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<document id="D2E09F6633A01643940EC598C2188CB2" ID-CLB-Dataset="8827" ID-DOI="10.5852/ejt.2021.731.1205" ID-GBIF-Dataset="d48c7b51-c9b4-49e7-8abf-1ecee8db36a1" ID-ISSN="2118-9773" ID-Zenodo-Dep="4446916" ID-ZooBank="CBF3D6E0-9896-4852-AA9F-6DD49C950795" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610975383020" checkinUser="plazi" docAuthor="Verhaegen, Gerlien &amp; Haase, Martin" docDate="2021" docId="03903223FD75FFDFFDFCD3C08996FBF2" docLanguage="en" docName="ejt-731_71-96.pdf" docOrigin="European Journal of Taxonomy 731" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Obtusopyrgus farri Verhaegen &amp; Haase 2021, sp. nov." docType="treatment" docUuid="2D7008F8-BB41-448D-94AF-969925CECDAB" docUuidSource="ZooBank" docVersion="9" lastPageNumber="91" masterDocId="FFA94A5BFD66FFCBFF8AD12E8D70FFB7" masterDocTitle="All-inclusive descriptions of new freshwater snail taxa of the hyperdiverse family Tateidae (Gastropoda, Caenogastropoda) from the South Island of New Zealand" masterLastPageNumber="96" masterPageNumber="71" pageNumber="90" updateTime="1698924542573" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="92CABDDC1883CDF23C7D97E31924E44E">All-inclusive descriptions of new freshwater snail taxa of the hyperdiverse family Tateidae (Gastropoda, Caenogastropoda) from the South Island of New Zealand</mods:title>
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<mods:namePart id="88FE93D2EF0EAFEAEB68C1B8D7A5885B">Verhaegen, Gerlien</mods:namePart>
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<mods:affiliation id="B3D8C45A3FCD8A5600D1495682D9E156">AG Vogelwarte, Zoological Institute and Museum, University of Greifswald, Soldmannstrasse 23, Greifswald, Germany; Present address: Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Institute for Extra-cutting-edge Science and Technology Avant-garde Research (X-star), Advanced Science-Technology Research (ASTER) Program, 2 - 15, Natsushimacho, Yokosuka, Kanagawa 237 - 0061, Japan.</mods:affiliation>
<mods:nameIdentifier id="0E34655FE153686F5F115183B4052DFA" type="email">gerlienverhaegen@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="8B93419FE4FA4DA00E410784071C5861">Haase, Martin</mods:namePart>
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<mods:affiliation id="FC9FD2D4FFB3B1491B1BA8B49D246156">Vogelwarte, Zoological Institute and Museum, University of Greifswald, Soldmannstrasse 23, Greifswald, Germany.</mods:affiliation>
<mods:nameIdentifier id="5A33DBAB08F8D7ABB53E0B0C831F7CAD" type="email">martin.haase@uni-greifswald.de</mods:nameIdentifier>
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<mods:title id="39CF823CB8CB6EC42A3953C7CF9B440A">European Journal of Taxonomy</mods:title>
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<mods:part id="B25A31D36825BD6D3C9565EB2039345D">
<mods:date id="52212FD38222B0863EC1D6452457F826">2021</mods:date>
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<mods:number id="2C2CF75AB23D19259A0545EBA51DE11B">2021-01-18</mods:number>
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<treatment id="03903223FD75FFDFFDFCD3C08996FBF2" ID-DOI="http://doi.org/10.5281/zenodo.4448204" ID-GBIF-Taxon="177006639" ID-Zenodo-Dep="4448204" LSID="urn:lsid:zoobank.org:act:2D7008F8-BB41-448D-94AF-969925CECDAB" httpUri="http://treatment.plazi.org/id/03903223FD75FFDFFDFCD3C08996FBF2" lastPageId="20" lastPageNumber="91" pageId="19" pageNumber="90">
<subSubSection id="C323D0BEFD75FFD8FDFCD3C089BAFC9B" pageId="19" pageNumber="90" type="nomenclature">
<paragraph id="8B868335FD75FFD8FDFCD3C08ECEFCBE" blockId="19.[361,1226,750,848]" box="[630,958,750,777]" pageId="19" pageNumber="90">
<heading id="D0CE3459FD75FFD8FDFCD3C08ECEFCBE" box="[630,958,750,777]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="90" reason="2">
<taxonomicName id="4C39F8B6FD75FFD8FDFCD3C08E2CFCBF" ID-CoL="7SD2B" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[630,860,750,777]" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="90" phylum="Mollusca" rank="species" species="farri" status="sp. nov.">
<emphasis id="B94D5F27FD75FFD8FDFCD3C08E2CFCBF" bold="true" box="[630,860,750,777]" italics="true" pageId="19" pageNumber="90">Obtusopyrgus farri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD75FFD8FCEED3C18ECEFCBE" box="[868,958,751,777]" pageId="19" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD75FFD8FD98D21B8952FCF8" box="[530,1058,821,848]" pageId="19" pageNumber="90" type="description">
<paragraph id="8B868335FD75FFD8FD98D21B8952FCF8" blockId="19.[361,1226,750,848]" box="[530,1058,821,848]" pageId="19" pageNumber="90">
<figureCitation id="13029FB0FD75FFD8FD98D21B8F2BFCF8" box="[530,603,821,847]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[189,1398,1043,1871]" captionTargetId="figure-163@9.[189,1400,1043,1871]" captionTargetPageId="9" captionText="Fig. 4. Types. A–B. Catapyrgus jami sp. nov. C–D. Opacuincola lisannea sp. nov. (D from northern locality). E–F. Op. gretathunbergae sp. nov. G–H. Op. mete kahurangi subsp. nov. I–J. Obtusopyrgus farri sp. nov. A, C, E, G, I, holotypes, rest paratypes." figureDoi="http://doi.org/10.5281/zenodo.4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="19" pageNumber="90">Figs 4</figureCitation>
I–J,
<figureCitation id="13029FB0FD75FFD8FD07D21B8FEAFCE7" box="[653,666,821,848]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,962,1876]" captionTargetId="figure-214@10.[189,1399,962,1876]" captionTargetPageId="10" captionText="Fig. 5. Shells. SEM photographs of paratypes. A. Catapyrgus jami sp. nov. B–C. Opacuincola lisannea sp. nov. (B, topotype; C, northern locality).D–E. Op. gretathunbergae sp. nov.F–G. Op. mete kahurangi subsp. nov. H–I. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446932" httpUri="https://zenodo.org/record/4446932/files/figure.png" pageId="19" pageNumber="90">5</figureCitation>
H–I,
<figureCitation id="13029FB0FD75FFD8FD52D21B8F96FCF8" box="[728,742,821,847]" captionStart="Fig" captionStartId="12.[189,232,1017,1043]" captionTargetBox="[190,1399,266,946]" captionTargetId="figure-15@12.[190,1399,266,946]" captionTargetPageId="12" captionText="Fig. 6. Protoconch. SEM photographs of paratypes. A. Catapyrgus jami sp. nov. B–C. Opacuincola lisannea sp. nov. (B, topotype; C, northern locality). D. Op. gretathunbergae sp. nov. E. Op. mete kahurangi subsp. nov. F. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446934" httpUri="https://zenodo.org/record/4446934/files/figure.png" pageId="19" pageNumber="90">6</figureCitation>
F,
<figureCitation id="13029FB0FD75FFD8FC89D21B8E61FCF8" box="[771,785,821,847]" captionStart="Fig" captionStartId="13.[189,232,1964,1990]" captionTargetBox="[189,1398,1079,1914]" captionTargetId="figure-275@13.[189,1398,1079,1914]" captionTargetPageId="13" captionText="Fig. 8. Radula. SEM photographs. A. Catapyrgus jami sp. nov. B. Opacuincola lisannea sp. nov. (from type locality). C. Op. gretathunbergae sp. nov. D. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446940" httpUri="https://zenodo.org/record/4446940/files/figure.png" pageId="19" pageNumber="90">8</figureCitation>
D,
<figureCitation id="13029FB0FD75FFD8FCBCD21B8E22FCF8" box="[822,850,821,847]" captionStart="Fig" captionStartId="16.[188,231,1927,1953]" captionTargetBox="[227,1361,975,1851]" captionTargetId="figure-267@16.[227,1361,975,1851]" captionTargetPageId="16" captionText="Fig. 11. Penis. SEM photographs. A. Opacuincola lisannea sp. nov. (from type locality). B. Op. gretathunbergae sp. nov.C. Op. mete kahurangi subsp. nov.D. Obtusopyrgus farri sp. nov.Abbreviations: e = eye; l = penial lobe; s = snout; t = tentacle." figureDoi="http://doi.org/10.5281/zenodo.4446946" httpUri="https://zenodo.org/record/4446946/files/figure.png" pageId="19" pageNumber="90">11</figureCitation>
D,
<figureCitation id="13029FB0FD75FFD8FCFCD21B8EE6FCF8" box="[886,918,821,847]" captionStart="Fig" captionStartId="20.[189,232,1928,1954]" captionTargetBox="[485,1118,1291,1849]" captionTargetId="figure-321@20.[406,1162,1251,1866]" captionTargetPageId="20" captionText="Fig. 14. Female genitalia of Obtusopyrgus farri sp. nov. µCT scans. Abbreviations: ag = albumen gland; bc = bursa copulatrix; cg = capsule gland; od = oviduct; ov = ovary; rs = receptaculum seminis; vc = ventral channel. Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4446952" httpUri="https://zenodo.org/record/4446952/files/figure.png" pageId="19" pageNumber="90">14</figureCitation>
; Tables
<tableCitation id="C6BBB68EFD75FFD8FC7CD21B8970FCF8" box="[1014,1024,821,847]" captionStart-0="Table 1" captionStart-1="Table 1" captionStartId-0="5.[189,258,265,291]" captionStartId-1="6.[189,258,265,291]" captionTargetBox-0="[200,1288,480,1976]" captionTargetBox-1="[200,1288,362,975]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Table 1 (continued on next page). Morphometrics of new taxa. A–E following the names refer to the localities in Figure 1. Abbreviations: AH = aperture height; AW = aperture width; BWW = bodywhorl width; CV = coefficient of variation corrected for unequal sample sizes; max = maximum; min = minimum; SD = standard deviation; SH = shell height; SW = shell width; W = number of whorls. Measurements in mm." captionText-1="Table 1. (continued)." httpUri-0="http://table.plazi.org/id/DF46D3BDFD63FFCEFF37D0278CB5FE07" httpUri-1="http://table.plazi.org/id/DF46D3BDFD60FFCDFF37D0278CC6FE93" pageId="19" pageNumber="90" tableUuid-0="DF46D3BDFD63FFCEFF37D0278CB5FE07" tableUuid-1="DF46D3BDFD60FFCDFF37D0278CC6FE93">1</tableCitation>
–
<tableCitation id="C6BBB68EFD75FFD8FB98D21B8952FCF8" box="[1042,1058,821,847]" captionStart="Table 2" captionStartId="7.[189,258,265,291]" captionTargetBox="[200,1382,478,1976]" captionTargetPageId="7" captionText="Table 2. Diagnostic alignment positions. All characters are of type 1, i.e., they are fixed in a particular taxon. Positions refer to the alignments of COI and 16S rRNA in Electronic Supplements 1 and 2, respectively. Position 382 in bold is the only diagnostic position for Opacuincola gretathunbergae across all samples of the genus. Therefore, it was also compared to its sister species Op. ngatapuna. In Ob. farri, sequencing of COI failed." httpUri="http://table.plazi.org/id/DF46D3BDFD61FFCCFF37D0278F14FE07" pageId="19" pageNumber="90" tableUuid="DF46D3BDFD61FFCCFF37D0278F14FE07">2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD75FFD8FF37D25889DFFBFA" pageId="19" pageNumber="90" type="diagnosis">
<paragraph id="8B868335FD75FFD8FF37D2588C46FC27" blockId="19.[189,310,886,912]" box="[189,310,886,912]" pageId="19" pageNumber="90">
<heading id="D0CE3459FD75FFD8FF37D2588C46FC27" bold="true" box="[189,310,886,912]" fontSize="11" level="3" pageId="19" pageNumber="90" reason="3">
<emphasis id="B94D5F27FD75FFD8FF37D2588C46FC27" bold="true" box="[189,310,886,912]" pageId="19" pageNumber="90">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD75FFD8FF37D28889DFFBFA" blockId="19.[189,1400,933,1101]" pageId="19" pageNumber="90">
In the new species, the central tooth of the radula has more cusps on the edge and less on the basis than in the only other known representative of the genus,
<taxonomicName id="4C39F8B6FD75FFD8FCB2D2E78EB1FC54" authorityName="Haase" authorityYear="2008" box="[824,961,969,995]" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="90" phylum="Mollusca" rank="species" species="alpinus">
<emphasis id="B94D5F27FD75FFD8FCB2D2E78EB1FC54" box="[824,961,969,995]" italics="true" pageId="19" pageNumber="90">Ob. alpinus</emphasis>
</taxonomicName>
. The bursa copulatrix is smaller and more elongate compared to the larger, more globular one in the latter. As a consequence, the seminal receptacle reaches far behind the bursa in
<taxonomicName id="4C39F8B6FD75FFD8FD49D5218E5FFB9D" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[707,815,1039,1066]" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="90" phylum="Mollusca" rank="species" species="farri" status="sp. nov.">
<emphasis id="B94D5F27FD75FFD8FD49D5218E5FFB9D" box="[707,815,1039,1066]" italics="true" pageId="19" pageNumber="90">Ob. farri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD75FFD8FCB0D53E8EEAFB9D" box="[826,922,1040,1066]" pageId="19" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
, whereas in
<taxonomicName id="4C39F8B6FD75FFD8FBBDD52189B3FB9E" authorityName="Haase" authorityYear="2008" box="[1079,1219,1039,1065]" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="90" phylum="Mollusca" rank="species" species="alpinus">
<emphasis id="B94D5F27FD75FFD8FBBDD52189B3FB9E" box="[1079,1219,1039,1065]" italics="true" pageId="19" pageNumber="90">Ob. alpinus</emphasis>
</taxonomicName>
it lies entirely against the bursa. These species differ at five diagnostic alignment positions of
<typeStatus id="54823D97FD75FFD8FBD5D51D89E2FBFA" box="[1119,1170,1075,1101]" pageId="19" pageNumber="90">type</typeStatus>
1.
</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD75FFD8FF37D55B8F91FA90" pageId="19" pageNumber="90" type="etymology">
<paragraph id="8B868335FD75FFD8FF37D55B8C34FB38" blockId="19.[189,324,1141,1167]" box="[189,324,1141,1167]" pageId="19" pageNumber="90">
<heading id="D0CE3459FD75FFD8FF37D55B8C34FB38" bold="true" box="[189,324,1141,1167]" fontSize="11" level="3" pageId="19" pageNumber="90" reason="3">
<emphasis id="B94D5F27FD75FFD8FF37D55B8C34FB38" bold="true" box="[189,324,1141,1167]" pageId="19" pageNumber="90">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD75FFD8FF37D58D8F91FA90" blockId="19.[189,1401,1187,1320]" pageId="19" pageNumber="90">
<taxonomicName id="4C39F8B6FD75FFD8FF37D58D8CD3FB0A" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[189,419,1187,1213]" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="90" phylum="Mollusca" rank="species" species="farri" status="sp. nov.">
<emphasis id="B94D5F27FD75FFD8FF37D58D8CD3FB0A" box="[189,419,1187,1213]" italics="true" pageId="19" pageNumber="90">Obtusopyrgus farri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD75FFD8FE25D58A8F7EFB09" box="[431,526,1188,1214]" pageId="19" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
is named after Gareth Farr, acclaimed
<collectingCountry id="F32EC3A5FD75FFD8FC72D58A89EAFB0A" box="[1016,1178,1187,1214]" name="New Zealand" pageId="19" pageNumber="90">New Zealand</collectingCountry>
percussionist and composer integrating non-European music styles including Maori music into Western classical music resulting in the most fascinating and colorful synthesis of different musical expressions. His alter ego, the drag queen Lilith LaCroix, is also colorful.
</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD75FFD8FF37D46189C3F9D0" pageId="19" pageNumber="90" type="materials_examined">
<paragraph id="8B868335FD75FFD8FF37D4618CDAFADE" blockId="19.[189,426,1359,1385]" box="[189,426,1359,1385]" pageId="19" pageNumber="90">
<heading id="D0CE3459FD75FFD8FF37D4618CDAFADE" bold="true" box="[189,426,1359,1385]" fontSize="11" level="3" pageId="19" pageNumber="90" reason="3">
<emphasis id="B94D5F27FD75FFD8FF37D4618CDAFADE" bold="true" box="[189,426,1359,1385]" pageId="19" pageNumber="90">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD75FFD8FF60D4508910F9B4" blockId="19.[189,1398,1406,1539]" pageId="19" pageNumber="90">
<materialsCitation id="3B518968FD75FFD8FF60D450892CF9B4" ID-GBIF-Occurrence="3019435309" collectedFrom="on leaves, stones, woody debris in small stream crossing track" collectingDate="2016-03-13" collectionCode="NMNZ" collectorName="G. Verhaegen &amp; M. Haase" country="New Zealand" county="Lakes National Park" latitude="-41.825195" location="Lake Rotoiti" longLatPrecision="1" longitude="172.83058" municipality="Lake Rotoiti" pageId="19" pageNumber="90" specimenCode="NMNZ.M.330195" specimenCount="1" stateProvince="Nelson Lakes National Park" typeStatus="holotype">
<emphasis id="B94D5F27FD75FFD8FF60D4508C2CFA2F" bold="true" box="[234,348,1406,1432]" pageId="19" pageNumber="90">
<typeStatus id="54823D97FD75FFD8FF60D4508C2CFA2F" box="[234,348,1406,1432]" pageId="19" pageNumber="90" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13029FB0FD75FFD8FEE6D4508CCDFA2F" box="[364,445,1406,1432]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[189,1398,1043,1871]" captionTargetId="figure-163@9.[189,1400,1043,1871]" captionTargetPageId="9" captionText="Fig. 4. Types. A–B. Catapyrgus jami sp. nov. C–D. Opacuincola lisannea sp. nov. (D from northern locality). E–F. Op. gretathunbergae sp. nov. G–H. Op. mete kahurangi subsp. nov. I–J. Obtusopyrgus farri sp. nov. A, C, E, G, I, holotypes, rest paratypes." figureDoi="http://doi.org/10.5281/zenodo.4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="19" pageNumber="90">Fig. 4I</figureCitation>
)
<collectingCountry id="F32EC3A5FD75FFD8FF37D48C8CE2FA0C" box="[189,402,1441,1468]" name="New Zealand" pageId="19" pageNumber="90">NEW ZEALAND</collectingCountry>
•
<collectingRegion id="49FD4DD7FD75FFD8FE3AD48F8E72FA0C" box="[432,770,1441,1467]" pageId="19" pageNumber="90">Nelson Lakes National Park</collectingRegion>
,
<location id="8EE6D5EEFD75FFD8FC9AD48F8ED8FA0C" LSID="urn:lsid:plazi:treatment:03903223FD75FFDFFDFCD3C08996FBF2:8EE6D5EEFD75FFD8FC9AD48F8ED8FA0C" box="[784,936,1441,1467]" country="New Zealand" county="Lakes National Park" latitude="-41.825195" longLatPrecision="1" longitude="172.83058" municipality="Lake Rotoiti" name="Lake Rotoiti" pageId="19" pageNumber="90" stateProvince="Nelson Lakes National Park">Lake Rotoiti</location>
,
<locationDeviation id="9A349A17FD75FFD8FC3FD48F8CEDFA68" pageId="19" pageNumber="90">small stream crossing Lakeside Track close to lake shore</locationDeviation>
;
<geoCoordinate id="EE0DE5F2FD75FFD8FE20D4EB8F3DFA68" box="[426,589,1475,1503]" degrees="41" direction="south" minutes="49" orientation="latitude" pageId="19" pageNumber="90" precision="1" seconds="30.7" value="-41.825195">41°49ʹ30.7ʺ S</geoCoordinate>
,
<geoCoordinate id="EE0DE5F2FD75FFD8FDD0D4ED8E60FA68" box="[602,784,1475,1503]" degrees="172" direction="east" minutes="49" orientation="longitude" pageId="19" pageNumber="90" precision="1" seconds="50.1" value="172.83058">172°49ʹ50.1ʺ E</geoCoordinate>
;
<date id="FF87A5F5FD75FFD8FC97D4EB8ECFFA68" box="[797,959,1477,1503]" pageId="19" pageNumber="90" value="2016-03-13">
<collectingDate id="EFC35C1DFD75FFD8FC97D4EB8ECFFA68" box="[797,959,1477,1503]" pageId="19" pageNumber="90" value="2016-03-13">13 Mar. 2016</collectingDate>
</date>
;
<collectorName id="26CCE6E3FD75FFD8FC46D4EB891BFA68" box="[972,1131,1477,1503]" pageId="19" pageNumber="90">G. Verhaegen</collectorName>
and
<collectorName id="26CCE6E3FD75FFD8FB2FD4EB8866FA68" box="[1189,1302,1477,1503]" pageId="19" pageNumber="90">M. Haase</collectorName>
leg.;
<collectedFrom id="3825C0C8FD75FFD8FAD3D4EB8E05F9B5" pageId="19" pageNumber="90">on leaves, stones, woody debris in small stream crossing track</collectedFrom>
;
<specimenCode id="DB9F2B4EFD75FFD8FC0BD4C6892CF9B4" box="[897,1116,1512,1539]" pageId="19" pageNumber="90">NMNZ.M.330195</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B868335FD75FFD8FF60D70489C3F9D0" blockId="19.[189,1203,1577,1640]" pageId="19" pageNumber="90">
<materialsCitation id="3B518968FD75FFD8FF60D70489DFF9D0" ID-GBIF-Occurrence="3019435305" collectionCode="NMNZ" country="New Zealand" county="Lakes National Park" location="NEW ZEALAND" municipality="Lake Rotoiti" pageId="19" pageNumber="90" specimenCode="NMNZ.M.330196" specimenCount="21" stateProvince="Nelson" typeStatus="paratype">
<emphasis id="B94D5F27FD75FFD8FF60D7048C17F9F3" bold="true" box="[234,359,1578,1604]" pageId="19" pageNumber="90">
<typeStatus id="54823D97FD75FFD8FF60D7048C17F9F3" box="[234,359,1578,1604]" pageId="19" pageNumber="90" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<figureCitation id="13029FB0FD75FFD8FEFDD7048CA0F9F3" box="[375,464,1578,1604]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[189,1398,1043,1871]" captionTargetId="figure-163@9.[189,1400,1043,1871]" captionTargetPageId="9" captionText="Fig. 4. Types. A–B. Catapyrgus jami sp. nov. C–D. Opacuincola lisannea sp. nov. (D from northern locality). E–F. Op. gretathunbergae sp. nov. G–H. Op. mete kahurangi subsp. nov. I–J. Obtusopyrgus farri sp. nov. A, C, E, G, I, holotypes, rest paratypes." figureDoi="http://doi.org/10.5281/zenodo.4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="19" pageNumber="90">Figs 4J</figureCitation>
;
<figureCitation id="13029FB0FD75FFD8FE56D7078C99F9F3" box="[476,489,1577,1604]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,962,1876]" captionTargetId="figure-214@10.[189,1399,962,1876]" captionTargetPageId="10" captionText="Fig. 5. Shells. SEM photographs of paratypes. A. Catapyrgus jami sp. nov. B–C. Opacuincola lisannea sp. nov. (B, topotype; C, northern locality).D–E. Op. gretathunbergae sp. nov.F–G. Op. mete kahurangi subsp. nov. H–I. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446932" httpUri="https://zenodo.org/record/4446932/files/figure.png" pageId="19" pageNumber="90">5</figureCitation>
H–I)
<collectingCountry id="F32EC3A5FD75FFD8FF37D7608CE1F9D0" box="[189,401,1613,1640]" name="New Zealand" pageId="19" pageNumber="90">NEW ZEALAND</collectingCountry>
•
<specimenCount id="9D3F48BCFD75FFD8FE20D7638CB7F9D0" box="[426,455,1613,1639]" pageId="19" pageNumber="90">21</specimenCount>
specs; same collection data as for holotype;
<specimenCode id="DB9F2B4EFD75FFD8FC5ED76389DFF9D0" box="[980,1199,1613,1639]" pageId="19" pageNumber="90">NMNZ.M.330196</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD75FFDFFF37D7A08E77FCF0" lastPageId="20" lastPageNumber="91" pageId="19" pageNumber="90" type="description">
<paragraph id="8B868335FD75FFD8FF37D7A08C3EF91F" blockId="19.[189,334,1678,1704]" box="[189,334,1678,1704]" pageId="19" pageNumber="90">
<heading id="D0CE3459FD75FFD8FF37D7A08C3EF91F" bold="true" box="[189,334,1678,1704]" fontSize="11" level="3" pageId="19" pageNumber="90" reason="3">
<emphasis id="B94D5F27FD75FFD8FF37D7A08C3EF91F" bold="true" box="[189,334,1678,1704]" pageId="19" pageNumber="90">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD75FFD8FF37D79089A5F8F5" blockId="19.[189,1401,1725,1858]" pageId="19" pageNumber="90">
SHELL (
<figureCitation id="13029FB0FD75FFD8FE9FD7938C10F96F" box="[277,352,1725,1752]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[189,1398,1043,1871]" captionTargetId="figure-163@9.[189,1400,1043,1871]" captionTargetPageId="9" captionText="Fig. 4. Types. A–B. Catapyrgus jami sp. nov. C–D. Opacuincola lisannea sp. nov. (D from northern locality). E–F. Op. gretathunbergae sp. nov. G–H. Op. mete kahurangi subsp. nov. I–J. Obtusopyrgus farri sp. nov. A, C, E, G, I, holotypes, rest paratypes." figureDoi="http://doi.org/10.5281/zenodo.4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="19" pageNumber="90">Figs 4</figureCitation>
I–J, 5H–I). Blunt-ovate to pupiform, about 1.7 times as high as than wide, light brown, translucent with brown periostracum; protoconch almost smooth, comprising up to 1 whorl (
<figureCitation id="13029FB0FD75FFD8FA85D7CF881AF94B" box="[1295,1386,1761,1788]" captionStart="Fig" captionStartId="12.[189,232,1017,1043]" captionTargetBox="[190,1399,266,946]" captionTargetId="figure-15@12.[190,1399,266,946]" captionTargetPageId="12" captionText="Fig. 6. Protoconch. SEM photographs of paratypes. A. Catapyrgus jami sp. nov. B–C. Opacuincola lisannea sp. nov. (B, topotype; C, northern locality). D. Op. gretathunbergae sp. nov. E. Op. mete kahurangi subsp. nov. F. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446934" httpUri="https://zenodo.org/record/4446934/files/figure.png" pageId="19" pageNumber="90">Fig. 6F</figureCitation>
); entire shell with 3.625 to 4.25 whorls, teleoconch initially with very fine longitudinal ridges, then without structure apart from growth lines; umbilicus narrow; aperture orthocline, as high as wide.
</paragraph>
<paragraph id="8B868335FD75FFD8FF37D64789F7F833" blockId="19.[189,1159,1897,1924]" box="[189,1159,1897,1924]" pageId="19" pageNumber="90">OPERCULUM. Yellow to light orange, paucispiral; nucleus submarginal, without peg.</paragraph>
<paragraph id="8B868335FD75FFD8FF37D6858EFBF85F" blockId="19.[189,1399,1963,2024]" pageId="19" pageNumber="90">
EXTERNAL FEATURES (
<figureCitation id="13029FB0FD75FFD8FE25D6858C89F872" box="[431,505,1963,1989]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[189,1398,1043,1871]" captionTargetId="figure-163@9.[189,1400,1043,1871]" captionTargetPageId="9" captionText="Fig. 4. Types. A–B. Catapyrgus jami sp. nov. C–D. Opacuincola lisannea sp. nov. (D from northern locality). E–F. Op. gretathunbergae sp. nov. G–H. Op. mete kahurangi subsp. nov. I–J. Obtusopyrgus farri sp. nov. A, C, E, G, I, holotypes, rest paratypes." figureDoi="http://doi.org/10.5281/zenodo.4446930" httpUri="https://zenodo.org/record/4446930/files/figure.png" pageId="19" pageNumber="90">Figs 4</figureCitation>
I–J). Epidermis with irregular, large pigment blotches; eyes well developed and entirely pigmented; tentacles without particular ciliation.
</paragraph>
<paragraph id="8B868335FD72FFDFFF37D0278837FE94" blockId="20.[189,1351,265,291]" box="[189,1351,265,291]" pageId="20" pageNumber="91">MANTLE CAVITY (n = 4). 8–11 ctenidial filaments; osphradium ovate-elongate, behind middle of gill.</paragraph>
<paragraph id="8B868335FD72FFDFFF37D0658C4FFE1C" blockId="20.[189,1399,330,427]" pageId="20" pageNumber="91">
DIGESTIVE SYSTEM. Radula has formula R 5 1 5/2–3 2-3, L 5–7 1 5–7, M1 23–28, M2 25–28 (
<figureCitation id="13029FB0FD72FFDFFA83D0648818FED2" box="[1289,1384,330,357]" captionStart="Fig" captionStartId="13.[189,232,1964,1990]" captionTargetBox="[189,1398,1079,1914]" captionTargetId="figure-275@13.[189,1398,1079,1914]" captionTargetPageId="13" captionText="Fig. 8. Radula. SEM photographs. A. Catapyrgus jami sp. nov. B. Opacuincola lisannea sp. nov. (from type locality). C. Op. gretathunbergae sp. nov. D. Obtusopyrgus farri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4446940" httpUri="https://zenodo.org/record/4446940/files/figure.png" pageId="20" pageNumber="91">Fig. 8D</figureCitation>
); stomach without caecum; rectal loop pointing left in roof of mantle cavity, in males more distinct than in females.
</paragraph>
<paragraph id="8B868335FD72FFDFFF37D0FD8E9AFD2A" blockId="20.[188,1400,466,669]" pageId="20" pageNumber="91">
FEMALE GENITALIA (n = 4;
<figureCitation id="13029FB0FD72FFDFFE7AD0FC8F39FE5B" box="[496,585,466,493]" captionStart="Fig" captionStartId="20.[189,232,1928,1954]" captionTargetBox="[485,1118,1291,1849]" captionTargetId="figure-321@20.[406,1162,1251,1866]" captionTargetPageId="20" captionText="Fig. 14. Female genitalia of Obtusopyrgus farri sp. nov. µCT scans. Abbreviations: ag = albumen gland; bc = bursa copulatrix; cg = capsule gland; od = oviduct; ov = ovary; rs = receptaculum seminis; vc = ventral channel. Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4446952" httpUri="https://zenodo.org/record/4446952/files/figure.png" pageId="20" pageNumber="91">Fig. 14</figureCitation>
). Ovary lobate sac starting&gt; 1.5 whorls below apex, comprising ca 0.3 whorl and not reaching stomach; renal oviduct first coiling 180° clockwise, then 270° counterclockwise; one distal, large receptaculum seminis reaching far behind bursa copulatrix; bursa copulatrix an elongate sac lying behind albumen gland, bursal duct entering anteriorly; ovoviviparous, brooding at least three embryos in pallial oviduct, pallial oviduct as brood pouch, albumen gland has about third of length of capsule gland, the latter histologically uniform in CT scans.
</paragraph>
<paragraph id="8B868335FD72FFDFFF37D3EA8E77FCF0" blockId="20.[189,1401,708,839]" pageId="20" pageNumber="91">
MALE GENITALIA (n = 1). Testis lobate sac, starting ca 1 whorl below apex, comprising up to 1 whorl, reaching stomach; vesicula seminalis coils along anterior half of testis; proximal vas deferens inserts close to middle of kidney-shaped prostate, distal vas deferens leaving anteriorly; penis massive, broad, short, pointed tip, without appendages (
<figureCitation id="13029FB0FD72FFDFFD04D2038F89FCF0" box="[654,761,813,839]" captionStart="Fig" captionStartId="16.[188,231,1927,1953]" captionTargetBox="[227,1361,975,1851]" captionTargetId="figure-267@16.[227,1361,975,1851]" captionTargetPageId="16" captionText="Fig. 11. Penis. SEM photographs. A. Opacuincola lisannea sp. nov. (from type locality). B. Op. gretathunbergae sp. nov.C. Op. mete kahurangi subsp. nov.D. Obtusopyrgus farri sp. nov.Abbreviations: e = eye; l = penial lobe; s = snout; t = tentacle." figureDoi="http://doi.org/10.5281/zenodo.4446946" httpUri="https://zenodo.org/record/4446946/files/figure.png" pageId="20" pageNumber="91">Fig. 11D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C323D0BEFD72FFDFFF37D2418996FBF2" pageId="20" pageNumber="91" type="discussion">
<paragraph id="8B868335FD72FFDFFF37D2418C5FFC3E" blockId="20.[189,303,879,905]" box="[189,303,879,905]" pageId="20" pageNumber="91">
<heading id="D0CE3459FD72FFDFFF37D2418C5FFC3E" bold="true" box="[189,303,879,905]" fontSize="11" level="3" pageId="20" pageNumber="91" reason="3">
<emphasis id="B94D5F27FD72FFDFFF37D2418C5FFC3E" bold="true" box="[189,303,879,905]" pageId="20" pageNumber="91">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B868335FD72FFDFFF37D2B38996FBF2" blockId="20.[189,1399,925,1093]" pageId="20" pageNumber="91">
The new species is slightly larger than
<taxonomicName id="4C39F8B6FD72FFDFFD00D2B38E63FC00" authorityName="Haase" authorityYear="2008" box="[650,787,925,951]" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="91" phylum="Mollusca" rank="species" species="alpinus">
<emphasis id="B94D5F27FD72FFDFFD00D2B38E63FC00" box="[650,787,925,951]" italics="true" pageId="20" pageNumber="91">Ob. alpinus</emphasis>
</taxonomicName>
(shell height: Mann-Whitney U-test: z = 2.869, p = 0.004), but in terms of shape, they cannot be distinguished (shell height/shell width: Mann-Whitney U-test: z = 0.764, p = 0.445). The genetic and phylogenetic distinction of both species of
<taxonomicName id="4C39F8B6FD72FFDFFB59D2CA8807FC49" authorityName="Haase" authorityYear="2008" box="[1235,1399,996,1022]" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="91" phylum="Mollusca" rank="genus">
<emphasis id="B94D5F27FD72FFDFFB59D2CA8807FC49" box="[1235,1399,996,1022]" italics="true" pageId="20" pageNumber="91">Obtusopyrgus</emphasis>
</taxonomicName>
was only based on 16S as sequencing of COI failed (
<figureCitation id="13029FB0FD72FFDFFCC7D5298EE9FB96" box="[845,921,1031,1057]" captionStart="Fig" captionStartId="4.[189,232,1927,1953]" captionTargetBox="[223,1363,820,1877]" captionTargetId="figure-161@4.[186,1401,746,1891]" captionTargetPageId="4" captionText="Fig. 2. Phylogenetic analysis. Maximum likelihood tree with bootstrap support&gt; 50 / posterior probabilities from Bayesian analysis&gt; 0.50 for nodes. New taxa in bold. Letters A-E after new taxa refer to localities in Figure 1." figureDoi="http://doi.org/10.5281/zenodo.4446924" httpUri="https://zenodo.org/record/4446924/files/figure.png" pageId="20" pageNumber="91">Fig. 2</figureCitation>
). There were five diagnostic characters (
<tableCitation id="C6BBB68EFD72FFDFFF4FD5048C50FBF3" box="[197,288,1066,1092]" captionStart="Table 2" captionStartId="7.[189,258,265,291]" captionTargetPageId="7" captionText="Table 2. Diagnostic alignment positions. All characters are of type 1, i.e., they are fixed in a particular taxon. Positions refer to the alignments of COI and 16S rRNA in Electronic Supplements 1 and 2, respectively. Position 382 in bold is the only diagnostic position for Opacuincola gretathunbergae across all samples of the genus. Therefore, it was also compared to its sister species Op. ngatapuna. In Ob. farri, sequencing of COI failed." httpUri="http://table.plazi.org/id/DF46D3BDFD61FFCCFF37D0278F14FE07" pageId="20" pageNumber="91" tableUuid="DF46D3BDFD61FFCCFF37D0278F14FE07">Table 2</tableCitation>
). The well-developed eyes indicate that
<taxonomicName id="4C39F8B6FD72FFDFFD73D5048E12FBF3" authority="Verhaegen &amp; Haase, 2021" authorityName="Verhaegen &amp; Haase" authorityYear="2021" box="[761,866,1066,1092]" class="Gastropoda" family="Tateidae" genus="Obtusopyrgus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="91" phylum="Mollusca" rank="species" species="farri" status="sp. nov.">
<emphasis id="B94D5F27FD72FFDFFD73D5048E12FBF3" box="[761,866,1066,1092]" italics="true" pageId="20" pageNumber="91">Ob. farri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EE25CFD72FFDFFCE3D5058EB4FBF2" box="[873,964,1067,1093]" pageId="20" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
inhabits epigean waters.
</paragraph>
</subSubSection>
</treatment>
</document>
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