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<document id="7F4B07A0051003F2272CF41BEDDD2A3B" ID-CLB-Dataset="45917" ID-DOI="10.5281/zenodo.279260" ID-GBIF-Dataset="ffbcaa56-6a77-4df4-b944-508fb074171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="279260" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460413266680" checkinUser="plazi" docAuthor="Shear, William A." docDate="2011" docId="03C47A60FFD0724760C058FA7A587AF8" docLanguage="en" docName="zt03109p038.pdf" docOrigin="Zootaxa 3109" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pseudotremia loomisi Shear, 2011, n. sp." docType="treatment" docVersion="9" lastPageNumber="6" masterDocId="FFFD0218FFD5724260575C087F167D21" masterDocTitle="Cave millipeds of the United States. X. New species and records of the genus Pseudotremia Cope. 2. Species from Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="6" updateTime="1698301493022" updateUser="ExternalLinkService">
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<mods:title id="1EF9125EE2FB53F86B37E12A17E7554F">Cave millipeds of the United States. X. New species and records of the genus Pseudotremia Cope. 2. Species from Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae)</mods:title>
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<mods:namePart id="8A3DC35700D3A7D7DC0C5625A594FA31">Shear, William A.</mods:namePart>
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<mods:date id="874EF43A8669CDD6BC16203C7A2DF13A">2011</mods:date>
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<paragraph id="8BD2CB76FFD0724760C058FA7EFB782D" blockId="5.[151,493,1266,1325]" box="[151,493,1266,1292]" pageId="5" pageNumber="6">
<heading id="D09A7C1AFFD0724760C058FA7EFB782D" bold="true" box="[151,493,1266,1292]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="B9191764FFD0724760C058FA7EFB782D" bold="true" box="[151,493,1266,1292]" pageId="5" pageNumber="6">
<taxonomicName id="4C6DB0F5FFD0724760C058FA7E8A782D" box="[151,412,1266,1292]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="loomisi" status="sp. nov.">
<emphasis id="B9191764FFD0724760C058FA7E8A782D" bold="true" box="[151,412,1266,1292]" italics="true" pageId="5" pageNumber="6">Pseudotremia loomisi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22AAA1FFFD0724761FC58FA7EFB782D" box="[427,493,1266,1292]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="8BD2CB76FFD0724760C0591D7FE9780C" blockId="5.[151,493,1266,1325]" box="[151,255,1301,1325]" pageId="5" pageNumber="6">
<figureCitation id="1356D7F3FFD0724760C0591D7FE9780C" box="[151,255,1301,1325]" captionStart="FIGURES 1 8" captionStartId="6.[151,264,1777,1800]" captionTargetBox="[151,1436,193,1752]" captionTargetId="figure@6.[151,1436,193,1754]" captionTargetPageId="6" captionText="FIGURES 1 8. New species of Pseudotremia. Figs. 1 7, P. loomisi, n. sp. Fig. 1, ocelli. Fig. 2, metazonite, dorsal view. Fig. 3, diplosegment, lateral view. Fig. 4, gonopods, ventral view. Fig. 5, gonopods, anterior view. Fig. 6, gonopods, lateral view. Fig. 7, left ninth leg, posterior view. Fig. 8, ocelli of P. c o n t or ta, n. sp." httpUri="https://zenodo.org/record/279261/files/figure.png" pageId="5" pageNumber="6">Figs. 17</figureCitation>
</paragraph>
<paragraph id="8BD2CB76FFD0724760C059557EB878B8" blockId="5.[151,1436,1373,2010]" pageId="5" pageNumber="6">
<emphasis id="B9191764FFD0724760C059557FF57854" bold="true" box="[151,227,1373,1397]" pageId="5" pageNumber="6">
<typeStatus id="54D675D4FFD0724760C059557FCC7854" box="[151,218,1373,1397]" pageId="5" pageNumber="6">Types</typeStatus>
:
</emphasis>
Male
<typeStatus id="54D675D4FFD07247616459557E8E7854" box="[307,408,1373,1397]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
, male and female
<typeStatus id="54D675D4FFD07247622759567DCD7857" box="[624,731,1374,1398]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
from Clarks Cave,
<emphasis id="B9191764FFD07247639C59557B2E7854" box="[971,1080,1373,1397]" italics="true" pageId="5" pageNumber="6">Bath Co.,</emphasis>
VIRGINIA, collected
<date id="FFD3EDB6FFD07247651459557FD978B8" pageId="5" pageNumber="6" value="1997-05-21">21 May 1997</date>
by David Hubbard.
</paragraph>
</subSubSection>
<subSubSection id="C37798FDFFD07247609059AD7A1B78C0" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BD2CB76FFD07247609059AD7A1B78C0" blockId="5.[151,1436,1373,2010]" pageId="5" pageNumber="6">
<emphasis id="B9191764FFD07247609059AD7E55789F" bold="true" box="[199,323,1445,1470]" pageId="5" pageNumber="6">Diagnosis:</emphasis>
The apical and subapical spines of the MAP are both missing and there are no colpocoxite processes, while the ninth legpair telopodite appears two-segmented; a unique combination of characters.
</paragraph>
</subSubSection>
<subSubSection id="C37798FDFFD07247609059E57DB57B6C" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BD2CB76FFD07247609059E57DB57B6C" blockId="5.[151,1436,1373,2010]" pageId="5" pageNumber="6">
<emphasis id="B9191764FFD07247609059E57E477B24" bold="true" box="[199,337,1517,1541]" pageId="5" pageNumber="6">Etymology:</emphasis>
The species epithet honors the late H. F. Loomis, who, in a series of papers some 70 to 80 years ago, described many cave millipeds from the Appalachians, including species of
<taxonomicName id="4C6DB0F5FFD0724764485A197BAC7B08" box="[1055,1210,1553,1577]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9191764FFD0724764485A197BAC7B08" box="[1055,1210,1553,1577]" italics="true" pageId="5" pageNumber="6">Pseudotremia</emphasis>
</taxonomicName>
. Suggested vernacular name: Loomiss Rough-backed Millipede.
</paragraph>
</subSubSection>
<subSubSection id="C37798FDFFD0724760905A517A587AF8" pageId="5" pageNumber="6" type="description">
<paragraph id="8BD2CB76FFD0724760905A517C217BFC" blockId="5.[151,1436,1373,2010]" pageId="5" pageNumber="6">
<emphasis id="B9191764FFD0724760905A517E4D7B53" bold="true" box="[199,347,1625,1650]" pageId="5" pageNumber="6">Description:</emphasis>
Male
<typeStatus id="54D675D4FFD0724761F65A517D157B50" box="[417,515,1625,1649]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
about
<quantity id="4C956693FFD0724762195A517D8A7B50" box="[590,668,1625,1649]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="6" unit="mm" value="27.0">27 mm</quantity>
long,
<quantity id="4C956693FFD0724762B55A517C217B50" box="[738,823,1625,1649]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="6" unit="mm" value="2.3">2.3 mm</quantity>
wide, third antennal segment
<quantity id="4C956693FFD0724764D15A517BFF7B50" box="[1158,1257,1625,1649]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="5" pageNumber="6" unit="mm" value="1.45">1.45 mm</quantity>
long. Ocelli 19 on each side, well-formed, contiguous, darkly pigmented (fig. 1). Segmental shoulders prominent; midbody metazonites with 35 longitudinal rugae medial to shoulders, middorsally smooth; 1213 strong lateral striations (figs. 2, 3). Color purplish gray-brown, mottled darker anteriorly.
</paragraph>
<paragraph id="8BD2CB76FFD0724760875AE17C107AB0" blockId="5.[151,1436,1373,2010]" pageId="5" pageNumber="6">Gonopods (figs. 46) robust, angiocoxites deeply divided on each side by U-shaped sinus; MAPs relatively narrow in lateral view, lacking apical and subapical spines; LAPs long, distally divided with dorsal branch much smaller, lateral branches curve inward and cross in midline. Colpoxites deeply divided but not conspicuously mitten-shaped. Colpocoxite processes absent. Ninth legs (fig. 7) with distal segment shorter than basal, basal segment (coxoprefemur) distally swollen, proximal knobs small.</paragraph>
<paragraph id="8BD2CB76FFD0724760905B957EC67A94" blockId="5.[151,1436,1373,2010]" box="[199,464,1949,1973]" pageId="5" pageNumber="6">Female similar to male.</paragraph>
<paragraph id="8BD2CB76FFD0724760905BCA7A587AF8" blockId="5.[151,1436,1373,2010]" box="[199,1358,1985,2010]" pageId="5" pageNumber="6">
<emphasis id="B9191764FFD0724760905BCA7E057AFB" bold="true" box="[199,275,1986,2010]" pageId="5" pageNumber="6">Notes:</emphasis>
This species is evidently a troglophile. The specimens are depigmented from long preservation.
</paragraph>
</subSubSection>
</treatment>
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