255 lines
36 KiB
XML
255 lines
36 KiB
XML
<document id="3DD57150F67B338912A829A8EE3ED459" ID-DOI="10.11646/zootaxa.3620.1.2" ID-GBIF-Dataset="60ae7d68-88b7-4da4-acae-6e4cda073ea0" ID-ISSN="1175-5326" ID-Zenodo-Dep="283400" ID-ZooBank="3870AC9F-A4EE-4E36-86C7-E1D6967A2A2E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460578993957" checkinUser="plazi" docAuthor="Wu, Min & Xu, Qin" docDate="2013" docId="03B8B70EFF9EFFD3FF05FC2AFED5B6BB" docLanguage="en" docName="zt03620p066.pdf" docOrigin="Zootaxa 3620 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Serina egressa Sturany 1900" docType="treatment" docVersion="5" lastPageNumber="51" masterDocId="FF81CF76FF98FFDBFF92FFF2FFB8B51F" masterDocTitle="Serina Gredler (Gastropoda, Stylommatophora: Enidae), the continuous-peristomed mountain snails endemic to the eastern slope of the Qinghai-Xizang Plateau" masterLastPageNumber="66" masterPageNumber="43" pageNumber="49" updateTime="1698323809859" updateUser="plazi">
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<mods:title id="9C80BC2892C1AB22A74DB5D4ED95BEB6">Serina Gredler (Gastropoda, Stylommatophora: Enidae), the continuous-peristomed mountain snails endemic to the eastern slope of the Qinghai-Xizang Plateau</mods:title>
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<mods:namePart id="CABE457E0A591607F71B1208CDE1C0E4">Wu, Min</mods:namePart>
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<mods:roleTerm id="0122265ED4A297EAFFF81106BE2AB9CC">Author</mods:roleTerm>
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<mods:namePart id="E8E6BFE9821719F1BD55DABCD8ED0162">Xu, Qin</mods:namePart>
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<mods:typeOfResource id="45E4007C25F9370BAD3044B210864E2F">text</mods:typeOfResource>
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<mods:title id="7C0A11D9561C7E21745081BB07D3A963">Zootaxa</mods:title>
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<mods:part id="71904E920DFB3F74A682D97B62269BC0">
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<mods:date id="38CE20E82E308074759C64CF5BDD27C8">2013</mods:date>
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<mods:number id="DD9A83C829E7FFA90A0A163DF63F1E8B">3620</mods:number>
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<treatment id="03B8B70EFF9EFFD3FF05FC2AFED5B6BB" ID-DOI="http://doi.org/10.5281/zenodo.6162550" ID-GBIF-Taxon="127663370" ID-Zenodo-Dep="6162550" LSID="urn:lsid:plazi:treatment:03B8B70EFF9EFFD3FF05FC2AFED5B6BB" httpUri="http://treatment.plazi.org/id/03B8B70EFF9EFFD3FF05FC2AFED5B6BB" lastPageId="8" lastPageNumber="51" pageId="6" pageNumber="49">
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<subSubSection id="C30B5593FF9EFFDDFF05FC2AFC95B146" pageId="6" pageNumber="49" type="nomenclature">
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<paragraph id="8BAE0618FF9EFFDDFF05FC2AFDAFB6ED" blockId="6.[151,535,984,1043]" box="[151,535,984,1011]" pageId="6" pageNumber="49">
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<heading id="D0E6B174FF9EFFDDFF05FC2AFDAFB6ED" bold="true" box="[151,535,984,1011]" fontSize="11" level="1" pageId="6" pageNumber="49" reason="1">
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<emphasis id="B965DA0AFF9EFFDDFF05FC2AFDAFB6ED" bold="true" box="[151,535,984,1011]" pageId="6" pageNumber="49">
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3.
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<taxonomicName id="4C117D9BFF9EFFDDFF26FC2BFDAFB6ED" ID-CoL="7T4SK" authority="Sturany, 1900" authorityName="Sturany" authorityYear="1900" box="[180,535,984,1011]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="49" phylum="Mollusca" rank="species" species="egressa">
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<emphasis id="B965DA0AFF9EFFDDFF26FC2BFEDDB6EC" bold="true" box="[180,357,985,1011]" italics="true" pageId="6" pageNumber="49">Serina egressa</emphasis>
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Sturany, 1900
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAE0618FF9EFFDDFF05FC09FDBDB10C" blockId="6.[151,535,984,1043]" box="[151,517,1019,1043]" pageId="6" pageNumber="49">
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<figureCitation id="132A1A9DFF9EFFDDFF05FC09FF5BB10C" box="[151,227,1019,1043]" captionStart="FIGURE 1" captionStartId="2.[151,250,1303,1326]" captionTargetBox="[153,1432,201,1255]" captionTargetId="figure@2.[151,1436,171,1279]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Serina species. S. soluta inflata Yen, 1939 is also indicated although synonymized in this paper. S. soluta stenochila (Möllendorff, 1901) is not mapped due to imprecise locality information." httpUri="https://zenodo.org/record/283401/files/figure.png" pageId="6" pageNumber="49">Figs. 1</figureCitation>
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,
