368 lines
48 KiB
XML
368 lines
48 KiB
XML
<document id="8DF09B253B895872E67A9B0BF1FDF5B6" ID-CLB-Dataset="182445" ID-DOI="10.11646/zootaxa.5169.5.5" ID-GBIF-Dataset="ba51e6d4-299a-45d5-bf7c-6df3b8f967f5" ID-ISSN="1175-5326" ID-Zenodo-Dep="6952676" ID-ZooBank="77339EEB-28FC-4D5A-AB22-9AACA4DF3050" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659440969926" checkinUser="plazi" docAuthor="Czaja, Alexander, Covich, Alan P., Neubauer, Thomas A., Estrada-Rodríguez, José Luis, Ávila-Rodríguez, Verónica & Mata, Jorge Sáenz" docDate="2022" docId="03C4F212A303FF81A6ADFA8D9267D628" docLanguage="en" docName="zootaxa.5169.5.5.pdf" docOrigin="Zootaxa 5169 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spinopyrgus luismaedai Czaja, Covich, Neubauer & Estrada-Rodriguez 2022, n. gen. et n. sp." docType="treatment" docVersion="7" lastPageNumber="476" masterDocId="FFFD8A6AA301FF85A63AFFBD9050D17C" masterDocTitle="A new freshwater snail genus and species (Gastropoda: Caenogastropoda, Cochliopidae) with extremely spinous shells from sub-recent spring deposits in northeastern Mexico" masterLastPageNumber="480" masterPageNumber="472" pageNumber="474" updateTime="1699363345607" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0CD3E73FDE3B724284BC053136A1D073">A new freshwater snail genus and species (Gastropoda: Caenogastropoda, Cochliopidae) with extremely spinous shells from sub-recent spring deposits in northeastern Mexico</mods:title>
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<mods:namePart id="B9E1461C8C2F7DF202124B36AD8951A4">Czaja, Alexander</mods:namePart>
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<mods:affiliation id="D54458EB00724F6C040860AE711E074A">drczaja @ ujed. mx; https: // orcid. org / 0000 - 0001 - 9157 - 4509</mods:affiliation>
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<mods:nameIdentifier id="A214EE510B0A099D4B63407ACBA1FFA4" type="email">drczaja@ujed.mx</mods:nameIdentifier>
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<mods:namePart id="3B31A19EFDD67B24ED41B4593F7B725F">Covich, Alan P.</mods:namePart>
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<mods:namePart id="3E3A6B03D77623E95763EBF73B7D1742">Neubauer, Thomas A.</mods:namePart>
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<mods:affiliation id="7C49CFA37EEBF06C0C9E6D024D173271">Department of Animal Ecology and Systematics, Justus Liebig University, 35392 Giessen, Germany & Naturalis Biodiversity Center, 2300 RA Leiden, The Netherlands & SNSB-Bavarian State Collection for Paleontology and Geology, 80333 Munich, Germany.</mods:affiliation>
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<mods:namePart id="47FE2542DD907B14A9A833BA11B7A99F">Estrada-Rodríguez, José Luis</mods:namePart>
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<mods:namePart id="DD6333B95F891FDF5D20C909F8161915">Ávila-Rodríguez, Verónica</mods:namePart>
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<mods:affiliation id="BAACEC8DB7FB2E873EE1C605054AA842">vavilar @ gmail. com; https: // orcid. org / 0000 - 0002 - 2999 - 5764</mods:affiliation>
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<mods:nameIdentifier id="6F3910AC58D5B38B39EA041B12F0C5C6" type="email">vavilar@gmail.com</mods:nameIdentifier>
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<mods:namePart id="81B5A49EC142C01C7D8D382ABA5E6EF8">Mata, Jorge Sáenz</mods:namePart>
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<mods:affiliation id="6DA3B67E0B214E12CD5DECF769A0EF59">jsaenz _ mata @ ujed. mx; https: // orcid. org / 0000 - 0001 - 9958 - 9676</mods:affiliation>
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<mods:nameIdentifier id="1F9CFAE10CC1F45BEC30EBBD1DA69308" type="email">jsaenz_mata@ujed.mx</mods:nameIdentifier>
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<mods:title id="9F6EF814BF676C1C79A370E774DD86F3">Zootaxa</mods:title>
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<mods:part id="BA22D002846DC4B2333B268AEDCD963F">
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<mods:date id="2C4BF527A04C372DB0205242314E80D5">2022</mods:date>
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<mods:number id="7111E693BB6DD402FF03C7FC4A9339DA">2022-08-02</mods:number>
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<mods:number id="FF4B8A304C55EA890E40322DF0819ED4">5169</mods:number>
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<treatment id="03C4F212A303FF81A6ADFA8D9267D628" ID-DOI="http://doi.org/10.5281/zenodo.7305277" ID-GBIF-Taxon="204252355" ID-Zenodo-Dep="7305277" LSID="urn:lsid:plazi:treatment:03C4F212A303FF81A6ADFA8D9267D628" httpUri="http://treatment.plazi.org/id/03C4F212A303FF81A6ADFA8D9267D628" lastPageId="4" lastPageNumber="476" pageId="2" pageNumber="474">
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<heading id="D09AF468A303FF87A6ADFA8D9484D436" bold="true" box="[151,1236,1327,1354]" fontSize="11" level="1" pageId="2" pageNumber="474" reason="1">
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<taxonomicName id="4C6D3887A303FF87A6ADFA8D9443D436" ID-CoL="9GJFD" authority="Czaja, Covich, Neubauer & Estrada-Rodriguez" authorityName="Czaja, Covich, Neubauer & Estrada-Rodriguez" authorityYear="2022" box="[151,1043,1327,1354]" class="Gastropoda" family="Cochliopidae" genus="Spinopyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="474" phylum="Mollusca" rank="species" species="luismaedai" status="n. gen. et n. sp.">
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<emphasis id="B9199F16A303FF87A6ADFA8D91EAD436" bold="true" box="[151,442,1327,1354]" italics="true" pageId="2" pageNumber="474">Spinopyrgus luismaedai</emphasis>
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Czaja, Covich, Neubauer & Estrada-Rodríguez
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</taxonomicName>
