treatments-xml/data/03/DA/87/03DA879D0379FFC2A6F9F935B2EEFE35.xml

<document id="DA16A00E35E8F304EC9DBDE67C123EC7" ID-CLB-Dataset="31992" ID-DOI="10.11646/zootaxa.4324.1.12" ID-GBIF-Dataset="9a9d34ed-1eea-499b-b688-432e911e9ecc" ID-ISSN="1175-5326" ID-Zenodo-Dep="996997" ID-ZooBank="Fbe20203-F19F-42E6-Bdb5-22A1D04Fa427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506405916695" checkinUser="plazi" docAuthor="Phan, Quoc Toan" docDate="2017" docId="03DA879D0379FFC2A6F9F935B2EEFE35" docLanguage="en" docName="zootaxa.4324.1.12.pdf" docOrigin="Zootaxa 4324 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeliccia duytan Phan 2017" docType="treatment" docVersion="7" lastPageNumber="200" masterDocId="FFE3FFE50379FFC7A66EFFE3B358FF94" masterDocTitle="Coeliccia duytan sp. nov. from the Central Highlands of Vietnam (Odonata: Platycnemididae)" masterLastPageNumber="200" masterPageNumber="195" pageNumber="195" updateTime="1698468973748" updateUser="ExternalLinkService">
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<mods:title id="00CD7B5220BA2020C79B8F9911E52BDC">Coeliccia duytan sp. nov. from the Central Highlands of Vietnam (Odonata: Platycnemididae)</mods:title>
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<mods:namePart id="A4793E01F8AA5E83317CFC93BD303916">Phan, Quoc Toan</mods:namePart>
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<mods:date id="CDF0710CC1A65A89C9BD624F9F2E2F4B">2017</mods:date>
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<treatment id="03DA879D0379FFC2A6F9F935B2EEFE35" ID-DOI="http://doi.org/10.5281/zenodo.6015841" ID-GBIF-Taxon="134840093" ID-Zenodo-Dep="6015841" LSID="urn:lsid:plazi:treatment:03DA879D0379FFC2A6F9F935B2EEFE35" httpUri="http://treatment.plazi.org/id/03DA879D0379FFC2A6F9F935B2EEFE35" lastPageId="5" lastPageNumber="200" pageId="0" pageNumber="195">
<subSubSection id="C36965000379FFC7A6F9F935B1E4F964" box="[151,700,1750,1776]" pageId="0" pageNumber="195" type="nomenclature">
<paragraph id="8BCC368B0379FFC7A6F9F935B1E4F964" blockId="0.[151,700,1750,1776]" box="[151,700,1750,1776]" pageId="0" pageNumber="195">
<heading id="D08481E70379FFC7A6F9F935B1E4F964" bold="true" box="[151,700,1750,1776]" fontSize="11" level="1" pageId="0" pageNumber="195" reason="1">
<emphasis id="B907EA990379FFC7A6F9F935B1E4F964" bold="true" box="[151,700,1750,1776]" pageId="0" pageNumber="195">
<emphasis id="B907EA990379FFC7A6F9F935B22BF964" bold="true" box="[151,371,1750,1776]" italics="true" pageId="0" pageNumber="195">
<taxonomicName id="4C734D080379FFC7A6F9F935B239F964" ID-CoL="9WYQ9" authorityName="Phan" authorityYear="2017" box="[151,353,1750,1776]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="195" phylum="Arthropoda" rank="species" species="duytan">Coeliccia duytan</taxonomicName>
s
</emphasis>
p. nov. (
<figureCitation id="13482A0E0379FFC7A7BBF935B168F964" box="[469,560,1750,1776]" captionStart="FIGURE 1" captionStartId="1.[151,244,1952,1974]" captionTargetBox="[188,1397,295,1930]" captionTargetId="figure@1.[188,1398,295,1930]" captionTargetPageId="1" captionText="FIGURE 1. Coeliccia spp., head and thorax, ♂. (1 a) Coeliccia duytan sp. nov.; (1 b) Coeliccia hayashii." httpUri="https://zenodo.org/record/996999/files/figure.png" pageId="0" pageNumber="195">Figs 1a</figureCitation>
, 
<figureCitation id="13482A0E0379FFC7A450F935B121F964" box="[574,633,1750,1776]" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="0" pageNumber="195">2a–c</figureCitation>
, 
<figureCitation id="13482A0E0379FFC7A4E8F935B1EAF964" box="[646,690,1750,1776]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="3.[151,244,1850,1872]" captionStartId-1="4.[151,244,1949,1971]" captionStartId-2="5.[151,244,1692,1714]" captionTargetBox-0="[188,1398,642,1829]" captionTargetBox-1="[153,1433,677,1928]" captionTargetBox-2="[153,1433,818,1671]" captionTargetId-0="figure@3.[188,1398,642,1829]" captionTargetId-1="figure@4.[153,1433,677,1929]" captionTargetId-2="figure@5.[153,1433,818,1671]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 3. Coeliccia duytan sp. nov. (3 a) head male; (3 b) head female; (3 c) prothorax female in lateral view; (3 d) prothorax female in dorsal view; (3 e) tip of abdomen, female." captionText-1="FIGURE 4. Coeliccia duytan sp. nov. in nature (4 a) male; (4 b) female." captionText-2="FIGURE 5. Coeliccia duytan sp. nov. in copulation." httpUri-0="https://zenodo.org/record/997003/files/figure.png" httpUri-1="https://zenodo.org/record/997005/files/figure.png" httpUri-2="https://zenodo.org/record/997007/files/figure.png" pageId="0" pageNumber="195">3–5</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36965000379FFC7A6F9F8FFB268F80A" pageId="0" pageNumber="195" type="materials_examined">
<paragraph id="8BCC368B0379FFC7A6F9F8FFB26EF8C1" blockId="0.[151,1436,1820,2027]" pageId="0" pageNumber="195">
<materialsCitation id="3B1B3CD60379FFC7A6F9F8FFB26EF8C1" ID-GBIF-Occurrence="1656006054" collectingDate="2017-05-19" collectorName="Quoc Toan Phan" country="Vietnam" location="Chu Mom Ray National Park" pageId="0" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Kon Tum Province" typeStatus="holotype">
<emphasis id="B907EA990379FFC7A6F9F8FFB25DF8A7" bold="true" box="[151,261,1820,1843]" pageId="0" pageNumber="195">
<typeStatus id="54C888290379FFC7A6F9F8FFB259F8A7" box="[151,257,1820,1843]" pageId="0" pageNumber="195" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D75FD020379FFC7A760F8FFB27CF8A7" box="[270,292,1820,1843]" pageId="0" pageNumber="195" type="male">♂</specimenCount>
, 
<location id="8EAC60500379FFC7A75CF8FFB12FF8A7" LSID="urn:lsid:plazi:treatment:03DA879D0379FFC2A6F9F935B2EEFE35:8EAC60500379FFC7A75CF8FFB12FF8A7" box="[306,631,1820,1843]" country="Vietnam" name="Chu Mom Ray National Park" pageId="0" pageNumber="195" stateProvince="Kon Tum Province">Chu Mom Ray National Park</location>
, 
<collectingRegion id="49B7F8690379FFC7A4EAF8FFB00FF8A7" box="[644,855,1820,1843]" country="Vietnam" name="Kon Tum" pageId="0" pageNumber="195">
Kon 
<collectingRegion id="49B7F8690379FFC7A4D5F8FFB00FF8A7" box="[699,855,1820,1843]" inferredBySuffix="true" pageId="0" pageNumber="195">Tum Province</collectingRegion>
</collectingRegion>
, the Central Highlands of 
<collectingCountry id="F364761B0379FFC7A215F8FFB780F8A7" box="[1147,1240,1820,1843]" name="Vietnam" pageId="0" pageNumber="195">Vietnam</collectingCountry>
, 
<collectingDate id="EF89E9A30379FFC7A28BF8FFB60CF8A7" box="[1253,1364,1820,1843]" pageId="0" pageNumber="195" value="2017-05-19">
<date id="FFCD104B0379FFC7A28BF8FFB60CF8A7" box="[1253,1364,1820,1843]" pageId="0" pageNumber="195" value="2017-05-19">
19.V. 