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<figureCitation id="132A1A9DFF9EFFDDFF63FC09FF47B10C" box="[241,255,1019,1043]" captionStart="FIGURE 2. A" captionStartId="4.[151,250,1761,1784]" captionTargetBox="[151,1435,195,1740]" captionTargetId="figure@4.[151,1436,195,1740]" captionTargetPageId="4" captionText="FIGURE 2. A. Serina belae (Hilber, 1883), SMF- 42069, paratype, Bangda. B, C. Serina cathaica Gredler, 1898: B. SMF- 42046, between Yue-lin-kuan and Wen-hsien. C. NHM- 1902.5.13.19. D. Clausiliopsis szechenyi (Böttger, 1883), syntype of B. sobrina Preston, NHM- 1912.3.26.9, Gansu. E – J. Serina egressa Sturany, 1900. E. NHM- 1902.5.13.21, Gansu. F. HBUMM- 05426 – specimen 21, Wenxian. G. HBUMM- 05426 – specimen 23, Wenxian. H. SMF- 42038 (from type locality, between Yuelin-kuan and Wen-hsien). I. HBUMM- 5673 – specimen 22, Lianghekou. J. HBUMM- 5673 – specimen 23, Lianghekou. K – O. Serina prostoma (Ancey, 1884). K. NHM- 1912.6.27.69 – 71. L. A paratype of Buliminus (Holcauchen) tubios Annandale, 1924, NHM- 1923.5.24.17 – 36, Gorge of Mekong above Yangtze. M. SMF- 42050, paratype, Dshy-kow. N. HBUMM- 04296 – specimen 25, Yanjing. O. HBUMM- 04296 – specimen 24, Yanjing." httpUri="https://zenodo.org/record/283402/files/figure.png" pageId="6" pageNumber="49">2</figureCitation>
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E–J, 3, 9;
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<tableCitation id="C69333A3FF9EFFDDFEE2FC09FE53B10C" box="[368,491,1019,1043]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="5.[151,235,152,173]" captionStartId-1="5.[151,235,1295,1316]" captionStartId-2="6.[151,235,298,319]" captionTargetBox-0="[151,1436,323,1223]" captionTargetBox-1="[151,1435,1374,1654]" captionTargetBox-2="[159,1399,347,882]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="TABLE 1. Shell measurements of Serina species (specimen lots: Serina belae, SMF- 42069; S. cathaica, SMF- 42046; S. egressa, HBUMM- 05426 + 05464 + 05538 + 05563 + 05648 + 0 5673, SMF- 42038; S. prostoma, NHM- 1912.6.27.69 - 71 + 1923.5.24.17 - 36, SMF- 42050, HBUMM- 04287 - + 04296; S. ser, SMF- 42036; S. subser, HBUMM- 00287 + 05796 + 05709 + 0 5392, SMF- 42041; S. soluta, SMF- 42095 + 42096 (S. soluta “ inflata ”), SMF- 42048 + 42049 (S. soluta “ stenochila ”), SMF 42047, HBUMM- 04318; S. vincentii, SMF 42032; S. denticulata n. sp., HBUMM- 00281)." captionText-1="TABLE 2. Measurements of genitalia of some Serina species (unit: mm; measured specimens: S. egressa, 2; S. prostoma, 2; S. subser, 2; S. denticulata n. sp., 1)." captionText-2="TABLE 3. Diagnostics of Serina species. Anatomy data from this study and Schileyko, 1998 (Serina ser Gredler, 1898)." httpUri-0="http://table.plazi.org/id/DF6E5690FF9DFFDEFF05FF6AFBD5B436" httpUri-1="http://table.plazi.org/id/DF6E5690FF9DFFDEFF05FAFDFE4AB05B" httpUri-2="http://table.plazi.org/id/DF6E5690FF9EFFDDFF05FED8FAE5B45F" pageId="6" pageNumber="49" tableUuid-0="DF6E5690FF9DFFDEFF05FF6AFBD5B436" tableUuid-1="DF6E5690FF9DFFDEFF05FAFDFE4AB05B" tableUuid-2="DF6E5690FF9EFFDDFF05FED8FAE5B45F">Tables 1–3</tableCitation>
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,
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<tableCitation id="C69333A3FF9EFFDDFE65FC09FDBDB10C" box="[503,517,1019,1043]" captionStart="TABLE 6" captionStartId="22.[151,235,293,314]" captionTargetBox="[156,1416,936,1454]" captionTargetPageId="22" captionText="TABLE 6. Data set containing 18 characters (0 - 17) and 11 taxa used in the phylogenetic analyses. Information of the data contributors – S. ser (Gredler, 1898): shell characters from SMF- 42036, genitalian characters after Schileyko (1998, Fig. 235 B); S. egressa Sturany, 1900: HBUMM- 05426, 0 5464, 0 5538, 0 5563, 0 5648, 0 5673, SMF- 42038; S. subser Gredler, 1898: HBUMM- 00287, 0 5796, 0 5709, 05392; S. prostoma (Ancey, 1884): HBUMM- 04287, 0 4296, SMF- 042050; S. denticulata n. sp.: HBUMM- 0 0 281, types; Pupinidius latilabrum (Annandale, 1924): HBUMM- 04328; Pupinidius melinostoma (Möllendorff, 1901): HBUMM- 05480; Pupinidius nanpingensis (Möllendorff, 1901): HBUMM- 05497; Pupinidius wenxian Wu & Zheng, 2009: HBUMM- 05432; Pupopsis pupopsis (Gredler, 1898): HBUMM- 04298; Clausiliopsis szechenyi (Böttger, 1883): HBUMM- 05478; Holcauchen sulcata (Möllendorff, 1901): HBUMM- 05706. Measurements are from 334 shells of above species. Description of character and character states: Character 0, whorl number of embryonic shell; Character 1, whorl number; Character 2, shell height; Character 3, shell breadth; Character 4, aperture height; Character 5, aperture width; Character 6, ratio of shell height to shell breadth; Character 7, ratio of shell height to aperture height. For Characters 0 ~ 7, grouped by cluster center 1 – 1, grouped by cluster center 2 – 2, grouped by cluster center 1 or 2 – 3 (the clustering centers for character states 1 and 2 of Characters 0 - 7 see in Table 5). Character 8, peristome continuous with a channel – 1, aperture not continuous – 0; Character 9, palatal