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<taxonomicNameLabel id="A22A226DA303FF87A223FA8D9484D436" box="[1049,1236,1328,1354]" pageId="2" pageNumber="474" rank="species">n. gen. et n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C377108FA303FF87A6ADFAE99554D4EF" pageId="2" pageNumber="474" type="description">
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<paragraph id="8BD24304A303FF87A6ADFAE99136D412" blockId="2.[151,1284,1327,1427]" box="[151,358,1364,1391]" pageId="2" pageNumber="474">
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(
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<figureCitation id="13565F81A303FF87A6A5FAE9914CD412" box="[159,284,1364,1390]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="2" pageNumber="474">Figures 2A</figureCitation>
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to 3P)
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</paragraph>
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<paragraph id="8BD24304A303FF87A6ADFAC59554D4EF" blockId="2.[151,1284,1327,1427]" box="[151,1284,1400,1427]" pageId="2" pageNumber="474">
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ZooBank registration number:
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<uri id="FFFC4F06A303FF87A7CBFAC59554D4EF" box="[497,1284,1400,1427]" pageId="2" pageNumber="474">
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urn:lsid:zoobank.org:act:
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<uuid id="FFCB79D1A303FF87A53FFAC59554D4EF" box="[773,1284,1400,1427]" pageId="2" pageNumber="474">385187F6-986B-4200-84D1-91147F22F4D1</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C377108FA303FF87A6ADFA7D9246D482" pageId="2" pageNumber="474" type="etymology">
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<paragraph id="8BD24304A303FF87A6ADFA7D9246D482" blockId="2.[151,1437,1472,2039]" pageId="2" pageNumber="474">
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<emphasis id="B9199F16A303FF87A6ADFA7D914ED4A6" bold="true" box="[151,286,1472,1498]" pageId="2" pageNumber="474">Etymology.</emphasis>
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The new species is named in honor of Dr. Luis Maeda Villalobos, Torreón,
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<collectingRegion id="49A98DE6A303FF87A250FA7D9482D4A6" box="[1130,1234,1472,1498]" country="Mexico" name="Coahuila de Zaragoza" pageId="2" pageNumber="474">Coahuila</collectingRegion>
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, who was the first collector of fossil snails at Viesca.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C377108FA303FF87A6FDF9B59138D6F6" pageId="2" pageNumber="474" type="materials_examined">
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<paragraph id="8BD24304A303FF87A6FDF9B5929CD75E" blockId="2.[151,1437,1472,2039]" box="[199,716,1544,1571]" pageId="2" pageNumber="474">
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<emphasis id="B9199F16A303FF87A6FDF9B5916BD75E" bold="true" box="[199,315,1544,1570]" pageId="2" pageNumber="474">
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<typeStatus id="54D6FDA6A303FF87A6FDF9B59166D75E" box="[199,310,1544,1570]" pageId="2" pageNumber="474" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Specimen UJMC-250 (
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<figureCitation id="13565F81A303FF87A47FF9B592EED75E" box="[581,702,1544,1570]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="2" pageNumber="474">Fig. 2A, B</figureCitation>
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).
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||
</paragraph>
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||
<paragraph id="8BD24304A303FF87A6FDF99194D7D716" blockId="2.[151,1437,1472,2039]" pageId="2" pageNumber="474">
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<emphasis id="B9199F16A303FF87A6FDF9919136D73A" bold="true" box="[199,358,1580,1606]" pageId="2" pageNumber="474">Type locality.</emphasis>
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Superficial sediments in the desiccated Juan Guerra spring, ca.
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<quantity id="4C95EEE1A303FF87A20FF991943ED73A" box="[1077,1134,1580,1606]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="474" unit="km" value="1.0">1 km</quantity>
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south of Viesca,
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||
<collectingRegion id="49A98DE6A303FF87A30AF99195C8D73A" box="[1328,1432,1580,1606]" country="Mexico" name="Coahuila de Zaragoza" pageId="2" pageNumber="474">Coahuila</collectingRegion>
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,
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<collectingCountry id="F37A0394A303FF87A6ADF9ED90A0D716" box="[151,240,1616,1642]" name="Mexico" pageId="2" pageNumber="474">Mexico</collectingCountry>
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. Collected by Alexander Czaja and José Luis Estrada-Rodríguez, 2014 (
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<figureCitation id="13565F81A303FF87A22DF9ED9426D716" box="[1047,1142,1616,1642]" captionStart="FIGURE 1" captionStartId="1.[151,250,1865,1889]" captionTargetBox="[153,1435,852,1840]" captionTargetId="figure-285@1.[151,1436,848,1841]" captionTargetPageId="1" captionText="FIGURE 1. Study area showing the locations of the Viesca springs. The scale bar refers to the map with the springs marked. The topography is illustrated using HillShade in ArcGIS." figureDoi="http://doi.org/10.5281/zenodo.6952678" httpUri="https://zenodo.org/record/6952678/files/figure.png" pageId="2" pageNumber="474">Figure 1</figureCitation>
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).
|
||
</paragraph>
|
||
<paragraph id="8BD24304A303FF87A6FDF9C99151D7AA" blockId="2.[151,1437,1472,2039]" pageId="2" pageNumber="474">
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<emphasis id="B9199F16A303FF87A6FDF9C99126D7F2" bold="true" box="[199,374,1652,1678]" pageId="2" pageNumber="474">Type material.</emphasis>
|
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<materialsCitation id="3B054959A303FF87A745F9C993CAD7CF" ID-GBIF-Occurrence="3962323302" collectionCode="UJMC" latitude="25.32139" longLatPrecision="21" longitude="-102.818886" pageId="2" pageNumber="474" specimenCode="UJMC 251, UJMC 252" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54D6FDA6A303FF87A745F9C991B7D7F2" box="[383,487,1652,1678]" pageId="2" pageNumber="474" type="holotype">Holotype</typeStatus>
|
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(
|
||
<figureCitation id="13565F81A303FF87A7CDF9C99224D7F2" box="[503,628,1652,1678]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="2" pageNumber="474">Fig. 2A, B</figureCitation>
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), UJMC-250 from Juan Guerra spring, Viesca,
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<geoCoordinate id="EE5925C3A303FF87A2AAF9C9957ED7F2" box="[1168,1326,1652,1679]" degrees="25" direction="north" minutes="19" orientation="latitude" pageId="2" pageNumber="474" precision="15" seconds="17" value="25.32139">25° 19’ 17’’N</geoCoordinate>
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,
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<geoCoordinate id="EE5925C3A303FF87A301F9C9908CD7CF" degrees="102" direction="west" minutes="49" orientation="longitude" pageId="2" pageNumber="474" precision="15" seconds="08" value="-102.818886">102° 49’ 08’’W</geoCoordinate>
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,
|
||
<typeStatus id="54D6FDA6A303FF87A6D0F9249105D7CF" box="[234,341,1689,1715]" pageId="2" pageNumber="474" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<figureCitation id="13565F81A303FF87A75FF92591B6D7CE" box="[357,486,1688,1714]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="2" pageNumber="474">Figs. 2C–F</figureCitation>
|
||
),
|
||
<specimenCode id="DBCBEB7FA303FF87A7C0F9259228D7CF" box="[506,632,1688,1715]" collectionCode="UJMC" pageId="2" pageNumber="474">UJMC 251</specimenCode>
|
||
,
|
||
<specimenCode id="DBCBEB7FA303FF87A4BEF9259353D7CF" box="[644,771,1688,1715]" collectionCode="UJMC" pageId="2" pageNumber="474">UJMC 252</specimenCode>
|
||
,
|
||
<specimenCode id="DBCBEB7FA303FF87A534F92593DDD7CF" box="[782,909,1688,1715]" collectionCode="UJMC" pageId="2" pageNumber="474">UJMC 252</specimenCode>
|
||
a
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="54D6FDA6A303FF87A59FF9249442D7CF" box="[933,1042,1689,1715]" pageId="2" pageNumber="474" type="paratype">Paratypes</typeStatus>
|
||
1, 2 and 3 from the same lot as the
|
||
<typeStatus id="54D6FDA6A303FF87A6ADF90190ADD7AA" box="[151,253,1724,1750]" pageId="2" pageNumber="474" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD24304A303FF87A6FDF95D923CD786" blockId="2.[151,1437,1472,2039]" box="[199,620,1760,1786]" pageId="2" pageNumber="474">
|
||
<emphasis id="B9199F16A303FF87A6FDF95D913CD786" bold="true" box="[199,364,1760,1786]" pageId="2" pageNumber="474">Type horizon.</emphasis>
|
||
Holocene (sub-fossil).
|
||
</paragraph>
|
||
<paragraph id="8BD24304A303FF87A6FDF8B99504D63F" blockId="2.[151,1437,1472,2039]" pageId="2" pageNumber="474">
|
||
<emphasis id="B9199F16A303FF87A6FDF8B991C6D662" bold="true" box="[199,406,1796,1822]" pageId="2" pageNumber="474">Studied material.</emphasis>
|
||
<materialsCitation id="3B054959A303FF87A7A6F8B99416D663" ID-GBIF-Occurrence="3962323304" box="[412,1094,1796,1823]" collectionCode="UJMC" pageId="2" pageNumber="474" specimenCode="UJMC 250-252" specimenCount="154">
|
||
<specimenCount id="9D6B888DA303FF87A7A6F8B99211D662" box="[412,577,1796,1823]" pageId="2" pageNumber="474" type="generic">154 specimens</specimenCount>
|
||
from the
|
||
<typeStatus id="54D6FDA6A303FF87A496F8B8928CD663" box="[684,732,1797,1823]" pageId="2" pageNumber="474">type</typeStatus>
|
||
locality (
|
||
<specimenCode id="DBCBEB7FA303FF87A57EF8B993A7D663" box="[836,1015,1796,1823]" collectionCode="UJMC" pageId="2" pageNumber="474">UJMC 250-252</specimenCode>
|
||
b, 254)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B054959A303FF87A26BF8B991B1D63E" ID-GBIF-Occurrence="3962323303" collectionCode="UJMC" collectorName="El Molino" pageId="2" pageNumber="474" specimenCode="UJMC 253-253" specimenCount="45">
|
||
<specimenCount id="9D6B888DA303FF87A26BF8B994B8D662" box="[1105,1256,1796,1823]" pageId="2" pageNumber="474" type="generic">45 specimens</specimenCount>
|
||
from
|
||
<collectorName id="269826D2A303FF87A311F8B995CCD662" box="[1323,1436,1796,1822]" pageId="2" pageNumber="474">El Molino</collectorName>
|
||
spring (
|
||
<specimenCode id="DBCBEB7FA303FF87A6D6F89591CFD63F" box="[236,415,1832,1859]" collectionCode="UJMC" pageId="2" pageNumber="474">UJMC 253-253</specimenCode>
|
||
c) and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B054959A303FF87A7D3F8959500D63F" ID-GBIF-Occurrence="3962323301" box="[489,1360,1832,1859]" collectionCode="UJMC" location="Hacienda de Hornos" pageId="2" pageNumber="474" specimenCode="UJMC 257-257" specimenCount="50">
|
||
<specimenCount id="9D6B888DA303FF87A7D3F89592D1D63E" box="[489,641,1832,1859]" pageId="2" pageNumber="474" type="generic">50 specimens</specimenCount>
|
||
from
|
||
<location id="8EB215DFA303FF87A4FFF89593F9D63F" LSID="urn:lsid:plazi:treatment:03C4F212A303FF81A6ADFA8D9267D628:8EB215DFA303FF87A4FFF89593F9D63F" box="[709,937,1832,1859]" name="Hacienda de Hornos" pageId="2" pageNumber="474">Hacienda de Hornos</location>
|
||
y Carranza spring (
|
||
<specimenCode id="DBCBEB7FA303FF87A2BCF895956BD63F" box="[1158,1339,1832,1859]" collectionCode="UJMC" pageId="2" pageNumber="474">UJMC 257-257</specimenCode>
|
||
c)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD24304A303FF87A6FDF8F19138D6F6" blockId="2.[151,1437,1472,2039]" pageId="2" pageNumber="474">
|
||
<emphasis id="B9199F16A303FF87A6FDF8F1910AD61A" bold="true" box="[199,346,1868,1894]" pageId="2" pageNumber="474">Occurrence.</emphasis>
|
||
Endemic to Juan Guerra, Hacienda de Hornos y Carranza and El Molino springs near Viesca,
|
||
<collectingRegion id="49A98DE6A303FF87A6ADF8CD90AFD6F6" box="[151,255,1904,1930]" country="Mexico" name="Coahuila de Zaragoza" pageId="2" pageNumber="474">Coahuila</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37A0394A303FF87A730F8CD9133D6F6" box="[266,355,1904,1930]" name="Mexico" pageId="2" pageNumber="474">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377108FA303FF87A6FDF82992C3D68A" pageId="2" pageNumber="474" type="diagnosis">
|
||
<paragraph id="8BD24304A303FF87A6FDF82992C3D68A" blockId="2.[151,1437,1472,2039]" pageId="2" pageNumber="474">
|
||
<emphasis id="B9199F16A303FF87A6FDF8299112D6D2" bold="true" box="[199,322,1940,1966]" pageId="2" pageNumber="474">Diagnosis.</emphasis>
|
||
Shell medium-sized, usually light brown or (when blanched) white, narrow-conic, variably umbilicate to imperforate. Aperture round polygonal, inner lip adnate. Teleoconch whorls having one to three distinct keels sculptured with broad, strong, conical spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF12138CA302FF86A6ADFB2D908ED420" ID-DOI="http://doi.org/10.5281/zenodo.6952682" ID-Zenodo-Dep="6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="3" pageNumber="475" startId="3.[151,250,1168,1192]" targetBox="[151,1436,181,1144]" targetPageId="3">
|
||
<paragraph id="8BD24304A302FF86A6ADFB2D908ED420" blockId="3.[151,1437,1168,1372]" pageId="3" pageNumber="475">
|
||
<emphasis id="B9199F16A302FF86A6ADFB2D9142D5D4" bold="true" box="[151,274,1168,1192]" pageId="3" pageNumber="475">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6D3887A302FF86A72DFB2C9259D5D4" authority="Czaja & Covich & Neubauer & Estrada-Rodríguez & Ávila-Rodríguez & Mata, 2022" authorityName="Czaja & Covich & Neubauer & Estrada-Rodríguez & Ávila-Rodríguez & Mata" authorityYear="2022" box="[279,521,1168,1192]" class="Gastropoda" family="Cochliopidae" genus="Spinopyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="475" phylum="Mollusca" rank="species" species="luismaedai" status="n. gen. et n. sp.">
|
||
<emphasis id="B9199F16A302FF86A72DFB2C9259D5D4" box="[279,521,1168,1192]" italics="true" pageId="3" pageNumber="475">Spinopyrgus luismaedai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22A226DA302FF86A435FB2D92FAD5D4" box="[527,682,1168,1192]" pageId="3" pageNumber="475" rank="species">
|
||
<emphasis id="B9199F16A302FF86A435FB2D9205D5D4" bold="true" box="[527,597,1168,1192]" pageId="3" pageNumber="475">n. gen.</emphasis>
|
||
et
|
||
<emphasis id="B9199F16A302FF86A44BFB2D92FAD5D4" bold="true" box="[625,682,1168,1192]" pageId="3" pageNumber="475">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="B9199F16A302FF86A48DFB2D92BAD5D4" bold="true" box="[695,746,1168,1192]" pageId="3" pageNumber="475">A–K</emphasis>
|
||
).
|
||
<emphasis id="B9199F16A302FF86A4C4FB2D9364D5D4" bold="true" box="[766,820,1168,1192]" pageId="3" pageNumber="475">A–B.</emphasis>
|
||
Holotype (UJMC-250) from Juan Guerra spring.
|
||
<emphasis id="B9199F16A302FF86A31CFB2D956ED5D4" bold="true" box="[1318,1342,1168,1192]" pageId="3" pageNumber="475">C.</emphasis>
|
||
Paratype 1 (UJMC-251) from Juan Guerra spring.
|
||
<emphasis id="B9199F16A302FF86A407FB099206D5B0" bold="true" box="[573,598,1204,1228]" pageId="3" pageNumber="475">D.</emphasis>
|
||
Paratype 2 (UJMC-252) from Juan Guerra spring.
|
||
<emphasis id="B9199F16A302FF86A258FB0994C3D5B0" bold="true" box="[1122,1171,1204,1228]" pageId="3" pageNumber="475">E, F.</emphasis>
|
||
Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels.
|
||
<emphasis id="B9199F16A302FF86A489FB659297D58C" bold="true" box="[691,711,1240,1264]" pageId="3" pageNumber="475">G</emphasis>
|
||
. Specimen (UJMC-253) from El Molino spring.
|
||
<emphasis id="B9199F16A302FF86A286FB659486D58C" bold="true" box="[1212,1238,1240,1264]" pageId="3" pageNumber="475">H.</emphasis>
|
||
Specimen (UJMC- 254) from Juan Guerra spring.
|
||
<emphasis id="B9199F16A302FF86A7E9FB4191B3D468" bold="true" box="[467,483,1276,1300]" pageId="3" pageNumber="475">I.</emphasis>
|
||
Specimen (UJMC-253) from El Molino spring, protoconch view.
|
||
<emphasis id="B9199F16A302FF86A2BCFB4194CAD468" bold="true" box="[1158,1178,1276,1300]" pageId="3" pageNumber="475">J.</emphasis>
|
||
Paratype 3 (UJMC-251), details of the carina with spines.
|
||
<emphasis id="B9199F16A302FF86A7D1FA9D9255D444" bold="true" box="[491,517,1312,1336]" pageId="3" pageNumber="475">K.</emphasis>
|
||
Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C377108FA302FF81A6FDFA1390BBD748" lastPageId="4" lastPageNumber="476" pageId="3" pageNumber="475" type="description">
|
||
<paragraph id="8BD24304A302FF86A6FDFA1394AFD6BC" blockId="3.[151,1437,1454,1985]" pageId="3" pageNumber="475">
|
||
<emphasis id="B9199F16A302FF86A6FDFA139109D4B4" bold="true" box="[199,345,1454,1480]" pageId="3" pageNumber="475">Description.</emphasis>
|
||
Shell medium-sized, thickened, imperforate, and conical, up to 7.25 whorls, the
|
||
<typeStatus id="54D6FDA6A302FF86A2E2FA13956AD4B4" box="[1240,1338,1454,1480]" pageId="3" pageNumber="475" type="holotype">holotype</typeStatus>
|
||
has 6.25 whorls, first two rounded. The apex is acute and inclined, protoconch consisting of about one whorl, smooth or slightly wrinkled, measuring 160–180 µm in diameter. A shoulder frequently develops on early teleoconch whorls. The slightly corrugate nucleus is narrow and immersed and has 40–50 μm in diameter. Protoconch-teleoconch boundary is clearly discernable, characterized by the onset of fine, axial growth lines, which are prosocline on first whorls and later become almost orthocline. Teleoconch whorls have fine or often more pronounced irregular axial ribs (
|
||
<figureCitation id="13565F81A302FF86A734F93B910DD7DD" box="[270,349,1670,1697]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="3" pageNumber="475">Fig. 2J</figureCitation>
|
||
,
|
||
<figureCitation id="13565F81A302FF86A753F93A91DDD7DD" box="[361,397,1671,1697]" captionStart="FIGURE 3" captionStartId="4.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1143]" captionTargetId="figure-19@4.[151,1436,181,1143]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of Spinopyrgus luismaedai n. gen. et n. sp. (L–P). L. Specimen (UJMC-255) from El Molino spring with strong spines. M. Specimen (UJMC-255) apical view. N. Specimen (UJMC-256), protoconch. O. Specimen (UJMC-255), details of the strong axial elements of the body whorl. P. Specimen (UJMC-256) from Juan Guerra spring, protoconch. Q-T. Pyrgophorus parvulus (Guilding) from Río Sabinas, Coahuila, Mexico (from Czaja et al. 2022, Fig. 5A). Q, R. Specimen (UJMC 460a) from Río Sabinas from both sides. S. Same specimen with details of the spiral elements of the body whorl. T. Same specimen (UJMC 460a), protoconch." figureDoi="http://doi.org/10.5281/zenodo.6952686" httpUri="https://zenodo.org/record/6952686/files/figure.png" pageId="3" pageNumber="475">3O</figureCitation>
|
||
), as well as two to three spiral carinae. Because of the strong carinae the whorls appear more angular than rounded (
|
||
<figureCitation id="13565F81A302FF86A7A3F9179246D7B9" box="[409,534,1706,1733]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="3" pageNumber="475">Fig. 2A–H</figureCitation>
|
||
,
|
||
<figureCitation id="13565F81A302FF86A418F9169215D7B9" box="[546,581,1707,1733]" captionStart="FIGURE 3" captionStartId="4.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1143]" captionTargetId="figure-19@4.[151,1436,181,1143]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of Spinopyrgus luismaedai n. gen. et n. sp. (L–P). L. Specimen (UJMC-255) from El Molino spring with strong spines. M. Specimen (UJMC-255) apical view. N. Specimen (UJMC-256), protoconch. O. Specimen (UJMC-255), details of the strong axial elements of the body whorl. P. Specimen (UJMC-256) from Juan Guerra spring, protoconch. Q-T. Pyrgophorus parvulus (Guilding) from Río Sabinas, Coahuila, Mexico (from Czaja et al. 2022, Fig. 5A). Q, R. Specimen (UJMC 460a) from Río Sabinas from both sides. S. Same specimen with details of the spiral elements of the body whorl. T. Same specimen (UJMC 460a), protoconch." figureDoi="http://doi.org/10.5281/zenodo.6952686" httpUri="https://zenodo.org/record/6952686/files/figure.png" pageId="3" pageNumber="475">3L</figureCitation>
|
||
). Along the central carina, many axial ribs extend into strong spines on the periphery. The spines are regularly spaced and always in the middle part of the whorl. The last three or four whorls are geniculate, angled at location of carina. Body whorl large, almost always having a second carina with spines on the lower side, sometimes with a third carina (
|
||
<figureCitation id="13565F81A302FF86A4A4F8AB9343D64D" box="[670,787,1814,1841]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="3" pageNumber="475">Fig. 2E, F</figureCitation>
|
||
). Whorls mostly with very pointed, shovel-shaped spines, with up to 30 spines on the body whorls, spines very pointed. The spines like all other shell ornamentations are part of the calcareous shell material. Suture slightly impressed. Many shells conserve still their thin light-brown periostracal layer (
|
||
<figureCitation id="13565F81A302FF86A756F83F91BCD6E1" box="[364,492,1922,1949]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="3" pageNumber="475">Figs. 2C–J</figureCitation>
|
||
). Aperture polygonal, mostly pentagonal in shape (
|
||
<figureCitation id="13565F81A302FF86A20FF83F94B1D6E1" box="[1077,1249,1922,1949]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="3" pageNumber="475">Figs. 2A, E, G</figureCitation>
|
||
), slightly longer than wide. Parietal lip adnate, the basal and outer lip rounded and thin. Opercula were not preserved.
|
||
</paragraph>
|
||
<caption id="DF12138CA305FF81A6ADFB2D921ED420" ID-DOI="http://doi.org/10.5281/zenodo.6952686" ID-Zenodo-Dep="6952686" httpUri="https://zenodo.org/record/6952686/files/figure.png" pageId="4" pageNumber="476" startId="4.[151,250,1168,1192]" targetBox="[151,1436,181,1143]" targetPageId="4">
|
||
<paragraph id="8BD24304A305FF81A6ADFB2D921ED420" blockId="4.[151,1437,1168,1372]" pageId="4" pageNumber="476">
|
||
<emphasis id="B9199F16A305FF81A6ADFB2D9143D5D4" bold="true" box="[151,275,1168,1192]" pageId="4" pageNumber="476">FIGURE 3.</emphasis>
|
||
SEM images of
|
||
<taxonomicName id="4C6D3887A305FF81A787FB2D92E0D5D4" authority="Czaja & Covich & Neubauer & Estrada-Rodríguez & Ávila-Rodríguez & Mata, 2022" authorityName="Czaja & Covich & Neubauer & Estrada-Rodríguez & Ávila-Rodríguez & Mata" authorityYear="2022" box="[445,688,1168,1192]" class="Gastropoda" family="Cochliopidae" genus="Spinopyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="476" phylum="Mollusca" rank="species" species="luismaedai" status="n. gen. et n. sp.">
|
||
<emphasis id="B9199F16A305FF81A787FB2D92E0D5D4" box="[445,688,1168,1192]" italics="true" pageId="4" pageNumber="476">Spinopyrgus luismaedai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22A226DA305FF81A48DFB2D9306D5D4" box="[695,854,1168,1192]" pageId="4" pageNumber="476" rank="species">
|
||
<emphasis id="B9199F16A305FF81A48DFB2D92AED5D4" bold="true" box="[695,766,1168,1192]" pageId="4" pageNumber="476">n. gen.</emphasis>
|
||
et
|
||
<emphasis id="B9199F16A305FF81A527FB2D9306D5D4" bold="true" box="[797,854,1168,1192]" pageId="4" pageNumber="476">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="B9199F16A305FF81A55FFB2D93EED5D4" bold="true" box="[869,958,1168,1192]" pageId="4" pageNumber="476">L–P). L.</emphasis>
|
||
Specimen (UJMC-255) from El Molino spring with strong spines.
|
||
<emphasis id="B9199F16A305FF81A766FB09912AD5B0" bold="true" box="[348,378,1204,1228]" pageId="4" pageNumber="476">M.</emphasis>
|
||
Specimen (UJMC-255) apical view.
|
||
<emphasis id="B9199F16A305FF81A4C8FB099354D5B0" bold="true" box="[754,772,1204,1228]" pageId="4" pageNumber="476">N</emphasis>
|
||
. Specimen (UJMC-256), protoconch.
|
||
<emphasis id="B9199F16A305FF81A2B3FB0994F3D5B0" bold="true" box="[1161,1187,1204,1228]" pageId="4" pageNumber="476">O.</emphasis>
|
||
Specimen (UJMC-255), details of the strong axial elements of the body whorl.
|
||
<emphasis id="B9199F16A305FF81A4F4FB6592B1D58C" bold="true" box="[718,737,1240,1264]" pageId="4" pageNumber="476">P.</emphasis>
|
||
Specimen (UJMC-256) from Juan Guerra spring, protoconch.
|
||
<emphasis id="B9199F16A305FF81A351FB6595CCD58C" bold="true" box="[1387,1436,1240,1264]" pageId="4" pageNumber="476">Q-T.</emphasis>
|
||
<taxonomicName id="4C6D3887A305FF81A6ADFB4191A1D468" authority="(Guilding)" baseAuthorityName="Guilding" baseAuthorityYear="1828" box="[151,497,1276,1300]" class="Gastropoda" family="Cochliopidae" genus="Pyrgophorus" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="476" phylum="Mollusca" rank="species" species="parvulus">
|
||
<emphasis id="B9199F16A305FF81A6ADFB41912DD468" box="[151,381,1276,1300]" italics="true" pageId="4" pageNumber="476">Pyrgophorus parvulus</emphasis>
|
||
(Guilding)
|
||
</taxonomicName>
|
||
from Río Sabinas, Coahuila, Mexico (from
|
||
<bibRefCitation id="EFFC3EF5A305FF81A5FCFB41942DD468" author="Czaja A. & Jorge Luis Becerra-Lopez, J. L. & Estrada-Rodriguez, J. L. & Romero-Mendez, U. & Cardoza-Martinez, G. F. & Saenz-Mata, J. & Estrada-Arellano, J. R. & Garza-Martinez, M. & Hernandez-Teran, F. & Cerano-Paredes, J." box="[966,1149,1276,1300]" pageId="4" pageNumber="476" pagination="1 - 22" refId="ref5104" refString="Czaja A., Jorge Luis Becerra-Lopez, J. L., Estrada-Rodriguez, J. L., Romero-Mendez, U., Cardoza-Martinez, G. F., Saenz-Mata, J., Estrada-Arellano, J. R., Garza-Martinez, M. A, Hernandez-Teran, F. & Cerano-Paredes, J. (2022) The Sabinas River basin in Coahuila, a new hotspot of molluscan biodiversity near Cuatro Cienegas, Chihuahuan Desert, northern Mexico. Revista Mexicana de Biodiversidad, 93, 1 - 22. https: // doi. org / 10.22201 / ib. 20078706 e. 2022.93.3588" type="journal article" year="2022">
|
||
Czaja
|
||
<emphasis id="B9199F16A305FF81A233FB409469D468" box="[1033,1081,1276,1300]" italics="true" pageId="4" pageNumber="476">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
, Fig. 5A).
|
||
<emphasis id="B9199F16A305FF81A2C9FB41957FD468" bold="true" box="[1267,1327,1276,1300]" pageId="4" pageNumber="476">Q, R.</emphasis>
|
||
Specimen (UJMC 460a) from Río Sabinas from both sides.
|
||
<emphasis id="B9199F16A305FF81A4ADFA9D92FBD444" bold="true" box="[663,683,1312,1336]" pageId="4" pageNumber="476">S.</emphasis>
|
||
Same specimen with details of the spiral elements of the body whorl.
|
||
<emphasis id="B9199F16A305FF81A3BDFA9D95CDD444" bold="true" box="[1415,1437,1312,1336]" pageId="4" pageNumber="476">T.</emphasis>
|
||
Same specimen (UJMC 460a), protoconch.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD24304A305FF81A6FDFA13931FD490" blockId="4.[151,1437,1454,1877]" pageId="4" pageNumber="476">
|
||
<emphasis id="B9199F16A305FF81A6FDFA1391E6D4B4" bold="true" box="[199,438,1454,1480]" pageId="4" pageNumber="476">Shell measurements</emphasis>
|
||
(mean ± standard deviation in parentheses; n = 20). SH 4.64 (0.46) mm, SW 2.19 (0.31) mm, WN 6.05 (0.49) whorls (shells from Juan Guerra spring).
|
||
</paragraph>
|
||
<paragraph id="8BD24304A305FF81A6FDFA4B90BBD748" blockId="4.[151,1437,1454,1877]" pageId="4" pageNumber="476">
|
||
<emphasis id="B9199F16A305FF81A6FDFA4B9240D76C" bold="true" box="[199,528,1526,1552]" pageId="4" pageNumber="476">
|
||
Measurements of
|
||
<typeStatus id="54D6FDA6A305FF81A7A6FA4B925BD76C" box="[412,523,1526,1552]" pageId="4" pageNumber="476" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
SH
|
||
<quantity id="4C95EEE1A305FF81A479FA4B92F9D76D" box="[579,681,1526,1553]" metricMagnitude="-3" metricUnit="m" metricValue="4.91" pageId="4" pageNumber="476" unit="mm" value="4.91">4.91 mm</quantity>
|
||
, SW
|
||
<quantity id="4C95EEE1A305FF81A4DCFA4B931CD76D" box="[742,844,1526,1553]" metricMagnitude="-3" metricUnit="m" metricValue="2.72" pageId="4" pageNumber="476" unit="mm" value="2.72">2.72 mm</quantity>
|
||
, AH
|
||
<quantity id="4C95EEE1A305FF81A5BCFA4B93BDD76D" box="[902,1005,1526,1553]" metricMagnitude="-3" metricUnit="m" metricValue="1.89" pageId="4" pageNumber="476" unit="mm" value="1.89">1.89 mm</quantity>
|
||
, AW
|
||
<quantity id="4C95EEE1A305FF81A210FA4B94C1D76D" box="[1066,1169,1526,1553]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="4" pageNumber="476" unit="mm" value="1.83">1.83 mm</quantity>
|
||
, AH/SH 0.39, WN 6.25 whorls.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377108FA305FF81A6FDF9839267D628" pageId="4" pageNumber="476" type="discussion">
|
||
<paragraph id="8BD24304A305FF81A6FDF9839267D628" blockId="4.[151,1437,1454,1877]" pageId="4" pageNumber="476">
|
||
<emphasis id="B9199F16A305FF81A6FDF983916BD724" bold="true" box="[199,315,1598,1624]" pageId="4" pageNumber="476">Remarks.</emphasis>
|
||
The prominent spine ornamentations of
|
||
<taxonomicName id="4C6D3887A305FF81A522F98393E3D724" authorityName="Czaja & Covich & Neubauer & Estrada-Rodríguez & Ávila-Rodríguez & Mata" authorityYear="2022" box="[792,947,1598,1625]" class="Gastropoda" family="Cochliopidae" genus="Spinopyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="476" phylum="Mollusca" rank="species" species="luismaedai">
|
||
<emphasis id="B9199F16A305FF81A522F9839376D724" box="[792,806,1598,1624]" italics="true" pageId="4" pageNumber="476">S</emphasis>
|
||
.
|
||
<emphasis id="B9199F16A305FF81A502F98393E3D724" box="[824,947,1598,1624]" italics="true" pageId="4" pageNumber="476">luismaedai</emphasis>
|
||
</taxonomicName>
|
||
with extremely long and almost “marinelike” spines are not seen in any other North American gastropod species. All spines are built completely of shell material and are not made of a combination of shell material and periostracal projections, like in some other spiny
|
||
<taxonomicName id="4C6D3887A305FF81A6ADF917927FD7B8" authority="Ancey" authorityName="Ancey" authorityYear="1888" box="[151,559,1706,1732]" class="Gastropoda" family="Cochliopidae" genus="Pyrgophorus" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="476" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9199F16A305FF81A6ADF9179178D7B8" box="[151,296,1706,1732]" italics="true" pageId="4" pageNumber="476">Pyrgophorus</emphasis>
|
||
Ancey (Cochliopidae)
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C6D3887A305FF81A464F91693A2D7B8" authority="Stimpson" authorityName="Stimpson" authorityYear="1865" box="[606,1010,1706,1732]" class="Gastropoda" family="Tateidae" genus="Potamopyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="476" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9199F16A305FF81A464F9169350D7B8" box="[606,768,1707,1732]" italics="true" pageId="4" pageNumber="476">Potamopyrgus</emphasis>
|
||
Stimpson (Tateidae)
|
||
</taxonomicName>
|
||
species. The variability of the shell ornamentations is remarkable especially regarding the number and form of the spines (
|
||
<figureCitation id="13565F81A305FF81A2B8F9739550D794" box="[1154,1280,1742,1768]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="4" pageNumber="476">Fig. 2C–K</figureCitation>
|
||
). Most of the shells are turriform with 5–7 elongated whorls (
|
||
<figureCitation id="13565F81A305FF81A491F94F9377D670" box="[683,807,1778,1804]" captionStart="FIGURE 2" captionStartId="3.[151,250,1168,1192]" captionTargetBox="[151,1436,181,1144]" captionTargetId="figure-25@3.[151,1436,181,1144]" captionTargetPageId="3" captionText="FIGURE 2. Spinopyrgus luismaedai n. gen. et n. sp. (A–K). A–B. Holotype (UJMC-250) from Juan Guerra spring. C. Paratype 1 (UJMC-251) from Juan Guerra spring. D. Paratype 2 (UJMC-252) from Juan Guerra spring. E, F. Paratype 3 (UJMC-252a) from Juan Guerra spring, specimen with three keels. G. Specimen (UJMC-253) from El Molino spring. H. Specimen (UJMC- 254) from Juan Guerra spring. I. Specimen (UJMC-253) from El Molino spring, protoconch view. J. Paratype 3 (UJMC-251), details of the carina with spines. K. Specimen (UJMC-253b) from El Molino spring, details of the carina with shovel-shaped spines." figureDoi="http://doi.org/10.5281/zenodo.6952682" httpUri="https://zenodo.org/record/6952682/files/figure.png" pageId="4" pageNumber="476">Fig. 2A–H</figureCitation>
|
||
), but there are also in all springs a sub-population with a more rounded general form and less than 5 whorls. This variability could be an indication of different ecophenotypes within the
|
||
<taxonomicName id="4C6D3887A305FF81A735F88791F6D628" authorityName="Czaja & Covich & Neubauer & Estrada-Rodríguez & Ávila-Rodríguez & Mata" authorityYear="2022" box="[271,422,1850,1877]" class="Gastropoda" family="Cochliopidae" genus="Spinopyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="476" phylum="Mollusca" rank="species" species="luismaedai">
|
||
<emphasis id="B9199F16A305FF81A735F887914DD628" box="[271,285,1850,1876]" italics="true" pageId="4" pageNumber="476">S</emphasis>
|
||
.
|
||
<emphasis id="B9199F16A305FF81A711F88791F6D628" box="[299,422,1850,1876]" italics="true" pageId="4" pageNumber="476">luismaedai</emphasis>
|
||
</taxonomicName>
|
||
populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |