treatments-xml/data/6E/30/87/6E30879DFFC7FFC333B99342EF98FB50.xml
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<mods:title id="B987B23C63EB468ED1D3AF9AE61285D6">A new species of the genus Clastopus Fairmaire, 1898 (Coleoptera: Tenebrionidae: Pedinini) from Madagascar</mods:title>
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<mods:namePart id="3308C6DEEEE0FE37B7CC84B88E079F36">Iwan, Dariusz</mods:namePart>
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Genus
<taxonomicName id="21994D08FFC7FFC333C39342EF48FE53" ID-CoL="3QJP" ID-ENA="457787" authority="Fairmaire, 1898" authorityName="Fairmaire" authorityYear="1898" box="[237,564,365,391]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="genus">
<emphasis id="D4EDEA99FFC7FFC333C39342EC18FE53" bold="true" box="[237,356,365,391]" italics="true" pageId="1" pageNumber="61">Clastopus</emphasis>
<bibRefCitation id="82084B7AFFC7FFC332429342EF48FE53" author="Fairmaire" box="[364,564,365,391]" pageId="1" pageNumber="61" refString="Fairmaire, L. (1898) Descriptions de Coleopteres de d'Asie et de Malaisie. Annales de la Societe Entomologique de France, 67: 382 - 400." type="journal article" year="1898">Fairmaire, 1898</bibRefCitation>
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<taxonomicName id="21994D08FFC7FFC333B99399ECA6FE1F" ID-CoL="3QJP" authority="Fairmaire, 1898: 407" authorityName="Fairmaire" authorityPageNumber="407" authorityYear="1898" box="[151,474,437,460]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="genus">
<emphasis id="D4EDEA99FFC7FFC333B99399ED81FE18" box="[151,253,438,460]" italics="true" pageId="1" pageNumber="61">Clastopus</emphasis>
<treatmentCitation id="6738109AFFC7FFC3322C939AECA6FE1F" author="Fairmaire" box="[258,474,437,459]" page="407" pageId="1" pageNumber="61" year="1898">
<bibRefCitation id="82084B7AFFC7FFC3322C939AECDBFE1F" author="Fairmaire" box="[258,423,437,459]" pageId="1" pageNumber="61" refString="Fairmaire, L. (1898) Descriptions de Coleopteres de d'Asie et de Malaisie. Annales de la Societe Entomologique de France, 67: 382 - 400." type="journal article" year="1898">Fairmaire, 1898</bibRefCitation>
: 407
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</taxonomicName>
.—
<treatmentCitation id="6738109AFFC7FFC332D4939AEFCDFE1F" author="Gebien" box="[506,689,437,459]" page="272" pageId="1" pageNumber="61" year="1910">
<bibRefCitation id="82084B7AFFC7FFC332D4939AEF03FE1F" author="Gebien" box="[506,639,437,459]" pageId="1" pageNumber="61" refString="Gebien, H. (1910) Tenebrionidae I. In: Junk, W. &amp; Schenkling, S. (eds.) Coleopterorum Catalogus, 18, pp. 167 - 354, Berlin." type="journal article" year="1910">Gebien 1910</bibRefCitation>
: 272
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, 1938: 291,
<treatmentCitation id="6738109AFFC7FFC330009399EEB3FE1F" author="Iwan" box="[814,975,437,459]" page="500" pageId="1" pageNumber="61" year="2001">
<bibRefCitation id="82084B7AFFC7FFC330009399EEE1FE1F" author="Iwan" box="[814,925,438,459]" pageId="1" pageNumber="61" refString="Iwan, D. (2001) Systematics of the Madagascan genera Clastopus Fairmaire, 1898 and Lechius Iwan, 1995 (Coleoptera: Tenebrionidae: Platynotini). Annales Zoologici, 51 (4), 497 - 504." type="journal article" year="2001">Iwan 2001</bibRefCitation>
: 500
</treatmentCitation>
, 2002a: 60, 2002b: 259, 2004: 739, 2005: 23, 2010: 50.
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</paragraph>
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<treatmentCitationGroup id="C68911A5FFC7FFC333B993D9EEFEFDD8" box="[151,898,501,524]" pageId="1" pageNumber="61">
<taxonomicName id="21994D08FFC7FFC333B993D9ECADFDDF" authority="Iwan, 1996: 390" authorityName="Iwan" authorityPageNumber="390" authorityYear="1996" box="[151,465,501,524]" class="Insecta" family="Tenebrionidae" genus="Hovademulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="D4EDEA99FFC7FFC333B993D9EC5EFDD8" box="[151,290,502,524]" italics="true" pageId="1" pageNumber="61">Hovademulus</emphasis>
<treatmentCitation id="6738109AFFC7FFC3320693D9ECADFDDF" author="Iwan" box="[296,465,501,523]" page="390" pageId="1" pageNumber="61" year="1996">
<bibRefCitation id="82084B7AFFC7FFC3320693D9ECE2FDDF" author="Iwan" box="[296,414,501,523]" pageId="1" pageNumber="61" refString="Iwan, D. (1996) Revision of the genera of the newly-established group of Madagascan melanocratoid Platynotina (Coleoptera: Tenebrionidae: Platynotini). Genus, 7 (3), 379 - 449." type="journal article" year="1996">Iwan, 1996</bibRefCitation>
: 390
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</taxonomicName>
.—
<treatmentCitation id="6738109AFFC7FFC332DE93D9EFEDFDDF" author="Iwan" box="[496,657,502,523]" page="331" pageId="1" pageNumber="61" year="1998">
<bibRefCitation id="82084B7AFFC7FFC332DE93D9EF23FDDF" author="Iwan" box="[496,607,502,523]" pageId="1" pageNumber="61" refString="Iwan, D. (1998) Hovademus madagascariensis sp. nov. from the Madagascar (Coleoptera: Tenebrionidae: Platynotini). Genus, 9 (3), 331 - 335." type="journal article" year="1998">Iwan 1998</bibRefCitation>
: 331
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,
<treatmentCitation id="6738109AFFC7FFC331B593D9EE46FDDF" author="Iwan" box="[667,826,501,523]" page="500" pageId="1" pageNumber="61" year="2001">
<bibRefCitation id="82084B7AFFC7FFC331B593D9EE77FDDF" author="Iwan" box="[667,779,502,523]" pageId="1" pageNumber="61" refString="Iwan, D. (2001) Systematics of the Madagascan genera Clastopus Fairmaire, 1898 and Lechius Iwan, 1995 (Coleoptera: Tenebrionidae: Platynotini). Annales Zoologici, 51 (4), 497 - 504." type="journal article" year="2001">Iwan 2001</bibRefCitation>
: 500
</treatmentCitation>
(syn.).
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</paragraph>
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<typeStatus id="39228829FFC7FFC333B99015EDB0FD9B" box="[151,204,570,591]" pageId="1" pageNumber="61">Type</typeStatus>
species.
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<taxonomicName id="21994D08FFC7FFC332079016EF46FD9B" authority="Fairmaire, 1898" authorityName="Fairmaire" authorityYear="1898" box="[297,570,569,591]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="genus">
<emphasis id="D4EDEA99FFC7FFC332079016ECF2FD9B" box="[297,398,569,591]" italics="true" pageId="1" pageNumber="61">Clastopus</emphasis>
<bibRefCitation id="82084B7AFFC7FFC332BB9016EF46FD9B" author="Fairmaire" box="[405,570,569,591]" pageId="1" pageNumber="61" refString="Fairmaire, L. (1898) Descriptions de Coleopteres de d'Asie et de Malaisie. Annales de la Societe Entomologique de France, 67: 382 - 400." type="journal article" year="1898">Fairmaire, 1898</bibRefCitation>
</taxonomicName>
:
<taxonomicName id="21994D08FFC7FFC3316A9016EEA2FD9B" authority="Fairmaire, 1898" authorityName="Fairmaire" authorityYear="1898" box="[580,990,569,591]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="species" species="eurynotoides">
<emphasis id="D4EDEA99FFC7FFC3316A9016EE4EFD9B" box="[580,818,569,591]" italics="true" pageId="1" pageNumber="61">Clastopus eurynotoides</emphasis>
<bibRefCitation id="82084B7AFFC7FFC330179016EEA2FD9B" author="Fairmaire" box="[825,990,569,591]" pageId="1" pageNumber="61" refString="Fairmaire, L. (1898) Descriptions de Coleopteres de d'Asie et de Malaisie. Annales de la Societe Entomologique de France, 67: 382 - 400." type="journal article" year="1898">Fairmaire, 1898</bibRefCitation>
</taxonomicName>
; designated by monotypy.
<taxonomicName id="21994D08FFC7FFC333B99077ECE1FDBA" authority="Iwan, 1996" authorityName="Iwan" authorityYear="1996" box="[151,413,600,622]" class="Insecta" family="Tenebrionidae" genus="Hovademulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="D4EDEA99FFC7FFC333B99077EC5EFDBA" box="[151,290,600,622]" italics="true" pageId="1" pageNumber="61">Hovademulus</emphasis>
<bibRefCitation id="82084B7AFFC7FFC332069076ECE1FDBA" author="Iwan" box="[296,413,600,622]" pageId="1" pageNumber="61" refString="Iwan, D. (1996) Revision of the genera of the newly-established group of Madagascan melanocratoid Platynotina (Coleoptera: Tenebrionidae: Platynotini). Genus, 7 (3), 379 - 449." type="journal article" year="1996">Iwan, 1996</bibRefCitation>
</taxonomicName>
:
<taxonomicName id="21994D08FFC7FFC332889077EE62FDB9" authority="Fairmaire, 1902" authorityName="Fairmaire" authorityYear="1902" box="[422,798,600,622]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="61" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="D4EDEA99FFC7FFC332889077EF0AFDBA" box="[422,630,600,622]" italics="true" pageId="1" pageNumber="61">Selinus punctipennis</emphasis>
Fairmaire, 1902
</taxonomicName>
(=
<taxonomicName id="21994D08FFC7FFC3306E9077E9A4FDBA" authority="Fairmaire, 1898" authorityName="Fairmaire" authorityYear="1898" box="[832,1240,600,622]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="species" species="eurynotoides">
<emphasis id="D4EDEA99FFC7FFC3306E9077E952FDBA" box="[832,1070,600,622]" italics="true" pageId="1" pageNumber="61">Clastopus eurynotoides</emphasis>
<bibRefCitation id="82084B7AFFC7FFC3371A9077E9A4FDBA" author="Fairmaire" box="[1076,1240,600,622]" pageId="1" pageNumber="61" refString="Fairmaire, L. (1898) Descriptions de Coleopteres de d'Asie et de Malaisie. Annales de la Societe Entomologique de France, 67: 382 - 400." type="journal article" year="1898">Fairmaire, 1898</bibRefCitation>
</taxonomicName>
); by original designation.
</paragraph>
<paragraph id="E626368BFFC7FFC333B99096E84DFC94" blockId="1.[151,1436,697,1156]" pageId="1" pageNumber="61">
<emphasis id="D4EDEA99FFC7FFC333B99096ECDEFD06" bold="true" box="[151,418,697,722]" pageId="1" pageNumber="61">Diagnostic characters.</emphasis>
According to the taxonomic hypothesis proposed by
<bibRefCitation id="82084B7AFFC7FFC3372A9095E9F3FD05" author="Iwan" box="[1028,1167,697,721]" pageId="1" pageNumber="61" refString="Iwan, D. (2001) Systematics of the Madagascan genera Clastopus Fairmaire, 1898 and Lechius Iwan, 1995 (Coleoptera: Tenebrionidae: Platynotini). Annales Zoologici, 51 (4), 497 - 504." type="journal article" year="2001">Iwan (2001)</bibRefCitation>
this genus is characterized by: (1) deep circumocular depression; (2) widened mid part of mentum; (3) basal border of pronotum interrupted medially; (4) posterior angles of pronotum not protruding beyond the base; (5) 5th abdominal ventrite unmarginated; (6) protibia in both sexes slender and (7) male protibia having concavity on the inner side.