tooth present – 1, palatal tooth absent – 0; Character 10, parietal tooth present – 1, parietal tooth absent – 0; Character 11, with 2 columellar teeth – 2, with 1 columellar tooth – 1, columellar tooth absent – 0; Character 12, epiphallus twisted – 1, epiphallus straight – 0; Character 13, epiphallic caecum present – 1, epiphallic caecum absent – 0; Character 14, penial caecum present – 1, penical caecum absent – 0; Character 15, bursa copulatrix duct twisted – 1, bursa copulatrix duct straight – 0; Character 16, diverticle absent – 0, diverticle not terminally expanded – 1, diverticle terminally expanded – 2; Character 17, free oviduct longer than vagina – 1, free oviduct subequal to or shorter than vagina – 0." httpUri="http://table.plazi.org/id/DF6E5690FF8EFFCDFF05FED7FB38B67C" pageId="6" pageNumber="49" tableUuid="DF6E5690FF8EFFCDFF05FED7FB38B67C">6</tableCitation>
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</paragraph>
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<paragraph id="8BAE0618FF9EFFDDFF05FBB6FC95B146" blockId="6.[151,1209,1091,1175]" box="[151,813,1091,1114]" pageId="6" pageNumber="49">
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<taxonomicName id="4C117D9BFF9EFFDDFF05FBB6FE8EB145" box="[151,310,1092,1114]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="49" phylum="Mollusca" rank="species" species="cathaica">
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<emphasis id="B965DA0AFF9EFFDDFF05FBB6FE8EB145" box="[151,310,1092,1114]" italics="true" pageId="6" pageNumber="49">Serina cathaica</emphasis>
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</taxonomicName>
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<taxonomicName id="4C117D9BFF9EFFDDFEAFFBB7FDD7B146" authority="Sturany, 1900: 35" authorityName="Sturany" authorityPageNumber="35" authorityYear="1900" box="[317,623,1091,1114]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="49" phylum="Mollusca" rank="variety" species="cathaica" variety="egressa">
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var.
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<emphasis id="B965DA0AFF9EFFDDFEFAFBB7FE0CB145" box="[360,436,1093,1114]" italics="true" pageId="6" pageNumber="49">egressa</emphasis>
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Sturany, 1900: 35
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</taxonomicName>
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, pl. 3, figs. 14–16.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B5593FF9EFFDDFF05FB90FB01B188" pageId="6" pageNumber="49" type="description">
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<paragraph id="8BAE0618FF9EFFDDFF05FB90FB01B188" blockId="6.[151,1209,1091,1175]" pageId="6" pageNumber="49">
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<taxonomicName id="4C117D9BFF9EFFDDFF05FB90FEEEB167" authority="Serina" authorityName="Serina" box="[151,342,1122,1144]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="49" phylum="Mollusca" rank="genus">
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<emphasis id="B965DA0AFF9EFFDDFF05FB90FF46B167" box="[151,254,1122,1144]" italics="true" pageId="6" pageNumber="49">Buliminus</emphasis>
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(
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<emphasis id="B965DA0AFF9EFFDDFE9EFB91FEF6B167" box="[268,334,1123,1144]" italics="true" pageId="6" pageNumber="49">Serina</emphasis>
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)
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</taxonomicName>
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<emphasis id="B965DA0AFF9EFFDDFECFFB91FE57B167" box="[349,495,1123,1144]" italics="true" pageId="6" pageNumber="49">ser egressus—</emphasis>
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Möllendorff, 1901: 359. Kobelt, 1902: 859, pl. 124, figs. 28–30.
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<emphasis id="B965DA0AFF9EFFDDFF05FB70FED0B188" box="[151,360,1154,1175]" italics="true" pageId="6" pageNumber="49">Serina ser egressa—</emphasis>
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Yen, 1939: 86, pl. 8, fig. 17. Yen, 1942: 255. Chen, Zhou, Luo & Zhang, 2003: 442.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B5593FF9EFFDDFF05FB36FB13B3CB" pageId="6" pageNumber="49" type="materials_examined">
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<paragraph id="8BAE0618FF9EFFDDFF05FB36FAB7B1C4" blockId="6.[151,1436,1219,2000]" box="[151,1295,1219,1244]" pageId="6" pageNumber="49">
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<emphasis id="B965DA0AFF9EFFDDFF05FB36FE8DB1C3" bold="true" box="[151,309,1219,1244]" pageId="6" pageNumber="49">
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<typeStatus id="54AAB8BAFF9EFFDDFF05FB36FF6AB1C3" box="[151,210,1220,1244]" pageId="6" pageNumber="49">Type</typeStatus>
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locality.
