354 lines
44 KiB
XML
354 lines
44 KiB
XML
<document id="B684C3C97162703A24DD5FBBAB5D4EE0" ID-DOI="10.11646/zootaxa.4012.1.9" ID-GBIF-Dataset="4c0d8376-71b9-42d1-8dbc-08e0ff16338a" ID-ISSN="1175-5326" ID-Zenodo-Dep="244632" ID-ZooBank="55822610-6FE6-4218-A412-A080F3218F51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461239579310" checkinUser="plazi" docAuthor="Keith, Denis, Sabatinelli, Guido & Uliana, Marco" docDate="2015" docId="03DA87884C3EFFE4FF0FB543FAC36BF6" docLanguage="en" docName="zt04012p180.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pharaonus (Pharaonus) adelphus Keith, Sabatinelli" docType="treatment" docVersion="8" lastPageNumber="171" masterDocId="FFE3FFF04C3FFFE0FF98B26BFF826853" masterDocTitle="Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa" masterLastPageNumber="180" masterPageNumber="167" pageNumber="168" updateTime="1698605019073" updateUser="plazi">
|
||
<mods:mods id="F67B8F19DBEEF167D5879E744D1430A4" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="C1F56C9AF6162CA6B1506764D8CE1D98">
|
||
<mods:title id="22D4F0ED3EBC7711D4689512F64E6882">Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="5B120B755F51CCB572A05480D6033057" type="personal">
|
||
<mods:role id="BF8855483A2D80472C0E199E5FA41404">
|
||
<mods:roleTerm id="F9FACF26B517CAAA84007E457DCD7A21">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A062003090EFECC7B8B51AFDB5A489A2">Keith, Denis</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="445E8F11C493BB1F235F2B5EA61E7016" type="personal">
|
||
<mods:role id="D0D94174D61D26CAEF43860123C1D24A">
|
||
<mods:roleTerm id="E77B27E81B0B5BDB45EFAE50A6FC4C5D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F2689E949CFC595F05E78CADAC67B695">Sabatinelli, Guido</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="CA4EB5659E057B52B83CA7B677997C95" type="personal">
|
||
<mods:role id="23E1848A5CD24B5EC0EB8FA76256B4B3">
|
||
<mods:roleTerm id="7338EC6BC60F1135B4FB8B1E8341CD8C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="96A9427B35D307D058BA7AC200609827">Uliana, Marco</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="661C7683490CD0FD0789578CD3192D34">text</mods:typeOfResource>
|
||
<mods:relatedItem id="36F14E4A041ABCE42E1CA5B71C71AC3D" type="host">
|
||
<mods:titleInfo id="7FED6FC34121BF850F20B49E69B71BDF">
|
||
<mods:title id="0BDA140F6DBF7A5F41A289FA72ECC24D">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="94B368161D2D949D37BCE559ED8625C6">
|
||
<mods:date id="349BD171D8FAB89DA34ADEB7076284BA">2015</mods:date>
|
||
<mods:detail id="D4FF88EBE7DDF1A5BDBE4195FF07FDC5" type="volume">
|
||
<mods:number id="A7893DE5775ED1094F54EBF31793CFBA">4012</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="BFDAC044B249B62FD1352710D7E0126A" type="issue">
|
||
<mods:number id="182300D3D326D1222D4D25CE1DCF08E1">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="A621122C2F33FBA46DE793230FC1B63F" unit="page">
|
||
<mods:start id="A908369B8B9950FD264E60EE5FA2AE6F">167</mods:start>
|
||
<mods:end id="045D718FCBD180E57D08D61DA41E345A">180</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="4EE585822D7D3651876E3107C5D69762">journal article</mods:classification>
|
||
<mods:identifier id="3805C17AD3278EFF15A711439FB86A33" type="DOI">10.11646/zootaxa.4012.1.9</mods:identifier>
|
||
<mods:identifier id="0956837F54A28D072565428EFAC99D4D" type="GBIF-Dataset">4c0d8376-71b9-42d1-8dbc-08e0ff16338a</mods:identifier>
|
||
<mods:identifier id="553BDFB769059FD5F0CD706BFF02BFBC" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="56BD6DF08381DE45F66EC08E0C1CA8F4" type="Zenodo-Dep">244632</mods:identifier>
|
||
<mods:identifier id="21FFD9D869844FD26650504CAE5796BD" type="ZooBank">55822610-6FE6-4218-A412-A080F3218F51</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03DA87884C3EFFE4FF0FB543FAC36BF6" ID-DOI="http://doi.org/10.5281/zenodo.5694702" ID-GBIF-Taxon="119632615" ID-Zenodo-Dep="5694702" LSID="urn:lsid:plazi:treatment:03DA87884C3EFFE4FF0FB543FAC36BF6" httpUri="http://treatment.plazi.org/id/03DA87884C3EFFE4FF0FB543FAC36BF6" lastPageId="4" lastPageNumber="171" pageId="1" pageNumber="168">
|
||
<subSubSection id="C36965154C3EFFE1FF0FB543FE126F37" pageId="1" pageNumber="168" type="nomenclature">
|
||
<paragraph id="8BCC369E4C3EFFE1FF0FB543FBCB6F11" blockId="1.[151,1097,1832,1892]" box="[151,1097,1832,1858]" pageId="1" pageNumber="168">
|
||
<heading id="D08481F24C3EFFE1FF0FB543FBCB6F11" bold="true" box="[151,1097,1832,1858]" fontSize="11" level="1" pageId="1" pageNumber="168" reason="1">
|
||
<emphasis id="B907EA8C4C3EFFE1FF0FB543FBCB6F11" bold="true" box="[151,1097,1832,1858]" pageId="1" pageNumber="168">
|
||
<taxonomicName id="4C734D1D4C3EFFE1FF0FB543FC966F11" authority="Keith, Sabatinelli" authorityName="Keith, Sabatinelli" box="[151,788,1832,1858]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="species" species="adelphus" subGenus="Pharaonus">
|
||
<emphasis id="B907EA8C4C3EFFE1FF0FB543FE9F6F11" bold="true" box="[151,285,1832,1858]" italics="true" pageId="1" pageNumber="168">Pharaonus</emphasis>
|
||
(
|
||
<emphasis id="B907EA8C4C3EFFE1FEB5B543FE306F11" bold="true" box="[301,434,1832,1858]" italics="true" pageId="1" pageNumber="168">Pharaonus</emphasis>
|
||
)
|
||
<emphasis id="B907EA8C4C3EFFE1FE5CB543FDB36F11" bold="true" box="[452,561,1832,1858]" italics="true" pageId="1" pageNumber="168">adelphus</emphasis>
|
||
Keith, Sabatinelli
|
||
</taxonomicName>
|
||
, and Uliana, new species
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C3EFFE1FF0FB520FE126F37" blockId="1.[151,1097,1832,1892]" box="[151,400,1867,1892]" pageId="1" pageNumber="168">
|
||
(
|
||
<figureCitation id="13482A1B4C3EFFE1FF07B520FE886F37" box="[159,266,1867,1892]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1953,1975]" captionTargetBox="[212,1376,196,1915]" captionTargetId="figure@3.[201,1386,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Habitus of Pharaonus (Pharaonus) species. 3: P. adelphus, male paratype (Kerman, Iran). 4: P. adelphus, female paratype (Kerman, Bakram-djord [= Bahramjerd], Iran). 5: P. caucasicus, male (Gorovan sands, Armenia). 6: P. farsensis, male paratype (Darab, Iran)." httpUri="https://zenodo.org/record/244634/files/figure.png" pageId="1" pageNumber="168">Figs. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C3EFFE1FE8EB527FEB66F37" box="[278,308,1868,1892]" captionStart="FIGURES 13 – 14" captionStartId="7.[151,264,1550,1572]" captionTargetBox="[169,1432,195,1512]" captionTargetId="figure@7.[155,1432,195,1528]" captionTargetPageId="7" captionText="FIGURES 13 – 14. Different elytral setation in evidence. 13: P. adelphus. 14: P. farsensis." httpUri="https://zenodo.org/record/244637/files/figure.png" pageId="1" pageNumber="168">13</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C3EFFE1FED8B520FE056F37" box="[320,391,1867,1892]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="1" pageNumber="168">15, 21</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C3EFFE2FF0FB5F8FD9E6886" lastPageId="2" lastPageNumber="169" pageId="1" pageNumber="168" type="diagnosis">
|
||
<paragraph id="8BCC369E4C3EFFE2FF0FB5F8FD9E6886" blockId="1.[151,1436,1939,2036]" lastBlockId="2.[151,1437,151,2015]" lastPageId="2" lastPageNumber="169" pageId="1" pageNumber="168">
|
||
<emphasis id="B907EA8C4C3EFFE1FF0FB5F8FE936FFF" bold="true" box="[151,273,1939,1964]" pageId="1" pageNumber="168">Diagnosis.</emphasis>
|
||
A species of
|
||
<taxonomicName id="4C734D1D4C3EFFE1FE26B5F8FDBB6FFF" box="[446,569,1939,1964]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907EA8C4C3EFFE1FE26B5F8FDBB6FFF" box="[446,569,1939,1964]" italics="true" pageId="1" pageNumber="168">Pharaonus</emphasis>
|
||
</taxonomicName>
|
||
with dark body integuments, except for yellowish-brown elytra. Clypeus strongly transverse (2.2–2.5 times as wide as long), with right angles. Head and pronotum with long, erect pilosity; elytra glabrous except for on narrow basal-humeral area. Pronotum impressed along the midline, punctation dense and coarse. Pygidium with a large, impunctate area. Antennal club of the male 0.91–0.94 times as long as the clypeus width. Protibiae bidentate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C3DFFE2FF5FB2B4FAEB6936" pageId="2" pageNumber="169" type="materials_examined">
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB2B4FAEB6936" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">
|
||
<emphasis id="B907EA8C4C3DFFE2FF5FB2B4FE3468AB" bold="true" box="[199,438,223,248]" pageId="2" pageNumber="169">Examined material.</emphasis>
|
||
<typeStatus id="54C8883C4C3DFFE2FE27B2B4FDAE68AB" box="[447,556,223,248]" pageId="2" pageNumber="169" type="holotype">Holotype</typeStatus>
|
||
: [
|
||
<collectingCountry id="F364760E4C3DFFE2FDD8B28BFDF668A4" box="[576,628,224,247]" name="Iran" pageId="2" pageNumber="169">Iran</collectingCountry>
|
||
], Kerman,
|
||
<date id="FFCD105E4C3DFFE2FD6DB28BFCAC68AB" box="[757,814,224,248]" pageId="2" pageNumber="169">2.VI.</date>
|
||
[19]51, [leg.] Salavatian, ♂ (
|
||
<collectionCode id="ED62AE5B4C3DFFE2FB1CB28BFB6268AB" box="[1156,1248,224,248]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="169">MHNG</collectionCode>
|
||
); aedeagus (
|
||
<figureCitation id="13482A1B4C3DFFE2FAF7B2B4FF31694E" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="2" pageNumber="169">Fig. 15</figureCitation>
|
||
d) lost after being photographed.
|
||
<typeStatus id="54C8883C4C3DFFE2FDA8B36EFD23694E" box="[560,673,261,285]" pageId="2" pageNumber="169" type="paratype">Paratypes</typeStatus>
|
||
: same data as the
|
||
<typeStatus id="54C8883C4C3DFFE2FCE9B36FFC54694E" box="[881,982,260,285]" pageId="2" pageNumber="169" type="holotype">holotype</typeStatus>
|
||
,
|
||
<specimenCount id="9D75FD174C3DFFE2FC7BB36FFB8C694F" box="[995,1038,260,284]" pageId="2" pageNumber="169" type="generic">2 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C3DFFE2FB87B36FFBF1694F" box="[1055,1139,260,284]" httpUri="http://grbio.org/cool/gj99-xj2c" name="Jardí Botànic Marimurtra" pageId="2" pageNumber="169">HMIM</collectionCode>
|
||
,
|
||
<collectionCode id="ED62AE5B4C3DFFE2FB18B36FFB5A694F" box="[1152,1240,260,284]" httpUri="http://grbio.org/cool/vphy-ev73" name="Mie University Culture Collection (Culture Collection, Laboratory of Plant Pathology)" pageId="2" pageNumber="169">MUCC</collectionCode>
|
||
); [
|
||
<collectingCountry id="F364760E4C3DFFE2FB6BB36EFAA5694F" box="[1267,1319,261,284]" name="Iran" pageId="2" pageNumber="169">Iran</collectingCountry>
|
||
], Kerman, Bakram-djord [= Bahramjerd,
|
||
<emphasis id="B907EA8C4C3DFFE2FE76B342FDAE6913" box="[494,556,297,320]" italics="true" pageId="2" pageNumber="169">sensu</emphasis>
|
||
<bibRefCitation id="EFE24B6F4C3DFFE2FDABB34CFD3F6913" author="Barari" box="[563,701,295,320]" pageId="2" pageNumber="169" refString="Barari, H. (1998) The first report of two species of scarab beetles from Iran. (Coleoptera: Scarabaeoidea). Journal of the Entomological Society of Iran, 16 - 17, 39 - 40 (in English), 73 - 74 (in Persian)." type="book chapter" year="1998">Barari 1998</bibRefCitation>
|
||
],
|
||
<date id="FFCD105E4C3DFFE2FD48B34CFC956913" box="[720,791,295,320]" pageId="2" pageNumber="169">17.IV.</date>
|
||
[19]51, [leg.] Salavatian leg, 1 ♀ (
|
||
<collectionCode id="ED62AE5B4C3DFFE2FB0DB343FB636913" box="[1173,1249,296,320]" pageId="2" pageNumber="169">GSCP</collectionCode>
|
||
). [
|
||
<collectingCountry id="F364760E4C3DFFE2FB63B343FAAD696C" box="[1275,1327,296,319]" name="Iran" pageId="2" pageNumber="169">Iran</collectingCountry>
|
||
],
|
||
<collectingCountry id="F364760E4C3DFFE2FADAB34CFA1A6913" box="[1346,1432,295,320]" name="Iran" pageId="2" pageNumber="169">Persien</collectingCountry>
|
||
, Prov. Kerman,
|
||
<date id="FFCD105E4C3DFFE2FED8B327FEEC6936" box="[320,366,332,357]" pageId="2" pageNumber="169">7.V.</date>
|
||
[19]64, and a more recent label “
|
||
<collectingCountry id="F364760E4C3DFFE2FD44B327FCA76936" box="[732,805,332,357]" name="Iran" pageId="2" pageNumber="169">Persia</collectingCountry>
|
||
, Prov. Kerniam”[sic!], 1 ♀ (
|
||
<collectionCode id="ED62AE5B4C3DFFE2FBFAB327FB226937" box="[1122,1184,332,356]" httpUri="http://grbio.org/cool/fx6k-enrh" name="Natal Museum" pageId="2" pageNumber="169">NMP</collectionCode>
|
||
, ex coll. D. Král).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C3DFFE2FF5FB304FBE46F8C" pageId="2" pageNumber="169" type="description">
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB304FF7669FF" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">
|
||
<emphasis id="B907EA8C4C3DFFE2FF5FB304FDCD69DB" bold="true" box="[199,591,367,392]" pageId="2" pageNumber="169">
|
||
Description of the male
|
||
<typeStatus id="54C8883C4C3DFFE2FE70B304FDCD69DB" box="[488,591,367,392]" pageId="2" pageNumber="169" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Length (from the anterior margin of the clypeus to the apex of the elytra)
|
||
<quantity id="4C8B9B7B4C3DFFE2FF0FB3FFFF7269FF" box="[151,240,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="2" pageNumber="169" unit="mm" value="8.1">8.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB3D1FB1969A4" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">Head, pronotum and underside black, antenna light brown with apex of 1st article and club darker, legs black with brown tarsi, elytra yellowish brown, darker along the suture and the epipleural margin.</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB069FB756B44" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">Head: clypeus very transverse (2.5 times as wide as long), almost rectangular, with margins clearly raised. Anterior margin straight, anterior angles right, broadly rounded. Integument glabrous; with large, shallow, and irregular punctures defining a coarse and reticulate sculpture. Frontoclypeal suture almost invisible, but marked by a dense tuft of long, yellowish-gray, erect pilosity. Frons with sculpture similar to that of the clypeus, and with pilosity becoming shorter and sparser going from frontoclypeal suture towards vertex. Ocular canthus strong, eyes small, not very protruding. Antennae with 9 antennomeres; club with 3 antennomeres, 1.8 times longer than the funicle (antennomeres 2–6) and 0.93 times as long as clypeal width. Scape noticeably enlarged at the apex; antennomere 2 wider than long; antennomeres 3, 4, 5 increasing in length; antennomere 6 discoidal.</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB148FE896C41" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">Pronotum transverse, 1.35 times wider than long. Sides subparallel in the basal two thirds, strongly convergent in the anterior third, slightly concave in both regions and most notably in the basal region. Anterior angles acute; posterior angles rounded and right. Base broadly truncate in front of scutellum. Anterior margin inconspicuous, lateral and basal margins distinct, the basal margin broader laterally. Punctation dense, forming irregular series of rough punctures, open dorsally. Surface with long, greyish-yellow, erect pilosity. Median longitudinal furrow weakly defined over the entire length. Scutellum semi-elliptical, with sculpture and pilosity similar to those of the pronotum.</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB675FAD36D5C" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">Elytra quite flat with 7 poorly defined striae of punctures, with sparse punctures over entire surface, punctures less dense than on pronotum. First (sutural) interval slightly raised in the distal half, flat in the basal half. Humeral callus strong, more finely punctate than the surrounding integument. Apical callus also evident, but weaker. Surface glabrous but for some sparse setae along the basal and humeral area. Epipleural margin broad and inflated under the humeral callus, then flattened and narrowing until the middle of the elytra; from the middle of the elytra to the apex again with relief in the form of a narrow border. Membranous margin of the epipleura gradually widening from the basal third to the apex, where it is conspicuous. Suture prolonged in a short, apical spine.</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB771FF736DCC" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">Propygidium punctate all over, without apical fringe of pilosity. Pygidium convex, with a glabrous central band, which is smooth at the base and sparsely punctate at apical half. Distinct punctate triangular area on either side of central band bearing a patch of almost adpressed, greyish-white, long pilosity. Apical margin with erect pilosity.</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB7C1FBF16E00" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">Protibiae bidentate, both teeth acute and directed forwards; apical tooth slightly curved. Apical inner spur short, inconspicuous. Protarsomeres 1–4 slightly and gradually widened. Protarsomere 5 approximately of the same length as the 4 preceding tarsomeres. Inner claw broad, bifid; outer claw narrow. Mesotibiae clearly tumefied in the middle. Claws of the mesolegs simple and elongate. Metatibiae elongate, slightly enlarged in the middle, with 2 notches bearing spiniform setae. Claws of the metalegs identical to the mesolegs.</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB435FED26EC8" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">Labrum appearing slightly bilobate. Terminal maxillary palpomere elongate, fusiform, with a sulcus in the basal two thirds.</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB4CDFE4C6EEC" blockId="2.[151,1437,151,2015]" box="[199,462,1702,1727]" pageId="2" pageNumber="169">
|
||
Aedeagus as in
|
||
<figureCitation id="13482A1B4C3DFFE2FEEFB4CDFE486EEC" box="[375,458,1702,1727]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="2" pageNumber="169">Fig. 15</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB4A1FE886F1C" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">
|
||
<emphasis id="B907EA8C4C3DFFE2FF5FB4A1FE9C6EB0" bold="true" box="[199,286,1738,1763]" pageId="2" pageNumber="169">Female</emphasis>
|
||
(
|
||
<figureCitation id="13482A1B4C3DFFE2FEAEB4A1FE006EB0" box="[310,386,1738,1763]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1953,1975]" captionTargetBox="[212,1376,196,1915]" captionTargetId="figure@3.[201,1386,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Habitus of Pharaonus (Pharaonus) species. 3: P. adelphus, male paratype (Kerman, Iran). 4: P. adelphus, female paratype (Kerman, Bakram-djord [= Bahramjerd], Iran). 5: P. caucasicus, male (Gorovan sands, Armenia). 6: P. farsensis, male paratype (Darab, Iran)." httpUri="https://zenodo.org/record/244634/files/figure.png" pageId="2" pageNumber="169">Fig. 4</figureCitation>
|
||
) with the following sexual dimorphism: slightly larger, squatter; pronotal longitudinal impression less evident; pygidium flat; legs lighter in color (brown or reddish brown), less elongate; prolegs with distal femoral tooth larger and inner anterior claw not modified; antennal club distinctly shorter. Genital sclerites as in
|
||
<figureCitation id="13482A1B4C3DFFE2FF2CB55DFE846F1C" box="[180,262,1846,1871]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="2" pageNumber="169">Fig. 21</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C3DFFE2FF5FB531FBE46F8C" blockId="2.[151,1437,151,2015]" pageId="2" pageNumber="169">
|
||
<emphasis id="B907EA8C4C3DFFE2FF5FB531FECF6F20" bold="true" box="[199,333,1882,1907]" pageId="2" pageNumber="169">Variability.</emphasis>
|
||
Size of males ranging from
|
||
<quantity id="4C8B9B7B4C3DFFE2FD13B530FC966F21" box="[651,788,1883,1907]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="8.4" metricValueMin="8.1" pageId="2" pageNumber="169" unit="mm" value="8.25" valueMax="8.4" valueMin="8.1">8.1–8.4 mm</quantity>
|
||
, size of females from
|
||
<quantity id="4C8B9B7B4C3DFFE2FB95B531FB156F21" box="[1037,1175,1882,1907]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="2" pageNumber="169" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.5–9.5 mm</quantity>
|
||
. Clypeus 2.2–2.5 times as wide as long, antennal club of the male 0.91–0.94 as long as the clypeus width. Sides of the pronotum either slightly concave in the distal portion or straight. Shape of parameres (
|
||
<figureCitation id="13482A1B4C3DFFE2FC28B5C9FB806FE8" box="[944,1026,1954,1979]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="2" pageNumber="169">Fig. 15</figureCitation>
|
||
) variable, especially in lateral view. No other relevant variation was observed on the small number of examined specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0C66164C3CFFE3FF0FB5CAFD9F6FA7" httpUri="https://zenodo.org/record/244634/files/figure.png" pageId="3" pageNumber="170" targetBox="[212,1376,196,1915]" targetPageId="3">
|
||
<paragraph id="8BCC369E4C3CFFE3FF0FB5CAFD9F6FA7" blockId="3.[151,1435,1953,2036]" pageId="3" pageNumber="170">
|
||
<emphasis id="B907EA8C4C3CFFE3FF0FB5CAFEBF6FE4" bold="true" box="[151,317,1953,1975]" pageId="3" pageNumber="170">FIGURES 3–6.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C734D1D4C3CFFE3FE24B5CAFD376FE4" box="[444,693,1953,1975]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="170" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907EA8C4C3CFFE3FE24B5CAFDA96FE4" box="[444,555,1953,1975]" italics="true" pageId="3" pageNumber="170">Pharaonus</emphasis>
|
||
(
|
||
<emphasis id="B907EA8C4C3CFFE3FDA5B5CAFD2E6FE4" box="[573,684,1953,1975]" italics="true" pageId="3" pageNumber="170">Pharaonus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species. 3:
|
||
<taxonomicName id="4C734D1D4C3CFFE3FCACB5C9FC2E6FE4" box="[820,940,1953,1975]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="170" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C3CFFE3FCACB5C9FCC26FE4" box="[820,832,1954,1975]" italics="true" pageId="3" pageNumber="170">P</emphasis>
|
||
.
|
||
<emphasis id="B907EA8C4C3CFFE3FCC8B5CAFC2E6FE4" box="[848,940,1953,1975]" italics="true" pageId="3" pageNumber="170">adelphus</emphasis>
|
||
</taxonomicName>
|
||
, male paratype (Kerman, Iran). 4:
|
||
<taxonomicName id="4C734D1D4C3CFFE3FA86B5C9FA176FE4" box="[1310,1429,1953,1975]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="170" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C3CFFE3FA86B5C9FAA96FE4" box="[1310,1323,1954,1975]" italics="true" pageId="3" pageNumber="170">P</emphasis>
|
||
.
|
||
<emphasis id="B907EA8C4C3CFFE3FAA2B5CAFA176FE4" box="[1338,1429,1953,1975]" italics="true" pageId="3" pageNumber="170">adelphus</emphasis>
|
||
</taxonomicName>
|
||
, female paratype (Kerman, Bakram-djord [= Bahramjerd], Iran). 5:
|
||
<emphasis id="B907EA8C4C3CFFE3FCFDB5AAFC7B6F85" box="[869,1017,1985,2006]" italics="true" pageId="3" pageNumber="170">
|
||
<taxonomicName id="4C734D1D4C3CFFE3FCFDB5AAFC776F85" box="[869,1013,1985,2006]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="170" phylum="Arthropoda" rank="species" species="caucasicus">P. caucasicus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
male (Gorovan sands, Armenia). 6:
|
||
<taxonomicName id="4C734D1D4C3CFFE3FA11B5AAFF736FA7" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="170" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="B907EA8C4C3CFFE3FA11B5AAFF736FA7" italics="true" pageId="3" pageNumber="170">P. farsensis</emphasis>
|
||
</taxonomicName>
|
||
, male paratype (Darab, Iran).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36965154C3BFFE4FF5FB2FCFDB76886" pageId="4" pageNumber="171" type="etymology">
|
||
<paragraph id="8BCC369E4C3BFFE4FF5FB2FCFDB76886" blockId="4.[151,1436,151,248]" pageId="4" pageNumber="171">
|
||
<emphasis id="B907EA8C4C3BFFE4FF5FB2FCFECC68E3" bold="true" box="[199,334,151,176]" pageId="4" pageNumber="171">Etymology.</emphasis>
|
||
From the Greek ἀδελφός, meaning “brother”, with reference to the similarities to
|
||
<emphasis id="B907EA8C4C3BFFE4FB46B2F2FAE668E3" box="[1246,1380,152,176]" italics="true" pageId="4" pageNumber="171">
|
||
<taxonomicName id="4C734D1D4C3BFFE4FB46B2F2FAE268E3" box="[1246,1376,152,176]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="farsensis">P. farsensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with which this species initially confused.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C3BFFE4FF5FB2B4FD4968A4" box="[199,715,223,248]" pageId="4" pageNumber="171" type="discussion">
|
||
<paragraph id="8BCC369E4C3BFFE4FF5FB2B4FD4968A4" blockId="4.[151,1436,151,248]" box="[199,715,223,248]" pageId="4" pageNumber="171">
|
||
<emphasis id="B907EA8C4C3BFFE4FF5FB2B4FECE68AB" bold="true" box="[199,332,223,248]" pageId="4" pageNumber="171">Discussion.</emphasis>
|
||
See discussion under
|
||
<taxonomicName id="4C734D1D4C3BFFE4FDDCB28AFD4668AB" box="[580,708,224,248]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="B907EA8C4C3BFFE4FDDCB28AFD4668AB" box="[580,708,224,248]" italics="true" pageId="4" pageNumber="171">P. farsensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C3BFFE4FF0FB320FEA269DB" pageId="4" pageNumber="171" type="reference_group">
|
||
<paragraph id="8BCC369E4C3BFFE4FF0FB320FEA269DB" blockId="4.[151,735,331,392]" pageId="4" pageNumber="171">
|
||
<taxonomicName id="4C734D1D4C3BFFE4FF0FB320FD5D6937" authority="Reitter, 1888" authorityName="Reitter" authorityYear="1888" box="[151,735,331,357]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="caucasicus" subGenus="Pharaonus">
|
||
<emphasis id="B907EA8C4C3BFFE4FF0FB320FDB76937" bold="true" box="[151,565,331,357]" pageId="4" pageNumber="171">
|
||
<emphasis id="B907EA8C4C3BFFE4FF0FB320FE956937" bold="true" box="[151,279,331,356]" italics="true" pageId="4" pageNumber="171">Pharaonus</emphasis>
|
||
(
|
||
<emphasis id="B907EA8C4C3BFFE4FEBFB320FE246937" bold="true" box="[295,422,331,356]" italics="true" pageId="4" pageNumber="171">Pharaonus</emphasis>
|
||
)
|
||
<emphasis id="B907EA8C4C3BFFE4FE2FB327FDB76937" bold="true" box="[439,565,332,356]" italics="true" pageId="4" pageNumber="171">caucasicus</emphasis>
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE24B6F4C3BFFE4FDDBB327FD556937" author="Reitter" box="[579,727,332,357]" pageId="4" pageNumber="171" refString="Reitter, E. (1888) Ubersicht der Arten der Coleopteren-Gattung Phyllopertha Kirby aus Europa, den angrenzenden Landern, Sibirien mit Central-Asien. Entomologische Nachrichten, 14, 289 - 294." type="journal article" year="1888">Reitter, 1888</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13482A1B4C3BFFE4FF07B304FF6F69DB" box="[159,237,367,392]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1953,1975]" captionTargetBox="[212,1376,196,1915]" captionTargetId="figure@3.[201,1386,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Habitus of Pharaonus (Pharaonus) species. 3: P. adelphus, male paratype (Kerman, Iran). 4: P. adelphus, female paratype (Kerman, Bakram-djord [= Bahramjerd], Iran). 5: P. caucasicus, male (Gorovan sands, Armenia). 6: P. farsensis, male paratype (Darab, Iran)." httpUri="https://zenodo.org/record/244634/files/figure.png" pageId="4" pageNumber="171">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C3BFFE4FF62B304FE9569DB" box="[250,279,367,392]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="4" pageNumber="171">16</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C3BFFE4FF0FB3DCFB216A33" pageId="4" pageNumber="171" type="materials_examined">
|
||
<paragraph id="8BCC369E4C3BFFE4FF0FB3DCFB216A33" blockId="4.[151,1437,439,933]" pageId="4" pageNumber="171">
|
||
<emphasis id="B907EA8C4C3BFFE4FF0FB3DCFE046983" bold="true" box="[151,390,439,464]" pageId="4" pageNumber="171">Examined material.</emphasis>
|
||
<collectingCountry id="F364760E4C3BFFE4FE08B3DCFE756983" box="[400,503,439,464]" name="Armenia" pageId="4" pageNumber="171">Armenia</collectingCountry>
|
||
:
|
||
<specimenCount id="9D75FD174C3BFFE4FD9CB3D3FD136983" box="[516,657,439,464]" pageId="4" pageNumber="171" type="generic">2 specimens</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C3BFFE4FD3BB3D3FD7D6983" LSID="urn:lsid:biocol.org:col:34988" box="[675,767,440,464]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="171">MNHN</collectionCode>
|
||
);
|
||
<collectingCountry id="F364760E4C3BFFE4FC8DB3DCFC466983" box="[789,964,439,464]" name="Armenia" pageId="4" pageNumber="171">Armenian SSR</collectingCountry>
|
||
, Vedi,
|
||
<date id="FFCD105E4C3BFFE4FB8FB3D3FBDD6983" box="[1047,1119,440,464]" pageId="4" pageNumber="171">19.VI.</date>
|
||
[19]82, leg. Kalashjan:
|
||
<specimenCount id="9D75FD174C3BFFE4FAF7B3D3FA1E699C" box="[1391,1436,440,464]" pageId="4" pageNumber="171" type="generic">2 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C3BFFE4FF07B3B7FF6969A7" box="[159,235,476,500]" pageId="4" pageNumber="171">GSCP</collectionCode>
|
||
);
|
||
<collectingCountry id="F364760E4C3BFFE4FE98B3B7FEE769A6" box="[256,357,476,501]" name="Armenia" pageId="4" pageNumber="171">Armenia</collectingCountry>
|
||
, Vedi, Gorovan,
|
||
<date id="FFCD105E4C3BFFE4FDBFB3B7FD0F69A6" box="[551,653,476,501]" pageId="4" pageNumber="171" value="1984-06-16">16.VI.84</date>
|
||
, leg. Kalashian:
|
||
<specimenCount id="9D75FD174C3BFFE4FCD5B3B7FCFB69A7" box="[845,889,476,500]" pageId="4" pageNumber="171" type="generic">4 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C3BFFE4FC11B3B7FC5F69A7" box="[905,989,476,500]" pageId="4" pageNumber="171">DKCC</collectionCode>
|
||
);
|
||
<collectingCountry id="F364760E4C3BFFE4FC6AB3B7FBD569A6" box="[1010,1111,476,501]" name="Armenia" pageId="4" pageNumber="171">Armenia</collectingCountry>
|
||
, Vedi distr., Gorovan Sands,
|
||
<date id="FFCD105E4C3BFFE4FF0FB06BFE9B6A4B" box="[151,281,512,536]" pageId="4" pageNumber="171" value="1992-06-22">22.VI.1992</date>
|
||
, vendit I. Osipov:
|
||
<specimenCount id="9D75FD174C3BFFE4FE7EB06BFD926A44" box="[486,528,512,536]" pageId="4" pageNumber="171" type="generic">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C3BFFE4FD86B06BFDE86A4B" box="[542,618,512,536]" pageId="4" pageNumber="171">GSCP</collectionCode>
|
||
);
|
||
<collectingCountry id="F364760E4C3BFFE4FDE5B394FD606A4B" box="[637,738,511,536]" name="Armenia" pageId="4" pageNumber="171">Armenia</collectingCountry>
|
||
, Vedi, Garavanskie peski [=Gorovan Sands],
|
||
<date id="FFCD105E4C3BFFE4FB45B394FAEA6A4B" box="[1245,1384,511,536]" pageId="4" pageNumber="171" value="1994-06-06" valueMax="1994-06-08" valueMin="1994-06-06">6-8.VI.1994</date>
|
||
, leg. P. Kazarjan:
|
||
<specimenCount id="9D75FD174C3BFFE4FEB6B04FFEDE6A6F" box="[302,348,548,572]" pageId="4" pageNumber="171" type="generic">2 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C3BFFE4FEE8B04FFE4B6A6F" box="[368,457,548,572]" httpUri="http://grbio.org/cool/vphy-ev73" name="Mie University Culture Collection (Culture Collection, Laboratory of Plant Pathology)" pageId="4" pageNumber="171">MUCC</collectionCode>
|
||
);
|
||
<collectingCountry id="F364760E4C3BFFE4FE78B04FFDC76A6E" box="[480,581,548,573]" name="Armenia" pageId="4" pageNumber="171">Armenia</collectingCountry>
|
||
, Ararat prov., Goravan village subr., semi-desert,
|
||
<date id="FFCD105E4C3BFFE4FB09B04FFABA6A6F" box="[1169,1336,548,572]" pageId="4" pageNumber="171" value="2013-06-03" valueMax="2013-06-08" valueMin="2013-06-03">03-08.VI.2013</date>
|
||
, Leg. T. Ghrejyan:
|
||
<specimenCount id="9D75FD174C3BFFE4FE96B023FEBA6A0C" box="[270,312,584,608]" pageId="4" pageNumber="171" type="generic">2 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C3BFFE4FEDFB023FE226A33" box="[327,416,584,608]" httpUri="http://grbio.org/cool/vphy-ev73" name="Mie University Culture Collection (Culture Collection, Laboratory of Plant Pathology)" pageId="4" pageNumber="171">MUCC</collectionCode>
|
||
);
|
||
<collectingCountry id="F364760E4C3BFFE4FE2BB02CFD9A6A33" box="[435,536,583,608]" name="Armenia" pageId="4" pageNumber="171">Armenia</collectingCountry>
|
||
, Vedi, Gorovan 17-
|
||
<date id="FFCD105E4C3BFFE4FD76B02CFCE76A33" box="[750,869,583,608]" pageId="4" pageNumber="171" value="2013-06-20">20.6.2013</date>
|
||
, leg. Skoupy:
|
||
<specimenCount id="9D75FD174C3BFFE4FB9DB023FBAD6A0C" box="[1029,1071,584,608]" pageId="4" pageNumber="171" type="generic">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C3BFFE4FBA6B023FB156A33" box="[1086,1175,584,608]" httpUri="http://grbio.org/cool/vphy-ev73" name="Mie University Culture Collection (Culture Collection, Laboratory of Plant Pathology)" pageId="4" pageNumber="171">MUCC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C3BFFE4FF5FB007FDAF6AD6" box="[199,557,620,645]" pageId="4" pageNumber="171" type="distribution">
|
||
<paragraph id="8BCC369E4C3BFFE4FF5FB007FDAF6AD6" blockId="4.[151,1437,439,933]" box="[199,557,620,645]" pageId="4" pageNumber="171">
|
||
<emphasis id="B907EA8C4C3BFFE4FF5FB007FEE26AD6" bold="true" box="[199,352,620,645]" pageId="4" pageNumber="171">Distribution.</emphasis>
|
||
<collectingCountry id="F364760E4C3BFFE4FEFFB007FE3A6AD6" box="[359,440,620,645]" name="Turkey" pageId="4" pageNumber="171">Turkey</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C3BFFE4FE5BB007FDAB6AD6" box="[451,553,620,645]" name="Armenia" pageId="4" pageNumber="171">Armenia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C3BFFE4FF5FB0E4FAC36BF6" pageId="4" pageNumber="171" type="discussion">
|
||
<paragraph id="8BCC369E4C3BFFE4FF5FB0E4FEDE6AA3" blockId="4.[151,1437,439,933]" pageId="4" pageNumber="171">
|
||
<emphasis id="B907EA8C4C3BFFE4FF5FB0E4FEB96AFB" bold="true" box="[199,315,655,680]" pageId="4" pageNumber="171">Remarks.</emphasis>
|
||
The record for
|
||
<collectingCountry id="F364760E4C3BFFE4FE67B0FBFDAE6AF4" box="[511,556,656,679]" name="Iran" pageId="4" pageNumber="171">Iran</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFE24B6F4C3BFFE4FDD8B0E4FD4F6AFB" author="Barari" box="[576,717,655,680]" pageId="4" pageNumber="171" refString="Barari, H. (1998) The first report of two species of scarab beetles from Iran. (Coleoptera: Scarabaeoidea). Journal of the Entomological Society of Iran, 16 - 17, 39 - 40 (in English), 73 - 74 (in Persian)." type="book chapter" year="1998">Barari 1998</bibRefCitation>
|
||
) is based on the specimens from Kerman, Bahramjerd, here recognized as belonging to the new species
|
||
<taxonomicName id="4C734D1D4C3BFFE4FD1EB0DEFC856A9E" box="[646,775,692,717]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C3BFFE4FD1EB0DEFC856A9E" box="[646,775,692,717]" italics="true" pageId="4" pageNumber="171">P. adelphus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C734D1D4C3BFFE4FC8EB0DFFB916A9F" box="[790,1043,692,717]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="caucasicus">
|
||
<emphasis id="B907EA8C4C3BFFE4FC8EB0DFFB916A9F" box="[790,1043,692,717]" italics="true" pageId="4" pageNumber="171">Pharaonus caucasicus</emphasis>
|
||
</taxonomicName>
|
||
should therefore be removed from the Iranian fauna.
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C3BFFE4FF5FB097FAC36BF6" blockId="4.[151,1437,439,933]" pageId="4" pageNumber="171">
|
||
The record for
|
||
<collectingCountry id="F364760E4C3BFFE4FEE9B097FE436B46" box="[369,449,764,789]" name="Turkey" pageId="4" pageNumber="171">Turkey</collectingCountry>
|
||
is apparently based only on the
|
||
<typeStatus id="54C8883C4C3BFFE4FCB7B096FCDD6B46" box="[815,863,765,789]" pageId="4" pageNumber="171">type</typeStatus>
|
||
locality (Aralik) by
|
||
<bibRefCitation id="EFE24B6F4C3BFFE4FBD0B097FB6B6B47" author="Reitter" box="[1096,1257,764,789]" pageId="4" pageNumber="171" refString="Reitter, E. (1888) Ubersicht der Arten der Coleopteren-Gattung Phyllopertha Kirby aus Europa, den angrenzenden Landern, Sibirien mit Central-Asien. Entomologische Nachrichten, 14, 289 - 294." type="journal article" year="1888">Reitter (1888)</bibRefCitation>
|
||
, which requires confirmation. However, Aralik is close to the boundary with
|
||
<collectingCountry id="F364760E4C3BFFE4FCD8B174FC276B6B" box="[832,933,799,824]" name="Armenia" pageId="4" pageNumber="171">Armenia</collectingCountry>
|
||
, just about
|
||
<quantity id="4C8B9B7B4C3BFFE4FBBCB174FBE96B6B" box="[1060,1131,799,824]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="171" unit="km" value="15.0">15 km</quantity>
|
||
from the Armenian area of occurrence. The present-day distribution seems limited to sandy areas of the Goravan State Nature Reservation, while in the past the species was recorded from nearby sites from which it has apparently disappeared (Iablokov-
|
||
<bibRefCitation id="EFE24B6F4C3BFFE4FF0FB1E7FED26BF6" author="Khnzorian" box="[151,336,908,933]" pageId="4" pageNumber="171" refString="Iablokov-Khnzorian, S. (1967) Nasekomye zhestkokrylye. Plastincatousye. Fauna Armjanskoj SSR. Izvestiya Akademii Nauk Armyanskai SSR, 6, 1 - 226." type="journal article" year="1967">Khnzorian 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE24B6F4C3BFFE4FEC3B1E7FD336BF7" author="Kalashian" box="[347,689,908,933]" pageId="4" pageNumber="171" refString="Kalashian, M. & Karagyan, G. (2010) Pharaonus caucasicus (Reitter, 1893). In: Aghasyan, A. L. & Kalashyan, M. J. (Eds.), Red Data Book of Armenia. Invertebrate and Vertebrate Animals. Ministry of Nature Protection, Yerevan, Armenia, 81 pp. Available from: http: // www. mnp. am / red _ book _ fauna / index. htm (accessed 14 April 2015)" type="book" year="2010">Kalashian and Karagyan 2010</bibRefCitation>
|
||
; T. Ghrejyan [Gavar,
|
||
<collectingCountry id="F364760E4C3BFFE4FC3BB1E7FB896BF6" box="[931,1035,908,933]" name="Armenia" pageId="4" pageNumber="171">Armenia</collectingCountry>
|
||
], personal communication).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |