220 lines
23 KiB
XML
220 lines
23 KiB
XML
<document id="ED254576601C8D417EB35EAF838BD758" ID-DOI="10.11646/zootaxa.4012.1.9" ID-GBIF-Dataset="4c0d8376-71b9-42d1-8dbc-08e0ff16338a" ID-ISSN="1175-5326" ID-Zenodo-Dep="244632" ID-ZooBank="55822610-6FE6-4218-A412-A080F3218F51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461239579310" checkinUser="plazi" docAuthor="Keith, Denis, Sabatinelli, Guido & Uliana, Marco" docDate="2015" docId="03DA87884C34FFEBFF0FB2F3FD126C5A" docLanguage="en" docName="zt04012p180.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pharaonus (Tamerlanius) lederi Reitter 1888" docType="treatment" docVersion="8" lastPageNumber="178" masterDocId="FFE3FFF04C3FFFE0FF98B26BFF826853" masterDocTitle="Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa" masterLastPageNumber="180" masterPageNumber="167" pageNumber="178" updateTime="1698605019073" updateUser="plazi">
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<mods:titleInfo id="7A0A62B27942D0BC2BB51A396D52D07E">
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<mods:title id="D39A0C4BCEF776D42254E1F3767AA44C">Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa</mods:title>
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<mods:name id="857B80B025AA9A1864E1D3AD20375056" type="personal">
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<mods:role id="2564B21ED9884E5671BAFDDEE7F22859">
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<mods:roleTerm id="74A01DCBDE790724C734E548A3B2DC3B">Author</mods:roleTerm>
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<mods:namePart id="AD753FD67A087EA898F5EB026DEFBF77">Keith, Denis</mods:namePart>
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<mods:namePart id="F9D6C31EDDEAB3F3BD87A06EFE775D2E">Sabatinelli, Guido</mods:namePart>
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</mods:name>
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<mods:roleTerm id="5F13D0E9DC687D28DF7F1D3F2D34DF92">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2AC1AF84A5EF7189E53EE817D519DB75">Uliana, Marco</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2A42D8068E7AFBB9218F1AA649B7F4A3">text</mods:typeOfResource>
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<mods:title id="96139040FD23A5B09F9E0DF93E051B0F">Zootaxa</mods:title>
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<mods:part id="C8F2059937336996459D5317F64D88C4">
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<mods:date id="A4D8932A96E6582A814486E8E494DD0A">2015</mods:date>
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<mods:detail id="4E019BDDB65E6BD3B91EBBEB775815C0" type="volume">
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<mods:number id="1CEDE6835CC958112A88287220A53294">4012</mods:number>
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</mods:detail>
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<mods:detail id="D2674BB3C517999864656B138C3CD55A" type="issue">
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<mods:number id="91537889637F39D6FD319F0D0D56A676">1</mods:number>
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<mods:classification id="F081AE3BDF0C5109913AC53AE6ADFAEC">journal article</mods:classification>
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<mods:identifier id="DDF5865F8B5044A7E9682EB22D5FF366" type="DOI">10.11646/zootaxa.4012.1.9</mods:identifier>
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<treatment id="03DA87884C34FFEBFF0FB2F3FD126C5A" ID-DOI="http://doi.org/10.5281/zenodo.5694714" ID-GBIF-Taxon="119632627" ID-Zenodo-Dep="5694714" LSID="urn:lsid:plazi:treatment:03DA87884C34FFEBFF0FB2F3FD126C5A" httpUri="http://treatment.plazi.org/id/03DA87884C34FFEBFF0FB2F3FD126C5A" lastPageNumber="178" pageId="11" pageNumber="178">
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<subSubSection id="C36965154C34FFEBFF0FB2F3FEAF6880" pageId="11" pageNumber="178" type="nomenclature">
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<paragraph id="8BCC369E4C34FFEBFF0FB2F3FD5668E1" blockId="11.[151,724,152,211]" box="[151,724,152,178]" pageId="11" pageNumber="178">
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<heading id="D08481F24C34FFEBFF0FB2F3FD5668E1" bold="true" box="[151,724,152,178]" fontSize="11" level="1" pageId="11" pageNumber="178" reason="1">
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||
<taxonomicName id="4C734D1D4C34FFEBFF0FB2F3FD5668E1" authority="Reitter, 1888" authorityName="Reitter" authorityYear="1888" box="[151,724,152,178]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="species" species="lederi" subGenus="Tamerlanius">
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||
<emphasis id="B907EA8C4C34FFEBFF0FB2F3FD5668E1" bold="true" box="[151,724,152,178]" pageId="11" pageNumber="178">
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<emphasis id="B907EA8C4C34FFEBFF0FB2F3FE9F68E1" bold="true" box="[151,285,152,178]" italics="true" pageId="11" pageNumber="178">Pharaonus</emphasis>
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||
(
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<emphasis id="B907EA8C4C34FFEBFEB5B2F3FE4768E1" bold="true" box="[301,453,152,178]" italics="true" pageId="11" pageNumber="178">Tamerlanius</emphasis>
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||
)
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<emphasis id="B907EA8C4C34FFEBFE4EB2F3FD9868E1" bold="true" box="[470,538,152,178]" italics="true" pageId="11" pageNumber="178">lederi</emphasis>
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||
(
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<bibRefCitation id="EFE24B6F4C34FFEBFDB2B2F3FD4E68E1" author="Reitter" box="[554,716,152,178]" pageId="11" pageNumber="178" refString="Reitter, E. (1888) Ubersicht der Arten der Coleopteren-Gattung Phyllopertha Kirby aus Europa, den angrenzenden Landern, Sibirien mit Central-Asien. Entomologische Nachrichten, 14, 289 - 294." type="journal article" year="1888">Reitter, 1888</bibRefCitation>
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)
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BCC369E4C34FFEBFF0FB2D1FEAF6880" blockId="11.[151,724,152,211]" box="[151,301,186,211]" pageId="11" pageNumber="178">
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||
(
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<figureCitation id="13482A1B4C34FFEBFF07B2D1FF796880" box="[159,251,186,211]" captionStart="FIGURES 11 – 12" captionStartId="6.[151,264,1129,1151]" captionTargetBox="[159,1427,207,1105]" captionTargetId="figure@6.[150,1437,195,1118]" captionTargetPageId="6" captionText="FIGURES 11 – 12. Habitus of Pharaonus (Tamerlanius) species. 11: P. l e de r i, male (Koron 15 km Ekoc. Dzejran, Bukhara, Uzbekistan). 12: P. semenowi, male (Katta-kum, Turkmenkuduk, Uzbekistan)." httpUri="https://zenodo.org/record/244636/files/figure.png" pageId="11" pageNumber="178">Figs. 11</figureCitation>
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,
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<figureCitation id="13482A1B4C34FFEBFE9FB2D0FEA66880" box="[263,292,187,211]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="11" pageNumber="178">22</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36965154C34FFEBFF0FB368FC93694B" box="[151,785,258,280]" pageId="11" pageNumber="178" type="reference_group">
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||
<paragraph id="8BCC369E4C34FFEBFF0FB368FC93694B" blockId="11.[151,785,258,280]" box="[151,785,258,280]" pageId="11" pageNumber="178">
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<emphasis id="B907EA8C4C34FFEBFF0FB368FF2A694B" box="[151,168,259,280]" italics="true" pageId="11" pageNumber="178">=</emphasis>
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||
<taxonomicName id="4C734D1D4C34FFEBFF36B369FDD9694B" authority="Ohaus, 1941" authorityName="Ohaus" authorityYear="1941" box="[174,603,258,280]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subSpecies" species="lederi" subSpecies="turcmenius">
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<emphasis id="B907EA8C4C34FFEBFF36B369FE56694B" box="[174,468,258,280]" italics="true" pageId="11" pageNumber="178">Pharaonus lederi turcmenius</emphasis>
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<bibRefCitation id="EFE24B6F4C34FFEBFE43B369FDD9694B" author="Ohaus" box="[475,603,258,280]" pageId="11" pageNumber="178" refString="Ohaus, F. (1941) Neue palaearktische Rutelinen. Entomologische Blatter, 37, 111 - 115." type="journal article" year="1941">Ohaus, 1941</bibRefCitation>
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</taxonomicName>
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<emphasis id="B907EA8C4C34FFEBFDF9B369FC93694B" bold="true" box="[609,785,258,280]" pageId="11" pageNumber="178">(new synonymy)</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36965154C34FFEBFF0FB32EFAB26A42" pageId="11" pageNumber="178" type="materials_examined">
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||
<paragraph id="8BCC369E4C34FFEBFF0FB32EFB0A69F5" blockId="11.[151,1437,325,1034]" pageId="11" pageNumber="178">
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||
<emphasis id="B907EA8C4C34FFEBFF0FB32EFE07690D" bold="true" box="[151,389,325,350]" pageId="11" pageNumber="178">Examined material.</emphasis>
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||
<typeStatus id="54C8883C4C34FFEBFE15B32DFE3F690D" box="[397,445,326,350]" pageId="11" pageNumber="178">type</typeStatus>
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specimens:
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<taxonomicName id="4C734D1D4C34FFEBFDD3B32EFC03690D" authority="Reitter" authorityName="Reitter" box="[587,897,325,350]" class="Insecta" family="Rutelidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="species" species="lederi">
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<emphasis id="B907EA8C4C34FFEBFDD3B32EFCA5690D" box="[587,807,325,350]" italics="true" pageId="11" pageNumber="178">Phyllopertha lederi</emphasis>
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||
Reitter
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||
</taxonomicName>
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: Turkmenia, Askabat [= Ashgabat], Leder:
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||
<specimenCount id="9D75FD174C34FFEBFAE8B32DFA1E690E" box="[1392,1436,326,350]" pageId="11" pageNumber="178" type="generic">1 ♂</specimenCount>
|
||
(
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||
<typeStatus id="54C8883C4C34FFEBFF07B302FE8569D1" box="[159,263,361,386]" pageId="11" pageNumber="178" type="holotype">holotype</typeStatus>
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||
); same data of
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<typeStatus id="54C8883C4C34FFEBFE2CB302FD9869D1" box="[436,538,361,386]" pageId="11" pageNumber="178" type="holotype">holotype</typeStatus>
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||
:
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||
<specimenCount id="9D75FD174C34FFEBFDBEB302FDD169D2" box="[550,595,361,385]" pageId="11" pageNumber="178" type="generic">2 ♂</specimenCount>
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||
, 2 ♀ (
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||
<typeStatus id="54C8883C4C34FFEBFD04B302FC8C69D2" box="[668,782,361,385]" pageId="11" pageNumber="178" type="paratype">paratypes</typeStatus>
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||
); Turkmenia, Leder:
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<specimenCount id="9D75FD174C34FFEBFC64B302FBAA69D2" box="[1020,1064,361,385]" pageId="11" pageNumber="178" type="generic">1 ♂</specimenCount>
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(
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<typeStatus id="54C8883C4C34FFEBFBA0B302FB1D69D2" box="[1080,1183,361,385]" pageId="11" pageNumber="178" type="paratype">paratype</typeStatus>
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||
). All from
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<collectionCode id="ED62AE5B4C34FFEBFA85B301FAF769D2" LSID="urn:lsid:biocol.org:col:34991" box="[1309,1397,362,385]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="11" pageNumber="178">HNHM</collectionCode>
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, ex coll. Reitter.
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<taxonomicName id="4C734D1D4C34FFEBFEBFB3E6FD4269F5" authority="Ohaus" authorityName="Ohaus" box="[295,704,397,422]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subSpecies" species="lederi" subSpecies="turcmenius">
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<emphasis id="B907EA8C4C34FFEBFEBFB3E6FDEC69F6" box="[295,622,397,422]" italics="true" pageId="11" pageNumber="178">Pharaonus lederi turcmenius</emphasis>
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Ohaus
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</taxonomicName>
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: Turkmen, Merw,
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<specimenCount id="9D75FD174C34FFEBFC15B3E5FC3B69F6" box="[909,953,398,422]" pageId="11" pageNumber="178" type="generic">1 ♂</specimenCount>
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||
,
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||
<typeStatus id="54C8883C4C34FFEBFC5DB3E6FBA569F5" box="[965,1063,397,422]" pageId="11" pageNumber="178" type="holotype">holotype</typeStatus>
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||
(MNB).
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||
</paragraph>
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<paragraph id="8BCC369E4C34FFEBFF0FB3DAFAB26A42" blockId="11.[151,1437,325,1034]" pageId="11" pageNumber="178">
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Other specimens:
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<collectingCountry id="F364760E4C34FFEBFEFAB3DAFE616999" box="[354,483,433,458]" name="Uzbekistan" pageId="11" pageNumber="178">Uzbekistan</collectingCountry>
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||
, Buchara, Koron 15 Km Ekoc. Dzejran,
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<date id="FFCD105E4C34FFEBFC36B3DAFB85699A" box="[942,1031,433,457]" pageId="11" pageNumber="178" value="1992-05-23">23.V.92</date>
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||
, Snizek leg.,
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<specimenCount id="9D75FD174C34FFEBFB06B3DAFAB56999" box="[1182,1335,433,458]" pageId="11" pageNumber="178" type="generic">16 specimens</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C34FFEBFADEB3DAFA1A699A" box="[1350,1432,433,457]" pageId="11" pageNumber="178">DKCC</collectionCode>
|
||
,
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||
<collectionCode id="ED62AE5B4C34FFEBFF0FB3BDFF6D69BD" box="[151,239,470,494]" httpUri="http://grbio.org/cool/vphy-ev73" name="Mie University Culture Collection (Culture Collection, Laboratory of Plant Pathology)" pageId="11" pageNumber="178">MUCC</collectionCode>
|
||
); Buchara,
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||
<specimenCount id="9D75FD174C34FFEBFEE1B3BDFE7D69BD" box="[377,511,469,494]" pageId="11" pageNumber="178" type="generic">1 specimen</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C34FFEBFD8CB3BDFDF269BD" LSID="urn:lsid:biocol.org:col:34988" box="[532,624,470,494]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="178">MNHN</collectionCode>
|
||
); Buchara, Repetek, 5.1900, Coll. Hauser, 3 ex (
|
||
<collectionCode id="ED62AE5B4C34FFEBFB26B3BDFA9469BD" box="[1214,1302,470,494]" httpUri="http://grbio.org/cool/vphy-ev73" name="Mie University Culture Collection (Culture Collection, Laboratory of Plant Pathology)" pageId="11" pageNumber="178">MUCC</collectionCode>
|
||
), Bucharia, Repetek (
|
||
<collectionCode id="ED62AE5B4C34FFEBFE99B392FED96A42" box="[257,347,505,529]" pageId="11" pageNumber="178">MSNM</collectionCode>
|
||
, ex. coll. Turati):
|
||
<specimenCount id="9D75FD174C34FFEBFDBAB392FDCD6A42" box="[546,591,505,529]" pageId="11" pageNumber="178" type="generic">1 ♂</specimenCount>
|
||
; [
|
||
<collectingCountry id="F364760E4C34FFEBFDF9B392FC806A41" box="[609,770,505,530]" name="Turkmenistan" pageId="11" pageNumber="178">Turkmenistan</collectingCountry>
|
||
] Transcaspien, Tedschen [=Tejen]:
|
||
<specimenCount id="9D75FD174C34FFEBFB17B392FB386A42" box="[1167,1210,505,529]" pageId="11" pageNumber="178" type="generic">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE5B4C34FFEBFB50B392FAA66A42" box="[1224,1316,505,529]" pageId="11" pageNumber="178">MSNM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C34FFEBFF5FB076FBBB6A65" box="[199,1081,541,566]" pageId="11" pageNumber="178" type="distribution">
|
||
<paragraph id="8BCC369E4C34FFEBFF5FB076FBBB6A65" blockId="11.[151,1437,325,1034]" box="[199,1081,541,566]" pageId="11" pageNumber="178">
|
||
<emphasis id="B907EA8C4C34FFEBFF5FB076FEE26A65" bold="true" box="[199,352,541,566]" pageId="11" pageNumber="178">Distribution.</emphasis>
|
||
<collectingCountry id="F364760E4C34FFEBFEFFB076FE716A65" box="[359,499,541,566]" name="Afghanistan" pageId="11" pageNumber="178">Afghanistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C34FFEBFE66B076FD066A65" box="[510,644,541,566]" name="Kazakhstan" pageId="11" pageNumber="178">Kazakhstan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C34FFEBFD16B076FD7D6A65" box="[654,767,541,566]" name="Tajikistan" pageId="11" pageNumber="178">Tajikistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C34FFEBFC93B076FC2B6A65" box="[779,937,541,566]" name="Turkmenistan" pageId="11" pageNumber="178">Turkmenistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C34FFEBFC2BB076FBB76A65" box="[947,1077,541,566]" name="Uzbekistan" pageId="11" pageNumber="178">Uzbekistan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36965154C34FFEBFF5FB02AFD126C5A" pageId="11" pageNumber="178" type="discussion">
|
||
<paragraph id="8BCC369E4C34FFEBFF5FB02AFD126C5A" blockId="11.[151,1437,325,1034]" pageId="11" pageNumber="178">
|
||
<emphasis id="B907EA8C4C34FFEBFF5FB02AFEB96A09" bold="true" box="[199,315,577,602]" pageId="11" pageNumber="178">Remarks.</emphasis>
|
||
The subspecies
|
||
<taxonomicName id="4C734D1D4C34FFEBFE6DB029FD546A0A" box="[501,726,577,602]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subSpecies" species="lederi" subSpecies="turcmenius">
|
||
<emphasis id="B907EA8C4C34FFEBFE6DB029FD816A0A" box="[501,515,578,601]" italics="true" pageId="11" pageNumber="178">P</emphasis>
|
||
.
|
||
<emphasis id="B907EA8C4C34FFEBFD89B02AFD546A0A" box="[529,726,577,602]" italics="true" pageId="11" pageNumber="178">lederi turcmenius</emphasis>
|
||
</taxonomicName>
|
||
(cited with the incorrect spelling
|
||
<taxonomicName id="4C734D1D4C34FFEBFBCBB029FAC36A0A" box="[1107,1345,577,602]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subSpecies" species="lederi" subSpecies="turcmenicus">
|
||
<emphasis id="B907EA8C4C34FFEBFBCBB029FBE36A0A" box="[1107,1121,578,601]" italics="true" pageId="11" pageNumber="178">P</emphasis>
|
||
.
|
||
<emphasis id="B907EA8C4C34FFEBFBE8B02AFAC36A0A" box="[1136,1345,577,602]" italics="true" pageId="11" pageNumber="178">lederi turcmenicus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFE24B6F4C34FFEBFAFDB029FF536A2D" author="Zorn" pageId="11" pageNumber="178" refString="Zorn, C. (2006) Scarabaeidae: Rutelinae: Anomalini. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Apollo Books, Stenstrup, pp. 251 - 276." type="book" year="2006">Zorn 2006</bibRefCitation>
|
||
) was described by
|
||
<bibRefCitation id="EFE24B6F4C34FFEBFE21B00EFDD56A2D" author="Ohaus" box="[441,599,613,638]" pageId="11" pageNumber="178" refString="Ohaus, F. (1941) Neue palaearktische Rutelinen. Entomologische Blatter, 37, 111 - 115." type="journal article" year="1941">Ohaus (1941)</bibRefCitation>
|
||
based on a single male from Merw (
|
||
<collectingCountry id="F364760E4C34FFEBFB8EB00EFB3A6A2D" box="[1046,1208,613,638]" name="Turkmenistan" pageId="11" pageNumber="178">Turkmenistan</collectingCountry>
|
||
). The status of this subspecies was never reassessed: it was apparently overlooked in the comprehensive works by
|
||
<bibRefCitation id="EFE24B6F4C34FFEBFB4BB0E2FA1E6AF1" author="Medvedev" box="[1235,1436,649,674]" pageId="11" pageNumber="178" refString="Medvedev, S. I. (1949) Fauna USSR: Coleoptera 10 Nr. 3: Lamellicornia (Scarabaeidae), subfamily Rutelinae. Izdatel'stvo Akademii Nauk SSSR, Moscow, Russia, pp. 1 - 371." type="journal article" year="1949">Medvedev (1949)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFE24B6F4C34FFEBFF5FB0C6FE086A95" author="Nikolajev" box="[199,394,685,710]" pageId="11" pageNumber="178" refString="Nikolajev, G. V. (1987) Scarabaeoidea from Kazakhstan and Central Asia. Academy of Sciences of Kazakhstan, Alma Ata, Kazakhstan, 232 pp." type="book" year="1987">Nikolajev (1987)</bibRefCitation>
|
||
, and remained unknown to
|
||
<bibRefCitation id="EFE24B6F4C34FFEBFD5BB0C6FCC86A95" author="Kral" box="[707,842,685,710]" pageId="11" pageNumber="178" refString="Kral, D. (2012) Pharaonus saharicus sp. nov. (Coleoptera: Scarabaeidae: Rutelinae) from the Central Sahara (Algeria). Klapalekiana, 48, 127 - 136." type="journal article" year="2012">Král (2012)</bibRefCitation>
|
||
. After the examination of both the
|
||
<typeStatus id="54C8883C4C34FFEBFB4EB0C5FA846A95" box="[1238,1286,686,710]" pageId="11" pageNumber="178">type</typeStatus>
|
||
series and of additional specimens, we could not find significant differences between
|
||
<taxonomicName id="4C734D1D4C34FFEBFC25B0B9FB196ABA" box="[957,1179,721,746]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subSpecies" species="lederi" subSpecies="turcmenius">
|
||
<emphasis id="B907EA8C4C34FFEBFC25B0B9FB196ABA" box="[957,1179,721,746]" italics="true" pageId="11" pageNumber="178">P. lederi turcmenius</emphasis>
|
||
</taxonomicName>
|
||
and the nominotypical subspecies
|
||
<taxonomicName id="4C734D1D4C34FFEBFE8DB09DFE3A6B5D" box="[277,440,757,782]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subSpecies" species="lederi" subSpecies="lederi">
|
||
<emphasis id="B907EA8C4C34FFEBFE8DB09DFE3A6B5D" box="[277,440,757,782]" italics="true" pageId="11" pageNumber="178">P. lederi lederi</emphasis>
|
||
</taxonomicName>
|
||
: therefore, we consider them synonyms. The differences mentioned by
|
||
<bibRefCitation id="EFE24B6F4C34FFEBFB4EB09EFAEC6B5D" author="Ohaus" box="[1238,1390,757,782]" pageId="11" pageNumber="178" refString="Ohaus, F. (1941) Neue palaearktische Rutelinen. Entomologische Blatter, 37, 111 - 115." type="journal article" year="1941">Ohaus (1941)</bibRefCitation>
|
||
can be confidently referred either to the poor condition of the specimen (almost complete absence of setae from the pronotum of
|
||
<taxonomicName id="4C734D1D4C34FFEBFEB2B155FD886B06" box="[298,522,829,854]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="subSpecies" species="lederi" subSpecies="turcmenius">
|
||
<emphasis id="B907EA8C4C34FFEBFEB2B155FD886B06" box="[298,522,829,854]" italics="true" pageId="11" pageNumber="178">P. lederi turcmenius</emphasis>
|
||
</taxonomicName>
|
||
), or to intraspecific variation (density of punctures on the pronotum, color of the legs). With reference to the latter character, we could verify that the color of the legs varies among specimens coming from the same area. Within the
|
||
<typeStatus id="54C8883C4C34FFEBFDFEB1EDFD146BCD" box="[614,662,902,926]" pageId="11" pageNumber="178">type</typeStatus>
|
||
series of
|
||
<emphasis id="B907EA8C4C34FFEBFC93B1EDFCE66BCE" box="[779,868,901,926]" italics="true" pageId="11" pageNumber="178">P. l e de r i</emphasis>
|
||
, one female from Ashgabat has dark femora but reddish-brown tibiae and tarsi, while the legs of the remaining specimens are uniformly dark brown. We also observed that the legs (including femora) are blackish brown to reddish brown (including femora, see
|
||
<figureCitation id="13482A1B4C34FFEBFAD9B1A6FA116BB5" box="[1345,1427,973,998]" captionStart="FIGURES 11 – 12" captionStartId="6.[151,264,1129,1151]" captionTargetBox="[159,1427,207,1105]" captionTargetId="figure@6.[150,1437,195,1118]" captionTargetPageId="6" captionText="FIGURES 11 – 12. Habitus of Pharaonus (Tamerlanius) species. 11: P. l e de r i, male (Koron 15 km Ekoc. Dzejran, Bukhara, Uzbekistan). 12: P. semenowi, male (Katta-kum, Turkmenkuduk, Uzbekistan)." httpUri="https://zenodo.org/record/244636/files/figure.png" pageId="11" pageNumber="178">Fig. 11</figureCitation>
|
||
) among the specimens from the Buchara area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |