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<bibCitation id="5E31D231FF8DFFD1FDEFF94BAE8FED77" author="León &amp; Monsalve" box="[566,936,1780,1806]" page="166" pageId="40" pageNumber="41" year="2006">
<bibRefCitation id="EF8AA7ACFF8DFFD1FDEFF94BAE84ED77" author="Leon &amp; Monsalve" box="[566,931,1780,1806]" firstAuthor="Leon" page="166" pageId="40" pageNumber="41" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">
<collectingRegion id="49DF14BFFF8DFFD1FDEFF94BAF55ED77" box="[566,626,1780,1806]" country="Nicaragua" name="Leon" pageId="40" pageNumber="41">León</collectingRegion>
&amp; Monsalve (2006: 166)
</bibRefCitation>
;
</bibCitation>
<bibCitation id="5E31D231FF8DFFD1FC76F94BA83BED76" author="Wasshausen et al." box="[943,1308,1780,1807]" page="385" pageId="40" pageNumber="41" year="2014">
Wasshausen 
<emphasis id="B96F064FFF8DFFD1FB9FF94AA9A6ED77" box="[1094,1153,1780,1806]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
(2014: 385);
</bibCitation>
<bibCitation id="5E31D231FF8DFFD1FAFDF94BAC4AED4B" author="Tebbitt" page="88" pageId="40" pageNumber="41" year="2020">
<bibRefCitation id="EF8AA7ACFF8DFFD1FAFDF94BAC4EED4B" author="Tebbitt M. C." journalOrPublisher="American Begonia Society, Sacramento" pageId="40" pageNumber="41" refId="ref232580" refString="Tebbitt M. C. 2020. Tuberous Begonias: a Monograph of Begonia Section Australes. American Begonia Society, Sacramento." title="Tuberous Begonias: a Monograph of Begonia Section Australes" type="book" year="2020">Tebbitt (2020: 88)</bibRefCitation>
.
</bibCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF8DFFD1FF64F8E6A9C5ED91" pageId="40" pageNumber="41" type="etymology">
<paragraph id="8BA4DA5DFF8DFFD1FF64F8E6AC62ED0A" blockId="40.[189,325,1881,1907]" box="[189,325,1881,1907]" pageId="40" pageNumber="41">
<heading id="D0EC6D31FF8DFFD1FF64F8E6AC62ED0A" bold="true" box="[189,325,1881,1907]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B96F064FFF8DFFD1FF64F8E6AC62ED0A" bold="true" box="[189,325,1881,1907]" pageId="40" pageNumber="41">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF8DFFD1FF64F837A9C5ED91" blockId="40.[189,1399,1928,2025]" pageId="40" pageNumber="41">
The epithet means ‘Herrera’s 
<taxonomicName id="4C1BA1DEFF8DFFD1FDF9F837AFA6EDD8" box="[544,641,1928,1953]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="B96F064FFF8DFFD1FDF9F837AFA6EDD8" box="[544,641,1928,1953]" italics="true" pageId="40" pageNumber="41">Begonia</emphasis>
</taxonomicName>
’. The protologue does not mention any specific 
<collectingRegion id="49DF14BFFF8DFFD1FB1DF837A839EDDB" box="[1220,1310,1928,1954]" country="Panama" name="Herrera" pageId="40" pageNumber="41">Herrera</collectingRegion>
but we believe this refers to Fortunato L. 
<collectingRegion id="49DF14BFFF8DFFD1FD91F813AF82EDBF" box="[584,677,1964,1990]" country="Panama" name="Herrera" pageId="40" pageNumber="41">Herrera</collectingRegion>
, a Peruvian botanist who made significant contributions to the knowledge of the flora of 
<collectingRegion id="49DF14BFFF8DFFD1FE34F871AFB4ED91" box="[493,659,1998,2024]" country="Peru" name="Cusco" pageId="40" pageNumber="41">Cusco Region</collectingRegion>
and who died the year 
<taxonomicName id="4C1BA1DEFF8DFFD1FC7DF870A90BED91" box="[932,1068,1998,2024]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="herrerae">
<emphasis id="B96F064FFF8DFFD1FC7DF870A90BED91" box="[932,1068,1998,2024]" italics="true" pageId="40" pageNumber="41">B. herrerae</emphasis>
</taxonomicName>
was published.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF8CFFD2FF64FEB7AEDCEB3E" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42" type="description">
<paragraph id="8BA4DA5DFF8CFFD0FF64FEB7AC68EB5B" blockId="41.[189,335,264,290]" box="[189,335,264,290]" pageId="41" pageNumber="42">
<heading id="D0EC6D31FF8CFFD0FF64FEB7AC68EB5B" bold="true" box="[189,335,264,290]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="B96F064FFF8CFFD0FF64FEB7AC68EB5B" bold="true" box="[189,335,264,290]" pageId="41" pageNumber="42">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF8CFFD2FF64FE88AEDCEB3E" blockId="41.[187,1400,311,1118]" lastBlockId="43.[189,1398,265,327]" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
Caulescent, tuberous herb, to 
<quantity id="4CE377B8FF8CFFD0FDFBFE88AF4AEB2B" box="[546,621,311,338]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="41" pageNumber="42" unit="cm" value="15.0">15 cm</quantity>
high. 
<emphasis id="B96F064FFF8CFFD0FD67FE88AE24EB28" box="[702,771,311,337]" italics="true" pageId="41" pageNumber="42">Tuber</emphasis>
globose, ca 2 × 
<quantity id="4CE377B8FF8CFFD0FC15FE88A906EB2B" box="[972,1057,311,338]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="41" pageNumber="42" unit="cm" value="1.2">1.2 cm</quantity>
, with 1 growing point. 
<emphasis id="B96F064FFF8CFFD0FAE6FE87A85FEB2B" box="[1343,1400,312,338]" italics="true" pageId="41" pageNumber="42">Stem</emphasis>
erect, rarely branching; internodes to 
<quantity id="4CE377B8FF8CFFD0FDA3FEE4AFECEB0C" box="[634,715,347,374]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="41" pageNumber="42" unit="cm" value="4.5">4.5 cm</quantity>
long, to 
<quantity id="4CE377B8FF8CFFD0FCEEFEE4AE5BEB0C" box="[823,892,347,374]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="41" pageNumber="42" unit="mm" value="5.0">5 mm</quantity>
thick, succulent, colour unknown, densely hispid. 
<emphasis id="B96F064FFF8CFFD0FEC1FEC1AC51EBE1" box="[280,374,382,408]" italics="true" pageId="41" pageNumber="42">Stipules</emphasis>
late deciduous, triangular, 5–7 × 
<quantity id="4CE377B8FF8CFFD0FCC0FEC1AEA6EBE0" box="[793,897,382,409]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="41" pageNumber="42" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
, apex acute, translucent, brown, sparsely hispid, margin serrate, long-ciliate. 
<emphasis id="B96F064FFF8CFFD0FDAAFE1DAFE2EBC2" box="[627,709,418,443]" italics="true" pageId="41" pageNumber="42">Leaves</emphasis>
4–6, alternate, basifixed; petiole 
<quantity id="4CE377B8FF8CFFD0FBBCFE1EA9F0EBC5" box="[1125,1239,417,444]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="5.8" metricValueMin="2.0" pageId="41" pageNumber="42" unit="cm" value="3.9" valueMax="5.8" valueMin="2.0">2–5.8 cm</quantity>
long, colour unknown, densely hispid; blade asymmetric, ovate, to 12 × 
<quantity id="4CE377B8FF8CFFD0FC4AFE7AAED0EB99" box="[915,1015,453,480]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="41" pageNumber="42" unit="cm" value="10.5">10.5 cm</quantity>
, membranaceous, apex acute to acuminate, base obliquely cordate, basal lobes overlapping, sinus to 
<quantity id="4CE377B8FF8CFFD0FC2EFE57A96BE87A" box="[1015,1100,488,515]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="41" pageNumber="42" unit="mm" value="25.0">25 mm</quantity>
deep, margin irregularly dentate, ciliate, upper surface green, hispid, lower surface pale green, densely hispid on the veins, hispid on the lamina, veins palmate but with 1 primary vein, 6–8 veined from the base, with 1–3 secondary veins on the larger side, 1–2 secondary veins on the smaller side. 
<emphasis id="B96F064FFF8CFFD0FC61FDECA947E814" box="[952,1120,595,621]" italics="true" pageId="41" pageNumber="42">Inflorescences</emphasis>
1–4 per stem, bisexual, axillary, erect, cymose, with 1–2 branches, bearing up to 4 staminate flowers and 2 pistillate flowers, protandrous; peduncle to 
<quantity id="4CE377B8FF8CFFD0FE3FFD25AF09E8CD" box="[486,558,666,692]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="41" pageNumber="42" unit="cm" value="18.0">18 cm</quantity>
long, colour unknown, densely hispid, bracts deciduous, triangular, 5–6 × 
<quantity id="4CE377B8FF8CFFD0FF0DFD02AC1FE8A1" box="[212,312,701,728]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="41" pageNumber="42" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
, translucent, colour unknown, sparsely hispid, apex acute, margin lacerate, ciliate. 
<emphasis id="B96F064FFF8CFFD0FAD9FD01AC35E882" italics="true" pageId="41" pageNumber="42">Staminate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FF8CFFD0FE7EFD5FACDDE882" box="[423,506,736,763]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="41" pageNumber="42" unit="mm" value="15.0">15 mm</quantity>
long, hispid; tepals 4, spreading, outer 2 elliptic, ovate, or obovate, 12–22 × 
<quantity id="4CE377B8FF8CFFD0FF64FCBBAC09E966" box="[189,302,772,799]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.9" metricValueMin="0.6" pageId="41" pageNumber="42" unit="mm" value="12.5" valueMax="19.0" valueMin="6.0">6–19 mm</quantity>
, apex acute, orange or red, glabrous, margin serrate, ciliate, inner 2 elliptic to obovate, 12–25 × 
<quantity id="4CE377B8FF8CFFD0FF65FC97AC0AE93B" box="[188,301,808,834]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.7" metricValueMin="0.4" pageId="41" pageNumber="42" unit="mm" value="10.5" valueMax="17.0" valueMin="4.0">4–17 mm</quantity>
, apex acute, orange or red, glabrous, margin entire, aciliate; stamens 13–25, spreading, yellow, filaments ca 
<quantity id="4CE377B8FF8CFFD0FE89FCF4ACB2E91F" box="[336,405,843,870]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="41" pageNumber="42" unit="mm" value="1.0">1 mm</quantity>
long, free, anthers cuboid, ca 1 × 
<quantity id="4CE377B8FF8CFFD0FCFCFCF4AEA5E91F" box="[805,898,843,870]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="41" pageNumber="42" unit="mm" value="0.5">0.5 mm</quantity>
, dehiscing via lateral slits, connectives not extended, filaments symmetrically fixed to the back of the stamens. 
<emphasis id="B96F064FFF8CFFD0FC14FCD1A9B3E9F1" box="[973,1172,878,904]" italics="true" pageId="41" pageNumber="42">Pistillate flowers</emphasis>
: pedicels to 
<quantity id="4CE377B8FF8CFFD0FAFCFCD1A850E9F0" box="[1317,1399,878,905]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="41" pageNumber="42" unit="mm" value="15.0">15 mm</quantity>
long; bracteoles lacking; tepals 5, subequal, deciduous in fruit, spreading, the largest lanceolate, 10–15 × 
<quantity id="4CE377B8FF8CFFD0FF65FC0AAC38E9A9" box="[188,287,949,976]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="41" pageNumber="42" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
, apex acute, orange or red, sparsely hispid, margin serrate, ciliate, the smallest elliptic, 10–16 × 
<quantity id="4CE377B8FF8CFFD0FF65FC66AC09E98A" box="[188,302,985,1012]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="41" pageNumber="42" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
, apex rounded, sparsely hispid, margin serrulate, ciliate; ovary body obovoid, 5–8.5 × 
<quantity id="4CE377B8FF8CFFD0FAC9FC66A855E98A" box="[1296,1394,985,1012]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="41" pageNumber="42" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
, colour unknown, glabrous, unequally 3-winged, wings triangular, largest 3–6.5 × 
<quantity id="4CE377B8FF8CFFD0FBB7FC43A9F6EE6E" box="[1134,1233,1020,1047]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="41" pageNumber="42" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">2–7 mm</quantity>
, smallest 2.5– 6 × 
<quantity id="4CE377B8FF8CFFD0FF32FB9FAC41EE43" box="[235,358,1056,1083]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="41" pageNumber="42" unit="mm" value="1.25" valueMax="2.0" valueMin="0.5">0.5–2 mm</quantity>
; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free, 
<quantity id="4CE377B8FF8CFFD0FF25FBFCAC50EE27" box="[252,375,1091,1118]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="41" pageNumber="42" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
long, once-divided, stigmatic papillae in a spirally twisted band. 
<emphasis id="B96F064FFF8CFFD0FB4BFBFBA871EE24" box="[1170,1366,1091,1117]" italics="true" pageId="41" pageNumber="42">Fruiting pedicel</emphasis>
to 
<quantity id="4CE377B8FF8EFFD2FF64FEB6AC36EB5D" box="[189,273,265,292]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="mm" value="20.0">20 mm</quantity>
long. 
<emphasis id="B96F064FFF8EFFD2FE86FEB5ACF9EB5A" box="[351,478,265,291]" italics="true" pageId="43" pageNumber="44">Fruit body</emphasis>
ovoid, to 8.5 × 
<quantity id="4CE377B8FF8EFFD2FD47FEB6AFC0EB5D" box="[670,743,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="43" pageNumber="44" unit="mm" value="9.0">9 mm</quantity>
, drying light brown, wings same shape as in ovary, the largest expanding to 11 × 
<quantity id="4CE377B8FF8EFFD2FE37FE92AF63EB3E" box="[494,580,301,327]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="43" pageNumber="44" unit="mm" value="21.0">21 mm</quantity>
, the smallest expanding to 8 × 
<quantity id="4CE377B8FF8EFFD2FC69FE93AED0EB3E" box="[944,1015,300,327]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="44" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF648AD5FF8CFFD0FF64F8FEAF24ED91" ID-DOI="http://doi.org/10.5281/zenodo.8178316" ID-Zenodo-Dep="8178316" httpUri="https://zenodo.org/record/8178316/files/figure.png" pageId="41" pageNumber="42" startId="41.[189,232,1857,1883]" targetBox="[194,1392,1193,1812]" targetPageId="41">
<paragraph id="8BA4DA5DFF8CFFD0FF64F8FEAF24ED91" blockId="41.[189,1399,1857,2024]" pageId="41" pageNumber="42">
<emphasis id="B96F064FFF8CFFD0FF64F8FEAC07ED22" bold="true" box="[189,288,1857,1883]" pageId="41" pageNumber="42">Fig. 17.</emphasis>
Distribution of 
<taxonomicName id="4C1BA1DEFF8CFFD0FE36F8FDA91BED22" authority="L. B. Sm. &amp; B. G. Schub." authorityName="L. B. Sm. &amp; B. G. Schub." box="[495,1084,1857,1884]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="section" section="Australes">
<emphasis id="B96F064FFF8CFFD0FE36F8FDAF75ED22" box="[495,594,1858,1883]" italics="true" pageId="41" pageNumber="42">Begonia</emphasis>
sect. 
<emphasis id="B96F064FFF8CFFD0FD7BF8FEAE35ED22" box="[674,786,1857,1883]" italics="true" pageId="41" pageNumber="42">Australes</emphasis>
L.B.Sm. &amp; B.G.Schub.
</taxonomicName>
in Peru and surrounding countries. 
<emphasis id="B96F064FFF8CFFD0FEE0F8DBAC69ED07" bold="true" box="[313,334,1892,1918]" pageId="41" pageNumber="42">A</emphasis>
. 
<taxonomicName id="4C1BA1DEFF8CFFD0FE86F8D9AE2DED06" authority="L. B. Sm. &amp; B. G. Schub." authorityName="L. B. Sm. &amp; B. G. Schub." box="[351,778,1893,1919]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="herrerae">
<emphasis id="B96F064FFF8CFFD0FE86F8D9ACCDED06" box="[351,490,1893,1919]" italics="true" pageId="41" pageNumber="42">B. herrerae</emphasis>
L.B.Sm. &amp; B.G.Schub.
</taxonomicName>
(red) and 
<taxonomicName id="4C1BA1DEFF8CFFD0FC53F8D9A95DED06" authority="Tebbitt" authorityName="Tebbitt" box="[906,1146,1893,1919]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="species" species="heliantha">
<emphasis id="B96F064FFF8CFFD0FC53F8D9A93AED06" box="[906,1053,1893,1919]" italics="true" pageId="41" pageNumber="42">B. heliantha</emphasis>
Tebbitt
</taxonomicName>
(blue). 
<emphasis id="B96F064FFF8CFFD0FB03F8DAA9CBED06" bold="true" box="[1242,1260,1893,1919]" pageId="41" pageNumber="42">B</emphasis>
. 
<taxonomicName id="4C1BA1DEFF8CFFD0FB25F8D9AC97EDDB" authority="Hook. f. var. veitchii" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="variety" species="veitchii" variety="veitchii">
<emphasis id="B96F064FFF8CFFD0FB25F8D9A851ED06" box="[1276,1398,1893,1919]" italics="true" pageId="41" pageNumber="42">B. veitchii</emphasis>
Hook.f. var. 
<emphasis id="B96F064FFF8CFFD0FE80F837AC97EDDB" box="[345,432,1928,1954]" italics="true" pageId="41" pageNumber="42">veitchii</emphasis>
</taxonomicName>
(red) and 
<taxonomicName id="4C1BA1DEFF8CFFD0FDEEF836A908EDDB" authority="Tebbitt" authorityName="Tebbitt" box="[567,1071,1928,1954]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="41" pageNumber="42" phylum="Tracheophyta" rank="variety" species="veitchii" variety="machupicchuensis">
<emphasis id="B96F064FFF8CFFD0FDEEF836AF93EDDB" box="[567,692,1928,1954]" italics="true" pageId="41" pageNumber="42">B. veitchii</emphasis>
var. 
<emphasis id="B96F064FFF8CFFD0FD2EF837AEF7EDDB" box="[759,976,1928,1954]" italics="true" pageId="41" pageNumber="42">machupicchuensis</emphasis>
Tebbitt
</taxonomicName>
(red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation.
</paragraph>
</caption>
<caption id="DF648AD5FF8FFFD3FF64F837AF0EED91" ID-DOI="http://doi.org/10.5281/zenodo.8178320" ID-Zenodo-Dep="8178320" httpUri="https://zenodo.org/record/8178320/files/figure.png" pageId="42" pageNumber="43" startId="42.[189,232,1928,1954]" targetBox="[229,1335,488,1812]" targetPageId="42">
<paragraph id="8BA4DA5DFF8FFFD3FF64F837AF0EED91" blockId="42.[189,1399,1928,2025]" pageId="42" pageNumber="43">
<emphasis id="B96F064FFF8FFFD3FF64F837AC3BEDDB" bold="true" box="[189,284,1928,1954]" pageId="42" pageNumber="43">Fig. 18.</emphasis>
<taxonomicName id="4C1BA1DEFF8FFFD3FEFCF836A9CFEDDB" authority="L. B. Sm. &amp; B. G. Schub. A. Habit. B. Inflorescence. C. Stamen" box="[293,1256,1928,1954]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="herrerae">
<emphasis id="B96F064FFF8FFFD3FEFCF836ACDFEDDB" box="[293,504,1928,1954]" italics="true" pageId="42" pageNumber="43">Begonia herrerae</emphasis>
L.B.Sm. &amp; B.G.Schub. 
<emphasis id="B96F064FFF8FFFD3FCF9F837AE14EDDB" bold="true" box="[800,819,1928,1954]" pageId="42" pageNumber="43">A</emphasis>
. Habit. 
<emphasis id="B96F064FFF8FFFD3FC41F837AE8DEDDB" bold="true" box="[920,938,1928,1954]" pageId="42" pageNumber="43">B</emphasis>
. Inflorescence. 
<emphasis id="B96F064FFF8FFFD3FBBEF837A95BEDDB" bold="true" box="[1127,1148,1928,1954]" pageId="42" pageNumber="43">C</emphasis>
. Stamen
</taxonomicName>
, front view. 
<emphasis id="B96F064FFF8FFFD3FF66F814ADF5EDBC" bold="true" box="[191,210,1963,1989]" pageId="42" pageNumber="43">D</emphasis>
. Style and stigma, front view. Illustration adapted from 
<bibRefCitation id="EF8AA7ACFF8FFFD3FCAFF814A987EDBF" author="Smith &amp; Schubert" box="[886,1184,1963,1990]" firstAuthor="Smith" pageId="42" pageNumber="43" pagination="23 - 31" refId="ref231334" refString="Smith L. B. &amp; Schubert D. C. 1945. Studies in the Begoniaceae 1. Contributions from the Gray Herbarium of Harvard University 154: 23 - 31." type="journal article" year="1945">Smith &amp; Schubert (1945</bibRefCitation>
: fig. 15), digitally remastered by P.W. Moonlight.
</paragraph>
</caption>
<subSubSection id="C30189D6FF8EFFD2FF64FED0A852E83F" pageId="43" pageNumber="44" type="conservation">
<paragraph id="8BA4DA5DFF8EFFD2FF64FED0AF4AEBF0" blockId="43.[189,621,366,393]" box="[189,621,366,393]" pageId="43" pageNumber="44">
<heading id="D0EC6D31FF8EFFD2FF64FED0AF4AEBF0" bold="true" box="[189,621,366,393]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B96F064FFF8EFFD2FF64FED0AF4AEBF0" bold="true" box="[189,621,366,393]" pageId="43" pageNumber="44">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF8EFFD2FF64FE22A852E83F" blockId="43.[188,1399,413,582]" pageId="43" pageNumber="44">
Previously assessed as Data Deficient (DD) by 
<bibRefCitation id="EF8AA7ACFF8EFFD2FCDFFE22A966EBCE" author="Leon &amp; Monsalve" box="[774,1089,413,439]" firstAuthor="Leon" pageId="43" pageNumber="44" pagination="165 - 170" refId="ref229226" refString="Leon B. &amp; Monsalve C. 2006. Begoniaceae endemicas del Peru. Revista Peruana de Biologia 13 (2): 165 - 170. https: // doi. org / 10.15381 / rpb. v 13 i 2.1810" type="journal article" year="2006">León &amp; Monsalve (2006)</bibRefCitation>
, who cited only the 
<typeStatus id="54A064FFFF8EFFD2FA9DFE21A850EBC1" box="[1348,1399,414,440]" pageId="43" pageNumber="44">type</typeStatus>
specimen. A second collection has since been identified from ca 
<quantity id="4CE377B8FF8EFFD2FC14FE7EA90FEBA2" box="[973,1064,449,476]" metricMagnitude="5" metricUnit="m" metricValue="5.3" pageId="43" pageNumber="44" unit="km" value="530.0">530 km</quantity>
to the southwest in La Paz Province in 
<collectingCountry id="F30C9ACDFF8EFFD2FE95FE5BAC85EB87" box="[332,418,484,510]" name="Bolivia" pageId="43" pageNumber="44">Bolivia</collectingCountry>
(
<bibRefCitation id="EF8AA7ACFF8EFFD2FE6BFE5BAF68EB86" author="Tebbitt M. C." box="[434,591,484,511]" journalOrPublisher="American Begonia Society, Sacramento" pageId="43" pageNumber="44" refId="ref232580" refString="Tebbitt M. C. 2020. Tuberous Begonias: a Monograph of Begonia Section Australes. American Begonia Society, Sacramento." title="Tuberous Begonias: a Monograph of Begonia Section Australes" type="book" year="2020">Tebbitt 2020</bibRefCitation>
). We have not observed 
<taxonomicName id="4C1BA1DEFF8EFFD2FCA8FE5AAEDDEB87" box="[881,1018,484,510]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="herrerae">
<emphasis id="B96F064FFF8EFFD2FCA8FE5AAEDDEB87" box="[881,1018,484,510]" italics="true" pageId="43" pageNumber="44">B. herrerae</emphasis>
</taxonomicName>
in the wild and have no data on population sizes or trends. The lower montane forests between its two collections are poorly collected so there may be many uncollected populations in this area. We assess 
<taxonomicName id="4C1BA1DEFF8EFFD2FC16FD93A970E83F" box="[975,1111,556,582]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="herrerae">
<emphasis id="B96F064FFF8EFFD2FC16FD93A970E83F" box="[975,1111,556,582]" italics="true" pageId="43" pageNumber="44">B. herrerae</emphasis>
</taxonomicName>
as Data Deficient (DD).
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF8EFFD2FF64FDD2A9EFE8CF" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BA4DA5DFF8EFFD2FF64FDD2AC97E8F1" blockId="43.[189,432,621,648]" box="[189,432,621,648]" pageId="43" pageNumber="44">
<heading id="D0EC6D31FF8EFFD2FF64FDD2AC97E8F1" bold="true" box="[189,432,621,648]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B96F064FFF8EFFD2FF64FDD2AC97E8F1" bold="true" box="[189,432,621,648]" pageId="43" pageNumber="44">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF8EFFD2FF64FD22A9EFE8CF" blockId="43.[189,1224,668,695]" box="[189,1224,668,695]" pageId="43" pageNumber="44">
Most similar to 
<taxonomicName id="4C1BA1DEFF8EFFD2FEA1FD22AF2EE8CF" box="[376,521,668,694]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="heliantha">
<emphasis id="B96F064FFF8EFFD2FEA1FD22AF2EE8CF" box="[376,521,668,694]" italics="true" pageId="43" pageNumber="44">B. heliantha</emphasis>
</taxonomicName>
but differing in the serrated margin to its staminate flowers.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FF8EFFD2FF64FD61A9C0E9CC" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BA4DA5DFF8EFFD2FF64FD61ACD2E881" blockId="43.[189,501,734,760]" box="[189,501,734,760]" pageId="43" pageNumber="44">
<heading id="D0EC6D31FF8EFFD2FF64FD61ACD2E881" bold="true" box="[189,501,734,760]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B96F064FFF8EFFD2FF64FD61ACD2E881" bold="true" box="[189,501,734,760]" pageId="43" pageNumber="44">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFF8EFFD2FF64FCB1A9C0E9CC" blockId="43.[189,1399,781,950]" pageId="43" pageNumber="44">
Known from 
<collectingCountry id="F30C9ACDFF8EFFD2FE82FCB2AC96E95E" box="[347,433,781,807]" name="Bolivia" pageId="43" pageNumber="44">Bolivia</collectingCountry>
and 
<collectingCountry id="F30C9ACDFF8EFFD2FE32FCB1AF02E951" box="[491,549,782,808]" name="Peru" pageId="43" pageNumber="44">Peru</collectingCountry>
. Within 
<collectingCountry id="F30C9ACDFF8EFFD2FD5FFCB1AF98E951" box="[646,703,782,808]" name="Peru" pageId="43" pageNumber="44">Peru</collectingCountry>
, it has been collected in 
<collectingRegion id="49DF14BFFF8EFFD2FC07FCB2A9ABE95E" box="[990,1164,781,807]" country="Peru" name="Madre de Dios" pageId="43" pageNumber="44">Madre de Dios</collectingRegion>
and 
<collectingRegion id="49DF14BFFF8EFFD2FB1FFCB2A828E95E" box="[1222,1295,781,807]" country="Peru" name="Cusco" pageId="43" pageNumber="44">Cusco</collectingRegion>
Regions (
<figureCitation id="1320C6D8FF8EFFD2FF1FFC8EAC11E932" box="[198,310,817,843]" captionStart="Fig" captionStartId="41.[189,232,1857,1883]" captionTargetBox="[194,1392,1193,1812]" captionTargetId="figure-379@41.[189,1398,1183,1818]" captionTargetPageId="41" captionText="Fig. 17. Distribution of Begonia sect. Australes L.B.Sm. &amp; B.G.Schub. in Peru and surrounding countries. A. B. herrerae L.B.Sm. &amp; B.G.Schub. (red) and B. heliantha Tebbitt (blue). B. B. veitchii Hook.f. var. veitchii (red) and B. veitchii var. machupicchuensis Tebbitt (red). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178316" httpUri="https://zenodo.org/record/8178316/files/figure.png" pageId="43" pageNumber="44">Fig. 17A</figureCitation>
). Found in Amazonia at an elevation of 
<quantity id="4CE377B8FF8EFFD2FCC4FC8EAE82E932" box="[797,933,817,843]" metricMagnitude="2" metricUnit="m" metricValue="2.6" metricValueMax="3.2" metricValueMin="2.0" pageId="43" pageNumber="44" unit="m" value="260.0" valueMax="320.0" valueMin="200.0">200–320 m</quantity>
a.s.l. in habitats described as sandy or rocky hills. 
<taxonomicName id="4C1BA1DEFF8EFFD2FE91FCEAAF3EE917" box="[328,537,852,878]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="herrerae">
<emphasis id="B96F064FFF8EFFD2FE91FCEAAF3EE917" box="[328,537,852,878]" italics="true" pageId="43" pageNumber="44">Begonia herrerae</emphasis>
</taxonomicName>
is tuberous and likely dies back to its tuber at some point during the year. All known collections of the species have been collected from August to November, which represents the end of the dry season. It is unclear however whether this is the result of collection bias.
</paragraph>
</subSubSection>
</treatment>
</document>