treatments-xml/data/03/DA/87/03DA87B6FF904947FF597B2DFE6AFC2E.xml
2024-06-21 12:22:17 +02:00

935 lines
118 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="10.5252/geodiversitas2022v44a33" ID-GBIF-Dataset="0dfcd71d-bb24-4a0e-bc6d-15fbe04997d0" ID-ISSN="1638-9395" ID-Zenodo-Dep="7477657" ID-ZooBank="urn:lsid:zoobank.org:pub:4DA9802D-9500-4FD8-96F5-F4DD3BBF56A3" checkinTime="1671650606024" checkinUser="felipe" docAuthor="Unger, Tanja, Hildenbrand, Anne, Stinnesbeck, Wolfgang &amp; Austermann, Gregor" docDate="2022" docId="03DA87B6FF904947FF597B2DFE6AFC2E" docLanguage="en" docName="Geodiversitas.44.33.1051-1087.pdf" docOrigin="Geodiversitas 44 (33)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Agraulos ceticephalus" docType="treatment" docVersion="7" lastPageNumber="1058" masterDocId="FFE3FFCEFF97494CFFAA7878FFDCFFC1" masterDocTitle="Biostratigraphy and taxonomy of polymerid trilobites of the Manuels River Formation (Drumian, middle Cambrian), Newfoundland, Canada" masterLastPageNumber="1087" masterPageNumber="1051" pageNumber="1056" updateTime="1672773163721" updateUser="diego">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Biostratigraphy and taxonomy of polymerid trilobites of the Manuels River Formation (Drumian, middle Cambrian), Newfoundland, Canada</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Unger, Tanja</mods:namePart>
<mods:affiliation>Institut für Geowissenschaften, Universität Heidelberg, Im Neuenheimer Feld 234, 69120 Heidelberg (Germany) and Institut für Geologie, Mineralogie und Geophysik, Ruhr-Universität Bochum, 44801 Bochum (Germany)</mods:affiliation>
<mods:nameIdentifier type="email">tanja.unger@ruhr-uni-bochum.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hildenbrand, Anne</mods:namePart>
<mods:affiliation>Institut für Geowissenschaften, Universität Heidelberg, Im Neuenheimer Feld 234, 69120 Heidelberg (Germany)</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stinnesbeck, Wolfgang</mods:namePart>
<mods:affiliation>Institut für Geowissenschaften, Universität Heidelberg, Im Neuenheimer Feld 234, 69120 Heidelberg (Germany)</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Austermann, Gregor</mods:namePart>
<mods:affiliation>Institut für Geowissenschaften, Universität Heidelberg, Im Neuenheimer Feld 234, 69120 Heidelberg (Germany)</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Geodiversitas</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>44</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>33</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1051</mods:start>
<mods:end>1087</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/geodiversitas2022v44a33</mods:identifier>
<mods:identifier type="GBIF-Dataset">0dfcd71d-bb24-4a0e-bc6d-15fbe04997d0</mods:identifier>
<mods:identifier type="ISSN">1638-9395</mods:identifier>
<mods:identifier type="Zenodo-Dep">7477657</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:4DA9802D-9500-4FD8-96F5-F4DD3BBF56A3</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7477270" ID-GBIF-Taxon="207934064" ID-Zenodo-Dep="7477270" LSID="urn:lsid:plazi:treatment:03DA87B6FF904947FF597B2DFE6AFC2E" httpUri="http://treatment.plazi.org/id/03DA87B6FF904947FF597B2DFE6AFC2E" lastPageId="11" lastPageNumber="1058" pageId="7" pageNumber="1056">
<subSubSection box="[243,664,853,880]" pageId="7" pageNumber="1056" type="nomenclature">
<paragraph blockId="7.[243,664,853,880]" box="[243,664,853,880]" pageId="7" pageNumber="1056">
<heading box="[243,664,853,880]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="1056" reason="2">
<taxonomicName authority="(Barrande, 1846)" baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[243,664,853,880]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis bold="true" box="[243,472,853,880]" italics="true" pageId="7" pageNumber="1056">Agraulos ceticephalus</emphasis>
(
<bibRefCitation author="Barrande" box="[486,656,854,880]" firstAuthor="Barrande" pageId="7" pageNumber="1056" refId="ref23464" refString="BARRANDE J. 1846. - Notice preliminaire sur le systeme silurien et les Trilobites de Boheme. Hirschfeld, Leipzig, 96 p. https: // doi. org / 10.5962 / bhl. title. 9142" type="book" year="1846">Barrande, 1846</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[399,508,886,913]" pageId="7" pageNumber="1056" type="description">
<paragraph blockId="7.[399,508,886,913]" box="[399,508,886,913]" pageId="7" pageNumber="1056">
(
<figureCitation box="[408,471,886,912]" captionStart="FIG" captionStartId="8.[132,143,1849,1866]" captionTargetBox="[132,1455,215,1783]" captionTargetId="figure-27@8.[699,1268,767,1323]" captionTargetPageId="8" captionText="FIG. 4. — Agraulos ceticephalus (Barrande, 1846): A, internal cast of a cranidium attached to the thorax with preserved exoskeleton (NFM F-3476 from 5.97 m); B, internal cast of a cranidium attached to the thorax, partly covered with sediment, eye ridges and glabellar furrows slightly visible (NFM F-3951 from 7.10 m); C, internal cast of a cranidium attached to the thorax with preserved exoskeleton, eye ridges present (NFM F-3957 from 3.30 m); D, internal cast of a cephalon attached to the thorax, librigena attached, eye ridges present (NFM F-3621 from 9.08 m); E, mould of a cranidium attached to the thorax, spines on axial rings present,coarse ornamentation on thorax visible (NFM F-3476 from 5.98 m); F, mould of a cranidium attached to the thorax,occipital spine preserved (sedimentary filling laying in glabellar mould), coarse ornamentation on thorax visible (NFM F-3946 from 4.27 m); G, internal cast of a cranidium attached to the thorax, projection of axial spine preserved (NFM F-3831 from 7.43 m); H, internal cast of a cranidium attached to the thorax, eye ridges present (NFM F-2787 from 7.56 m); I, internal cast of a cranidium attached to first thoracic segments, eye ridges and glabellar furrows preserved (NFM F-3778 from 5.96 m). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7477665" httpUri="https://zenodo.org/record/7477665/files/figure.png" pageId="7" pageNumber="1056">Figs 4</figureCitation>
;
<figureCitation box="[485,499,886,913]" captionStart="FIG" captionStartId="10.[132,143,1872,1889]" captionTargetBox="[239,1349,215,1848]" captionTargetId="figure-209@10.[793,1350,768,1324]" captionTargetPageId="10" captionText="FIG. 5. — Agraulos ceticephalus (Barrande, 1846): A, internal cast of a cranidium, eye ridges present, two grooves visible (NFM F-2785 from 1.96 m); B, internal cast of a cranidium, one groove visible, occipital ring triangular in shape with projection of spine preserved (NFM F-3062 from 2.10 m); C, internal cast of a cranidium, occipital ring ending in a spine (NFM F-2841 from 2.40 m); D, internal cast of a cranidium, projection of occipital spine preserved (NFM F-2919 from 2.44 m); E, internal cast of a cranidium, eye ridges and occipital spine preserved (NFM F-2846 from 2.53 m); F, internal cast of a cranidium, eye ridges present, projection of occipital spine preserved (NFM F-3321 from 5.52 m); G, internal cast of a cranidium, eye ridges present, projection of occipital spine preserved (NFM F-2863 from 2.52 m); H, internal cast of a cranidium, two grooves visible,profile on the right shows that front is not as domed as it seems in overview (NFM F-3225 from 5.19 m). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7477667" httpUri="https://zenodo.org/record/7477667/files/figure.png" pageId="7" pageNumber="1056">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="1056" type="reference_group">
<paragraph blockId="7.[132,715,945,969]" box="[132,715,945,969]" pageId="7" pageNumber="1056">
<treatmentCitationGroup box="[132,715,945,969]" pageId="7" pageNumber="1056">
<treatmentCitation author="Barrande" box="[132,499,945,969]" page="12" pageId="7" pageNumber="1056" year="1846">
<taxonomicName authority="Barrande, 1846: 12" authorityName="Barrande" authorityPageNumber="12" authorityYear="1846" box="[132,499,945,969]" class="Gastropoda" family="Arionidae" genus="Arion" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="1056" phylum="Mollusca" rank="species" species="ceticephalus">
<emphasis box="[132,301,945,969]" italics="true" pageId="7" pageNumber="1056">Arion ceticephalus</emphasis>
<bibRefCitation author="Barrande" box="[307,499,945,969]" firstAuthor="Barrande" page="12" pageId="7" pageNumber="1056" refId="ref23464" refString="BARRANDE J. 1846. - Notice preliminaire sur le systeme silurien et les Trilobites de Boheme. Hirschfeld, Leipzig, 96 p. https: // doi. org / 10.5962 / bhl. title. 9142" type="book" year="1846">Barrande, 1846: 12</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 13;
<treatmentCitation author="Barrande" box="[548,711,945,969]" page="405 - 412" pageId="7" pageNumber="1056" year="1852">
<bibRefCitation author="Barrande" box="[548,711,945,969]" firstAuthor="Barrande" page="405 - 412" pageId="7" pageNumber="1056" pagination="1 - 935" refId="ref23595" refString="BARRANDE J. 1852 b. - Systeme silurien du Centre de la Boheme. 1 ere Partie: Texte. Crustaces: Trilobites. Recherches paleontologiques 1: 1 - 935. https: // www. biodiversitylibrary. org / page / 57395510" type="journal article" year="1852" yearSuffix="b">1852b: 405-412</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[131,776,994,1415]" pageId="7" pageNumber="1056">
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[132,329,994,1018]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[132,329,994,1018]" italics="true" pageId="7" pageNumber="1056">Agraulos ceticephalus</emphasis>
</taxonomicName>
<bibRefCitation author="Hawle &amp; Corda" box="[358,620,994,1019]" firstAuthor="Hawle" page="27" pageId="7" pageNumber="1056" pagination="117 - 292" refId="ref25827" refString="HAWLE I. &amp; CORDA A. J. C. 1847. - Prodrom einer Monographie der bohmischen Trilobiten. J. G. Calve'sche Buchhandlung, Prague, 176 p. (Reprinted in 1848 in Abhandlungen der Koniglichen Bohmischen Gesellschaft der Wissenschaften 5 (2): 117 - 292). https: // www. biodiversitylibrary. org / page / 45618120" type="book chapter" year="1847">Hawle &amp; Corda 1847: 27</bibRefCitation>
; 1848: 143. —
<bibRefCitation author="Miller" box="[134,303,1021,1045]" firstAuthor="Miller" page="527" pageId="7" pageNumber="1056" refId="ref28272" refString="MILLER S. A. 1889. - North American Geology and Paleontology for the Use of Amateurs, Students, and Scientists. Western Methodist book concern, Cincinnati, Ohio, 793 p. https: // doi. org / 10.5962 / bhl. title. 28778" type="book" year="1889">Miller 1889: 527</bibRefCitation>
. —
<bibRefCitation author="Pompeckj" box="[346,552,1021,1045]" firstAuthor="Pompeckj" page="548" pageId="7" pageNumber="1056" pagination="498 - 614" refId="ref28633" refString="POMPECKJ J. F. 1896. - Das Kambrium von Tejrovic und Skrej in Bohmen. Jahrbuch der Kaiserlich-Koniglichen Geologischen Reichsanstalt 94: 498 - 614. https: // www. biodiversitylibrary. org / page / 35578547" type="journal article" year="1896">Pompeckj 1896: 548</bibRefCitation>
, pl. 17, figs 12, 13. —
<bibRefCitation author="Gronwall" box="[133,331,1047,1072]" firstAuthor="Gronwall" page="158" pageId="7" pageNumber="1056" pagination="1 - 230" refId="ref25638" refString="GRONWALL K. A. 1902. - Bornholms Paradoxideslag og deres fauna. Danmarks geologiske Undersogelse, II Raekke 13 (1): 1 - 230." type="journal article" year="1902">Grönwall 1902: 158</bibRefCitation>
, 159, pl. 4, fig. 25. —
<bibRefCitation author="Lorenz" box="[548,670,1047,1071]" firstAuthor="Lorenz" pageId="7" pageNumber="1056" pagination="53 - 108" refId="ref27919" refString="LORENZ T. 1906. - Beitrage zur Geologie und Palaeontologie von Ostasien unter besonderer Berucksichtigung der Provinz Schantung in China: II. Palaeontologischer - Teil. Zeitschrift der Deutschen geologischen Gesellschaft 58 (1): 53 - 108. https: // www. biodiversitylibrary. org / page / 43913940" type="journal article" year="1906">Lorenz 1906</bibRefCitation>
: unnamed text fig. p. 67 upper right corner. —
<bibRefCitation author="Walcott" box="[499,631,1074,1098]" firstAuthor="Walcott" pageId="7" pageNumber="1056" pagination="3 - 277" refId="ref30768" refString="WALCOTT C. D. 1913. - The Cambrian faunas of China, in Walcott C. D. (ed.), Research in China vol. 3. Carnegie Institution Publication 54 (3): 3 - 277." type="journal article" year="1913">Walcott 1913</bibRefCitation>
: pl. 15, figs 1, 1a, b. —
<bibRefCitation author="Wurm" box="[219,377,1100,1124]" firstAuthor="Wurm" page="87" pageId="7" pageNumber="1056" pagination="71 - 93" refId="ref31321" refString="WURM A. 1925. - Ueber ein Vorkommen von Mittelcambrium (Paradoxidesschichten) im bayrischen Frankenwald bei Wildenstein sudlich Presseck. Neues Jahrbuch fur Mineralogie, Geognosie Geologie und Palaontologie 52 (1): 71 - 93." type="journal article" year="1925">Wurm 1925: 87</bibRefCitation>
, 88, pl. 3, figs 16, 17. —
<bibRefCitation author="Roch" box="[618,775,1100,1124]" firstAuthor="Roch" page="132" pageId="7" pageNumber="1056" refId="ref29177" refString="ROCH E. 1930. - Etudes geologiques dans la region Meridionale du Maroc occidental. Protat Freres imprimeurs, Macon, 542 p." type="book" year="1930">Roch 1930: 132</bibRefCitation>
(?). —
<bibRefCitation author="Thoral" box="[198,355,1127,1151]" firstAuthor="Thoral" page="50" pageId="7" pageNumber="1056" refId="ref30557" refString="THORAL M. 1935. - Contribution a l'etude paleontologique de l'Ordovicien inferieur de la Montagne Noire et revision sommaire de la faune cambrienne de la Montagne Noire. Unpublished PhD thesis, Universite de Paris, Montpellier, 362 p." type="book" year="1935">Thoral 1935: 50</bibRefCitation>
, 51, pl. 3, fig. 12. —
<bibRefCitation author="Prantl" box="[556,722,1127,1151]" firstAuthor="Prantl" page="262" pageId="7" pageNumber="1056" refId="ref28751" refString="PRANTL F. 1952. - Zivot ceskych pramori. Prirodovedecke Vydavatelstvi, Praze, 390 p." type="book" year="1952">Prantl 1952: 262</bibRefCitation>
, 263, unnamed fig. p. 264. —
<bibRefCitation author="Hupe" box="[369,479,1153,1177]" firstAuthor="Hupe" pageId="7" pageNumber="1056" pagination="44 - 246" refId="ref26509" refString="HUPE P. 1953. - Classe des Trilobites, in PIVETEAU J. (ed.), Traite de paleontologie. Tome III. Les forms ultimes d'Invertebres: morphologie et evolution. Onychophores, Arthropodes, Echniodermes, Stomocordes. Mason et Cie, Paris: 44 - 246." type="book chapter" year="1953">Hupé 1953</bibRefCitation>
: fig. 120.1; 1955: fig. 96.2. —
<bibRefCitation author="Snajdr" box="[131,357,1180,1207]" firstAuthor="Snajdr" page="174 - 177" pageId="7" pageNumber="1056" pagination="1 - 280" refId="ref30272" refString="SNAJDR M. 1958. - Trilobiti ceskeho stredniho kambria. Rozpravy Ustredniho ustavu geologickeho 24: 1 - 280." type="journal article" year="1958">Šnajdr 1958: 174-177</bibRefCitation>
, fig. 37; pl. 36, figs 1-13; 1990: 106, unnamed fig. p. 107. —
<bibRefCitation author="Horny &amp; Bastl" box="[342,539,1206,1230]" firstAuthor="Horny" pageId="7" pageNumber="1056" refId="ref26355" refString="HORNY R. &amp; BASTL F. 1970. - Type Specimens of Fossils in the National Museum Prague, Volume 1, Trilobita. Museum of National History, Prague, 354 p." type="book" year="1970">Horný &amp; Bastl 1970</bibRefCitation>
: pl. 4, fig. 9. —
<bibRefCitation author="Fletcher" firstAuthor="Fletcher" pageId="7" pageNumber="1056" refId="ref24893" refString="FLETCHER T. P. 1972 b. - Geology and Lower to Middle Cambrian Trilobite Faunas of the Southwest Avalon, Newfoundland, Part Two, Palaeontology &amp; Bibliography. Unpublished PhD thesis, University of Cambridge, 294 p." type="book" year="1972" yearSuffix="b">Fletcher 1972b</bibRefCitation>
: pl. 69, figs 3, 4, 5a-c, 6, pl. 70, figs 1a, b, 2; 2006: pl. 34, fig. 35.; 2017: 11-19, figs 5G-I, 6A-P, 7A-S, 8A-E, H-L, 16B, J, L (
<emphasis box="[140,203,1286,1309]" italics="true" pageId="7" pageNumber="1056">partim</emphasis>
). —
<bibRefCitation author="Whittington" box="[252,481,1285,1310]" firstAuthor="Whittington" page="594" pageId="7" pageNumber="1056" pagination="577 - 609" refId="ref31075" refString="WHITTINGTON H. B. 1988. - Hypostomes and ventral cephalic sutures in Cambrian trilobites. Palaeontology 31 (3): 577 - 609. https: // www. biodiversitylibrary. org / page / 49775869" type="journal article" year="1988">Whittington 1988: 594</bibRefCitation>
, text-fig. 14, pl. 55, figs 2, 4, 5, 8, 9; 1992: pls 7, 51. —
<bibRefCitation author="Fatka" box="[361,469,1312,1336]" firstAuthor="Fatka" pageId="7" pageNumber="1056" pagination="12 - 17" refId="ref24692" refString="FATKA O. 1990. - Das Kambrium von Skryje und Tyrovice, in WIEDER W. K. (ed). Klassissche Fundtsellen der Palaontologie. Band 2: 23. Fundgebiete und Aufschlusse in Danemark, Deutschland, England, Frankreich, Osterreich, Schweiz und Tschechoslowakei. Goldschenk-Verlag Korb, Germany: 12 - 17." type="book chapter" year="1990">Fatka 1990</bibRefCitation>
: unnamed fig. p. 13 upper right corner; 2011: fig. 17.6. —
<bibRefCitation author="Muller" box="[390,563,1338,1363]" firstAuthor="Muller" page="529" pageId="7" pageNumber="1056" refId="ref28408" refString="MULLER A. H. 1994. - Band II: Invertebraten, Teil 2: Mollusca 2 - Arthropoda 1, 4., neu bearb. und erw. Aufl. Gustav Fischer Verlag, Jena, 618 p." type="book" year="1994">Müller 1994: 529</bibRefCitation>
, fig. 619. —
<bibRefCitation author="Rudolph" firstAuthor="Rudolph" page="217" pageId="7" pageNumber="1056" refId="ref29287" refString="RUDOLPH F. 1994. - Die Trilobiten der mittelkambrischen Geschiebe: Systematik, Morphologie und Okologie. Verlag Frank Rudolph, Wankendorf, 309 p." type="book" year="1994">Rudolph 1994: 217</bibRefCitation>
, pl. 24, fig. 5. —
<bibRefCitation author="Cotton" box="[393,518,1365,1389]" firstAuthor="Cotton" pageId="7" pageNumber="1056" pagination="167 - 207" refId="ref24405" refString="COTTON T. J. 2001. - The phylogeny and systematics of blind Cambrian ptychoparioid trilobites. Palaeontology 44 (1): 167 - 207. https: // doi. org / 10.1111 / 1475 - 4983.00176" type="journal article" year="2001">Cotton 2001</bibRefCitation>
: pl. 2, figs 4, 5. —
<bibRefCitation author="Fletcher" etAl="et al." firstAuthor="Fletcher" page="330" pageId="7" pageNumber="1056" pagination="312 - 336" refId="ref25165" refString="FLETCHER T. P., THEOKRITOFF G., LORD G. S. &amp; ZEOLI G. 2005. - The early paradoxidid harlani trilobite fauna of Massachusetts and its correlatives in Newfoundland, Morocco, and Spain. Journal of Paleontology 79 (2): 312 - 336. https: // doi. org / bdcjsg" type="journal article" year="2005">
Fletcher
<emphasis box="[132,179,1391,1415]" italics="true" pageId="7" pageNumber="1056">et al.</emphasis>
2005: 330
</bibRefCitation>
, 331, figs 11.1-7. —
<bibRefCitation author="Fatka" box="[493,653,1391,1415]" etAl="et al." firstAuthor="Fatka" pageId="7" pageNumber="1056" pagination="89 - 101" refId="ref24770" refString="FATKA O., BUDIL P., CRONIER C., CUVELIER J., LAIBL L., OUDOIRE T., POLECHOVA M. &amp; FATKOVA L. 2015. - Cambrian fossils from the Barrandian area (Czech Republic) housed in the Musee d'Histoire naturelle de Lille. Carnets de geologie 15 (9): 89 - 101. https: // doi. org / 10.4267 / 2042 / 56878" type="journal article" year="2015">
Fatka
<emphasis box="[552,598,1391,1415]" italics="true" pageId="7" pageNumber="1056">et al.</emphasis>
2015
</bibRefCitation>
: figs 4C, G.
</paragraph>
<paragraph blockId="7.[131,775,1441,1518]" pageId="7" pageNumber="1056">
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[132,340,1441,1465]" class="Trilobita" family="Olenidae" genus="Arionellus" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[132,340,1441,1465]" italics="true" pageId="7" pageNumber="1056">Arionellus ceticephalus</emphasis>
</taxonomicName>
<bibRefCitation author="Barrande" box="[361,515,1441,1465]" firstAuthor="Barrande" pageId="7" pageNumber="1056" refId="ref23551" refString="BARRANDE J. 1852 a. - Systeme silurien du Centre de la Boheme. 1 ere Partie: Planches. Crustaces: Trilobites. Recherches paleontologiques 1: 49 pls. https: // www. biodiversitylibrary. org / page / 47135930" type="book" year="1852" yearSuffix="a">Barrande 1852a</bibRefCitation>
: pl. 10, figs 6, 8-21; pl. 11, fig. 7 (
<emphasis box="[198,262,1468,1491]" italics="true" pageId="7" pageNumber="1056">partim</emphasis>
). —
<bibRefCitation author="de Verneuil &amp; Barrande" box="[315,661,1467,1492]" firstAuthor="de Verneuil" page="526" pageId="7" pageNumber="1056" pagination="526 - 554" refId="ref24537" refString="DE VERNEUIL M. M. &amp; BARRANDE J. 1860. - Description des fossiles. Bulletin de la Societe geologique de France 17 (2): 526 - 554. https: // www. biodiversitylibrary. org / page / 54397903" type="journal article" year="1860">de Verneuil &amp; Barrande 1860: 526</bibRefCitation>
, 527, pl. 6, figs 13-17. —
<bibRefCitation author="Gurich" box="[274,441,1494,1518]" firstAuthor="Gurich" page="19" pageId="7" pageNumber="1056" refId="ref25667" refString="GURICH G. 1908. - Leitfossilien, ein Hilfsbuch zum Bestimmen von Versteinerungen bei geologischen Arbeiten in der Sammlung und im Felde; erste Lieferung: Kambrium und Silur. Berlin, 95 p. https: // doi. org / 10.1038 / 084200 a 0" type="book" year="1908">Gürich 1908: 19</bibRefCitation>
, pl. 5, fig. 2.
</paragraph>
<paragraph blockId="7.[132,708,1543,1568]" box="[132,708,1543,1568]" pageId="7" pageNumber="1056">
<treatmentCitationGroup box="[132,708,1543,1568]" pageId="7" pageNumber="1056">
<treatmentCitation author="Hicks" box="[132,535,1543,1568]" page="176" pageId="7" pageNumber="1056" year="1872">
<taxonomicName authority="Hicks, 1872: 176" authorityName="Hicks" authorityPageNumber="176" authorityYear="1872" box="[132,535,1543,1568]" class="Trilobita" family="Olenidae" genus="Arionellus" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[132,356,1543,1567]" italics="true" pageId="7" pageNumber="1056">Arionellus longicephalus</emphasis>
<bibRefCitation author="Hicks" box="[362,535,1543,1568]" firstAuthor="Hicks" page="176" pageId="7" pageNumber="1056" pagination="173 - 185" refId="ref26120" refString="HICKS H. 1872. - On some undescribed fossils from the Menevian Group. The Quarterly Journal of the Geological Society of London 28 (1): 173 - 185. https: // doi. org / 10.1144 / gsl. jgs. 1872.028.01 - 02.17" type="journal article" year="1872">Hicks, 1872: 176</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 5, figs 20-26.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[131,776,1593,1776]" pageId="7" pageNumber="1056">
<treatmentCitationGroup pageId="7" pageNumber="1056">
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[132,352,1593,1617]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[132,352,1593,1617]" italics="true" pageId="7" pageNumber="1056">Agraulos longicephalus</emphasis>
</taxonomicName>
<treatmentCitation author="Lake" box="[384,595,1593,1617]" page="157 - 159" pageId="7" pageNumber="1056" year="1932">
<bibRefCitation author="Lake" box="[384,595,1593,1617]" firstAuthor="Lake" page="157 - 159" pageId="7" pageNumber="1056" pagination="149 - 172" refId="ref27339" refString="LAKE P. 1932. - Monograph of the British Cambrian Trilobites, Part 7. Monograph of the Palaeontographical Society, London 84: 149 - 172. https: // doi. org / 10.1080 / 02693445.1932.12035620" type="journal article" year="1932">Lake 1932: 157-159</bibRefCitation>
</treatmentCitation>
, pl. 20, figs 2, 7, 10. —
<treatmentCitation author="Sdzuy" box="[198,409,1620,1644]" page="620 - 622" pageId="7" pageNumber="1056" year="1961">
<bibRefCitation author="Sdzuy" box="[198,409,1620,1644]" firstAuthor="Sdzuy" page="620 - 622" pageId="7" pageNumber="1056" pagination="503 - 594" refId="ref30076" refString="SDZUY K. 1961. - Das Kambrium Spaniens, Teil II: Trilobiten. 1 Akademie der Wissenschaften und der Literatur, Mainz, Abhandlungen der mathematisch-naturwissenschaftlichen Klasse 7: 503 - 594." type="journal article" year="1961">Sdzuy 1961: 620-622</bibRefCitation>
</treatmentCitation>
, figs 32, 33. —
<bibRefCitation author="Fletcher" box="[555,701,1620,1644]" firstAuthor="Fletcher" pageId="7" pageNumber="1056" refId="ref24893" refString="FLETCHER T. P. 1972 b. - Geology and Lower to Middle Cambrian Trilobite Faunas of the Southwest Avalon, Newfoundland, Part Two, Palaeontology &amp; Bibliography. Unpublished PhD thesis, University of Cambridge, 294 p." type="book" year="1972" yearSuffix="b">Fletcher 1972b</bibRefCitation>
: pl. 68, figs 5a-d, 7-11; pl. 69, figs 1, 2; 2006: pl. 34, fig. 34; 2017: figs 22A, B. —
<treatmentCitation author="Courtessole" box="[195,469,1672,1697]" page="138 - 140" pageId="7" pageNumber="1056" year="1973">
<bibRefCitation author="Courtessole" box="[195,469,1672,1697]" firstAuthor="Courtessole" page="138 - 140" pageId="7" pageNumber="1056" refId="ref24445" refString="COURTESSOLE R. 1973. - Le Cambrien moyen de la Montagne Noire: Biostratigraphie. Laboratoire de Geologie CEARN de la Faculte des Sciences de Toulouse, Toulouse, 248 p." type="book" year="1973">Courtessole 1973: 138-140</bibRefCitation>
</treatmentCitation>
, pl. 10, figs 5, 6, 8-10; pl. 16, figs 12, 13. —
<treatmentCitation author="Martin &amp; Dean" box="[278,530,1699,1723]" page="21" pageId="7" pageNumber="1056" year="1988">
<bibRefCitation author="Martin &amp; Dean" box="[278,530,1699,1723]" firstAuthor="Martin" page="21" pageId="7" pageNumber="1056" pagination="1 - 91" refId="ref27973" refString="MARTIN F. &amp; DEAN W. T. 1988. - Middle and upper Cambrian acritarch and trilobite zonation at Manuels River and Random Island, eastern Newfoundland. Geological Survey of Canada Bulletin 381 (1): 1 - 91." type="journal article" year="1988">Martin &amp; Dean 1988: 21</bibRefCitation>
</treatmentCitation>
, 22, pl. 3, figs 10, 12, 13 (
<emphasis box="[140,204,1726,1749]" italics="true" pageId="7" pageNumber="1056">partim</emphasis>
). —
<bibRefCitation author="Rees" box="[259,415,1725,1750]" etAl="et al." firstAuthor="Rees" pageId="7" pageNumber="1056" pagination="1 - 135" refId="ref28872" refString="REES A., THOMAS A., LEWIS M., HUGHES H. &amp; TURNER P. 2014. - Cambrian of Sw Wales: towards a united Avalonian Stratigraphy. Memoirs of the Geological Society London 42: 1 - 135. https: // doi. org / 10.1144 / m 42.0" type="journal article" year="2014">
Rees
<emphasis box="[310,358,1725,1749]" italics="true" pageId="7" pageNumber="1056">et al.</emphasis>
2014
</bibRefCitation>
: figs
<quantity box="[468,513,1726,1750]" metricMagnitude="-3" metricUnit="kg" metricValue="1.8" pageId="7" pageNumber="1056" unit="g" value="1.8">1.8g</quantity>
, h. —
<treatmentCitation author="Weidner &amp; Nielsen" page="47" pageId="7" pageNumber="1056" year="2014">
<bibRefCitation author="Weidner &amp; Nielsen" firstAuthor="Weidner" page="47" pageId="7" pageNumber="1056" pagination="23 - 92" refId="ref30854" refString="WEIDNER T. &amp; NIELSEN A. T. 2014. - A highly diverse trilobite fauna with Avalonian affinities from the middle Cambrian Acidusus atavus Zone (Drumian Stage) of Bornholm, Denmark. Journal of Systematic Palaeontology 12 (1): 23 - 92. https: // doi. org / 10.10 80 / 14772019.2012.740080" type="journal article" year="2014">Weidner &amp; Nielsen 2014: 47</bibRefCitation>
</treatmentCitation>
, 48, figs 41A-F; 2015: 5-7, figs 3, 4A-L.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[132,775,1801,1852]" pageId="7" pageNumber="1056">
<treatmentCitationGroup pageId="7" pageNumber="1056">
<taxonomicName box="[132,476,1801,1825]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="subSpecies" species="longicephalus" subSpecies="longicephalus">
<emphasis box="[132,476,1801,1825]" italics="true" pageId="7" pageNumber="1056">Agraulos longicephalus longicephalus</emphasis>
</taxonomicName>
<treatmentCitation author="Linan &amp; Gozalo" box="[507,771,1801,1826]" page="78" pageId="7" pageNumber="1056" year="1986">
<bibRefCitation author="Linan &amp; Gozalo" box="[507,771,1801,1826]" firstAuthor="Linan" page="78" pageId="7" pageNumber="1056" pagination="1 - 104" refId="ref27824" refString="LINAN E. &amp; GOZALO R. 1986. - Trilobites del Cambrico inferior y medo de murero (Cordillera Iberica). Memorias del Museo Paleontologico de la Universidad de Zaragoza 2: 1 - 104." type="journal article" year="1986">Liñan &amp; Gozalo 1986: 78</bibRefCitation>
</treatmentCitation>
, pl. 35, figs 11-14; pl. 36, figs 4, 5 (
<emphasis box="[474,538,1829,1852]" italics="true" pageId="7" pageNumber="1056">partim</emphasis>
).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[132,775,1878,1928]" pageId="7" pageNumber="1056">
<treatmentCitationGroup pageId="7" pageNumber="1056">
<taxonomicName box="[132,479,1878,1902]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="subSpecies" species="longicephalus" subSpecies="brevilimbarus">
<emphasis box="[132,479,1878,1902]" italics="true" pageId="7" pageNumber="1056">Agraulos longicephalus brevilimbarus</emphasis>
</taxonomicName>
<treatmentCitation author="Linan &amp; Gozalo" box="[509,771,1878,1902]" page="78" pageId="7" pageNumber="1056" year="1986">
<bibRefCitation author="Linan &amp; Gozalo" box="[509,771,1878,1902]" firstAuthor="Linan" page="78" pageId="7" pageNumber="1056" pagination="1 - 104" refId="ref27824" refString="LINAN E. &amp; GOZALO R. 1986. - Trilobites del Cambrico inferior y medo de murero (Cordillera Iberica). Memorias del Museo Paleontologico de la Universidad de Zaragoza 2: 1 - 104." type="journal article" year="1986">Liñan &amp; Gozalo 1986: 78</bibRefCitation>
</treatmentCitation>
, 79, pl. 35, figs 6-12; pl. 37, figs 1-8.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[132,775,1954,1978]" box="[132,775,1954,1978]" pageId="7" pageNumber="1056">
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[132,355,1954,1978]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="ceticephalus" subGenus="A.">
<emphasis box="[132,208,1954,1978]" italics="true" pageId="7" pageNumber="1056">Agraulos</emphasis>
(
<emphasis box="[219,241,1954,1977]" italics="true" pageId="7" pageNumber="1056">A.</emphasis>
)
<emphasis box="[252,355,1954,1978]" italics="true" pageId="7" pageNumber="1056">ceticephalus</emphasis>
</taxonomicName>
<bibRefCitation author="Schoenemann &amp; Clarkson" box="[373,671,1954,1978]" firstAuthor="Schoenemann" pageId="7" pageNumber="1056" pagination="45 - 50" refId="ref29980" refString="SCHOENEMANN B. &amp; CLARKSON E. N. K. 2011. - The eyes of Bohemian Trilobites. Geologicke vyzkumy na Morave a ve Slezsku 18 (1): 45 - 50." type="journal article" year="2011">Schoenemann &amp; Clarkson 2011</bibRefCitation>
: figs 5H, I.
</paragraph>
<paragraph blockId="7.[132,528,2003,2027]" box="[132,528,2003,2027]" pageId="7" pageNumber="1056">
<treatmentCitationGroup box="[132,528,2003,2027]" pageId="7" pageNumber="1056">
<taxonomicName authorityName="Billings" authorityYear="1872" box="[132,275,2003,2027]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="affinis">
<emphasis box="[132,275,2003,2027]" italics="true" pageId="7" pageNumber="1056">Agraulos affinis</emphasis>
</taxonomicName>
<bibRefCitation author="Fletcher" box="[300,436,2003,2027]" firstAuthor="Fletcher" pageId="7" pageNumber="1056" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017</bibRefCitation>
: fig.
<specimenCount box="[486,528,2003,2027]" pageId="7" pageNumber="1056" type="generic">11F.</specimenCount>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[813,1455,215,239]" box="[813,1455,215,239]" pageId="7" pageNumber="1056">
<treatmentCitationGroup box="[813,1455,215,239]" pageId="7" pageNumber="1056">
<treatmentCitation author="Fletcher" box="[813,1147,215,239]" page="21" pageId="7" pageNumber="1056" year="2017">
<taxonomicName authority="Fletcher, 2017: 19 - 21" authorityName="Fletcher" authorityPageNumber="19 - 21" authorityYear="2017" box="[813,1147,215,239]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis box="[813,942,215,239]" italics="true" pageId="7" pageNumber="1056">Agraulos lewisi</emphasis>
<bibRefCitation author="Fletcher" box="[946,1147,215,239]" firstAuthor="Fletcher" page="19 - 21" pageId="7" pageNumber="1056" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher, 2017: 19-21</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 3A-C, 8F, G (
<emphasis box="[1315,1376,216,239]" italics="true" pageId="7" pageNumber="1056">partim</emphasis>
),
<emphasis bold="true" box="[1393,1455,215,239]" pageId="7" pageNumber="1056">n. syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[813,1455,262,313]" pageId="7" pageNumber="1056">
<treatmentCitationGroup pageId="7" pageNumber="1056">
<taxonomicName authorityName="Billings" authorityYear="1872" box="[813,962,262,286]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="socialis">
<emphasis box="[813,962,262,286]" italics="true" pageId="7" pageNumber="1056">Agraulos socialis</emphasis>
</taxonomicName>
<treatmentCitation author="Fletcher" box="[987,1196,262,287]" page="23 - 27" pageId="7" pageNumber="1056" year="2017">
<bibRefCitation author="Fletcher" box="[987,1196,262,287]" firstAuthor="Fletcher" page="23 - 27" pageId="7" pageNumber="1056" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017: 23-27</bibRefCitation>
</treatmentCitation>
, figs 5J-L, 12A, K-Q, 13I- K, M, N, 14D, G, H, J, 16A, C.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[813,1305,336,361]" box="[813,1305,336,361]" pageId="7" pageNumber="1056">
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[813,892,337,361]" class="Trilobita" family="Agraulidae" genus="Agraulos" isUncertain="true" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="sp.1">
<emphasis box="[813,892,337,361]" italics="true" pageId="7" pageNumber="1056">Agraulos</emphasis>
</taxonomicName>
1
<bibRefCitation author="Fletcher" box="[936,1072,337,361]" firstAuthor="Fletcher" pageId="7" pageNumber="1056" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017</bibRefCitation>
: figs 13D, E, 14A,
<specimenCount box="[1263,1305,336,360]" pageId="7" pageNumber="1056" type="generic">16F.</specimenCount>
</paragraph>
<paragraph blockId="7.[813,1383,384,408]" box="[813,1383,384,408]" pageId="7" pageNumber="1056">
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[813,892,384,408]" class="Trilobita" family="Agraulidae" genus="Agraulos" isUncertain="true" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="sp.2">
<emphasis box="[813,892,384,408]" italics="true" pageId="7" pageNumber="1056">Agraulos</emphasis>
</taxonomicName>
2
<bibRefCitation author="Fletcher" box="[936,1072,384,408]" firstAuthor="Fletcher" pageId="7" pageNumber="1056" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017</bibRefCitation>
: figs 14I, 15A-C, 16E (
<emphasis box="[1304,1368,385,408]" italics="true" pageId="7" pageNumber="1056">partim</emphasis>
).
</paragraph>
<paragraph blockId="7.[813,1367,432,456]" box="[813,1367,432,456]" pageId="7" pageNumber="1056">
<taxonomicName authorityName="Fletcher" authorityYear="2017" box="[813,1044,432,456]" class="Trilobita" family="Agraulidae" genus="Skreiaspis" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis box="[813,1044,432,456]" italics="true" pageId="7" pageNumber="1056">Skreiaspis punctatissimus</emphasis>
</taxonomicName>
<bibRefCitation author="Fletcher" box="[1070,1206,432,456]" firstAuthor="Fletcher" pageId="7" pageNumber="1056" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017</bibRefCitation>
: figs 18S, U, W.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="1056" type="materials_examined">
<paragraph blockId="7.[811,1457,479,636]" pageId="7" pageNumber="1056">
<materialsCitation ID-GBIF-Occurrence="4003346302" collectionCode="SBNM" country="Czech Republic" location="Buchava Formation, Skreyje Member, Eccaparadoxides pusillus Zone from Skryje-Tyrovice Basin" pageId="7" pageNumber="1056" specimenCode="NM L 12581" specimenCount="1" typeStatus="lectotype">
<typeStatus box="[813,924,480,503]" pageId="7" pageNumber="1056">LECTOTYPE</typeStatus>
. —
<specimenCount box="[970,1065,479,503]" pageId="7" pageNumber="1056" type="generic">Specimen</specimenCount>
<specimenCode box="[1073,1204,479,503]" pageId="7" pageNumber="1056">NM L 12581</specimenCode>
(former ČC 345, No. 85 from
<collectionCode box="[865,934,506,530]" collectionName="SBNM" pageId="7" pageNumber="1056">SBNM</collectionCode>
collection Barrande), National Museum, Prague, Czech Republic, originally figured by
<bibRefCitation author="Barrande" box="[1126,1299,532,557]" firstAuthor="Barrande" pageId="7" pageNumber="1056" refId="ref23551" refString="BARRANDE J. 1852 a. - Systeme silurien du Centre de la Boheme. 1 ere Partie: Planches. Crustaces: Trilobites. Recherches paleontologiques 1: 49 pls. https: // www. biodiversitylibrary. org / page / 47135930" type="book" year="1852" yearSuffix="a">Barrande (1852a)</bibRefCitation>
and designated as the lectotype by
<bibRefCitation author="Snajdr" box="[1006,1144,559,586]" firstAuthor="Snajdr" pageId="7" pageNumber="1056" pagination="1 - 280" refId="ref30272" refString="SNAJDR M. 1958. - Trilobiti ceskeho stredniho kambria. Rozpravy Ustredniho ustavu geologickeho 24: 1 - 280." type="journal article" year="1958">Šnajdr (1958)</bibRefCitation>
. From the
<location LSID="urn:lsid:plazi:treatment:03DA87B6FF904947FF597B2DFE6AFC2E:8EAC607BFF90494BFB4D7A57FCBFFDBD" country="Czech Republic" name="Buchava Formation, Skreyje Member, Eccaparadoxides pusillus Zone from Skryje-Tyrovice Basin" pageId="7" pageNumber="1056">
Buchava Formation, Skreyje Member,
<taxonomicName authority="Zone" authorityName="Zone" box="[975,1252,585,609]" class="Trilobita" family="Paradoxididae" genus="Eccaparadoxides" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis box="[975,1195,585,609]" italics="true" pageId="7" pageNumber="1056">Eccaparadoxides pusillus</emphasis>
Zone
</taxonomicName>
from Skryje-Týřovice Basin
</location>
, Bohemia,
<collectingCountry box="[977,1134,612,636]" name="Czech Republic" pageId="7" pageNumber="1056">Czech Republic</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="1056" type="diagnosis">
<paragraph blockId="7.[811,1455,659,763]" pageId="7" pageNumber="1056">
DIAGNOSIS. — Portion in front of cranidium domed; front of glabella rounded to truncate; eyes distant at anterior half of glabella; pair of eye ridges expands from front of glabella towards eye lobes (based on
<bibRefCitation author="Miller" box="[845,962,739,763]" firstAuthor="Miller" pageId="7" pageNumber="1056" refId="ref28272" refString="MILLER S. A. 1889. - North American Geology and Paleontology for the Use of Amateurs, Students, and Scientists. Western Methodist book concern, Cincinnati, Ohio, 793 p. https: // doi. org / 10.5962 / bhl. title. 28778" type="book" year="1889">Miller 1889</bibRefCitation>
;
<bibRefCitation author="Fletcher" box="[974,1111,739,763]" firstAuthor="Fletcher" pageId="7" pageNumber="1056" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017</bibRefCitation>
, with modifications).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="1056" type="materials_examined">
<paragraph blockId="7.[811,1457,786,970]" pageId="7" pageNumber="1056">
MATERIAL EXAMINED. —
<materialsCitation ID-GBIF-Occurrence="4003346305" pageId="7" pageNumber="1056">
537 cranidia of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[1218,1413,786,810]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[1218,1413,786,810]" italics="true" pageId="7" pageNumber="1056">Agraulos ceticephalus</emphasis>
</taxonomicName>
(for
<collectionCode box="[813,870,813,836]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15126" lsid="urn:lsid:biocol.org:col:15126" name="The Rooms Corporation of Newfoundland and Labrador, Provincial Museum Division" pageId="7" pageNumber="1056" type="Museum">NFM</collectionCode>
numbers see Appendix 1)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="4003346314" pageId="7" pageNumber="1056">
<specimenCount box="[1137,1268,813,837]" pageId="7" pageNumber="1056" type="generic">24 specimens</specimenCount>
are attached to the thorax and seven have at least one librigena attached. The specimens are well to very well-preserved as internal casts and moulds. Some are pyritized and in a few the exoskeleton is preserved
</materialsCitation>
. All specimens range between 1.94 and
<quantity box="[1039,1106,919,943]" metricMagnitude="0" metricUnit="m" metricValue="9.11" pageId="7" pageNumber="1056" unit="m" value="9.11">9.11 m</quantity>
(
<figureCitation box="[1120,1172,919,943]" captionStart="FIG" captionStartId="5.[132,143,1986,2003]" captionTargetBox="[160,1402,216,1896]" captionTargetId="graphics-15@5.[160,1399,257,1895]" captionTargetPageId="5" captionText="FIG. 2. — Lithology of the type locality of the Manuels River Formation, range chart of the here described species. Agnostid biozones from Hildenbrand et al. (2021) and polymerid biozones from Howell (1925)." figureDoi="http://doi.org/10.5281/zenodo.7477661" httpUri="https://zenodo.org/record/7477661/files/figure.png" pageId="7" pageNumber="1056">Fig. 2</figureCitation>
) of the Manuels River Formation,
<typeStatus box="[863,905,946,970]" pageId="7" pageNumber="1056">type</typeStatus>
locality, Conception Bay South, Newfoundland,
<collectingCountry box="[1375,1452,945,969]" name="Canada" pageId="7" pageNumber="1056">Canada</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="1056" type="distribution">
<paragraph blockId="7.[811,1455,993,1308]" pageId="7" pageNumber="1056">
OCCURRENCE. —
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[982,1166,993,1017]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[982,1166,993,1017]" italics="true" pageId="7" pageNumber="1056">Agraulos ceticephalus</emphasis>
</taxonomicName>
is a common middle Cambrian trilobite and has been documented from southeastern
<collectingCountry box="[1306,1381,1019,1043]" name="Canada" pageId="7" pageNumber="1056">Canada</collectingCountry>
, eastern Newfoundland, in the
<taxonomicName box="[1022,1185,1046,1070]" class="Trilobita" family="Ptychagnostidae" genus="Tomagnostus" kingdom="Animalia" order="Agnostida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="fissus">
<emphasis box="[1022,1185,1046,1070]" italics="true" pageId="7" pageNumber="1056">Tomagnostus fissus</emphasis>
</taxonomicName>
to
<taxonomicName box="[1212,1398,1046,1070]" class="Trilobita" family="Ptychagnostidae" genus="Ptychagnostus" kingdom="Animalia" order="Agnostida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="atavus">
<emphasis box="[1212,1398,1046,1070]" italics="true" pageId="7" pageNumber="1056">Ptychagnostus atavus</emphasis>
</taxonomicName>
zones (
<bibRefCitation author="Fletcher" box="[821,1002,1072,1096]" etAl="et al." firstAuthor="Fletcher" pageId="7" pageNumber="1056" pagination="312 - 336" refId="ref25165" refString="FLETCHER T. P., THEOKRITOFF G., LORD G. S. &amp; ZEOLI G. 2005. - The early paradoxidid harlani trilobite fauna of Massachusetts and its correlatives in Newfoundland, Morocco, and Spain. Journal of Paleontology 79 (2): 312 - 336. https: // doi. org / bdcjsg" type="journal article" year="2005">
Fletcher
<emphasis box="[904,949,1072,1096]" italics="true" pageId="7" pageNumber="1056">et al.</emphasis>
2005
</bibRefCitation>
),
<taxonomicName box="[1022,1212,1072,1096]" class="Trilobita" family="Ptychagnostidae" genus="Ptychagnostus" kingdom="Animalia" order="Agnostida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="atavus">
<emphasis box="[1022,1212,1072,1096]" italics="true" pageId="7" pageNumber="1056">Ptychagnostus atavus</emphasis>
</taxonomicName>
to
<taxonomicName box="[1240,1454,1072,1096]" class="Trilobita" family="Spinagnostidae" genus="Hypagnostus" kingdom="Animalia" order="Agnostida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="parvifrons">
<emphasis box="[1240,1454,1072,1096]" italics="true" pageId="7" pageNumber="1056">Hypagnostus parvifrons</emphasis>
</taxonomicName>
zones (
<bibRefCitation author="Fletcher" box="[881,1017,1098,1123]" firstAuthor="Fletcher" pageId="7" pageNumber="1056" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017</bibRefCitation>
), and
<taxonomicName authority="Zone" authorityName="Zone" box="[1082,1309,1098,1122]" class="Trilobita" family="Paradoxididae" genus="Paradoxides" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="species" species="hicksi">
<emphasis box="[1082,1251,1098,1122]" italics="true" pageId="7" pageNumber="1056">Paradoxides hicksi</emphasis>
Zone
</taxonomicName>
(
<figureCitation box="[1324,1380,1098,1123]" captionStart="FIG" captionStartId="5.[132,143,1986,2003]" captionTargetBox="[160,1402,216,1896]" captionTargetId="graphics-15@5.[160,1399,257,1895]" captionTargetPageId="5" captionText="FIG. 2. — Lithology of the type locality of the Manuels River Formation, range chart of the here described species. Agnostid biozones from Hildenbrand et al. (2021) and polymerid biozones from Howell (1925)." figureDoi="http://doi.org/10.5281/zenodo.7477661" httpUri="https://zenodo.org/record/7477661/files/figure.png" pageId="7" pageNumber="1056">Fig. 2</figureCitation>
). It has also been reported from the
<collectingCountry box="[1087,1255,1125,1149]" name="United Kingdom" pageId="7" pageNumber="1056">United Kingdom</collectingCountry>
in Wales (
<taxonomicName authority="zone" authorityName="Zone" class="Trilobita" family="Spinagnostidae" genus="Hypagnostus" kingdom="Animalia" order="Agnostida" pageId="7" pageNumber="1078" phylum="Arthropoda" rank="species" species="parvifrons">
<emphasis italics="true" pageId="7" pageNumber="1056">Hypagnostus parvifrons</emphasis>
zone
</taxonomicName>
;
<bibRefCitation author="Rees" box="[1013,1171,1151,1176]" etAl="et al." firstAuthor="Rees" pageId="7" pageNumber="1056" pagination="1 - 135" refId="ref28872" refString="REES A., THOMAS A., LEWIS M., HUGHES H. &amp; TURNER P. 2014. - Cambrian of Sw Wales: towards a united Avalonian Stratigraphy. Memoirs of the Geological Society London 42: 1 - 135. https: // doi. org / 10.1144 / m 42.0" type="journal article" year="2014">
Rees
<emphasis box="[1064,1113,1151,1175]" italics="true" pageId="7" pageNumber="1056">et al.</emphasis>
2014
</bibRefCitation>
),
<collectingCountry box="[1194,1289,1151,1175]" name="Denmark" pageId="7" pageNumber="1056">Denmark</collectingCountry>
(
<taxonomicName authority="zone" authorityName="Zone" class="Trilobita" family="Ptychagnostidae" genus="Acidusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="7" pageNumber="1084" phylum="Arthropoda" rank="species" species="atavus">
<emphasis box="[1305,1455,1151,1175]" italics="true" pageId="7" pageNumber="1056">Acidusus atavus</emphasis>
zone
</taxonomicName>
;
<bibRefCitation author="Weidner &amp; Nielsen" box="[868,1109,1178,1202]" firstAuthor="Weidner" pageId="7" pageNumber="1056" pagination="23 - 92" refId="ref30854" refString="WEIDNER T. &amp; NIELSEN A. T. 2014. - A highly diverse trilobite fauna with Avalonian affinities from the middle Cambrian Acidusus atavus Zone (Drumian Stage) of Bornholm, Denmark. Journal of Systematic Palaeontology 12 (1): 23 - 92. https: // doi. org / 10.10 80 / 14772019.2012.740080" type="journal article" year="2014">Weidner &amp; Nielsen 2014</bibRefCitation>
,
<bibRefCitation author="Weidner &amp; Nielsen" box="[1121,1171,1178,1202]" firstAuthor="Weidner" pageId="7" pageNumber="1056" pagination="1 - 11" refId="ref30914" refString="WEIDNER T. &amp; NIELSEN A. T. 2015. - Agraulos longicephalus and Proampyx? depressus (Trilobita) from the Middle Cambrian of Bornholm, Denmark. Bulletin of the Geological Society of Denmark 63: 1 - 11. https: // doi. org / 10.37570 / bgsd- 2015 - 63 - 01" type="journal article" year="2015">2015</bibRefCitation>
), in Bornholm (
<taxonomicName authority="zone" authorityName="ZONE" class="Trilobita" family="Ptychagnostidae" genus="Ptychagnostus" kingdom="Animalia" order="Agnostida" pageId="7" pageNumber="1078" phylum="Arthropoda" rank="species" species="punctuosus">
<emphasis italics="true" pageId="7" pageNumber="1056">Ptychagnostus punctuosus</emphasis>
zone
</taxonomicName>
;
<bibRefCitation author="Rudolph" box="[978,1122,1204,1228]" firstAuthor="Rudolph" pageId="7" pageNumber="1056" refId="ref29287" refString="RUDOLPH F. 1994. - Die Trilobiten der mittelkambrischen Geschiebe: Systematik, Morphologie und Okologie. Verlag Frank Rudolph, Wankendorf, 309 p." type="book" year="1994">Rudolph 1994</bibRefCitation>
),
<collectingCountry box="[1142,1207,1205,1228]" name="France" pageId="7" pageNumber="1056">France</collectingCountry>
(
<bibRefCitation author="Thoral" box="[1221,1343,1204,1228]" firstAuthor="Thoral" pageId="7" pageNumber="1056" refId="ref30557" refString="THORAL M. 1935. - Contribution a l'etude paleontologique de l'Ordovicien inferieur de la Montagne Noire et revision sommaire de la faune cambrienne de la Montagne Noire. Unpublished PhD thesis, Universite de Paris, Montpellier, 362 p." type="book" year="1935">Thoral 1935</bibRefCitation>
),
<collectingCountry box="[1363,1455,1204,1228]" name="Germany" pageId="7" pageNumber="1056">Germany</collectingCountry>
(
<bibRefCitation author="Wurm" box="[821,942,1231,1255]" firstAuthor="Wurm" pageId="7" pageNumber="1056" pagination="71 - 93" refId="ref31321" refString="WURM A. 1925. - Ueber ein Vorkommen von Mittelcambrium (Paradoxidesschichten) im bayrischen Frankenwald bei Wildenstein sudlich Presseck. Neues Jahrbuch fur Mineralogie, Geognosie Geologie und Palaontologie 52 (1): 71 - 93." type="journal article" year="1925">Wurm 1925</bibRefCitation>
), the
<collectingCountry box="[1001,1154,1231,1255]" name="Czech Republic" pageId="7" pageNumber="1056">Czech Republic</collectingCountry>
(
<taxonomicName authority="zone" authorityName="Zone" box="[1170,1452,1231,1255]" class="Trilobita" family="Paradoxididae" genus="Eccaparadoxides" kingdom="Animalia" order="Redlichiida" pageId="7" pageNumber="1078" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis box="[1170,1397,1231,1255]" italics="true" pageId="7" pageNumber="1056">Eccaparadoxides pusillus</emphasis>
zone
</taxonomicName>
; e.g.,
<bibRefCitation author="Snajdr" box="[861,984,1257,1285]" firstAuthor="Snajdr" pageId="7" pageNumber="1056" refId="ref30293" refString="SNAJDR M. 1990. - Bohemian Trilobites. Geological Survey, Prague, 265 p." type="book" year="1990">Šnajdr 1990</bibRefCitation>
;
<bibRefCitation author="Fletcher" box="[998,1193,1257,1281]" etAl="et al." firstAuthor="Fletcher" pageId="7" pageNumber="1056" pagination="312 - 336" refId="ref25165" refString="FLETCHER T. P., THEOKRITOFF G., LORD G. S. &amp; ZEOLI G. 2005. - The early paradoxidid harlani trilobite fauna of Massachusetts and its correlatives in Newfoundland, Morocco, and Spain. Journal of Paleontology 79 (2): 312 - 336. https: // doi. org / bdcjsg" type="journal article" year="2005">
Fletcher
<emphasis box="[1086,1135,1257,1281]" italics="true" pageId="7" pageNumber="1056">et al.</emphasis>
2005
</bibRefCitation>
), and
<collectingCountry box="[1260,1315,1257,1281]" name="Spain" pageId="7" pageNumber="1056">Spain</collectingCountry>
(
<taxonomicName authorityName="Thoral" authorityYear="1947" box="[1331,1455,1257,1281]" class="Trilobita" family="Solenopleuridae" genus="Pardailhania" kingdom="Animalia" order="Agnostida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="genus">
<emphasis box="[1331,1455,1257,1281]" italics="true" pageId="7" pageNumber="1056">Pardailhania</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Thoral" authorityYear="1947" box="[855,1005,1284,1308]" class="Trilobita" family="Solenopleuridae" genus="Solenopleuropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Agnostida" pageId="7" pageNumber="1056" phylum="Arthropoda" rank="genus">
<emphasis box="[855,1005,1284,1308]" italics="true" pageId="7" pageNumber="1056">Solenopleuropsis</emphasis>
</taxonomicName>
zones;
<bibRefCitation author="Linan &amp; Gozalo" box="[1077,1295,1284,1308]" firstAuthor="Linan" pageId="7" pageNumber="1056" pagination="1 - 104" refId="ref27824" refString="LINAN E. &amp; GOZALO R. 1986. - Trilobites del Cambrico inferior y medo de murero (Cordillera Iberica). Memorias del Museo Paleontologico de la Universidad de Zaragoza 2: 1 - 104." type="journal article" year="1986">Liñan &amp; Gozalo 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="1056" type="description">
<paragraph blockId="7.[810,1457,1364,2028]" box="[813,957,1364,1390]" pageId="7" pageNumber="1056">DESCRIPTION</paragraph>
<paragraph blockId="7.[810,1457,1364,2028]" pageId="7" pageNumber="1056">The cranidia range from 3.0 mm to 16.0 mm in width and from 2.0 mm to 11.0 mm in length. The shape of the glabella varies from slightly trapezoidal to more equal-sided, with a rounded to truncate front. Three to four pairs of glabellar furrows are preserved mainly in larger-sized specimens.In small-sized specimens, the glabella is steeply domed, while it is flatter in largesized specimens.Nevertheless, the glabella is always more domed than the cheeks.Eye ridges, if preserved, initiate near the front of the glabella and connect with the eye lobes in a horizontal line, some line in a backwards angle towards the sides. The occipital ring points backwards roughly triangularly and the length of the spine varies with size. Some moulds of cephala connected to the thorax spare spines to nodes on up to five axial segments. These and an ornamentation are usually preserved on moulds. Several cephala show up to three variably pronounced grooves. One of these reaches from the front of the glabella to the anterior margin.The other two are developed towards the sides, varying in position. The front of the cranidium is highly domed but a lateral view and microscope measurements indicate that this is an optical illusion and it is still rather flattened domed.</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7477665" ID-Zenodo-Dep="7477665" httpUri="https://zenodo.org/record/7477665/files/figure.png" pageId="8" pageNumber="1057" startId="8.[132,143,1849,1866]" targetBox="[132,1455,215,1783]" targetPageId="8">
<paragraph blockId="8.[132,1456,1849,2025]" pageId="8" pageNumber="1057">
FIG. 4.
<emphasis bold="true" box="[188,207,1849,1866]" pageId="8" pageNumber="1057"></emphasis>
<taxonomicName authority="(Barrande, 1846)" baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[212,543,1849,1866]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="8" pageNumber="1057" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[212,396,1849,1866]" italics="true" pageId="8" pageNumber="1057">Agraulos ceticephalus</emphasis>
(
<bibRefCitation author="Barrande" box="[405,538,1849,1866]" firstAuthor="Barrande" pageId="8" pageNumber="1057" refId="ref23464" refString="BARRANDE J. 1846. - Notice preliminaire sur le systeme silurien et les Trilobites de Boheme. Hirschfeld, Leipzig, 96 p. https: // doi. org / 10.5962 / bhl. title. 9142" type="book" year="1846">Barrande, 1846</bibRefCitation>
)
</taxonomicName>
:
<emphasis bold="true" box="[550,563,1849,1866]" pageId="8" pageNumber="1057">A</emphasis>
, internal cast of a cranidium attached to the thorax with preserved exoskeleton (NFM F-3476 from 5.97 m);
<emphasis bold="true" box="[132,145,1872,1889]" pageId="8" pageNumber="1057">B</emphasis>
, internal cast of a cranidium attached to the thorax, partly covered with sediment, eye ridges and glabellar furrows slightly visible (NFM F-3951 from 7.10 m);
<emphasis bold="true" box="[132,146,1895,1912]" pageId="8" pageNumber="1057">C</emphasis>
, internal cast of a cranidium attached to the thorax with preserved exoskeleton, eye ridges present (NFM F-3957 from 3.30 m);
<emphasis bold="true" box="[1210,1224,1895,1912]" pageId="8" pageNumber="1057">D</emphasis>
, internal cast of a cephalon attached to the thorax, librigena attached, eye ridges present (NFM F-3621 from 9.08 m);
<emphasis bold="true" box="[882,894,1917,1934]" pageId="8" pageNumber="1057">E</emphasis>
, mould of a cranidium attached to the thorax, spines on axial rings present,coarse ornamentation on thorax visible (NFM F-3476 from 5.98 m);
<emphasis bold="true" box="[752,763,1940,1957]" pageId="8" pageNumber="1057">F</emphasis>
, mould of a cranidium attached to the thorax,occipital spine preserved (sedimentary filling laying in glabellar mould), coarse ornamentation on thorax visible (NFM F-3946 from 4.27 m);
<emphasis bold="true" box="[950,964,1963,1980]" pageId="8" pageNumber="1057">G</emphasis>
, internal cast of a cranidium attached to the thorax, projection of axial spine preserved (NFM F-3831 from 7.43 m);
<emphasis bold="true" box="[609,623,1986,2003]" pageId="8" pageNumber="1057">H</emphasis>
, internal cast of a cranidium attached to the thorax, eye ridges present (NFM F-2787 from 7.56 m);
<emphasis bold="true" box="[132,138,2008,2025]" pageId="8" pageNumber="1057">I</emphasis>
, internal cast of a cranidium attached to first thoracic segments, eye ridges and glabellar furrows preserved (NFM F-3778 from 5.96 m). Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection lastPageId="11" lastPageNumber="1060" pageId="9" pageNumber="1058" type="discussion">
<paragraph blockId="9.[130,777,216,2027]" box="[132,232,216,242]" pageId="9" pageNumber="1058">REMARKS</paragraph>
<paragraph blockId="9.[130,777,216,2027]" pageId="9" pageNumber="1058">
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[132,333,247,273]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[132,333,247,273]" italics="true" pageId="9" pageNumber="1058">Agraulos ceticephalus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[378,594,247,273]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[378,594,247,273]" italics="true" pageId="9" pageNumber="1058">Agraulos longicephalus</emphasis>
</taxonomicName>
are closely related species (e.g.,
<bibRefCitation author="Hicks" box="[264,385,279,305]" firstAuthor="Hicks" pageId="9" pageNumber="1058" pagination="173 - 185" refId="ref26120" refString="HICKS H. 1872. - On some undescribed fossils from the Menevian Group. The Quarterly Journal of the Geological Society of London 28 (1): 173 - 185. https: // doi. org / 10.1144 / gsl. jgs. 1872.028.01 - 02.17" type="journal article" year="1872">Hicks 1872</bibRefCitation>
;
<bibRefCitation author="Lake" box="[399,509,279,305]" firstAuthor="Lake" pageId="9" pageNumber="1058" pagination="149 - 172" refId="ref27339" refString="LAKE P. 1932. - Monograph of the British Cambrian Trilobites, Part 7. Monograph of the Palaeontographical Society, London 84: 149 - 172. https: // doi. org / 10.1080 / 02693445.1932.12035620" type="journal article" year="1932">Lake 1932</bibRefCitation>
;
<bibRefCitation author="Sdzuy" box="[521,645,279,305]" firstAuthor="Sdzuy" pageId="9" pageNumber="1058" pagination="503 - 594" refId="ref30076" refString="SDZUY K. 1961. - Das Kambrium Spaniens, Teil II: Trilobiten. 1 Akademie der Wissenschaften und der Literatur, Mainz, Abhandlungen der mathematisch-naturwissenschaftlichen Klasse 7: 503 - 594." type="journal article" year="1961">Sdzuy 1961</bibRefCitation>
;
<bibRefCitation author="Weidner &amp; Nielsen" firstAuthor="Weidner" pageId="9" pageNumber="1058" pagination="1 - 11" refId="ref30914" refString="WEIDNER T. &amp; NIELSEN A. T. 2015. - Agraulos longicephalus and Proampyx? depressus (Trilobita) from the Middle Cambrian of Bornholm, Denmark. Bulletin of the Geological Society of Denmark 63: 1 - 11. https: // doi. org / 10.37570 / bgsd- 2015 - 63 - 01" type="journal article" year="2015">Weidner &amp; Nielsen 2015</bibRefCitation>
) and both show a wide range of intraspecific variations, e.g. regarding the front of glabella or size and projection of the occipital spine (
<bibRefCitation author="Barrande" box="[408,577,374,401]" firstAuthor="Barrande" pageId="9" pageNumber="1058" pagination="1 - 935" refId="ref23595" refString="BARRANDE J. 1852 b. - Systeme silurien du Centre de la Boheme. 1 ere Partie: Texte. Crustaces: Trilobites. Recherches paleontologiques 1: 1 - 935. https: // www. biodiversitylibrary. org / page / 57395510" type="journal article" year="1852" yearSuffix="b">Barrande 1852b</bibRefCitation>
;
<bibRefCitation author="Lake" box="[589,699,375,401]" firstAuthor="Lake" pageId="9" pageNumber="1058" pagination="149 - 172" refId="ref27339" refString="LAKE P. 1932. - Monograph of the British Cambrian Trilobites, Part 7. Monograph of the Palaeontographical Society, London 84: 149 - 172. https: // doi. org / 10.1080 / 02693445.1932.12035620" type="journal article" year="1932">Lake 1932</bibRefCitation>
;
<bibRefCitation author="Sdzuy" firstAuthor="Sdzuy" pageId="9" pageNumber="1058" pagination="503 - 594" refId="ref30076" refString="SDZUY K. 1961. - Das Kambrium Spaniens, Teil II: Trilobiten. 1 Akademie der Wissenschaften und der Literatur, Mainz, Abhandlungen der mathematisch-naturwissenschaftlichen Klasse 7: 503 - 594." type="journal article" year="1961">Sdzuy 1961</bibRefCitation>
).
<bibRefCitation author="Barrande" box="[212,405,406,433]" firstAuthor="Barrande" pageId="9" pageNumber="1058" pagination="1 - 935" refId="ref23595" refString="BARRANDE J. 1852 b. - Systeme silurien du Centre de la Boheme. 1 ere Partie: Texte. Crustaces: Trilobites. Recherches paleontologiques 1: 1 - 935. https: // www. biodiversitylibrary. org / page / 57395510" type="journal article" year="1852" yearSuffix="b">Barrande (1852b)</bibRefCitation>
documented ontogenetic changes in
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[160,318,438,465]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[160,318,438,465]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. He emphasized an increase in depth of the occipital furrow and dorsal furrows, less domed genal regions, and an increase in the thickness of exoskeletons in adult forms. The latter was also identified in
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[496,679,534,560]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[496,679,534,560]" italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
by
<bibRefCitation author="Lake" firstAuthor="Lake" pageId="9" pageNumber="1058" pagination="149 - 172" refId="ref27339" refString="LAKE P. 1932. - Monograph of the British Cambrian Trilobites, Part 7. Monograph of the Palaeontographical Society, London 84: 149 - 172. https: // doi. org / 10.1080 / 02693445.1932.12035620" type="journal article" year="1932">Lake (1932)</bibRefCitation>
. Accordingly, the development of glabellar, axial and other furrows as well as the occipital ring are not considered to be reliable diagnostic characteristics. Also, specimens with a preserved exoskeleton may show morphological details not identified in internal casts (
<bibRefCitation author="Lake" box="[420,532,693,720]" firstAuthor="Lake" pageId="9" pageNumber="1058" pagination="149 - 172" refId="ref27339" refString="LAKE P. 1932. - Monograph of the British Cambrian Trilobites, Part 7. Monograph of the Palaeontographical Society, London 84: 149 - 172. https: // doi. org / 10.1080 / 02693445.1932.12035620" type="journal article" year="1932">Lake 1932</bibRefCitation>
;
<bibRefCitation author="Sdzuy" box="[545,671,693,719]" firstAuthor="Sdzuy" pageId="9" pageNumber="1058" pagination="503 - 594" refId="ref30076" refString="SDZUY K. 1961. - Das Kambrium Spaniens, Teil II: Trilobiten. 1 Akademie der Wissenschaften und der Literatur, Mainz, Abhandlungen der mathematisch-naturwissenschaftlichen Klasse 7: 503 - 594." type="journal article" year="1961">Sdzuy 1961</bibRefCitation>
). Despite the numerous descriptions and discussions about differences between
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[226,385,757,784]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[226,385,757,784]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[435,609,757,784]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[435,609,757,784]" italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
, differentiation is hence questionable and this work agrees with
<bibRefCitation author="Weidner &amp; Nielsen" firstAuthor="Weidner" pageId="9" pageNumber="1058" pagination="23 - 92" refId="ref30854" refString="WEIDNER T. &amp; NIELSEN A. T. 2014. - A highly diverse trilobite fauna with Avalonian affinities from the middle Cambrian Acidusus atavus Zone (Drumian Stage) of Bornholm, Denmark. Journal of Systematic Palaeontology 12 (1): 23 - 92. https: // doi. org / 10.10 80 / 14772019.2012.740080" type="journal article" year="2014">Weidner &amp; Nielsen (2014)</bibRefCitation>
that the genus
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[457,544,821,847]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="genus">
<emphasis box="[457,544,821,847]" italics="true" pageId="9" pageNumber="1058">Agraulos</emphasis>
</taxonomicName>
requires revision.
</paragraph>
<paragraph blockId="9.[130,777,216,2027]" pageId="9" pageNumber="1058">
<bibRefCitation author="Barrande" box="[155,344,853,880]" firstAuthor="Barrande" pageId="9" pageNumber="1058" refId="ref23551" refString="BARRANDE J. 1852 a. - Systeme silurien du Centre de la Boheme. 1 ere Partie: Planches. Crustaces: Trilobites. Recherches paleontologiques 1: 49 pls. https: // www. biodiversitylibrary. org / page / 47135930" type="book" year="1852" yearSuffix="a">Barrande (1852a)</bibRefCitation>
illustrated several specimens of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. The articulated specimens figured by the author on pl. 10, figs 1-5, and the cranidium on pl. 10, fig. 7, do not show the characteristic long preglabellar area of the genus
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[134,220,980,1006]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="genus">
<emphasis box="[134,220,980,1006]" italics="true" pageId="9" pageNumber="1058">Agraulos</emphasis>
</taxonomicName>
and are here excluded from
<emphasis box="[511,675,980,1007]" italics="true" pageId="9" pageNumber="1058">
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[511,671,980,1007]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">Ag. ceticephalus</taxonomicName>
,
</emphasis>
and even the genus. An enrolled specimen illustrated in lateral view of
<bibRefCitation author="Barrande" box="[133,313,1044,1071]" firstAuthor="Barrande" pageId="9" pageNumber="1058" refId="ref23551" refString="BARRANDE J. 1852 a. - Systeme silurien du Centre de la Boheme. 1 ere Partie: Planches. Crustaces: Trilobites. Recherches paleontologiques 1: 49 pls. https: // www. biodiversitylibrary. org / page / 47135930" type="book" year="1852" yearSuffix="a">Barrande (1852a</bibRefCitation>
: pl. 10, fig. 21) shows no diagnostic characteristics and assignment to
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[448,534,1076,1102]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="genus">
<emphasis box="[448,534,1076,1102]" italics="true" pageId="9" pageNumber="1058">Agraulos</emphasis>
</taxonomicName>
is questionable.
<bibRefCitation author="Hicks" firstAuthor="Hicks" pageId="9" pageNumber="1058" pagination="173 - 185" refId="ref26120" refString="HICKS H. 1872. - On some undescribed fossils from the Menevian Group. The Quarterly Journal of the Geological Society of London 28 (1): 173 - 185. https: // doi. org / 10.1144 / gsl. jgs. 1872.028.01 - 02.17" type="journal article" year="1872">Hicks (1872)</bibRefCitation>
first described
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[359,530,1108,1134]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[359,530,1108,1134]" italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
based on a longer form and a more domed genal region than seen in
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[611,768,1140,1166]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[611,768,1140,1166]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. However, all cranidia figured by
<bibRefCitation author="Hicks" box="[471,613,1172,1198]" firstAuthor="Hicks" pageId="9" pageNumber="1058" pagination="173 - 185" refId="ref26120" refString="HICKS H. 1872. - On some undescribed fossils from the Menevian Group. The Quarterly Journal of the Geological Society of London 28 (1): 173 - 185. https: // doi. org / 10.1144 / gsl. jgs. 1872.028.01 - 02.17" type="journal article" year="1872">Hicks (1872)</bibRefCitation>
, some attached to the thorax, are deformed, some elongated, and others compressed in length, as already mentioned by
<bibRefCitation author="Fletcher" box="[602,770,1236,1262]" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017)</bibRefCitation>
. The longer form of the cephalon is here interpreted as a result of tectonic deformation and the more domed genal region may also represent a preservational artefact. All
<specimenCount box="[663,777,1332,1358]" pageId="9" pageNumber="1058" type="generic" typeStatus="type">seven type</specimenCount>
specimens illustrated byHicks (1872) are here assigned to
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Roch" box="[263,398,1395,1421]" firstAuthor="Roch" pageId="9" pageNumber="1058" refId="ref29177" refString="ROCH E. 1930. - Etudes geologiques dans la region Meridionale du Maroc occidental. Protat Freres imprimeurs, Macon, 542 p." type="book" year="1930">Roch (1930)</bibRefCitation>
also referred to the similarity of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[292,459,1427,1453]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[292,459,1427,1453]" italics="true" pageId="9" pageNumber="1058">Ag.longicephalus</emphasis>
</taxonomicName>
. He compared specimens from
<collectingCountry box="[130,226,1459,1485]" name="Morocco" pageId="9" pageNumber="1058">Morocco</collectingCountry>
and
<collectingCountry box="[277,351,1459,1485]" name="Algeria" pageId="9" pageNumber="1058">Algeria</collectingCountry>
to
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[387,543,1459,1485]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[387,543,1459,1485]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
from the
<typeStatus box="[647,692,1459,1485]" pageId="9" pageNumber="1058">type</typeStatus>
locality in Central Europe. Nevertheless, the assignment of the material to
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[204,362,1523,1549]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[204,362,1523,1549]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
is questionable as no illustrations were presented byRoch (1930) and no information given regarding the number of studied specimens.
<bibRefCitation author="Lake" box="[495,625,1586,1613]" firstAuthor="Lake" pageId="9" pageNumber="1058" pagination="149 - 172" refId="ref27339" refString="LAKE P. 1932. - Monograph of the British Cambrian Trilobites, Part 7. Monograph of the Palaeontographical Society, London 84: 149 - 172. https: // doi. org / 10.1080 / 02693445.1932.12035620" type="journal article" year="1932">Lake (1932)</bibRefCitation>
also discussed the morphological differentiation of
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[516,689,1618,1645]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[516,689,1618,1645]" italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. According toLake (1932)
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[534,705,1650,1677]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[534,705,1650,1677]" italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
has an occipital ring ending in a triangular point and a more truncate glabella.However,
<bibRefCitation author="Sdzuy" box="[324,466,1714,1740]" firstAuthor="Sdzuy" pageId="9" pageNumber="1058" pagination="503 - 594" refId="ref30076" refString="SDZUY K. 1961. - Das Kambrium Spaniens, Teil II: Trilobiten. 1 Akademie der Wissenschaften und der Literatur, Mainz, Abhandlungen der mathematisch-naturwissenschaftlichen Klasse 7: 503 - 594." type="journal article" year="1961">Sdzuy (1961)</bibRefCitation>
suggested that the form of the occipital spine, or occipital node, is a variable characteristic and unsuited to define
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[322,407,1778,1804]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="genus">
<emphasis box="[322,407,1778,1804]" italics="true" pageId="9" pageNumber="1058">Agraulos</emphasis>
</taxonomicName>
. We here follow this latter view and interpret the more truncate glabella to fall within the intraspecific variation of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[307,463,1841,1868]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[307,463,1841,1868]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. The presented specimens are here assigned to this latter species.
</paragraph>
<paragraph blockId="9.[130,777,216,2027]" lastBlockId="9.[810,1457,215,2028]" pageId="9" pageNumber="1058">
<bibRefCitation author="Sdzuy" box="[155,296,1905,1931]" firstAuthor="Sdzuy" pageId="9" pageNumber="1058" pagination="503 - 594" refId="ref30076" refString="SDZUY K. 1961. - Das Kambrium Spaniens, Teil II: Trilobiten. 1 Akademie der Wissenschaften und der Literatur, Mainz, Abhandlungen der mathematisch-naturwissenschaftlichen Klasse 7: 503 - 594." type="journal article" year="1961">Sdzuy (1961)</bibRefCitation>
described a variable shape of the occipital ring of
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[158,325,1937,1964]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[158,325,1937,1964]" italics="true" pageId="9" pageNumber="1058">Ag.longicephalus</emphasis>
</taxonomicName>
, stating the shape as non-diagnostic. He followed
<bibRefCitation author="Lake" box="[198,322,1969,1995]" firstAuthor="Lake" pageId="9" pageNumber="1058" pagination="149 - 172" refId="ref27339" refString="LAKE P. 1932. - Monograph of the British Cambrian Trilobites, Part 7. Monograph of the Palaeontographical Society, London 84: 149 - 172. https: // doi. org / 10.1080 / 02693445.1932.12035620" type="journal article" year="1932">Lake (1932)</bibRefCitation>
in that
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[394,560,1969,1995]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[394,560,1969,1995]" italics="true" pageId="9" pageNumber="1058">Ag.longicephalus</emphasis>
</taxonomicName>
is closely allied to
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
but that the glabella of
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[499,670,2001,2027]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[499,670,2001,2027]" italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
is slightly more narrowed towards the front,and mentioned a more domed genal region. We rather suggest that the domed genal region of
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[811,983,279,305]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[811,983,279,305]" italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
results from preservational bias and that the narrowed front of the glabella ranges within the intraspecific variation of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[942,1097,343,369]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[942,1097,343,369]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. Hence, all cranidia illustrated by
<bibRefCitation author="Sdzuy" box="[814,958,375,401]" firstAuthor="Sdzuy" pageId="9" pageNumber="1058" pagination="503 - 594" refId="ref30076" refString="SDZUY K. 1961. - Das Kambrium Spaniens, Teil II: Trilobiten. 1 Akademie der Wissenschaften und der Literatur, Mainz, Abhandlungen der mathematisch-naturwissenschaftlichen Klasse 7: 503 - 594." type="journal article" year="1961">Sdzuy (1961)</bibRefCitation>
are here assigned to
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[1188,1347,375,401]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[1188,1347,375,401]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. Cranidia assigned to
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[923,1086,406,433]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[923,1086,406,433]" italics="true" pageId="9" pageNumber="1058">Ag.longicephalus</emphasis>
</taxonomicName>
byFletcher (1972b) andCourtessole (1973) also show the characteristics of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[1216,1372,438,465]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[1216,1372,438,465]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
and are here assigned to this species.
<bibRefCitation author="Linan &amp; Gozalo" box="[1104,1352,470,496]" firstAuthor="Linan" pageId="9" pageNumber="1058" pagination="1 - 104" refId="ref27824" refString="LINAN E. &amp; GOZALO R. 1986. - Trilobites del Cambrico inferior y medo de murero (Cordillera Iberica). Memorias del Museo Paleontologico de la Universidad de Zaragoza 2: 1 - 104." type="journal article" year="1986">Liñan &amp; Gozalo (1986)</bibRefCitation>
described two subspecies of
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[999,1170,502,529]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[999,1170,502,529]" italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
based on a more extended, respectively narrower preglabellar area compared to the border. These variations are also interpreted here as an intraspecific variation of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[901,1057,598,624]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[901,1057,598,624]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
and both subspecies are considered to be synonyms of the latter species. This interpretation includes cranidia, some attached to the thorax, illustrated by
<bibRefCitation author="Linan &amp; Gozalo" firstAuthor="Linan" pageId="9" pageNumber="1058" pagination="1 - 104" refId="ref27824" refString="LINAN E. &amp; GOZALO R. 1986. - Trilobites del Cambrico inferior y medo de murero (Cordillera Iberica). Memorias del Museo Paleontologico de la Universidad de Zaragoza 2: 1 - 104." type="journal article" year="1986">Liñan &amp; Gozalo (1986</bibRefCitation>
: pl. 35, figs 11-14, pl. 36, fig. 4, pl. 36, fig. 5), whereas cranidia illustrated by the authors in plate 36, figs 1-3, do not show the long preglabellar field typical for
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[1324,1409,757,783]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="genus">
<emphasis box="[1324,1409,757,783]" italics="true" pageId="9" pageNumber="1058">Agraulos</emphasis>
</taxonomicName>
and are here excluded from the species and even the genus.Specimens documented by
<bibRefCitation author="Martin &amp; Dean" box="[990,1243,821,848]" firstAuthor="Martin" pageId="9" pageNumber="1058" pagination="1 - 91" refId="ref27973" refString="MARTIN F. &amp; DEAN W. T. 1988. - Middle and upper Cambrian acritarch and trilobite zonation at Manuels River and Random Island, eastern Newfoundland. Geological Survey of Canada Bulletin 381 (1): 1 - 91." type="journal article" year="1988">Martin &amp; Dean (1988)</bibRefCitation>
as
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[1281,1457,821,847]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[1281,1457,821,847]" italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
are here assigned to
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[1019,1174,853,879]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[1019,1174,853,879]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
, except for pl. 3, figs 9 and 11. These images display the same cranidium devoid of a wide preglabellar field and are therefore atypical for
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[1275,1359,917,943]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="genus">
<emphasis box="[1275,1359,917,943]" italics="true" pageId="9" pageNumber="1058">Agraulos</emphasis>
</taxonomicName>
.
<bibRefCitation author="Fletcher" firstAuthor="Fletcher" pageId="9" pageNumber="1058" refId="ref24935" refString="FLETCHER T. P. 2006. - Bedrock Geology of the Cape St. Mary's Peninsula, Southwest Avalon Peninsula, Newfoundland (includes parts of Nts map sheets 1 M / 1, 1 N / 4, 1 L / 16 and 1 K 13). Report, 06 - 02. Government of Newfoundland and Labrador, Geological Survey, Department of Natural Resources, St. John's, 117 p." type="book" year="2006">Fletcher (2006)</bibRefCitation>
displayed
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[988,1140,949,975]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[988,1140,949,975]" italics="true" pageId="9" pageNumber="1058">Ag.ceticephalus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[1189,1357,949,975]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[1189,1357,949,975]" italics="true" pageId="9" pageNumber="1058">Ag.longicephalus</emphasis>
</taxonomicName>
. The two cranidia figured by the author have distant eyes and a rounded to truncate glabella; the cranidium illustrated on pl. 34, fig. 34, is here assigned to
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[1021,1181,1044,1071]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[1021,1181,1044,1071]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. Specimens presented by
<bibRefCitation author="Rees" box="[814,992,1076,1103]" etAl="et al." firstAuthor="Rees" pageId="9" pageNumber="1058" pagination="1 - 135" refId="ref28872" refString="REES A., THOMAS A., LEWIS M., HUGHES H. &amp; TURNER P. 2014. - Cambrian of Sw Wales: towards a united Avalonian Stratigraphy. Memoirs of the Geological Society London 42: 1 - 135. https: // doi. org / 10.1144 / m 42.0" type="journal article" year="2014">
Rees
<emphasis box="[866,915,1076,1102]" italics="true" pageId="9" pageNumber="1058">et al.</emphasis>
(2014)
</bibRefCitation>
and
<bibRefCitation author="Weidner &amp; Nielsen" box="[1042,1314,1076,1102]" firstAuthor="Weidner" pageId="9" pageNumber="1058" pagination="23 - 92" refId="ref30854" refString="WEIDNER T. &amp; NIELSEN A. T. 2014. - A highly diverse trilobite fauna with Avalonian affinities from the middle Cambrian Acidusus atavus Zone (Drumian Stage) of Bornholm, Denmark. Journal of Systematic Palaeontology 12 (1): 23 - 92. https: // doi. org / 10.10 80 / 14772019.2012.740080" type="journal article" year="2014">Weidner &amp; Nielsen (2014</bibRefCitation>
,
<bibRefCitation author="Weidner &amp; Nielsen" box="[1327,1381,1076,1103]" firstAuthor="Weidner" pageId="9" pageNumber="1058" pagination="1 - 11" refId="ref30914" refString="WEIDNER T. &amp; NIELSEN A. T. 2015. - Agraulos longicephalus and Proampyx? depressus (Trilobita) from the Middle Cambrian of Bornholm, Denmark. Bulletin of the Geological Society of Denmark 63: 1 - 11. https: // doi. org / 10.37570 / bgsd- 2015 - 63 - 01" type="journal article" year="2015">2015</bibRefCitation>
) as
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis italics="true" pageId="9" pageNumber="1058">Ag. longicephalus</emphasis>
</taxonomicName>
show the characteristics of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[1220,1374,1108,1134]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[1220,1374,1108,1134]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
and are hence assigned to this species.
</paragraph>
<paragraph blockId="9.[810,1457,215,2028]" lastBlockId="11.[130,778,215,1007]" lastPageId="11" lastPageNumber="1060" pageId="9" pageNumber="1058">
<bibRefCitation author="Fletcher" box="[835,1001,1172,1198]" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017)</bibRefCitation>
described and illustrated several species of
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[814,899,1204,1230]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="genus">
<emphasis box="[814,899,1204,1230]" italics="true" pageId="9" pageNumber="1058">Agraulos</emphasis>
</taxonomicName>
. The cranidium illustrated in fig. 11F was assigned to
<taxonomicName authority="Billings, 1872" authorityName="Billings" authorityYear="1872" box="[842,1155,1236,1262]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="affinis">
<emphasis box="[842,996,1236,1262]" italics="true" pageId="9" pageNumber="1058">Agraulos affinis</emphasis>
<bibRefCitation author="Billings" box="[1003,1155,1236,1262]" firstAuthor="Billings" pageId="9" pageNumber="1058" pagination="465 - 479" refId="ref23812" refString="BILLINGS E. 1872. - On some fossils from the primordial rocks of Newfoundland. Canadian Naturalist and Quarterly Journal of Science 6: 465 - 479. https: // doi. org / 10.5962 / bhl. title. 38279" type="journal article" year="1872">Billings, 1872</bibRefCitation>
</taxonomicName>
, by
<bibRefCitation author="Fletcher" box="[1199,1365,1236,1262]" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017)</bibRefCitation>
, but the specimen has eyes distant to the glabella. This character disagrees with the diagnosis of
<taxonomicName authorityName="Billings" authorityYear="1872" box="[1087,1186,1300,1326]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="affinis">
<emphasis box="[1087,1186,1300,1326]" italics="true" pageId="9" pageNumber="1058">Ag. affinis</emphasis>
</taxonomicName>
, in which eyes are located close to the glabella. Instead, the position of the eyes matches that seen in
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[940,1097,1363,1390]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[940,1097,1363,1390]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
and the specimen is here assigned to this species. Specimens illustrated in fig. 9A-C as
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
, on the other hand, display cranidia too deformed to determine the species. Specimens figured by
<bibRefCitation author="Fletcher" box="[1293,1448,1459,1485]" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017</bibRefCitation>
: figs 9D-I) as
<taxonomicName authority="Fletcher, 2017" authorityName="Fletcher" authorityYear="2017" box="[949,1255,1491,1517]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis box="[949,1097,1491,1517]" italics="true" pageId="9" pageNumber="1058">Agraulos lewisi</emphasis>
<bibRefCitation author="Fletcher" box="[1104,1255,1491,1517]" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher, 2017</bibRefCitation>
</taxonomicName>
are also too poorly preserved to be assigned specifically.
<figureCitation box="[1218,1336,1523,1549]" captionStart="FIG" captionStartId="6.[478,489,791,808]" captionTargetBox="[164,1456,217,767]" captionTargetId="graphics-581@6.[187,818,240,721]" captionTargetPageId="6" captionText="FIG. 3. — Visualization of measurement terms in cranidia (A) and pygidia (B)." figureDoi="http://doi.org/10.5281/zenodo.7477663" httpUri="https://zenodo.org/record/7477663/files/figure.png" pageId="9" pageNumber="1058">Figures 3A</figureCitation>
and
<figureCitation box="[1395,1424,1523,1549]" captionStart="FIG" captionStartId="14.[813,824,657,674]" captionTargetBox="[816,1451,215,616]" captionTargetId="figure-719@14.[816,1452,214,616]" captionTargetPageId="14" captionText="FIG. 8. —Bailiella aequalis (Linnarsson,1882) internal cast of a cranidium (NFM F-3811 from 5.90 m). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7477671" httpUri="https://zenodo.org/record/7477671/files/figure.png" pageId="9" pageNumber="1058">8F</figureCitation>
of
<bibRefCitation author="Fletcher" box="[814,977,1554,1581]" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017)</bibRefCitation>
reillustrated the same articulate and distorted specimen figured previously by
<bibRefCitation author="Lake" box="[1162,1286,1586,1613]" firstAuthor="Lake" pageId="9" pageNumber="1058" pagination="149 - 172" refId="ref27339" refString="LAKE P. 1932. - Monograph of the British Cambrian Trilobites, Part 7. Monograph of the Palaeontographical Society, London 84: 149 - 172. https: // doi. org / 10.1080 / 02693445.1932.12035620" type="journal article" year="1932">Lake (1932</bibRefCitation>
: pl. 20, fig. 3).
<bibRefCitation author="Fletcher" box="[814,977,1618,1644]" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017)</bibRefCitation>
edited the images using Photoshop (
<bibRefCitation author="Fletcher" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017</bibRefCitation>
: figs 3B, C, 8G). The figures 3C and 8G show the same image. The single specimen illustrated in figs 3A-C and 8F, G as
<taxonomicName authorityName="Fletcher" authorityYear="2017" box="[837,932,1714,1740]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis box="[837,932,1714,1740]" italics="true" pageId="9" pageNumber="1058">Ag. lewisi</emphasis>
</taxonomicName>
clearly shows the characteristics of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[1291,1446,1714,1740]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[1291,1446,1714,1740]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. The diagnosis of
<taxonomicName authorityName="Fletcher" authorityYear="2017" box="[987,1083,1746,1772]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis box="[987,1083,1746,1772]" italics="true" pageId="9" pageNumber="1058">Ag. lewisi</emphasis>
</taxonomicName>
presented by
<bibRefCitation author="Fletcher" box="[1227,1391,1746,1772]" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017)</bibRefCitation>
is not unique to the species, and characteristics as a “librigenal spine extending back to the posterior end of the third thoracic pleura” and “prominent stout median spines on the succeeding five rings” (
<bibRefCitation author="Fletcher" box="[891,1039,1873,1900]" firstAuthor="Fletcher" pageId="9" pageNumber="1058" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017</bibRefCitation>
) cannot be applied to the illustrations. Another characteristic, i.e. the “sharply pointed occipital spine”, is undiagnostic as discussed above. Consequently,
<taxonomicName authorityName="Fletcher" authorityYear="2017" box="[1337,1432,1937,1964]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis box="[1337,1432,1937,1964]" italics="true" pageId="9" pageNumber="1058">Ag. lewisi</emphasis>
</taxonomicName>
is here interpreted to be a synonym of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[1192,1349,1969,1995]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[1192,1349,1969,1995]" italics="true" pageId="9" pageNumber="1058">Ag. ceticephalus</emphasis>
</taxonomicName>
. Cranidia attributed byFletcher (2017) to
<taxonomicName baseAuthorityName="Hicks" baseAuthorityYear="1872" box="[1136,1303,2001,2027]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="1058" phylum="Arthropoda" rank="species" species="longicephalus">
<emphasis box="[1136,1303,2001,2027]" italics="true" pageId="9" pageNumber="1058">Ag.longicephalus</emphasis>
</taxonomicName>
and illustrated in figs 22A, B show eye ridges characteristic of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[617,773,215,242]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[617,773,215,242]" italics="true" pageId="11" pageNumber="1060">Ag. ceticephalus</emphasis>
</taxonomicName>
and are here assigned to this species.
<bibRefCitation author="Fletcher's" box="[502,667,247,273]" firstAuthor="Fletcher's" pageId="11" pageNumber="1060" pagination="35 - 56" refId="ref25012" refString="FLETCHER T. P. 2007. - Correlating the zones of ' Paradoxides hicksii ' and ' Paradoxides davidis ' in Cambrian Series 3. Memoirs of the Association of Australasian Palaeontologists 33: 35 - 56. https: // doi. org / 10.1080 / 03115510701586855" type="journal article" year="2007">Fletchers (2007</bibRefCitation>
: figs 5J-L, 12A, K-Q, 13I-K, M, N, 14D, G, H, J, 16A, C) display cranidia, or cranidia attached to the thorax, and were attributed by the author to
<taxonomicName authority="Billings, 1872" authorityName="Billings" authorityYear="1872" box="[309,626,343,369]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="species" species="socialis">
<emphasis box="[309,469,343,369]" italics="true" pageId="11" pageNumber="1060">Agraulos socialis</emphasis>
<bibRefCitation author="Billings" box="[476,626,343,369]" firstAuthor="Billings" pageId="11" pageNumber="1060" pagination="465 - 479" refId="ref23812" refString="BILLINGS E. 1872. - On some fossils from the primordial rocks of Newfoundland. Canadian Naturalist and Quarterly Journal of Science 6: 465 - 479. https: // doi. org / 10.5962 / bhl. title. 38279" type="journal article" year="1872">Billings, 1872</bibRefCitation>
</taxonomicName>
. Nevertheless, these illustrated specimens morphologically agree with
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis italics="true" pageId="11" pageNumber="1060">Ag. ceticephalus</emphasis>
</taxonomicName>
and are here assigned to this species.A new species,
<emphasis box="[130,241,438,465]" italics="true" pageId="11" pageNumber="1060">
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[130,217,438,464]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="genus">Agraulos</taxonomicName>
1
</emphasis>
<bibRefCitation author="Fletcher" box="[250,407,438,465]" firstAuthor="Fletcher" pageId="11" pageNumber="1060" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher, 2017</bibRefCitation>
, was introduced without a formal diagnosis or description.
<bibRefCitation author="Fletcher" box="[400,565,470,496]" firstAuthor="Fletcher" pageId="11" pageNumber="1060" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017)</bibRefCitation>
only states that the “new species status is marked by the less trapezoidal aspect of the cranium with a longish occipital spine significantly different from associated similar-sized ceticephalus cranidia”. These differences are here interpreted as intraspecific variations of
<emphasis box="[134,299,630,656]" italics="true" pageId="11" pageNumber="1060">
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[134,295,630,656]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="species" species="ceticephalus">Ag. ceticephalus</taxonomicName>
.
</emphasis>
Therefore, the cranidia illustrated by
<bibRefCitation author="Fletcher" firstAuthor="Fletcher" pageId="11" pageNumber="1060" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017</bibRefCitation>
: figs 13D, E, 14A, 16F) match the characteristics of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis italics="true" pageId="11" pageNumber="1060">Ag. ceticephalus</emphasis>
</taxonomicName>
and
<emphasis box="[306,412,693,720]" italics="true" pageId="11" pageNumber="1060">
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[306,392,693,719]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="genus">Agraulos</taxonomicName>
1
</emphasis>
is here interpreted as synonym of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[133,288,725,752]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[133,288,725,752]" italics="true" pageId="11" pageNumber="1060">Ag. ceticephalus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Fletcher" box="[300,462,725,752]" firstAuthor="Fletcher" pageId="11" pageNumber="1060" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017)</bibRefCitation>
also described the new species
<emphasis box="[130,235,757,783]" italics="true" pageId="11" pageNumber="1060">
<taxonomicName authorityName="Hawle &amp; Corda" authorityYear="1847" box="[130,216,757,783]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="genus">Agraulos</taxonomicName>
2
</emphasis>
(
<bibRefCitation author="Fletcher" box="[251,396,757,783]" firstAuthor="Fletcher" pageId="11" pageNumber="1060" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher 2017</bibRefCitation>
: figs 14I, 15A-C, 16E) which is here also interpreted as an intraspecific variation of
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[611,766,789,816]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[611,766,789,816]" italics="true" pageId="11" pageNumber="1060">Ag. ceticephalus</emphasis>
</taxonomicName>
, while cranidia illustrated in figures 15D, E are too poorly preserved to be assigned to any species.
<bibRefCitation author="Fletcher" box="[506,670,853,879]" firstAuthor="Fletcher" pageId="11" pageNumber="1060" pagination="175 - 217" refId="ref25061" refString="FLETCHER T. P. 2017. - Agraulos ceticephalus and other Cambrian Trilobites in the subfamily Agraulinae from Bohemia, Newfoundland and Wales. Papers in Palaeontology 3 (2): 175 - 217. https: // doi. org / 10.1002 / spp 2.1071" type="journal article" year="2017">Fletcher (2017)</bibRefCitation>
described
<taxonomicName authorityName="Fletcher" authorityYear="2017" box="[131,378,885,911]" class="Trilobita" family="Agraulidae" genus="Skreiaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis box="[131,378,885,911]" italics="true" pageId="11" pageNumber="1060">Skreiaspis punctatissimus</emphasis>
</taxonomicName>
as a new species and illustrated several specimens.Cranidia illustrated in figures 18S, U and W do not show the short preglabellar field characteristic for
<taxonomicName authorityName="Ruzicka" authorityYear="1946" box="[631,726,949,975]" class="Trilobita" family="Agraulidae" genus="Skreiaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="genus">
<emphasis box="[631,726,949,975]" italics="true" pageId="11" pageNumber="1060">Skreiaspis</emphasis>
</taxonomicName>
, but rather match
<taxonomicName baseAuthorityName="Barrande" baseAuthorityYear="1846" box="[274,431,980,1007]" class="Trilobita" family="Agraulidae" genus="Agraulos" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="1060" phylum="Arthropoda" rank="species" species="ceticephalus">
<emphasis box="[274,431,980,1007]" italics="true" pageId="11" pageNumber="1060">Ag. ceticephalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>