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</emphasis>
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Kutupa (between Yulingguan and
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<taxonomicName id="4C117D9BFF9EFFDDFD28FB31FCA5B1C4" box="[698,797,1219,1243]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="wenxian">Wenxian</taxonomicName>
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County, now belongs to Nanping, Sichuan).
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</paragraph>
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<paragraph id="8BAE0618FF9EFFDDFF5BFB15FB13B3CB" blockId="6.[151,1436,1219,2000]" pageId="6" pageNumber="49">
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<emphasis id="B965DA0AFF9EFFDDFF5BFB15FE0CB01F" bold="true" box="[201,436,1255,1280]" pageId="6" pageNumber="49">Material examined.</emphasis>
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HBUMM-05426: 194 fma (24 for measurement and 2 for dissection),
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<specimenCount id="9D17CD91FF9EFFDDFB74FB15FA80B1E0" box="[1254,1336,1255,1279]" pageId="6" pageNumber="49" type="generic">41 juv.</specimenCount>
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;
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<quantity id="4CE9ABFDFF9EFFDDFAD7FB15FA24B1E0" box="[1349,1436,1255,1279]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="6" pageNumber="49" unit="m" value="1070.0">1070 m</quantity>
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a.s.l., 32°57ʹ35.5ʹ N, 104°40ʹ41.6ʹ E, foot of Yuxushan hill,
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<taxonomicName id="4C117D9BFF9EFFDDFCD6FAF9FC1FB03C" box="[836,935,1291,1315]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="wenxian">Wenxian</taxonomicName>
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||
County, Gansu,
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<collectingCountry id="F3064688FF9EFFDDFBF9FAF9FB09B03C" box="[1131,1201,1291,1315]" name="China" pageId="6" pageNumber="49">China</collectingCountry>
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,
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<date id="FFAF20D8FF9EFFDDFB2CFAF9FAF7B03C" box="[1214,1359,1291,1315]" pageId="6" pageNumber="49" value="2006-09-27">2006–IX–27</date>
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||
, colln. MW, JL, WZ & LG. HBUMM-05464: 26 fma (24 for measurement and 1 for dissection),
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<specimenCount id="9D17CD91FF9EFFDDFB2AFADDFB45B058" box="[1208,1277,1327,1351]" pageId="6" pageNumber="49" type="generic">5 juv.</specimenCount>
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||
;
|
||
<quantity id="4CE9ABFDFF9EFFDDFA9EFADDFAEEB058" box="[1292,1366,1327,1351]" metricMagnitude="2" metricUnit="m" metricValue="9.43" pageId="6" pageNumber="49" unit="m" value="943.0">943 m</quantity>
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||
a.s.l., 32°56ʹ33.5ʹ N, 104°40ʹ33.2ʹ E, bank of Bailongjiang River,
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<taxonomicName id="4C117D9BFF9EFFDDFCABFAA1FC27B074" box="[825,927,1363,1387]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="wenxian">Wenxian</taxonomicName>
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||
, Gansu,
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||
<date id="FFAF20D8FF9EFFDDFB91FAA1FB2BB074" box="[1027,1171,1363,1387]" pageId="6" pageNumber="49" value="2006-09-27">2006–IX–27</date>
|
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, colln. MW, JL, WZ & LG. HBUMM-05538: 115 fma (20 measurement and 1 dissected),
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||
<specimenCount id="9D17CD91FF9EFFDDFC2BFA85FBBAB090" box="[953,1026,1399,1423]" pageId="6" pageNumber="49" type="generic">4 juv.</specimenCount>
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; from
|
||
<quantity id="4CE9ABFDFF9EFFDDFBC5FA85FB1CB090" box="[1111,1188,1399,1423]" metricMagnitude="2" metricUnit="m" metricValue="8.29" pageId="6" pageNumber="49" unit="m" value="829.0">829 m</quantity>
|
||
a.s.l., 32°51ʹ44.7ʹ N, 104°50ʹ48.2ʹ E to
|
||
<quantity id="4CE9ABFDFF9EFFDDFEF0FA6EFE12B0AB" box="[354,426,1436,1460]" metricMagnitude="2" metricUnit="m" metricValue="7.79" pageId="6" pageNumber="49" unit="m" value="779.0">779 m</quantity>
|
||
a.s.l., 32°51ʹ49.4ʹ N, 104°50ʹ37.7ʹ E, Hendan,
|
||
<taxonomicName id="4C117D9BFF9EFFDDFC2BFA6EFBA7B0AB" box="[953,1055,1436,1460]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="wenxian">Wenxian</taxonomicName>
|
||
, Gansu,
|
||
<date id="FFAF20D8FF9EFFDDFB13FA6EFAA9B0AB" box="[1153,1297,1436,1460]" pageId="6" pageNumber="49" value="2006-09-29">2006–IX–29</date>
|
||
, colln. MW, JL, WZ & LG. HBUMM-05563: 132 fma (20 measured and 1 dissected),
|
||
<specimenCount id="9D17CD91FF9EFFDDFB95FA4DFBF5B0C8" box="[1031,1101,1471,1495]" pageId="6" pageNumber="49" type="generic">2 juv.</specimenCount>
|
||
;
|
||
<quantity id="4CE9ABFDFF9EFFDDFBCFFA4DFB10B0C8" box="[1117,1192,1471,1495]" metricMagnitude="2" metricUnit="m" metricValue="7.89" pageId="6" pageNumber="49" unit="m" value="789.0">789 m</quantity>
|
||
a.s.l., 32°51ʹ41.6ʹ N, 104°50ʹ43.1ʹ E, Hendan,
|
||
<taxonomicName id="4C117D9BFF9EFFDDFE27FA16FDA3B0E3" box="[437,539,1508,1532]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="wenxian">Wenxian</taxonomicName>
|
||
, Gansu,
|
||
<date id="FFAF20D8FF9EFFDDFD13FA16FCA9B0E3" box="[641,785,1508,1532]" pageId="6" pageNumber="49" value="2006-09-29">2006–IX–29</date>
|
||
, colln. MW, JL, WZ & LG. HBUMM-05648: 5 fma (all measured) and
|
||
<specimenCount id="9D17CD91FF9EFFDDFED0F9F5FE3BB300" box="[322,387,1543,1567]" pageId="6" pageNumber="49" type="generic">2 juv.</specimenCount>
|
||
; rocks of limestone, from the site
|
||
<quantity id="4CE9ABFDFF9EFFDDFD6DF9F5FCEBB300" box="[767,851,1543,1567]" metricMagnitude="3" metricUnit="m" metricValue="1.137" pageId="6" pageNumber="49" unit="m" value="1137.0">1137 m</quantity>
|
||
a.s.l., 33°34ʹ21.7ʹ N, 104°39ʹ0 0.3ʹ E to the site
|
||
<quantity id="4CE9ABFDFF9EFFDDFAF7F9F5FF15B35B" metricMagnitude="3" metricUnit="m" metricValue="1.151" pageId="6" pageNumber="49" unit="m" value="1151.0">1151 m</quantity>
|
||
a.s.l., 33°34ʹ28.3ʹ N, 104°39ʹ10.7ʹ E, Jiaogong Town, Wudu County, S. Gansu,
|
||
<collectingCountry id="F3064688FF9EFFDDFBA7F9DEFBC3B35B" box="[1077,1147,1580,1604]" name="China" pageId="6" pageNumber="49">China</collectingCountry>
|
||
,
|
||
<date id="FFAF20D8FF9EFFDDFB15F9DEFAB6B35B" box="[1159,1294,1580,1604]" pageId="6" pageNumber="49" value="2006-10-02">2006–X–02</date>
|
||
, colln. JL & WZ. HBUMM-05673: numerous fma (20 measured and 2 dissected) and
|
||
<specimenCount id="9D17CD91FF9EFFDDFC60F9BDFBFCB378" box="[1010,1092,1615,1639]" pageId="6" pageNumber="49" type="generic">11 juv.</specimenCount>
|
||
; eastern bank of Lianghekou, limestone,
|
||
<quantity id="4CE9ABFDFF9EFFDDFE80F986FEDEB393" box="[274,358,1652,1676]" metricMagnitude="3" metricUnit="m" metricValue="1.223" pageId="6" pageNumber="49" unit="m" value="1223.0">1223 m</quantity>
|
||
a.s.l., 33°41ʹ47.3ʹ N, 104°29ʹ10.3ʹ E, Zhouqu County, S. Gansu,
|
||
<collectingCountry id="F3064688FF9EFFDDFBA8F986FB38B393" box="[1082,1152,1652,1676]" name="China" pageId="6" pageNumber="49">China</collectingCountry>
|
||
,
|
||
<date id="FFAF20D8FF9EFFDDFB19F986FAAAB393" box="[1163,1298,1652,1676]" pageId="6" pageNumber="49" value="2006-10-02">2006–X–02</date>
|
||
, colln. JL & WZ. NHM-1902.5.13.21–22 Reg.: 2 fms. Gansu,
|
||
<collectingCountry id="F3064688FF9EFFDDFD59F965FCA9B3B0" box="[715,785,1687,1711]" name="China" pageId="6" pageNumber="49">China</collectingCountry>
|
||
. SMF-42035: 6 fms. Labelled with "
|
||
<emphasis id="B965DA0AFF9EFFDDFB20F96AFA9DB3AF" box="[1202,1317,1688,1712]" italics="true" pageId="6" pageNumber="49">Serina ser</emphasis>
|
||
Gredler".. SMF-42038: 1 fms, measured. SMF-104651: 2 fms. SMF-42039: 4 fms.. SMF-42040: 1 fms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B5593FF9EFFDDFF5BF92DFC40B3E8" box="[201,1016,1759,1784]" pageId="6" pageNumber="49" type="distribution">
|
||
<paragraph id="8BAE0618FF9EFFDDFF5BF92DFC40B3E8" blockId="6.[151,1436,1219,2000]" box="[201,1016,1759,1784]" pageId="6" pageNumber="49">
|
||
<emphasis id="B965DA0AFF9EFFDDFF5BF92DFEDAB3E7" bold="true" box="[201,354,1759,1784]" pageId="6" pageNumber="49">Distribution.</emphasis>
|
||
Gansu (
|
||
<taxonomicName id="4C117D9BFF9EFFDDFE2DF92DFD9EB3E8" box="[447,550,1759,1783]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="65" phylum="Mollusca" rank="species" species="wenxian">Wenxian</taxonomicName>
|
||
, Wudu, Zhouqu), Sichuan (
|
||
<typeStatus id="54AAB8BAFF9EFFDDFCC8F92DFC34B3E8" box="[858,908,1759,1783]" pageId="6" pageNumber="49">type</typeStatus>
|
||
locality).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B5593FF9EFFDCFF5BF8F1FCA5B796" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49" type="diagnosis">
|
||
<paragraph id="8BAE0618FF9EFFDDFF5BF8F1FBA3B298" blockId="6.[151,1436,1219,2000]" pageId="6" pageNumber="49">
|
||
<emphasis id="B965DA0AFF9EFFDDFF5BF8F1FEFBB203" bold="true" box="[201,323,1795,1820]" pageId="6" pageNumber="49">Diagnosis.</emphasis>
|
||
Shell with the most swollen part occurred at penultimate whorl and body whorl. Whorls with a narrow, defined zone below the suture,. Last whorl gradually ascending towards aperture, or ascending immediately near aperture; straight or concaved at periphery. Aperture duplicate; completely attached to body whorl. Columellar margin with one prominent tooth which is expanded inwards.
|
||
</paragraph>
|
||
<paragraph id="8BAE0618FF9EFFDCFF5BF861FCA5B796" blockId="6.[151,1436,1219,2000]" lastBlockId="7.[151,1436,151,649]" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49">
|
||
<emphasis id="B965DA0AFF9EFFDDFF5BF861FEB3B2B3" bold="true" box="[201,267,1939,1964]" pageId="6" pageNumber="49">Shell.</emphasis>
|
||
Ovate-turriform; with apex not abruptly pointed; dextral; with the most swollen part occurred at penultimate whorl to body whorl; height 10.1–11.5–14.1, diameter major 3.1–3.7–4.2, ratio of height /diameter major 2.62–3.09–3.66. Growth lines usually not very clear, or fine and clear. Whorls 8–9 1/8+–
|
||
<date id="FFAF20D8FF9FFFDCFB25FF68FB5EB5BA" box="[1207,1254,151,178]" pageId="7" pageNumber="50" value="2010-02-01">10 1/2</date>
|
||
, convex; with a narrow, defined zone below the suture. Embryonic shell 1 1/2–1 3/4—2 1/8 whorls. Postnuclear whorls smooth. Last whorl gradually ascending towards aperture, or ascending immediately near aperture; straight or concaved at periphery; with smoothed spiral peripheral depression. Aperture in a plane and straight; rounded-ovate; slightly oblique; completely attached to body whorl; armed; height 2.9–3.3–3.7, width 2.3–2.9–3.4, ratio of shell height / apertural height 2.91–3.41–4.14. Duplicate aperture indistinctly present. Palatal tooth absent. Palatal margin rounded; toothless. Peristome and parietal callus almost completely fusing except for a channel at upper insertion of peristome. Reflexed part of peristome straight and not curved backward. Parietal tooth absent. Abapertural peripheral depression on body whorl spirally extending for ca. 0.750 whorl. Palatal wall without deep depression or tooth of irregular shape. Columellar margin reflexed; with one prominent tooth which is expanded inwards; not sinuous. Columella arched. Umbilicus widely open. Shell bandless; uniformly coloured or multicoloured; chestnut at apex. Subsequent whorls evenly reddish brown or reddish brown with numerous whitish streaks. Aperture white or brownish white. Measured specimens: HBUMM-5426, HBUMM-5464, HBUMM-5538, HBUMM-5563, HBUMM-5648, HBUMM-5673, SMF-42038: 114 shells.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6E5690FF9FFFDCFF05F9CFFEAFB38E" httpUri="https://zenodo.org/record/283403/files/figure.png" pageId="7" pageNumber="50" targetBox="[178,1434,721,1537]" targetPageId="7">
|
||
<paragraph id="8BAE0618FF9FFFDCFF05F9CFFEAFB38E" blockId="7.[151,1436,1597,1681]" pageId="7" pageNumber="50">
|
||
<emphasis id="B965DA0AFF9FFFDCFF05F9CFFEACB34C" bold="true" box="[151,276,1597,1620]" pageId="7" pageNumber="50">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C117D9BFF9FFFDCFE8EF9CDFDF3B34C" authority="Sturany, 1900" authorityName="Sturany" authorityYear="1900" box="[284,587,1598,1620]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="50" phylum="Mollusca" rank="species" species="egressa">
|
||
<emphasis id="B965DA0AFF9FFFDCFE8EF9CDFE0BB34B" box="[284,435,1599,1620]" italics="true" pageId="7" pageNumber="50">Serina egressa</emphasis>
|
||
Sturany, 1900
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B965DA0AFF9FFFDCFDC4F9CCFDD7B34C" bold="true" box="[598,623,1598,1619]" pageId="7" pageNumber="50">A.</emphasis>
|
||
General view of genitalia, HBUMM-05563–specimen 21, Hendan.
|
||
<emphasis id="B965DA0AFF9FFFDCFABCF9CCFAFDB34C" bold="true" box="[1326,1349,1598,1619]" pageId="7" pageNumber="50">B.</emphasis>
|
||
Internal view of penis, HBUMM-05563–specimen 21, Hendan.
|
||
<emphasis id="B965DA0AFF9FFFDCFC97F9AEFCA6B36C" bold="true" box="[773,798,1628,1651]" pageId="7" pageNumber="50">C.</emphasis>
|
||
General view of genitalia, HBUMM-05673–specimen 2, Lianghekou.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30B5593FF9FFFD3FF5BF94FFED5B6BB" lastPageId="8" lastPageNumber="51" pageId="7" pageNumber="50" type="description">
|
||
<paragraph id="8BAE0618FF9FFFD3FF5BF94FFBE2B498" blockId="7.[151,1436,1725,2002]" lastBlockId="8.[151,1436,151,932]" lastPageId="8" lastPageNumber="51" pageId="7" pageNumber="50">
|
||
<emphasis id="B965DA0AFF9FFFDCFF5BF94FFE87B3C9" bold="true" box="[201,319,1725,1750]" pageId="7" pageNumber="50">Genitalia.</emphasis>
|
||
Vas deferens short; proximally swollen; entering epiphallus laterally with distinct demarcation. Epiphallus long; narrowed towards distal end; forming numerous loops; without epiphallic caecum. Flagellum short; conic; proximally normal; with tip blunt. Penis shortly clavate and distally enlarged; terminally entering epiphallus; thin-walled. Longitudinal pilaster more than two; fused at epiphallic pore forming velum; forming 2 Vshaped structures. V-shaped pilaster with proximal free end approaching at penial retractor insertion; without papilla formed by fused distal pilasters. Penial process absent. Penial appendix short; branched off from penis at some distance from atrium; with A-1 fused with A-2, and with distinct A-3, A-4 and A-5. A-1 long; without verge toward penial chamber. A-2 internally with longitudinal pilasters; near A-3 internally with a ring of papilla. A-3 opening into A-2 by a long needle-like papilla. A-5 short; straight. Penial retractor biramous; with arms arising from diaphragm separately; with penial branch attaching to medial penis; with appendical branch attaching to A- 1+A-2. Atrium short; with strong atrial retractor. Free oviduct moderately long; longer than vagina. Vagina short; straight. Bursa copulatrix duct very long; proximally twisted. Bursa copulatrix in normal size; well defined; shortly with stalk; without apical ligament; with short neck. Diverticle longer than reservoir; expanded. Bursa copulatrix and diverticle almost indistinguishable; forked more distally from their base. Measurement of genitalia: See
|
||
<tableCitation id="C69333A3FF90FFD3FACCFEBEFF1DB498" captionStart="TABLE 2" captionStartId="5.[151,235,1295,1316]" captionTargetBox="[151,1435,1374,1654]" captionTargetPageId="5" captionText="TABLE 2. Measurements of genitalia of some Serina species (unit: mm; measured specimens: S. egressa, 2; S. prostoma, 2; S. subser, 2; S. denticulata n. sp., 1)." httpUri="http://table.plazi.org/id/DF6E5690FF9DFFDEFF05FAFDFE4AB05B" pageId="8" pageNumber="51" tableUuid="DF6E5690FF9DFFDEFF05FAFDFE4AB05B">Table 2</tableCitation>
|
||
(HBUMM-05673–specimen 22 and HBUMM-05563–specimen 21 were measured).
|
||
</paragraph>
|
||
<paragraph id="8BAE0618FF90FFD3FF5BFE61FB6AB723" blockId="8.[151,1436,151,932]" pageId="8" pageNumber="51">
|
||
<emphasis id="B965DA0AFF90FFD3FF5BFE61FE79B4B3" bold="true" box="[201,449,403,428]" pageId="8" pageNumber="51">Taxonomic remarks.</emphasis>
|
||
Conchologically, this species most resembles
|
||
<emphasis id="B965DA0AFF90FFD3FC4AFE67FBF4B4B3" box="[984,1100,405,428]" italics="true" pageId="8" pageNumber="51">Serina ser</emphasis>
|
||
, under which
|
||
<taxonomicName id="4C117D9BFF90FFD3FB66FE67FADFB4B3" box="[1268,1383,405,428]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="51" phylum="Mollusca" rank="species" species="egressa">
|
||
<emphasis id="B965DA0AFF90FFD3FB66FE67FADFB4B3" box="[1268,1383,405,428]" italics="true" pageId="8" pageNumber="51">S. egressa</emphasis>
|
||
</taxonomicName>
|
||
was once treated as a subspecies. By principal component analysis (PCA analysis) it shows that aperture height contributes most in distinguishing
|
||
<taxonomicName id="4C117D9BFF90FFD3FD8DFE2FFD29B4EB" box="[543,657,477,500]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="51" phylum="Mollusca" rank="species" species="egressa">
|
||
<emphasis id="B965DA0AFF90FFD3FD8DFE2FFD95B4EB" box="[543,557,477,500]" italics="true" pageId="8" pageNumber="51">S</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF90FFD3FDAEFE2FFD29B4EB" box="[572,657,477,500]" italics="true" pageId="8" pageNumber="51">egressa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B965DA0AFF90FFD3FD5BFE2FFD6FB4EB" box="[713,727,477,500]" italics="true" pageId="8" pageNumber="51">S</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF90FFD3FD74FE2FFCB0B4EB" box="[742,776,477,500]" italics="true" pageId="8" pageNumber="51">ser</emphasis>
|
||
:
|
||
<taxonomicName id="4C117D9BFF90FFD3FC8AFE2FFC32B4EB" box="[792,906,477,500]" class="Gastropoda" family="Enidae" genus="Serina" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="51" phylum="Mollusca" rank="species" species="egressa">
|
||
<emphasis id="B965DA0AFF90FFD3FC8AFE2FFC9EB4EB" box="[792,806,477,500]" italics="true" pageId="8" pageNumber="51">S</emphasis>
|
||
.
|
||
<emphasis id="B965DA0AFF90FFD3FCA7FE2FFC32B4EB" box="[821,906,477,500]" italics="true" pageId="8" pageNumber="51">egressa</emphasis>
|
||
</taxonomicName>
|
||
has absolutely less aperture height than
|
||
<emphasis id="B965DA0AFF90FFD3FAC5FE2FFA2DB4EB" box="[1367,1429,477,500]" italics="true" pageId="8" pageNumber="51">S. ser</emphasis>
|
||
. Furthermore, the genitalian divergence, e.g. the presence or absence of ephiphallic caecum, suggests that they are two distinct species. Whether
|
||
<emphasis id="B965DA0AFF90FFD3FE75FDD7FD9DB723" box="[487,549,549,572]" italics="true" pageId="8" pageNumber="51">S. ser</emphasis>
|
||
has the columellar tooth is not yet known through this study.
|
||
</paragraph>
|
||
<paragraph id="8BAE0618FF90FFD3FF5BFDB5FE52B628" blockId="8.[151,1436,151,932]" pageId="8" pageNumber="51">
|
||
There are two
|
||
<typeStatus id="54AAB8BAFF90FFD3FEE1FDBAFE16B77F" box="[371,430,584,608]" pageId="8" pageNumber="51">types</typeStatus>
|
||
of shell (
|
||
<figureCitation id="132A1A9DFF90FFD3FDB2FDB5FDD7B740" box="[544,623,583,607]" captionStart="FIGURE 2. A" captionStartId="4.[151,250,1761,1784]" captionTargetBox="[151,1435,195,1740]" captionTargetId="figure@4.[151,1436,195,1740]" captionTargetPageId="4" captionText="FIGURE 2. A. Serina belae (Hilber, 1883), SMF- 42069, paratype, Bangda. B, C. Serina cathaica Gredler, 1898: B. SMF- 42046, between Yue-lin-kuan and Wen-hsien. C. NHM- 1902.5.13.19. D. Clausiliopsis szechenyi (Böttger, 1883), syntype of B. sobrina Preston, NHM- 1912.3.26.9, Gansu. E – J. Serina egressa Sturany, 1900. E. NHM- 1902.5.13.21, Gansu. F. HBUMM- 05426 – specimen 21, Wenxian. G. HBUMM- 05426 – specimen 23, Wenxian. H. SMF- 42038 (from type locality, between Yuelin-kuan and Wen-hsien). I. HBUMM- 5673 – specimen 22, Lianghekou. J. HBUMM- 5673 – specimen 23, Lianghekou. K – O. Serina prostoma (Ancey, 1884). K. NHM- 1912.6.27.69 – 71. L. A paratype of Buliminus (Holcauchen) tubios Annandale, 1924, NHM- 1923.5.24.17 – 36, Gorge of Mekong above Yangtze. M. SMF- 42050, paratype, Dshy-kow. N. HBUMM- 04296 – specimen 25, Yanjing. O. HBUMM- 04296 – specimen 24, Yanjing." httpUri="https://zenodo.org/record/283402/files/figure.png" pageId="8" pageNumber="51">Figs. 2</figureCitation>
|
||
E–G, H–J), each of which is only found in a certain population of this species. One is evenly reddish brown and another one is brown with whitish streaks (respectively "brown pattern" and "striated pattern" in the text). However, both the PCA analysis upon brown pattern shells (88 shells from HBUMM-05426, HBUMM-05464, HBUMM-05538 and HBUMM-05563) and the striated pattern shells (26 shells from HBUMM-05648, HBUMM-05673 and SMF-42038), and the same genitalia anatomy (
|
||
<figureCitation id="132A1A9DFF90FFD3FB6AFD25FAFDB7F0" box="[1272,1349,727,751]" captionStart="FIGURE 3" captionStartId="7.[151,250,1597,1620]" captionTargetBox="[178,1434,721,1537]" captionTargetId="figure@7.[151,1436,702,1576]" captionTargetPageId="7" captionText="FIGURE 3. Serina egressa Sturany, 1900. A. General view of genitalia, HBUMM- 05563 – specimen 21, Hendan. B. Internal view of penis, HBUMM- 05563 – specimen 21, Hendan. C. General view of genitalia, HBUMM- 05673 – specimen 2, Lianghekou." httpUri="https://zenodo.org/record/283403/files/figure.png" pageId="8" pageNumber="51">Figs. 3</figureCitation>
|
||
A–B of brown pattern shell and 3C of striated pattern shell) suggest that they are merely of the same species which shows divergence in shell coloration.
|
||
</paragraph>
|
||
<paragraph id="8BAE0618FF90FFD3FF5BFCB1FED5B6BB" blockId="8.[151,1436,151,932]" pageId="8" pageNumber="51">
|
||
<emphasis id="B965DA0AFF90FFD3FF5BFCB1FE00B643" bold="true" box="[201,440,835,860]" pageId="8" pageNumber="51">Ecological remarks.</emphasis>
|
||
The brown pattern specimens were found in the slate area with lower altitude range of 780 to1070 metres and the striated pattern specimens lived in the limestone regions with a higher altitude range of 1140–1230 metres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |