value="30.14166">N 30°8'29.98&quot;</geoCoordinate>
,
<geoCoordinate id="EE47503EFFA63977FA7CF9BB876C1BD0" direction="west" orientation="longitude" pageId="5" pageNumber="324" precision="1" value="-8.485211">W 8°29'6.76&quot;</geoCoordinate>
, [
<quantity id="4C8B9B1CFFA63977FEDBF999871C1BCF" box="[293,383,1655,1680]" metricMagnitude="3" metricUnit="m" metricValue="1.584" pageId="5" pageNumber="324" unit="m" value="1584.0">1584 m</quantity>
elev.], point 54, desert, S. Henriques leg.
<date id="FFCD1039FFA63977FC96F99682251BD0" box="[872,1094,1655,1680]" pageId="5" pageNumber="324" value="2011-09-29">29 September 2011</date>
(
<collectionCode id="ED62AE3CFFA63977FBA6F99682F51BD0" LSID="urn:lsid:biocol.org:col:34838" box="[1112,1174,1656,1680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="324">SMF</collectionCode>
);
<specimenCount id="9D75FD70FFA63977FB55F996829A1BD0" box="[1195,1273,1655,1680]" pageId="5" pageNumber="324" type="male">1 male</specimenCount>
(PJ 3397, SD 646), Anti-Atlas Range, between Sdass and Ouaoufenrha, ca. 98 air km E of
<collectingCountry id="F3647669FFA63977FBABF97282C61BF5" box="[1109,1189,1692,1717]" name="Morocco" pageId="5" pageNumber="324">Agadir</collectingCountry>
,
<geoCoordinate id="EE47503EFFA63977FB49F973830D1BF5" box="[1207,1390,1692,1717]" direction="north" orientation="latitude" pageId="5" pageNumber="324" precision="1" value="30.35404">N 30°21'14.54&quot;</geoCoordinate>
,
<geoCoordinate id="EE47503EFFA63977FA7CF97387781B98" direction="west" orientation="longitude" pageId="5" pageNumber="324" precision="1" value="-8.577489">W 8°34'38.96&quot;</geoCoordinate>
[
<quantity id="4C8B9B1CFFA63977FED5F92E87E61B97" box="[299,389,1728,1752]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="324" unit="m" value="1000.0">1000 m</quantity>
elev.], point 51, desert, S. Henriques leg.
<date id="FFCD1039FFA63977FC95F92E822B1B98" box="[875,1096,1727,1752]" pageId="5" pageNumber="324" value="2011-09-29">29 September 2011</date>
(
<collectionCode id="ED62AE3CFFA63977FBA7F92E82F41B98" LSID="urn:lsid:biocol.org:col:34838" box="[1113,1175,1728,1752]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="324">SMF</collectionCode>
);
<specimenCount id="9D75FD70FFA63977FB53F92E82991B98" box="[1197,1274,1727,1752]" pageId="5" pageNumber="324" type="male">1 male</specimenCount>
(PJ 3398, SD 647),
<specimenCount id="9D75FD70FFA63977FF26F90A87471BBD" box="[216,292,1764,1789]" pageId="5" pageNumber="324" type="male">1 male</specimenCount>
(PJ 3399, SD 648), Anti-Atlas Range, between Irherm and Armdaz, ca. 115 air km ESE of
<collectingCountry id="F3647669FFA63977FAD3F90A831F1BBD" box="[1325,1404,1764,1789]" name="Morocco" pageId="5" pageNumber="324">Agadir</collectingCountry>
,
<geoCoordinate id="EE47503EFFA63977FA76F90B87461A60" direction="north" orientation="latitude" pageId="5" pageNumber="324" precision="1" value="30.054869">N 30° 3'17.53&quot;</geoCoordinate>
,
<geoCoordinate id="EE47503EFFA63977FECEF8E687B51A60" box="[304,470,1799,1824]" direction="west" orientation="longitude" pageId="5" pageNumber="324" precision="1" value="-8.4636">W 8°27'48.96&quot;</geoCoordinate>
[
<quantity id="4C8B9B1CFFA63977FE1AF8E9845F1A5F" box="[484,572,1799,1824]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="5" pageNumber="324" unit="m" value="1640.0">1640 m</quantity>
elev.], point 48, desert, S. Henriques leg.