</paragraph>
<paragraph id="E626368BFFC7FFC333E79164EF24FBCC" blockId="1.[151,1436,697,1156]" pageId="1" pageNumber="61">
<emphasis id="D4EDEA99FFC7FFC333E79164ECC4FCB0" bold="true" box="[201,440,843,868]" pageId="1" pageNumber="61">Systematic position.</emphasis>
According to the results of
<bibRefCitation id="82084B7AFFC7FFC331D89163EEDEFCB0" box="[758,930,844,868]" pageId="1" pageNumber="61" refString="Iwan, D. (2002 a) Generic classification of the tribe Platynotini (Coleoptera: Tenebrionidae), with notes on phylogeny. Annales Zoologici, 52 (1), 1 - 149." type="journal article">Iwan`s (2002a)</bibRefCitation>
cladistic analysis genus
<taxonomicName id="21994D08FFC7FFC337949163E856FCB0" box="[1210,1322,844,868]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="genus">
<emphasis id="D4EDEA99FFC7FFC337949163E856FCB0" box="[1210,1322,844,868]" italics="true" pageId="1" pageNumber="61">Clastopus</emphasis>
</taxonomicName>
is closely related to
<emphasis id="D4EDEA99FFC7FFC33229915FECF7FC5C" box="[263,395,880,904]" italics="true" pageId="1" pageNumber="61">Hovademus</emphasis>
Ardoin, 1970 and
<taxonomicName id="21994D08FFC7FFC33170915FEE01FC5C" authority="Iwan, 1996" authorityName="Iwan" authorityYear="1996" box="[606,893,880,904]" class="Insecta" family="Tenebrionidae" genus="Pokryszkiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="D4EDEA99FFC7FFC33170915FEF89FC5C" box="[606,757,880,904]" italics="true" pageId="1" pageNumber="61">Pokryszkiella</emphasis>
<bibRefCitation id="82084B7AFFC7FFC331D2915EEE01FC5C" author="Iwan" box="[764,893,880,904]" pageId="1" pageNumber="61" refString="Iwan, D. (1996) Revision of the genera of the newly-established group of Madagascan melanocratoid Platynotina (Coleoptera: Tenebrionidae: Platynotini). Genus, 7 (3), 379 - 449." type="journal article" year="1996">Iwan, 1996</bibRefCitation>
</taxonomicName>
. These genera forms a monophiletic group characterised by the structure of pronotum (posterior angles not reaching the middle of the base and basal border interrupted medially) (Fig. 1).
<taxonomicName id="21994D08FFC7FFC332EF9197EF4DFC04" box="[449,561,952,976]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="genus">
<emphasis id="D4EDEA99FFC7FFC332EF9197EF4DFC04" box="[449,561,952,976]" italics="true" pageId="1" pageNumber="61">Clastopus</emphasis>
</taxonomicName>
differes from
<emphasis id="D4EDEA99FFC7FFC331F99197EE27FC04" box="[727,859,952,976]" italics="true" pageId="1" pageNumber="61">Hovademus</emphasis>
and
<taxonomicName id="21994D08FFC7FFC330BB9197E950FC04" box="[917,1068,952,976]" class="Insecta" family="Tenebrionidae" genus="Pokryszkiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="D4EDEA99FFC7FFC330BB9197E950FC04" box="[917,1068,952,976]" italics="true" pageId="1" pageNumber="61">Pokryszkiella</emphasis>
</taxonomicName>
by having a narrower border of prosternal proccess and short metasternum. Additionally, from
<taxonomicName id="21994D08FFC7FFC3307291F3EE8FFC20" box="[860,1011,988,1012]" class="Insecta" family="Tenebrionidae" genus="Pokryszkiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="D4EDEA99FFC7FFC3307291F3EE8FFC20" box="[860,1011,988,1012]" italics="true" pageId="1" pageNumber="61">Pokryszkiella</emphasis>
</taxonomicName>
is can be distinguished by the lack of horns on the anterior margin of clypeus.
</paragraph>
</subSubSection>
<subSubSection id="AE836500FFC7FFC333E7960CEF98FB50" pageId="1" pageNumber="61" type="distribution">
<paragraph id="E626368BFFC7FFC333E7960CEF1CFBE8" blockId="1.[151,1436,697,1156]" box="[201,608,1059,1084]" pageId="1" pageNumber="61">
<emphasis id="D4EDEA99FFC7FFC333E7960CEC1EFBE8" bold="true" box="[201,354,1059,1084]" pageId="1" pageNumber="61">Distribution.</emphasis>
<collectingCountry id="9E8E761BFFC7FFC33247960BEC8CFBE8" box="[361,496,1060,1084]" name="Madagascar" pageId="1" pageNumber="61">Madagascar</collectingCountry>
(Fig. 21).
</paragraph>
<paragraph id="E626368BFFC7FFC333E79667EF98FB50" blockId="1.[151,1436,697,1156]" pageId="1" pageNumber="61">
<emphasis id="D4EDEA99FFC7FFC333E79667ECC2FBB4" bold="true" box="[201,446,1095,1120]" pageId="1" pageNumber="61">Included species (5):</emphasis>
<taxonomicName id="21994D08FFC7FFC332E99666EFA0FBB4" authority="Iwan, 2005" authorityName="Iwan" authorityYear="2005" box="[455,732,1096,1120]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="species" species="aberlenci">
<emphasis id="D4EDEA99FFC7FFC332E99666EF2EFBB4" box="[455,594,1096,1120]" italics="true" pageId="1" pageNumber="61">C. aberlenci</emphasis>
<bibRefCitation id="82084B7AFFC7FFC331749666EFA0FBB4" author="Iwan" box="[602,732,1096,1120]" pageId="1" pageNumber="61" refString="Iwan, D. (2005) A new species of Clastopus Fairmaire, 1898 from Madagascar, with notes on the known first - stage larvae (Coleoptera: Tenebrionidae: Pedinini: Platynotina). Annales Zoologici, 55 (1), 23 - 27." type="journal article" year="2005">Iwan, 2005</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="21994D08FFC7FFC331C69666E925FBB4" authority="Fairmaire, 1898" authorityName="Fairmaire" authorityYear="1898" box="[744,1113,1096,1120]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="species" species="eurynotoides">
<emphasis id="D4EDEA99FFC7FFC331C69666EEE6FBB4" box="[744,922,1096,1120]" italics="true" pageId="1" pageNumber="61">C. eurynotoides</emphasis>
<bibRefCitation id="82084B7AFFC7FFC3308C9667E925FBB4" author="Fairmaire" box="[930,1113,1096,1120]" pageId="1" pageNumber="61" refString="Fairmaire, L. (1898) Descriptions de Coleopteres de d'Asie et de Malaisie. Annales de la Societe Entomologique de France, 67: 382 - 400." type="journal article" year="1898">Fairmaire, 1898</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="21994D08FFC7FFC3374A9666E8EBFBB4" authority="Iwan, 1996" authorityName="Iwan" authorityYear="1996" box="[1124,1431,1096,1120]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="species" species="ordinarius">
<emphasis id="D4EDEA99FFC7FFC3374A9666E980FBB4" box="[1124,1276,1096,1120]" italics="true" pageId="1" pageNumber="61">C. ordinarius</emphasis>
(
<bibRefCitation id="82084B7AFFC7FFC336259667E8ECFBB4" author="Iwan" box="[1291,1424,1096,1120]" pageId="1" pageNumber="61" refString="Iwan, D. (1996) Revision of the genera of the newly-established group of Madagascan melanocratoid Platynotina (Coleoptera: Tenebrionidae: Platynotini). Genus, 7 (3), 379 - 449." type="journal article" year="1996">Iwan, 1996</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="21994D08FFC7FFC333B99642ECBEFB50" authority="Iwan, 1996" authorityName="Iwan" authorityYear="1996" box="[151,450,1132,1156]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="species" species="tenuiculus">
<emphasis id="D4EDEA99FFC7FFC333B99642EC56FB50" box="[151,298,1132,1156]" italics="true" pageId="1" pageNumber="61">C. tenuiculus</emphasis>
(
<bibRefCitation id="82084B7AFFC7FFC332149643ECC6FB50" author="Iwan" box="[314,442,1132,1156]" pageId="1" pageNumber="61" refString="Iwan, D. (1996) Revision of the genera of the newly-established group of Madagascan melanocratoid Platynotina (Coleoptera: Tenebrionidae: Platynotini). Genus, 7 (3), 379 - 449." type="journal article" year="1996">Iwan, 1996</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="21994D08FFC7FFC332D79642EFFFFB50" box="[505,643,1132,1156]" class="Magnoliopsida" family="Brassicaceae" genus="Clastopus" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="61" phylum="Tracheophyta" rank="species" species="griswoldi" status="sp. nov.">
<emphasis id="D4EDEA99FFC7FFC332D79642EFFFFB50" box="[505,643,1132,1156]" italics="true" pageId="1" pageNumber="61">C. griswoldi</emphasis>
</taxonomicName>
<emphasis id="D4EDEA99FFC7FFC331A49643EF98FB50" bold="true" box="[650,740,1132,1156]" pageId="1" pageNumber="61">
<taxonomicNameLabel id="CFDE57E2FFC7FFC331A49643EF98FB50" box="[650,740,1132,1156]" pageId="1" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>