<docYear id="9315DA700379FFC7A374F8FFB60CF8A7" box="[1306,1364,1820,1843]" pageId="0" pageNumber="195">2017</docYear>
</date>
</collectingDate>
, 
<collectorName id="2686535D0379FFC7A30CF8FFB25FF8C1" pageId="0" pageNumber="195">Quoc Toan Phan</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BCC368B0379FFC7A6A9F887B268F80A" blockId="0.[151,1436,1820,2027]" pageId="0" pageNumber="195">
<emphasis id="B907EA990379FFC7A6A9F887B261F8EF" bold="true" box="[199,313,1892,1915]" pageId="0" pageNumber="195">Paratypes</emphasis>
. 8♂♂, 3♀♀, same date, location and collector as holotype; 
<materialsCitation id="3B1B3CD60379FFC7A5D4F887B268F80A" ID-GBIF-Occurrence="1656006053" collectingDate="2016-05-10" collectorName="Nguyen Dang Van" pageId="0" pageNumber="195" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D75FD020379FFC7A5D4F887B0A9F8E8" box="[954,1009,1892,1916]" pageId="0" pageNumber="195" type="male">2♂♂</specimenCount>
, 
<specimenCount id="9D75FD020379FFC7A593F887B746F8EF" box="[1021,1054,1892,1916]" pageId="0" pageNumber="195" type="female">1♀</specimenCount>
, same location, 
<collectingDate id="EF89E9A30379FFC7A2A3F887B664F8EF" box="[1229,1340,1892,1915]" pageId="0" pageNumber="195" value="2016-05-10">
<date id="FFCD104B0379FFC7A2A3F887B664F8EF" box="[1229,1340,1892,1915]" pageId="0" pageNumber="195" value="2016-05-10">10.V.2016</date>
</collectingDate>
, 
<collectorName id="2686535D0379FFC7A329F887B259F80A" pageId="0" pageNumber="195">Nguyen Dang Van</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36965000379FFC7A6A9F84EB602F850" box="[199,1370,1965,1988]" pageId="0" pageNumber="195" type="etymology">
<paragraph id="8BCC368B0379FFC7A6A9F84EB602F850" blockId="0.[151,1436,1820,2027]" box="[199,1370,1965,1988]" pageId="0" pageNumber="195">
<emphasis id="B907EA990379FFC7A6A9F84EB21FF850" bold="true" box="[199,327,1965,1988]" pageId="0" pageNumber="195">Etymology.</emphasis>
This specific epithet is derived from my office Duy Tan University; 
<taxonomicName id="4C734D080379FFC7A24AF84EB737F850" authorityName="Phan" authorityYear="2017" box="[1060,1135,1965,1988]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="195" phylum="Arthropoda" rank="species" species="duytan">duytan</taxonomicName>
, a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C36965000379FFC3A6A8F837B169FF45" lastPageId="4" lastPageNumber="199" pageId="0" pageNumber="195" type="description">
<paragraph id="8BCC368B0379FFC6A6A8F837B61FFF60" blockId="0.[151,1436,1820,2027]" lastBlockId="1.[151,1435,152,244]" lastPageId="1" lastPageNumber="196" pageId="0" pageNumber="195">
<emphasis id="B907EA990379FFC7A6A8F837B28AF87F" bold="true" box="[198,466,2004,2027]" pageId="0" pageNumber="195">
Description of 
<typeStatus id="54C888290379FFC7A700F837B296F87F" box="[366,462,2004,2027]" pageId="0" pageNumber="195" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="B907EA990379FFC7A7B2F830B14CF87F" box="[476,532,2003,2027]" italics="true" pageId="0" pageNumber="195">Head</emphasis>
(
<figureCitation id="13482A0E0379FFC7A44BF837B12EF87F" box="[549,630,2004,2027]" captionStart="FIGURE 1" captionStartId="1.[151,244,1952,1974]" captionTargetBox="[188,1397,295,1930]" captionTargetId="figure@1.[188,1398,295,1930]" captionTargetPageId="1" captionText="FIGURE 1. Coeliccia spp., head and thorax, ♂. (1 a) Coeliccia duytan sp. nov.; (1 b) Coeliccia hayashii." httpUri="https://zenodo.org/record/996999/files/figure.png" pageId="0" pageNumber="195">Figs 1a</figureCitation>
, 
<figureCitation id="13482A0E0379FFC7A4EBF837B1F8F87F" box="[645,672,2004,2027]" captionStart="FIGURE 3" captionStartId="3.[151,244,1850,1872]" captionTargetBox="[188,1398,642,1829]" captionTargetId="figure@3.[188,1398,642,1829]" captionTargetPageId="3" captionText="FIGURE 3. Coeliccia duytan sp. nov. (3 a) head male; (3 b) head female; (3 c) prothorax female in lateral view; (3 d) prothorax female in dorsal view; (3 e) tip of abdomen, female." httpUri="https://zenodo.org/record/997003/files/figure.png" pageId="0" pageNumber="195">3a</figureCitation>
). Anteclypeus pale blue except for two isolated black spots adjacent to labrum; labrum and mandibles matte black. Genae pale blue, extending to the level of scape (
<figureCitation id="13482A0E0378FFC6A2E2FF7BB783FF3B" box="[1164,1243,152,175]" captionStart="FIGURE 3" captionStartId="3.[151,244,1850,1872]" captionTargetBox="[188,1398,642,1829]" captionTargetId="figure@3.[188,1398,642,1829]" captionTargetPageId="3" captionText="FIGURE 3. Coeliccia duytan sp. nov. (3 a) head male; (3 b) head female; (3 c) prothorax female in lateral view; (3 d) prothorax female in dorsal view; (3 e) tip of abdomen, female." httpUri="https://zenodo.org/record/997003/files/figure.png" pageId="1" pageNumber="196">Fig. 3a</figureCitation>
). Antennae black, pedicel yellow. Top of head matt black with two spots posteriolateral of median ocellus and two streaks at the anterior corners of the lateral ocelli and a transverse wedged shaped pale yellow spot on just posterior to postoccipital lobes (
<figureCitation id="13482A0E0378FFC6A29EFF3EB661FF60" box="[1264,1337,221,244]" captionStart="FIGURE 1" captionStartId="1.[151,244,1952,1974]" captionTargetBox="[188,1397,295,1930]" captionTargetId="figure@1.[188,1398,295,1930]" captionTargetPageId="1" captionText="FIGURE 1. Coeliccia spp., head and thorax, ♂. (1 a) Coeliccia duytan sp. nov.; (1 b) Coeliccia hayashii." httpUri="https://zenodo.org/record/996999/files/figure.png" pageId="1" pageNumber="196">Fig. 1a</figureCitation>
).
</paragraph>
<caption id="DF0C66030378FFC6A6F9F843B7D8F821" box="[151,1152,1952,1974]" httpUri="https://zenodo.org/record/996999/files/figure.png" pageId="1" pageNumber="196" targetBox="[188,1397,295,1930]" targetPageId="1">
<paragraph id="8BCC368B0378FFC6A6F9F843B7D8F821" blockId="1.[151,1152,1952,1974]" box="[151,1152,1952,1974]" pageId="1" pageNumber="196">
<emphasis id="B907EA990378FFC6A6F9F843B255F822" bold="true" box="[151,269,1952,1974]" pageId="1" pageNumber="196">FIGURE 1.</emphasis>
<taxonomicName id="4C734D080378FFC6A77CF843B236F821" authorityName="Kirby" authorityYear="1890" box="[274,366,1952,1973]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B907EA990378FFC6A77CF843B236F821" box="[274,366,1952,1973]" italics="true" pageId="1" pageNumber="196">Coeliccia</emphasis>
</taxonomicName>
spp., head and thorax, ♂. (1a) 
<taxonomicName id="4C734D080378FFC6A4F9F843B062F821" authority="Phan, 2017" authorityName="Phan" authorityYear="2017" box="[663,826,1952,1973]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="duytan" status="sp. nov.">
<emphasis id="B907EA990378FFC6A4F9F843B062F821" box="[663,826,1952,1973]" italics="true" pageId="1" pageNumber="196">Coeliccia duytan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457E20378FFC6A52EF843B0D4F822" box="[832,908,1952,1974]" pageId="1" pageNumber="196" rank="species">
<emphasis id="B907EA990378FFC6A52EF843B0D4F822" bold="true" box="[832,908,1952,1974]" pageId="1" pageNumber="196">sp. nov.</emphasis>
</taxonomicNameLabel>
; (1b) 
<taxonomicName id="4C734D080378FFC6A5A9F843B722F821" authorityName="Phan &amp; Kompier" authorityYear="2016" box="[967,1146,1952,1973]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="B907EA990378FFC6A5A9F843B722F821" box="[967,1146,1952,1973]" italics="true" pageId="1" pageNumber="196">Coeliccia hayashii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BCC368B037BFFC5A6A9FF78B28BFE16" blockId="2.[151,1436,155,802]" pageId="2" pageNumber="197">
Thorax (
<figureCitation id="13482A0E037BFFC5A74AFF78B22AFF26" box="[292,370,155,178]" captionStart="FIGURE 1" captionStartId="1.[151,244,1952,1974]" captionTargetBox="[188,1397,295,1930]" captionTargetId="figure@1.[188,1398,295,1930]" captionTargetPageId="1" captionText="FIGURE 1. Coeliccia spp., head and thorax, ♂. (1 a) Coeliccia duytan sp. nov.; (1 b) Coeliccia hayashii." httpUri="https://zenodo.org/record/996999/files/figure.png" pageId="2" pageNumber="197">Fig. 1a</figureCitation>
). Anterior and posterior lobes of prothorax black, middle lobes pruinose blue. Most of propleuron matt black, lower part yellow. Synthorax largely black. Mesepisternum black with a pruinose spot occupying ventral fourth. Mesepimeron, metepisternum and mesinfraepisternum black except for a small pale yellow on mesinfraepisternum adjacent to mesocoxa and ventral fourth of metepisternum above coxa and encompassing metastigma. Metinfraepisternum yellow excluding a black spot adjacent to metepisternum. Large rectangular pruinose area extending from posterior half of mesepimeron to anterior third of metepisternum including metapleural suture. Metepimeron entirely yellow.
</paragraph>
<paragraph id="8BCC368B037BFFC5A6A9FE6DB1F2FE53" blockId="2.[151,1436,155,802]" pageId="2" pageNumber="197">
Legs (
<figureCitation id="13482A0E037BFFC5A760FE6DB207FE31" box="[270,351,398,421]" captionStart="FIGURE 1" captionStartId="1.[151,244,1952,1974]" captionTargetBox="[188,1397,295,1930]" captionTargetId="figure@1.[188,1398,295,1930]" captionTargetPageId="1" captionText="FIGURE 1. Coeliccia spp., head and thorax, ♂. (1 a) Coeliccia duytan sp. nov.; (1 b) Coeliccia hayashii." httpUri="https://zenodo.org/record/996999/files/figure.png" pageId="2" pageNumber="197">Fig. 1a</figureCitation>
). Coxae pale yellow. Femora and tibiae pale yellow with flexor surface of femora and extensor surfaces of tibiae black. Tarsi and armature black.
</paragraph>
<paragraph id="8BCC368B037BFFC5A6A9FE30B074FD99" blockId="2.[151,1436,155,802]" pageId="2" pageNumber="197">Wings hyaline, 17–18 and 17 postnodal crossveins in forewings and hindwings, respectively. Pterostigma brown with narrow pale margin, surmounting 1.5–2 underlying cells.</paragraph>
<paragraph id="8BCC368B037BFFC5A6A9FDFBB383FDE1" blockId="2.[151,1436,155,802]" pageId="2" pageNumber="197">
Abdomen. Black above with pale yellow markings as follows: S1 yellow, black dorsally; S2–8 black above, shading into pale yellow ventrally, black at apices of each segment; S9 black with posterior 1/6 yellow; S10 entirely yellow (
<figureCitation id="13482A0E037BFFC5A31EFDD8B396FDE1" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="2" pageNumber="197">Fig. 2a–b</figureCitation>
).
</paragraph>
<paragraph id="8BCC368B037BFFC5A6A9FD63B785FD03" blockId="2.[151,1436,155,802]" box="[199,1245,640,663]" pageId="2" pageNumber="197">
Genital ligula shown in 
<figureCitation id="13482A0E037BFFC5A7A9FD63B149FD03" box="[455,529,640,663]" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="2" pageNumber="197">Fig. 2c</figureCitation>
with terminal segment armed medially with a pair of long flagella.
</paragraph>
<paragraph id="8BCC368B037BFFC5A6A9FD40B022FD6B" blockId="2.[151,1436,155,802]" pageId="2" pageNumber="197">
Anal appendages (
<figureCitation id="13482A0E037BFFC5A7E2FD40B2ABFD2E" box="[396,499,675,698]" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="2" pageNumber="197">Fig. 2a–b</figureCitation>
) yellow, cerci short and robust, ca. length of S10, in lateral view bearing a small ventral tooth along swollen medial portion; in dorsal view, cercus narrowing distally and rounded at apex. Linear paraproct ca. 1/ 3 longer than cercus, its medially directed tip ending in a black tooth.
</paragraph>
<paragraph id="8BCC368B037BFFC5A6A9FCE8B09EFCB6" blockId="2.[151,1436,155,802]" box="[199,966,779,802]" pageId="2" pageNumber="197">
<emphasis id="B907EA99037BFFC5A6A9FCE8B236FCB6" bold="true" box="[199,366,779,802]" pageId="2" pageNumber="197">Measurements</emphasis>
(in mm). Hindwing 26; abdomen (incl. appendages) 41.
</paragraph>
<caption id="DF0C6603037BFFC5A6F9F914B64AF8BE" httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="2" pageNumber="197" targetBox="[189,1396,854,1749]" targetPageId="2">
<paragraph id="8BCC368B037BFFC5A6F9F914B64AF8BE" blockId="2.[151,1436,1783,1834]" pageId="2" pageNumber="197">
<emphasis id="B907EA99037BFFC5A6F9F914B255F899" bold="true" box="[151,269,1783,1805]" pageId="2" pageNumber="197">FIGURE 2.</emphasis>
<taxonomicName id="4C734D08037BFFC5A77AF91BB228F899" authorityName="Kirby" authorityYear="1890" box="[276,368,1784,1805]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B907EA99037BFFC5A77AF91BB228F899" box="[276,368,1784,1805]" italics="true" pageId="2" pageNumber="197">Coeliccia</emphasis>
</taxonomicName>
spp., ♂. [2a–c] 
<taxonomicName id="4C734D08037BFFC5A463F91BB1E9F899" authority="Phan, 2017" authorityName="Phan" authorityYear="2017" box="[525,689,1784,1805]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="duytan" status="sp. nov.">
<emphasis id="B907EA99037BFFC5A463F91BB1E9F899" box="[525,689,1784,1805]" italics="true" pageId="2" pageNumber="197">Coeliccia duytan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457E2037BFFC5A4D6F914B05DF899" box="[696,773,1783,1805]" pageId="2" pageNumber="197" rank="species">
<emphasis id="B907EA99037BFFC5A4D6F914B05DF899" bold="true" box="[696,773,1783,1805]" pageId="2" pageNumber="197">sp. nov.</emphasis>
</taxonomicNameLabel>
; [2d–f] 
<emphasis id="B907EA99037BFFC5A53BF91BB748F899" box="[853,1040,1784,1805]" italics="true" pageId="2" pageNumber="197">
<taxonomicName id="4C734D08037BFFC5A53BF91BB754F899" authorityName="Phan &amp; Kompier" authorityYear="2016" box="[853,1036,1784,1805]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="hayashii">Coeliccia hayashii</taxonomicName>
.
</emphasis>
(2a, 2d) appendages in dorsal view; (2b, 2e) appendages in lateral view; (2c, 2f) genital ligula in dorsal view (2f from Fig. 4C, p. 411 in Phan &amp; Kompier 2016).
</paragraph>
</caption>
<paragraph id="8BCC368B037BFFC5A6A9F8B6B29AF81B" blockId="2.[151,1435,1877,2004]" pageId="2" pageNumber="197">
<emphasis id="B907EA99037BFFC5A6A9F8B6B29AF8F8" bold="true" box="[199,450,1877,1900]" pageId="2" pageNumber="197">Description of female.</emphasis>
Head (
<figureCitation id="13482A0E037BFFC5A47FF8B6B139F8F8" box="[529,609,1877,1900]" captionStart="FIGURE 3" captionStartId="3.[151,244,1850,1872]" captionTargetBox="[188,1398,642,1829]" captionTargetId="figure@3.[188,1398,642,1829]" captionTargetPageId="3" captionText="FIGURE 3. Coeliccia duytan sp. nov. (3 a) head male; (3 b) head female; (3 c) prothorax female in lateral view; (3 d) prothorax female in dorsal view; (3 e) tip of abdomen, female." httpUri="https://zenodo.org/record/997003/files/figure.png" pageId="2" pageNumber="197">Fig. 3b</figureCitation>
). Head color pattern similar to that of male except yellow bands from lateral ocelli extending to the eyes.
</paragraph>
<paragraph id="8BCC368B037BFFC4A6A9F878B288FEAD" blockId="2.[151,1435,1877,2004]" lastBlockId="3.[151,1436,152,591]" lastPageId="3" lastPageNumber="198" pageId="2" pageNumber="197">
Thorax (
<figureCitation id="13482A0E037BFFC5A74CF878B2D6F826" box="[290,398,1947,1970]" captionStart="FIGURE 3" captionStartId="3.[151,244,1850,1872]" captionTargetBox="[188,1398,642,1829]" captionTargetId="figure@3.[188,1398,642,1829]" captionTargetPageId="3" captionText="FIGURE 3. Coeliccia duytan sp. nov. (3 a) head male; (3 b) head female; (3 c) prothorax female in lateral view; (3 d) prothorax female in dorsal view; (3 e) tip of abdomen, female." httpUri="https://zenodo.org/record/997003/files/figure.png" pageId="2" pageNumber="197">Figs 3c–d</figureCitation>
, 
<figureCitation id="13482A0E037BFFC5A7F4F878B2E0F826" box="[410,440,1947,1970]" captionStart="FIGURE 4" captionStartId="4.[151,244,1949,1971]" captionTargetBox="[153,1433,677,1928]" captionTargetId="figure@4.[153,1433,677,1929]" captionTargetPageId="4" captionText="FIGURE 4. Coeliccia duytan sp. nov. in nature (4 a) male; (4 b) female." httpUri="https://zenodo.org/record/997005/files/figure.png" pageId="2" pageNumber="197">4b</figureCitation>
). Prothorax black except except for two large, mediolateral separated yellow spots on middle lobe; propleuron yellow. Posterior lobe armed medially with a sconcave subrectangular lobe, its base constricted, dorsolaterally to medial lobe strongly concave in a semicircular fashion before terminating as a medially pointed lateral lobe (
<figureCitation id="13482A0E037AFFC4A6BAFF59B27BFF45" box="[212,291,186,209]" captionStart="FIGURE 3" captionStartId="3.[151,244,1850,1872]" captionTargetBox="[188,1398,642,1829]" captionTargetId="figure@3.[188,1398,642,1829]" captionTargetPageId="3" captionText="FIGURE 3. Coeliccia duytan sp. nov. (3 a) head male; (3 b) head female; (3 c) prothorax female in lateral view; (3 d) prothorax female in dorsal view; (3 e) tip of abdomen, female." httpUri="https://zenodo.org/record/997003/files/figure.png" pageId="3" pageNumber="198">Fig. 3c</figureCitation>
); a recessed obtuse tooth present below semicircular concavity as shown in 
<figureCitation id="13482A0E037AFFC4A234FF59B7F1FF45" box="[1114,1193,186,209]" captionStart="FIGURE 3" captionStartId="3.[151,244,1850,1872]" captionTargetBox="[188,1398,642,1829]" captionTargetId="figure@3.[188,1398,642,1829]" captionTargetPageId="3" captionText="FIGURE 3. Coeliccia duytan sp. nov. (3 a) head male; (3 b) head female; (3 c) prothorax female in lateral view; (3 d) prothorax female in dorsal view; (3 e) tip of abdomen, female." httpUri="https://zenodo.org/record/997003/files/figure.png" pageId="3" pageNumber="198">Fig. 3d</figureCitation>
. Synthorax black with yellow areas as follows: thin antehumeral stripe; mesepimeron with small isolated yellow spot above antehumeral stripe (
<figureCitation id="13482A0E037AFFC4A6F1FEE3B3A2FE83" box="[159,250,256,279]" captionStart="FIGURE 4" captionStartId="4.[151,244,1949,1971]" captionTargetBox="[153,1433,677,1928]" captionTargetId="figure@4.[153,1433,677,1929]" captionTargetPageId="4" captionText="FIGURE 4. Coeliccia duytan sp. nov. in nature (4 a) male; (4 b) female." httpUri="https://zenodo.org/record/997005/files/figure.png" pageId="3" pageNumber="198">Fig. 4b</figureCitation>
); ventral margin of mesinfraepisternum; metepisternum except for distinct black metapleural stripe; metepimeron entirely yellow.
</paragraph>
<paragraph id="8BCC368B037AFFC4A6A9FEA6B2FEFEEB" blockId="3.[151,1436,152,591]" pageId="3" pageNumber="198">Legs. Coxae and femora pale yellow, flexor and extensor surfaces black. Tibiae pale yellow, flexor surface black. Tarsi and armature black.</paragraph>
<paragraph id="8BCC368B037AFFC4A6A9FE69B1A5FE50" blockId="3.[151,1436,152,591]" pageId="3" pageNumber="198">Wings. Hyaline, 18 postnodal crossveins in forewings and 17–18 postnodal crossveins in hindwings. Pterostigma brown with narrow pale margin, surmounting 1.5–2 cells.</paragraph>
<paragraph id="8BCC368B037AFFC4A6A9FE33B72FFDB8" blockId="3.[151,1436,152,591]" pageId="3" pageNumber="198">
Abdomen. S1 largely black dorsally; S2–7 black dorsally and laterally becoming pale ventrally; S8 almost entirely yellow but basal 0.70 black dorsolaterally; S9 dark yellow, black ventrally; S10 black with anterior margin dark yellow. Cerci black, ovipositor black with small yellow spot anteriorly and dorsally at apex (
<figureCitation id="13482A0E037AFFC4A272FDF6B732FDB8" box="[1052,1130,533,556]" captionStart="FIGURE 3" captionStartId="3.[151,244,1850,1872]" captionTargetBox="[188,1398,642,1829]" captionTargetId="figure@3.[188,1398,642,1829]" captionTargetPageId="3" captionText="FIGURE 3. Coeliccia duytan sp. nov. (3 a) head male; (3 b) head female; (3 c) prothorax female in lateral view; (3 d) prothorax female in dorsal view; (3 e) tip of abdomen, female." httpUri="https://zenodo.org/record/997003/files/figure.png" pageId="3" pageNumber="198">Fig. 3e</figureCitation>
).
</paragraph>
<paragraph id="8BCC368B037AFFC4A6A9FDDBB095FDDB" blockId="3.[151,1436,152,591]" box="[199,973,568,591]" pageId="3" pageNumber="198">
<emphasis id="B907EA99037AFFC4A6A9FDDBB236FDDB" bold="true" box="[199,366,568,591]" pageId="3" pageNumber="198">Measurements</emphasis>
(in mm). Hind wing 25; abdomen (incl. appendages) 35.
</paragraph>
<caption id="DF0C6603037AFFC4A6F9F8D9B165F8F9" httpUri="https://zenodo.org/record/997003/files/figure.png" pageId="3" pageNumber="198" targetBox="[188,1398,642,1829]" targetPageId="3">
<paragraph id="8BCC368B037AFFC4A6F9F8D9B165F8F9" blockId="3.[151,1435,1850,1901]" pageId="3" pageNumber="198">
<emphasis id="B907EA99037AFFC4A6F9F8D9B254F8C4" bold="true" box="[151,268,1850,1872]" pageId="3" pageNumber="198">FIGURE 3.</emphasis>
<taxonomicName id="4C734D08037AFFC4A77DF8D8B2EDF8C4" authority="Phan, 2017" authorityName="Phan" authorityYear="2017" box="[275,437,1851,1872]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="duytan" status="sp. nov.">
<emphasis id="B907EA99037AFFC4A77DF8D8B2EDF8C4" box="[275,437,1851,1872]" italics="true" pageId="3" pageNumber="198">Coeliccia duytan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457E2037AFFC4A7D2F8D9B151F8C4" box="[444,521,1850,1872]" pageId="3" pageNumber="198" rank="species">
<emphasis id="B907EA99037AFFC4A7D2F8D9B151F8C4" bold="true" box="[444,521,1850,1872]" pageId="3" pageNumber="198">sp. nov.</emphasis>
</taxonomicNameLabel>
(3a) head male; (3b) head female; (3c) prothorax female in lateral view; (3d) prothorax female in dorsal view; (3e) tip of abdomen, female.
</paragraph>
</caption>
<paragraph id="8BCC368B037AFFC4A6A9F87BB068F846" blockId="3.[151,1436,1944,2002]" pageId="3" pageNumber="198">
<emphasis id="B907EA99037AFFC4A6A9F87BB17BF83B" bold="true" box="[199,547,1944,1967]" pageId="3" pageNumber="198">
Variation in 
<typeStatus id="54C88829037AFFC4A708F87BB294F83B" box="[358,460,1944,1967]" pageId="3" pageNumber="198" type="paratype">paratype</typeStatus>
males.
</emphasis>
Some 
<typeStatus id="54C88829037AFFC4A413F87BB181F83B" box="[637,729,1944,1967]" pageId="3" pageNumber="198" type="paratype">paratype</typeStatus>
males have a yellow spot near the posterior margin of the metepisternum below the rectangular pruinosity on the thorax.
</paragraph>
<paragraph id="8BCC368B037DFFC3A6A9FF7BB169FF45" blockId="4.[151,1436,152,625]" pageId="4" pageNumber="199">
<emphasis id="B907EA99037DFFC3A6A9FF7BB128FF3B" bold="true" box="[199,624,152,175]" pageId="4" pageNumber="199">
Variation in other 
<typeStatus id="54C88829037DFFC3A7C7FF7BB157FF3B" box="[425,527,152,175]" pageId="4" pageNumber="199" type="paratype">paratype</typeStatus>
females
</emphasis>
. 
<typeStatus id="54C88829037DFFC3A4EDFF7BB1B8FF3B" box="[643,736,152,175]" pageId="4" pageNumber="199" type="paratype">Paratype</typeStatus>
females shown little variation with the 
<typeStatus id="54C88829037DFFC3A2C5FF7BB65FFF3B" box="[1195,1287,152,175]" pageId="4" pageNumber="199" type="paratype">paratype</typeStatus>
used for the description of the female in this paper.
</paragraph>
</subSubSection>
<subSubSection id="C3696500037DFFC2A6A9FF3EB2EEFE35" lastPageId="5" lastPageNumber="200" pageId="4" pageNumber="199" type="diagnosis">
<paragraph id="8BCC368B037DFFC3A6A9FF3EB253FEC8" blockId="4.[151,1436,152,625]" pageId="4" pageNumber="199">
<emphasis id="B907EA99037DFFC3A6A9FF3EB299FF60" bold="true" box="[199,449,221,244]" pageId="4" pageNumber="199">Differential diagnosis.</emphasis>
Adult males of 
<taxonomicName id="4C734D08037DFFC3A405FF3EB18BFF60" authority="Phan, 2017" authorityName="Phan" authorityYear="2017" box="[619,723,220,244]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="duytan" status="sp. nov.">
<emphasis id="B907EA99037DFFC3A405FF3EB18BFF60" box="[619,723,220,244]" italics="true" pageId="4" pageNumber="199">C. duytan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457E2037DFFC3A4B6FF3EB075FF60" box="[728,813,221,244]" pageId="4" pageNumber="199" rank="species">
<emphasis id="B907EA99037DFFC3A4B6FF3EB075FF60" bold="true" box="[728,813,221,244]" pageId="4" pageNumber="199">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C734D08037DFFC3A50FFF3EB081FF60" authorityName="Phan &amp; Kompier" authorityYear="2016" box="[865,985,220,244]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="B907EA99037DFFC3A50FFF3EB081FF60" box="[865,985,220,244]" italics="true" pageId="4" pageNumber="199">C. hayashii</emphasis>
</taxonomicName>
can be easily separated from all described 
<taxonomicName id="4C734D08037DFFC3A6F9FF1CB3A5FE83" authorityName="Kirby" authorityYear="1890" box="[151,253,255,279]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B907EA99037DFFC3A6F9FF1CB3A5FE83" box="[151,253,255,279]" italics="true" pageId="4" pageNumber="199">Coeliccia</emphasis>
</taxonomicName>
by the presence of a large rectangular pruinose spot on the synthorax spreading out, but not fully covering, on both mesepimeron and metepisternum (
<figureCitation id="13482A0E037DFFC3A43FFEC1B1E6FEAD" box="[593,702,290,313]" captionStart="FIGURE 1" captionStartId="1.[151,244,1952,1974]" captionTargetBox="[188,1397,295,1930]" captionTargetId="figure@1.[188,1398,295,1930]" captionTargetPageId="1" captionText="FIGURE 1. Coeliccia spp., head and thorax, ♂. (1 a) Coeliccia duytan sp. nov.; (1 b) Coeliccia hayashii." httpUri="https://zenodo.org/record/996999/files/figure.png" pageId="4" pageNumber="199">Fig. 1a–b</figureCitation>
). 
<taxonomicName id="4C734D08037DFFC3A4B8FEC1B01CFEAD" authorityName="Phan" authorityYear="2017" box="[726,836,289,313]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="duytan">
<emphasis id="B907EA99037DFFC3A4B8FEC1B01CFEAD" box="[726,836,289,313]" italics="true" pageId="4" pageNumber="199">C. duytan</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C734D08037DFFC3A5B1FEC1B705FEAD" authorityName="Phan &amp; Kompier" authorityYear="2016" box="[991,1117,289,313]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="B907EA99037DFFC3A5B1FEC1B705FEAD" box="[991,1117,289,313]" italics="true" pageId="4" pageNumber="199">C. hayashii</emphasis>
</taxonomicName>
by the following diagnostic characters:
</paragraph>
<paragraph id="8BCC368B037DFFC3A6A9FE8BB141FE35" blockId="4.[151,1436,152,625]" pageId="4" pageNumber="199">
(1) Mesepisternum of 
<taxonomicName id="4C734D08037DFFC3A7D3FE8BB170FEEB" authorityName="Phan" authorityYear="2017" box="[445,552,359,383]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="duytan">
<emphasis id="B907EA99037DFFC3A7D3FE8BB170FEEB" box="[445,552,359,383]" italics="true" pageId="4" pageNumber="199">C. duytan</emphasis>
</taxonomicName>
with a large quadrate pruinose spot (
<figureCitation id="13482A0E037DFFC3A5A8FE8BB74DFEEB" box="[966,1045,360,383]" captionStart="FIGURE 1" captionStartId="1.[151,244,1952,1974]" captionTargetBox="[188,1397,295,1930]" captionTargetId="figure@1.[188,1398,295,1930]" captionTargetPageId="1" captionText="FIGURE 1. Coeliccia spp., head and thorax, ♂. (1 a) Coeliccia duytan sp. nov.; (1 b) Coeliccia hayashii." httpUri="https://zenodo.org/record/996999/files/figure.png" pageId="4" pageNumber="199">Fig. 1a</figureCitation>
) while 
<taxonomicName id="4C734D08037DFFC3A202FE8BB7B0FEEB" authorityName="Phan &amp; Kompier" authorityYear="2016" box="[1132,1256,359,383]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="B907EA99037DFFC3A202FE8BB7B0FEEB" box="[1132,1256,359,383]" italics="true" pageId="4" pageNumber="199">C. hayashii</emphasis>
</taxonomicName>
has a long blue narrow antehumeral stripe (
<figureCitation id="13482A0E037DFFC3A7D3FE69B154FE35" box="[445,524,394,417]" captionStart="FIGURE 1" captionStartId="1.[151,244,1952,1974]" captionTargetBox="[188,1397,295,1930]" captionTargetId="figure@1.[188,1398,295,1930]" captionTargetPageId="1" captionText="FIGURE 1. Coeliccia spp., head and thorax, ♂. (1 a) Coeliccia duytan sp. nov.; (1 b) Coeliccia hayashii." httpUri="https://zenodo.org/record/996999/files/figure.png" pageId="4" pageNumber="199">Fig. 1b</figureCitation>
).
</paragraph>
<paragraph id="8BCC368B037DFFC3A6A9FE4EB383FE73" blockId="4.[151,1436,152,625]" pageId="4" pageNumber="199">
(2) Apex of cercus in 
<taxonomicName id="4C734D08037DFFC3A7D8FE4EB178FE50" authorityName="Phan" authorityYear="2017" box="[438,544,428,452]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="duytan">
<emphasis id="B907EA99037DFFC3A7D8FE4EB178FE50" box="[438,544,428,452]" italics="true" pageId="4" pageNumber="199">C. duytan</emphasis>
</taxonomicName>
slightly inflated (
<figureCitation id="13482A0E037DFFC3A48FFE4EB013FE50" box="[737,843,429,452]" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="4" pageNumber="199">Fig. 2a–b</figureCitation>
), but tapering into a point apically in 
<taxonomicName id="4C734D08037DFFC3A288FE4EB638FE50" authorityName="Phan &amp; Kompier" authorityYear="2016" box="[1254,1376,428,452]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="B907EA99037DFFC3A288FE4EB638FE50" box="[1254,1376,428,452]" italics="true" pageId="4" pageNumber="199">C. hayashii</emphasis>
</taxonomicName>
(
<figureCitation id="13482A0E037DFFC3A31FFE4EB396FE73" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="4" pageNumber="199">Fig. 2c–d</figureCitation>
).
</paragraph>
<paragraph id="8BCC368B037DFFC3A6A9FE11B072FDB8" blockId="4.[151,1436,152,625]" pageId="4" pageNumber="199">
(3) S 
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<taxonomicName id="4C734D08037DFFC3A75CFE11B2C5FD9D" authorityName="Phan" authorityYear="2017" box="[306,413,497,521]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="duytan">
<emphasis id="B907EA99037DFFC3A75CFE11B2C5FD9D" box="[306,413,497,521]" italics="true" pageId="4" pageNumber="199">C. duytan</emphasis>
</taxonomicName>
is almost entirely black except for a narrow yellow apical ring posteriorly (
<figureCitation id="13482A0E037DFFC3A28DFE11B617FD9D" box="[1251,1359,498,521]" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="4" pageNumber="199">Fig. 2a–b</figureCitation>
). In 
<taxonomicName id="4C734D08037DFFC3A3EAFE11B3A9FDB8" authorityName="Phan &amp; Kompier" authorityYear="2016" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="B907EA99037DFFC3A3EAFE11B3A9FDB8" italics="true" pageId="4" pageNumber="199">C. hayashii</emphasis>
</taxonomicName>
the posterior half of S9 is largely yellow (
<figureCitation id="13482A0E037DFFC3A4DAFDF6B045FDB8" box="[692,797,533,556]" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="4" pageNumber="199">Fig. 2d–e</figureCitation>
).
</paragraph>
<paragraph id="8BCC368B037DFFC3A6A9FDDBB200FDE5" blockId="4.[151,1436,152,625]" pageId="4" pageNumber="199">
(4) Flagella of genital ligula gradually tapering with acute apices in 
<taxonomicName id="4C734D08037DFFC3A5F9FDDBB0A7FDDB" authorityName="Phan" authorityYear="2017" box="[919,1023,567,591]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="duytan">
<emphasis id="B907EA99037DFFC3A5F9FDDBB0A7FDDB" box="[919,1023,567,591]" italics="true" pageId="4" pageNumber="199">C. duytan</emphasis>
</taxonomicName>
(
<figureCitation id="13482A0E037DFFC3A262FDDBB702FDDB" box="[1036,1114,568,591]" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="4" pageNumber="199">Fig. 2c</figureCitation>
), but apical 1/5 spatulate in 
<taxonomicName id="4C734D08037DFFC3A3EAFDDBB3A9FDE5" authorityName="Phan &amp; Kompier" authorityYear="2016" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="B907EA99037DFFC3A3EAFDDBB3A9FDE5" italics="true" pageId="4" pageNumber="199">C. hayashii</emphasis>
</taxonomicName>
(
<figureCitation id="13482A0E037DFFC3A76EFDB9B214FDE5" box="[256,332,602,625]" captionStart="FIGURE 2" captionStartId="2.[151,244,1783,1805]" captionTargetBox="[189,1396,854,1749]" captionTargetId="figure@2.[188,1398,853,1762]" captionTargetPageId="2" captionText="FIGURE 2. Coeliccia spp., ♂. [2 a – c] Coeliccia duytan sp. nov.; [2 d – f] Coeliccia hayashii. (2 a, 2 d) appendages in dorsal view; (2 b, 2 e) appendages in lateral view; (2 c, 2 f) genital ligula in dorsal view (2 f from Fig. 4 C, p. 411 in Phan &amp; Kompier 2016)." httpUri="https://zenodo.org/record/997001/files/figure.png" pageId="4" pageNumber="199">Fig. 2f</figureCitation>
).
</paragraph>
<caption id="DF0C6603037DFFC3A6F9F87EB01EF827" box="[151,838,1949,1971]" httpUri="https://zenodo.org/record/997005/files/figure.png" pageId="4" pageNumber="199" targetBox="[153,1433,677,1928]" targetPageId="4">
<paragraph id="8BCC368B037DFFC3A6F9F87EB01EF827" blockId="4.[151,838,1949,1971]" box="[151,838,1949,1971]" pageId="4" pageNumber="199">
<emphasis id="B907EA99037DFFC3A6F9F87EB254F827" bold="true" box="[151,268,1949,1971]" pageId="4" pageNumber="199">FIGURE 4.</emphasis>
<taxonomicName id="4C734D08037DFFC3A77CF87DB2EDF827" authority="Phan, 2017" authorityName="Phan" authorityYear="2017" box="[274,437,1950,1971]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="duytan" status="sp. nov.">
<emphasis id="B907EA99037DFFC3A77CF87DB2EDF827" box="[274,437,1950,1971]" italics="true" pageId="4" pageNumber="199">Coeliccia duytan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457E2037DFFC3A7D5F87EB15FF827" box="[443,519,1949,1971]" pageId="4" pageNumber="199" rank="species">
<emphasis id="B907EA99037DFFC3A7D5F87EB15FF827" bold="true" box="[443,519,1949,1971]" pageId="4" pageNumber="199">sp. nov.</emphasis>
</taxonomicNameLabel>
in nature (4a) male; (4b) female.
</paragraph>
</caption>
<paragraph id="8BCC368B037CFFC2A6A9FF7BB000FF45" blockId="5.[151,1436,151,417]" pageId="5" pageNumber="200">
(5) The examined specimens of 
<taxonomicName id="4C734D08037CFFC2A457FF7BB1FEFF3B" authorityName="Phan" authorityYear="2017" box="[569,678,151,175]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="duytan">
<emphasis id="B907EA99037CFFC2A457FF7BB1FEFF3B" box="[569,678,151,175]" italics="true" pageId="5" pageNumber="200">C. duytan</emphasis>
</taxonomicName>
are smaller than 
<taxonomicName id="4C734D08037CFFC2A51AFF7BB0AAFF3B" authorityName="Phan &amp; Kompier" authorityYear="2016" box="[884,1010,151,175]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="B907EA99037CFFC2A51AFF7BB0AAFF3B" box="[884,1010,151,175]" italics="true" pageId="5" pageNumber="200">C. hayashii</emphasis>
</taxonomicName>
(hindwing 26 and 29, abdomen incl. appendages 41 and 
<quantity id="4C8B9B6E037CFFC2A706FF59B2C7FF45" box="[360,415,186,209]" metricMagnitude="0" metricUnit="m" metricValue="1.143" pageId="5" pageNumber="200" unit="in" value="45.0">45 in</quantity>
<taxonomicName id="4C734D08037CFFC2A7CBFF59B154FF45" authorityName="Phan" authorityYear="2017" box="[421,524,185,209]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="duytan">
<emphasis id="B907EA99037CFFC2A7CBFF59B154FF45" box="[421,524,185,209]" italics="true" pageId="5" pageNumber="200">C. duytan</emphasis>
</taxonomicName>
and 
<emphasis id="B907EA99037CFFC2A42EFF59B199FF45" box="[576,705,185,209]" italics="true" pageId="5" pageNumber="200">
<taxonomicName id="4C734D08037CFFC2A42EFF59B1E5FF45" authorityName="Phan &amp; Kompier" authorityYear="2016" box="[576,701,185,209]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="hayashii">C. hayashii</taxonomicName>
,
</emphasis>
respectively).
</paragraph>
<paragraph id="8BCC368B037CFFC2A6A9FF3EB20BFE83" blockId="5.[151,1436,151,417]" pageId="5" pageNumber="200">
The female of 
<taxonomicName id="4C734D08037CFFC2A70DFF3EB293FF60" authorityName="Phan" authorityYear="2017" box="[355,459,220,244]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="duytan">
<emphasis id="B907EA99037CFFC2A70DFF3EB293FF60" box="[355,459,220,244]" italics="true" pageId="5" pageNumber="200">C. duytan</emphasis>
</taxonomicName>
can be separated from most other congeners by the complex morphology of the prothoracic lobes (
<figureCitation id="13482A0E037CFFC2A6B0FEE3B21FFE83" box="[222,327,256,279]" captionStart="FIGURE 3" captionStartId="3.[151,244,1850,1872]" captionTargetBox="[188,1398,642,1829]" captionTargetId="figure@3.[188,1398,642,1829]" captionTargetPageId="3" captionText="FIGURE 3. Coeliccia duytan sp. nov. (3 a) head male; (3 b) head female; (3 c) prothorax female in lateral view; (3 d) prothorax female in dorsal view; (3 e) tip of abdomen, female." httpUri="https://zenodo.org/record/997003/files/figure.png" pageId="5" pageNumber="200">Fig. 3c–d</figureCitation>
).
</paragraph>
<paragraph id="8BCC368B037CFFC2A6A9FEC1B2EEFE35" blockId="5.[151,1436,151,417]" pageId="5" pageNumber="200">
<emphasis id="B907EA99037CFFC2A6A9FEC1B2E6FEAD" bold="true" box="[199,446,290,313]" pageId="5" pageNumber="200">Habitat and Ecology.</emphasis>
This is a shy species, usually perching ca. 
<quantity id="4C8B9B6E037CFFC2A5C5FEC1B740FEAD" box="[939,1048,290,313]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="200" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 cm</quantity>
high in dark forest underlain with decomposing leaf litter. It was found at streams (
<quantity id="4C8B9B6E037CFFC2A4F4FEA6B180FEC8" box="[666,728,325,348]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="5" pageNumber="200" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4–6m</quantity>
width) with large stones in pristine forest. At the 
<typeStatus id="54C88829037CFFC2A28BFEA6B64BFEC8" box="[1253,1299,325,348]" pageId="5" pageNumber="200">type</typeStatus>
locality, this species occurs with two other congeners 
<taxonomicName id="4C734D08037CFFC2A420FE8BB083FEEB" authority="Kompier &amp; Phan, 2017" authorityName="Kompier &amp; Phan" authorityYear="2017" box="[590,987,360,383]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="mientrung">
<emphasis id="B907EA99037CFFC2A420FE8BB183FEEB" box="[590,731,360,383]" italics="true" pageId="5" pageNumber="200">C. mientrung</emphasis>
Kompier &amp; Phan, 2017
</taxonomicName>
and 
<taxonomicName id="4C734D08037CFFC2A27FFE8BB64DFEEB" authority="(Selys, 1863)" authorityName="Selys" authorityYear="1863" baseAuthorityName="Selys" baseAuthorityYear="1863" box="[1041,1301,359,383]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="200" phylum="Arthropoda" rank="species" species="dydima">
<emphasis id="B907EA99037CFFC2A27FFE8BB726FEEB" box="[1041,1150,359,383]" italics="true" pageId="5" pageNumber="200">C. dydima</emphasis>
(Selys, 1863)
</taxonomicName>
(new record for 
<collectingCountry id="F364761B037CFFC2A6D2FE69B241FE35" box="[188,281,394,417]" name="Vietnam" pageId="5" pageNumber="200">Vietnam</collectingCountry>
, unpublished).
</paragraph>
</subSubSection>
</treatment>
</document>