<date id="FFCD1039FFA63977FBEFF8E682891A60" box="[1041,1258,1799,1824]" pageId="5" pageNumber="324" value="2011-09-29">29 September 2011</date>
(
<collectionCode id="ED62AE3CFFA63977FB07F8E683541A60" LSID="urn:lsid:biocol.org:col:34838" box="[1273,1335,1800,1824]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="324">SMF</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3696572FFA63975FF39F8C283031D95" lastPageId="7" lastPageNumber="326" pageId="5" pageNumber="324" type="description">
<paragraph id="8BCC36F9FFA63977FF39F8C284411A04" blockId="5.[151,1436,1583,2005]" box="[199,546,1836,1861]" pageId="5" pageNumber="324">
<emphasis id="B907EAEBFFA63977FF39F8C2873A1A05" bold="true" box="[199,345,1836,1861]" pageId="5" pageNumber="324">Description.</emphasis>
See
<bibRefCitation id="EFE24B08FFA63977FE70F8C3847D1A04" author="Jager" box="[398,542,1836,1861]" pageId="5" pageNumber="324" refString="Jager, P. (2000) The huntsman spider genus Cebrennus: four new species and a preliminary key to known species (Araneae: Sparassidae: Sparassinae). Revue Arachnologique, 13, 163 - 186." type="journal article" year="2000">Jäger (2000)</bibRefCitation>
.
</paragraph>
<paragraph id="8BCC36F9FFA63977FF39F8A183E91A95" blockId="5.[151,1436,1583,2005]" pageId="5" pageNumber="324">
<emphasis id="B907EAEBFFA63977FF39F8A187581A28" bold="true" box="[199,315,1871,1896]" pageId="5" pageNumber="324">Variation.</emphasis>
Males (n=4): PL 5.46.5, PW 4.75.5, AW 3.03.9, OL 5.37.2, OW 3.55.0; chelicerae with 2 anterior, 58 adnate posterior teeth, and 23 bristles at the posterior distal margin close to fang base. Structures of the copulatory organ correspond in most cases fully with those of the
<typeStatus id="54C8885BFFA63977FC3FF87682441AF1" box="[961,1063,1944,1969]" pageId="5" pageNumber="324" type="holotype">holotype</typeStatus>
. In the two males from between Irherm and Armdaz (
<figureCitation id="13482A7CFFA63977FE6FF85284731A94" box="[401,528,1980,2005]" captionStart="FIGURES 12 17. 12 13" captionStartId="6.[151,264,1915,1938]" captionTargetBox="[237,1386,200,1893]" captionTargetId="figure@6.[200,1387,193,1894]" captionTargetPageId="6" captionText="FIGURES 12 17. 12 13 Cebrennus rungsi Jäger, 2000, males from Anti-Atlas, Morocco; 14 Cebrennus spec. A, male from Beni Abbes Algeria; 15 17 Cebrennus castaneitarsis Simon, 1880, males (15 Algeria, 16 17 holotype male from Wahran, Algeria). Left male palp (12 16 ventral, 17 retrolateral). DE—distal part of embolus, K—embolic kink, PE—proximal part of embolus, RTA—retrolateral tibial apophysis." httpUri="https://zenodo.org/record/398876/files/figure.png" pageId="5" pageNumber="324">Figs 1213</figureCitation>
) the distal part of the embolus was distinctly shorter than that of the
<typeStatus id="54C8885BFFA63977FADEF85283E51A95" box="[1312,1414,1980,2005]" pageId="5" pageNumber="324" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DF0C6671FFA53974FF69F895843D1AAD" httpUri="https://zenodo.org/record/398876/files/figure.png" pageId="6" pageNumber="325" targetBox="[237,1386,200,1893]" targetPageId="6">
<paragraph id="8BCC36F9FFA53974FF69F895843D1AAD" blockId="6.[151,1436,1915,2029]" pageId="6" pageNumber="325">
<emphasis id="B907EAEBFFA53974FF69F89587361AD0" bold="true" box="[151,341,1915,1938]" pageId="6" pageNumber="325">FIGURES 1217.</emphasis>
1213
<taxonomicName id="4C734D7AFFA53974FE5DF89584B41AD1" authority="Jager, 2000" authorityName="Jager" authorityYear="2000" box="[419,727,1915,1937]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="rungsi">
<emphasis id="B907EAEBFFA53974FE5DF895843B1AD1" box="[419,600,1915,1937]" italics="true" pageId="6" pageNumber="325">Cebrennus rungsi</emphasis>
Jäger, 2000
</taxonomicName>
, males from Anti-Atlas, Morocco; 14
<taxonomicName id="4C734D7AFFA53974FB9CF89582AC1AD1" box="[1122,1231,1915,1937]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAEBFFA53974FB9CF89582AC1AD1" box="[1122,1231,1915,1937]" italics="true" pageId="6" pageNumber="325">Cebrennus</emphasis>
</taxonomicName>
spec. A, male from Beni Abbes Algeria; 1517
<taxonomicName id="4C734D7AFFA53974FE40F87485331AF0" authority="Simon, 1880" authorityName="Simon" authorityYear="1880" box="[446,848,1946,1968]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="325" phylum="Arthropoda" rank="species" species="castaneitarsis">
<emphasis id="B907EAEBFFA53974FE40F87484A21AF0" box="[446,705,1946,1968]" italics="true" pageId="6" pageNumber="325">Cebrennus castaneitarsis</emphasis>
Simon, 1880
</taxonomicName>
, males (15 Algeria, 1617 holotype male from Wahran, Algeria). Left male palp (1216 ventral, 17 retrolateral). DE—distal part of embolus, K—embolic kink, PE—proximal part of embolus, RTA—retrolateral tibial apophysis.
</paragraph>
</caption>
<paragraph id="8BCC36F9FFA43975FF69FF7983031D95" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="326">
Since all other characters (shape and size of RTA, shape of tegulum, reduced conductor and basal embolus, shape of cymbium) are congruent with the
<typeStatus id="54C8885BFFA43975FDCFFF5284F51D95" box="[561,662,188,213]" pageId="7" pageNumber="326" type="holotype">holotype</typeStatus>
, the shorter distal embolus is considered intraspecific variation.
</paragraph>
</subSubSection>
<subSubSection id="C3696572FFA43975FF39FF3185A51918" pageId="7" pageNumber="326" type="distribution">
<paragraph id="8BCC36F9FFA43975FF39FF3185F81C5C" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="326">
<emphasis id="B907EAEBFFA43975FF39FF3187031DB8" bold="true" box="[199,352,223,248]" pageId="7" pageNumber="326">Distribution.</emphasis>
All known records (including the
<typeStatus id="54C8885BFFA43975FD1BFF0E85761DB8" box="[741,789,224,248]" pageId="7" pageNumber="326">type</typeStatus>
locality Sous) lie in the West of the region Souss-Massa- Draâ in an altitudinal range from sea level to 1650 metres (
<figureCitation id="13482A7CFFA43975FCD2FEEA85ED1C5C" box="[812,910,260,285]" captionStart="FIGURE 173" captionStartId="35.[151,249,736,758]" captionTargetBox="[151,1436,225,688]" captionTargetId="figure@35.[151,1436,225,688]" captionTargetPageId="35" captionText="FIGURE 173. Distribution records of Cebrennus spp. (according to Jäger 2000 and references therein, this paper and Reimoser 1919). Note that the exact locality for C. laurae spec. nov. is unknown." httpUri="https://zenodo.org/record/398898/files/figure.png" pageId="7" pageNumber="326">Fig. 173</figureCitation>
).
</paragraph>
<paragraph id="8BCC36F9FFA43975FF69FE8185A51918" blockId="7.[151,1437,367,1112]" pageId="7" pageNumber="326">
The following four species (
<taxonomicName id="4C734D7AFFA43975FE2FFE9F840E1CC8" box="[465,621,369,392]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="castaneitarsis">
<emphasis id="B907EAEBFFA43975FE2FFE9F840E1CC8" box="[465,621,369,392]" italics="true" pageId="7" pageNumber="326">castaneitarsis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C734D7AFFA43975FD84FE9F84A01CC8" box="[634,707,369,392]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="wagae">
<emphasis id="B907EAEBFFA43975FD84FE9F84A01CC8" box="[634,707,369,392]" italics="true" pageId="7" pageNumber="326">wagae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C734D7AFFA43975FD2FFE8185311CC8" box="[721,850,367,392]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="B907EAEBFFA43975FD2FFE8185311CC8" box="[721,850,367,392]" italics="true" pageId="7" pageNumber="326">aethiopicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C734D7AFFA43975FCA1FE8185B61CC8" box="[863,981,367,392]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="flagellatus" status="sp. nov.">
<emphasis id="B907EAEBFFA43975FCA1FE8185B61CC8" box="[863,981,367,392]" italics="true" pageId="7" pageNumber="326">flagellatus</emphasis>
</taxonomicName>
<emphasis id="B907EAEBFFA43975FC22FE81822E1CC8" bold="true" box="[988,1101,367,392]" pageId="7" pageNumber="326">
<taxonomicNameLabel id="A2345790FFA43975FC22FE81822E1CC8" box="[988,1101,367,392]" pageId="7" pageNumber="326" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
) have very similar copulatory organs, differentiation of the species is in some cases difficult. Apparently, the male emboli have been elongated in the course of evolution as hypothesised by
<bibRefCitation id="EFE24B08FFA43975FD47FE5685311C90" author="Jager" box="[697,850,439,464]" pageId="7" pageNumber="326" refString="Jager, P. (2006) Lengthening of embolus and intromittent duct: a review of an evolutionary trend in the spider family Sparassidae (Arachnida: Araneae). In: Deltshev, C. &amp; Stoev, P. (Eds.), European Arachnology 2005. Acta zoologica bulgarica, Suppl. No 1, pp. 49 - 62." type="journal article" year="2006">Jäger (2006)</bibRefCitation>
. The same is true for the so-called “combined morphological changes” as the elongation of the RTA or the reduction of the palpal tibia length. In general, males are easier to identify than females. Easiest to distinguish is
<taxonomicName id="4C734D7AFFA43975FCCDFDEE85A91F58" box="[819,970,511,536]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="flagellatus" status="sp. nov.">
<emphasis id="B907EAEBFFA43975FCCDFDEE85A91F58" box="[819,970,511,536]" italics="true" pageId="7" pageNumber="326">C. flagellatus</emphasis>
</taxonomicName>
<emphasis id="B907EAEBFFA43975FC2CFE1182261F58" bold="true" box="[978,1093,511,536]" pageId="7" pageNumber="326">
<taxonomicNameLabel id="A2345790FFA43975FC2CFE1182261F58" box="[978,1093,511,536]" pageId="7" pageNumber="326" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(only known by the male sex) by its flagellum at the embolic kink and the distal RTA at right angles with the proximal part (
<figureCitation id="13482A7CFFA43975FB5BFDCA834B1F7D" box="[1189,1320,548,573]" captionStart="FIGURES 30 35. 30 31" captionStartId="14.[151,264,1777,1800]" captionTargetBox="[151,1384,238,1741]" captionTargetId="figure@14.[151,1436,195,1756]" captionTargetPageId="14" captionText="FIGURES 30 35. 30 31 Cebrennus aethiopicus Simon, 1880, male from Sudan; 32 35 Cebrennus flagellatus spec. nov., holotype male from Nangarhar Province, E Afghanistan. 30 33 Left male palp (30, 32 ventral, 31, 33 retrolateral); 34 Embolic kink showing flagellum, proximal; 35 Embolus tip, ventral. F—flagellum at embolus kink." httpUri="https://zenodo.org/record/398879/files/figure.png" pageId="7" pageNumber="326">Figs 3235</figureCitation>
). Males of
<taxonomicName id="4C734D7AFFA43975FF69FDA687371F20" box="[151,340,584,608]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="castaneitarsis">
<emphasis id="B907EAEBFFA43975FF69FDA687371F20" box="[151,340,584,608]" italics="true" pageId="7" pageNumber="326">C. castaneitarsis</emphasis>
</taxonomicName>
can be distinguished from those of the three other species by the shorter embolus, i.e. the arising point situated distinctly in prolateral position of the tegulum and the proximal part of the distal loop does not, or only barely so, extend beyond the tegulum proximally (
<figureCitation id="13482A7CFFA43975FCD0FD6185D61FE8" box="[814,949,655,680]" captionStart="FIGURES 12 17. 12 13" captionStartId="6.[151,264,1915,1938]" captionTargetBox="[237,1386,200,1893]" captionTargetId="figure@6.[200,1387,193,1894]" captionTargetPageId="6" captionText="FIGURES 12 17. 12 13 Cebrennus rungsi Jäger, 2000, males from Anti-Atlas, Morocco; 14 Cebrennus spec. A, male from Beni Abbes Algeria; 15 17 Cebrennus castaneitarsis Simon, 1880, males (15 Algeria, 16 17 holotype male from Wahran, Algeria). Left male palp (12 16 ventral, 17 retrolateral). DE—distal part of embolus, K—embolic kink, PE—proximal part of embolus, RTA—retrolateral tibial apophysis." httpUri="https://zenodo.org/record/398876/files/figure.png" pageId="7" pageNumber="326">Figs 1517</figureCitation>
); moreover the RTA is as short as in
<taxonomicName id="4C734D7AFFA43975FA7CFD7E876E1F8D" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="flagellatus" status="sp. nov.">
<emphasis id="B907EAEBFFA43975FA7CFD7E876E1F8D" italics="true" pageId="7" pageNumber="326">C. flagellatus</emphasis>
</taxonomicName>
<emphasis id="B907EAEBFFA43975FEE9FD5A87EF1F8D" bold="true" box="[279,396,692,717]" pageId="7" pageNumber="326">
<taxonomicNameLabel id="A2345790FFA43975FEE9FD5A87EF1F8D" box="[279,396,692,717]" pageId="7" pageNumber="326" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, but builds a larger angle with the distal tibia.
<taxonomicName id="4C734D7AFFA43975FC4EFD5A82181F8C" box="[944,1147,692,717]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="wagae">
<emphasis id="B907EAEBFFA43975FC4EFD5A82181F8C" box="[944,1147,692,717]" italics="true" pageId="7" pageNumber="326">Cebrennus wagae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D7AFFA43975FB49FD5B833F1F8D" box="[1207,1372,692,717]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="B907EAEBFFA43975FB49FD5B833F1F8D" box="[1207,1372,692,717]" italics="true" pageId="7" pageNumber="326">C. aethiopicus</emphasis>
</taxonomicName>
were differentiated by
<bibRefCitation id="EFE24B08FFA43975FEA8FD3687BE1FB0" author="Fage" box="[342,477,728,752]" pageId="7" pageNumber="326" refString="Fage, L. (1921) Remarques sur les araignees du genre Cebrennus suivies de la description de deux especes nouvelles. Bulletin de la Societe zoologique de France, 46, 157 - 166." type="journal article" year="1921">Fage (1921)</bibRefCitation>
through the absence (
<taxonomicName id="4C734D7AFFA43975FD2DFD37857F1FB0" box="[723,796,729,752]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="wagae">
<emphasis id="B907EAEBFFA43975FD2DFD37857F1FB0" box="[723,796,729,752]" italics="true" pageId="7" pageNumber="326">wagae</emphasis>
</taxonomicName>
) or presence (
<taxonomicName id="4C734D7AFFA43975FC45FD39825F1FB0" box="[955,1084,727,752]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="B907EAEBFFA43975FC45FD39825F1FB0" box="[955,1084,727,752]" italics="true" pageId="7" pageNumber="326">aethiopicus</emphasis>
</taxonomicName>
) of a distal spine on metatarsus III. Although there were two of three specimens of
<taxonomicName id="4C734D7AFFA43975FD08FD1385F81E55" box="[758,923,764,789]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="B907EAEBFFA43975FD08FD1385F81E55" box="[758,923,764,789]" italics="true" pageId="7" pageNumber="326">C. aethiopicus</emphasis>
</taxonomicName>
examined with a spination formula 3034 at metatarsus III, one had the usual 2024. Therefore, the spination cannot act as single diagnostic character.
<bibRefCitation id="EFE24B08FFA43975FAADFCF187311E1D" author="Simon" pageId="7" pageNumber="326" refString="Simon, E. (1880) Revision de la famille des Sparassidae (Arachnides). Actes de la Societe linneenne de Bordeaux, 34, 223 - 351." type="journal article" year="1880">Simon (1880: 331, key)</bibRefCitation>
used mainly the RTA as differentiating character and his distinctions are still valid:
<taxonomicName id="4C734D7AFFA43975FAFBFCAB830C1E1C" box="[1285,1391,837,860]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="wagae">
<emphasis id="B907EAEBFFA43975FAFBFCAB830C1E1C" box="[1285,1391,837,860]" italics="true" pageId="7" pageNumber="326">C. wagae</emphasis>
</taxonomicName>
has the longest RTA (
<figureCitation id="13482A7CFFA43975FE9DFC89878B1EC0" box="[355,488,871,896]" captionStart="FIGURES 18 23" captionStartId="10.[151,264,1960,1983]" captionTargetBox="[154,1436,193,1939]" captionTargetId="figure@10.[151,1436,193,1940]" captionTargetPageId="10" captionText="FIGURES 18 23. Cebrennus wagae (Simon, 1874), males (18 Libya, 19 20 Bou Saada, Algeria, 21 23 Chambi, Tunisia). Left male palp (18 19, 22 ventral, 20, 23 retrolateral, 21 prolateral). EB—embolus base." httpUri="https://zenodo.org/record/398877/files/figure.png" pageId="7" pageNumber="326">Figs 1823</figureCitation>
),
<taxonomicName id="4C734D7AFFA43975FE00FC8684DD1EC0" box="[510,702,872,896]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="castaneitarsis">
<emphasis id="B907EAEBFFA43975FE00FC8684DD1EC0" box="[510,702,872,896]" italics="true" pageId="7" pageNumber="326">C. castaneitarsis</emphasis>
</taxonomicName>
an almost straight RTA (
<figureCitation id="13482A7CFFA43975FC1DFC89820B1EC0" box="[995,1128,871,896]" captionStart="FIGURES 12 17. 12 13" captionStartId="6.[151,264,1915,1938]" captionTargetBox="[237,1386,200,1893]" captionTargetId="figure@6.[200,1387,193,1894]" captionTargetPageId="6" captionText="FIGURES 12 17. 12 13 Cebrennus rungsi Jäger, 2000, males from Anti-Atlas, Morocco; 14 Cebrennus spec. A, male from Beni Abbes Algeria; 15 17 Cebrennus castaneitarsis Simon, 1880, males (15 Algeria, 16 17 holotype male from Wahran, Algeria). Left male palp (12 16 ventral, 17 retrolateral). DE—distal part of embolus, K—embolic kink, PE—proximal part of embolus, RTA—retrolateral tibial apophysis." httpUri="https://zenodo.org/record/398876/files/figure.png" pageId="7" pageNumber="326">Figs 1517</figureCitation>
),
<taxonomicName id="4C734D7AFFA43975FB81FC8683401EC0" box="[1151,1315,871,896]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="B907EAEBFFA43975FB81FC8683401EC0" box="[1151,1315,871,896]" italics="true" pageId="7" pageNumber="326">C. aethiopicus</emphasis>
</taxonomicName>
exhibits a bend at the base of the RTA (especially in comparison to
<taxonomicName id="4C734D7AFFA43975FCE1FC6385BF1EE4" box="[799,988,909,932]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="castaneitarsis">
<emphasis id="B907EAEBFFA43975FCE1FC6385BF1EE4" box="[799,988,909,932]" italics="true" pageId="7" pageNumber="326">C. castaneitarsis</emphasis>
</taxonomicName>
) (
<figureCitation id="13482A7CFFA43975FC0BFC62821B1EE4" box="[1013,1144,908,933]" captionStart="FIGURES 24 29" captionStartId="12.[151,264,1964,1987]" captionTargetBox="[243,1405,210,1942]" captionTargetId="figure@12.[181,1405,193,1943]" captionTargetPageId="12" captionText="FIGURES 24 29. Cebrennus aethiopicus Simon, 1880, males (24 25 holotype male, Massaua, Eritrea, 26 27 Djibouti, 28 29 “ A Sefza ”). Left male palp (24, 26, 28 ventral, 25, 27, 29 retrolateral). EB—embolus base." httpUri="https://zenodo.org/record/398878/files/figure.png" pageId="7" pageNumber="326">Figs 2429</figureCitation>
). Females of this sub-unit of the
<taxonomicName id="4C734D7AFFA43975FF12FC5F87561E88" box="[236,309,945,968]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="wagae">
<emphasis id="B907EAEBFFA43975FF12FC5F87561E88" box="[236,309,945,968]" italics="true" pageId="7" pageNumber="326">wagae</emphasis>
</taxonomicName>
species-group are either not known (
<taxonomicName id="4C734D7AFFA43975FD01FC5E85FF1E88" box="[767,924,943,968]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="flagellatus" status="sp. nov.">
<emphasis id="B907EAEBFFA43975FD01FC5E85FF1E88" box="[767,924,943,968]" italics="true" pageId="7" pageNumber="326">C. flagellatus</emphasis>
</taxonomicName>
<emphasis id="B907EAEBFFA43975FC54FC4182471E88" bold="true" box="[938,1060,943,968]" pageId="7" pageNumber="326">
<taxonomicNameLabel id="A2345790FFA43975FC54FC4182471E88" box="[938,1060,943,968]" pageId="7" pageNumber="326" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
) or not clearly distinguishable. However, in
<taxonomicName id="4C734D7AFFA43975FED5FC3B87AD1EAD" box="[299,462,980,1005]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="B907EAEBFFA43975FED5FC3B87AD1EAD" box="[299,462,980,1005]" italics="true" pageId="7" pageNumber="326">C. aethiopicus</emphasis>
</taxonomicName>
the epigyne tends to be relatively (
<figureCitation id="13482A7CFFA43975FC9DFC3A85851EAD" box="[867,998,980,1005]" captionStart="FIGURES 55 62" captionStartId="18.[151,264,1848,1871]" captionTargetBox="[159,1436,195,1827]" captionTargetId="figure@18.[151,1436,195,1827]" captionTargetPageId="18" captionText="FIGURES 55 62. Cebrennus aethiopicus Simon, 1880, females (55 57 “ A Sefza ”, 58 60 Gheleb, Eritrea, 61 62 Eritrea). 55, 58, 61 Epigyne, ventral; 56, 59, 62 Vulva, dorsal; 57, 60 Schematic course of internal duct system, dorsal." httpUri="https://zenodo.org/record/398883/files/figure.png" pageId="7" pageNumber="326">Figs 5562</figureCitation>
) wider than in
<taxonomicName id="4C734D7AFFA43975FB68FC3B83621EAC" box="[1174,1281,981,1004]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="wagae">
<emphasis id="B907EAEBFFA43975FB68FC3B83621EAC" box="[1174,1281,981,1004]" italics="true" pageId="7" pageNumber="326">C. wagae</emphasis>
</taxonomicName>
(Figs 4454) and
<taxonomicName id="4C734D7AFFA43975FF37FC1687EB1950" box="[201,392,1016,1040]" class="Arachnida" family="Sparassidae" genus="Cebrennus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="326" phylum="Arthropoda" rank="species" species="castaneitarsis">
<emphasis id="B907EAEBFFA43975FF37FC1687EB1950" box="[201,392,1016,1040]" italics="true" pageId="7" pageNumber="326">C. castaneitarsis</emphasis>
</taxonomicName>
(
<figureCitation id="13482A7CFFA43975FE67FC19847C1950" box="[409,543,1015,1040]" captionStart="FIGURES 36 43. 36 38" captionStartId="15.[151,264,1920,1943]" captionTargetBox="[151,1435,193,1907]" captionTargetId="figure@15.[151,1435,193,1908]" captionTargetPageId="15" captionText="FIGURES 36 43. 36 38 Cebrennus castaneitarsis Simon, 1880, females from Wahran, Algeria or Casablanca, N Morocco (see note in text); 39 41 Cebrennus spec. B, female from Aouinet Torkoz, S Morocco; 42 44 Cebrennus wagae (Simon, 1874), female from Algier, Algeria. 36, 38 39, 42 Epigyne, ventral; 37, 40, 43 Vulva, dorsal; 41, 44 Schematic course of internal duct system, dorsal. GA—glandular appendages, LR—lateral ridge, MA—membranous atrium." httpUri="https://zenodo.org/record/398880/files/figure.png" pageId="7" pageNumber="326">Figs 3638</figureCitation>
). A distinct difference between the two latter species could not be found yet. Diagnoses contain a combination of characters, but few specimens (
<figureCitation id="13482A7CFFA43975FC6DFBF285891974" box="[915,1002,1052,1077]" captionStart="FIGURES 12 17. 12 13" captionStartId="6.[151,264,1915,1938]" captionTargetBox="[237,1386,200,1893]" captionTargetId="figure@6.[200,1387,193,1894]" captionTargetPageId="6" captionText="FIGURES 12 17. 12 13 Cebrennus rungsi Jäger, 2000, males from Anti-Atlas, Morocco; 14 Cebrennus spec. A, male from Beni Abbes Algeria; 15 17 Cebrennus castaneitarsis Simon, 1880, males (15 Algeria, 16 17 holotype male from Wahran, Algeria). Left male palp (12 16 ventral, 17 retrolateral). DE—distal part of embolus, K—embolic kink, PE—proximal part of embolus, RTA—retrolateral tibial apophysis." httpUri="https://zenodo.org/record/398876/files/figure.png" pageId="7" pageNumber="326">Figs 14</figureCitation>
,
<figureCitation id="13482A7CFFA43975FC09FBF2825D1974" box="[1015,1086,1052,1076]" captionStart="FIGURES 36 43. 36 38" captionStartId="15.[151,264,1920,1943]" captionTargetBox="[151,1435,193,1907]" captionTargetId="figure@15.[151,1435,193,1908]" captionTargetPageId="15" captionText="FIGURES 36 43. 36 38 Cebrennus castaneitarsis Simon, 1880, females from Wahran, Algeria or Casablanca, N Morocco (see note in text); 39 41 Cebrennus spec. B, female from Aouinet Torkoz, S Morocco; 42 44 Cebrennus wagae (Simon, 1874), female from Algier, Algeria. 36, 38 39, 42 Epigyne, ventral; 37, 40, 43 Vulva, dorsal; 41, 44 Schematic course of internal duct system, dorsal. GA—glandular appendages, LR—lateral ridge, MA—membranous atrium." httpUri="https://zenodo.org/record/398880/files/figure.png" pageId="7" pageNumber="326">3941</figureCitation>
) show that either transitions or even new species exist, which make a distinction in some cases difficult.
</paragraph>
</subSubSection>
</treatment>
</document>