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<mods:title id="D2203A7DD284E75938C665AA7B1A3D86">New data on the least known zelotines (Araneae, Gnaphosidae) of Greece and adjacent regions</mods:title>
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<treatment id="038F801CFFF33B1DFF448EAFFC8ACA2C" ID-DOI="http://doi.org/10.5281/zenodo.3510606" ID-GBIF-Taxon="119393570" ID-Zenodo-Dep="3510606" LSID="urn:lsid:plazi:treatment:038F801CFFF33B1DFF448EAFFC8ACA2C" httpUri="http://treatment.plazi.org/id/038F801CFFF33B1DFF448EAFFC8ACA2C" lastPageId="7" lastPageNumber="50" pageId="5" pageNumber="48">
<subSubSection id="C33C6281FFF33B1FFF448EAFFEB7CFEA" pageId="5" pageNumber="48" type="nomenclature">
<paragraph id="8B99310AFFF33B1FFF448EAFFD63CF8F" blockId="5.[151,735,151,212]" box="[151,735,151,177]" pageId="5" pageNumber="48">
<heading id="D0D18666FFF33B1FFF448EAFFD63CF8F" bold="true" box="[151,735,151,177]" fontSize="11" level="1" pageId="5" pageNumber="48" reason="1">
<taxonomicName id="4C264A89FFF33B1FFF448EAFFD63CF8F" ID-CoL="7CMJK" authority="Platnick &amp; Murphy, 1984" authorityName="Platnick &amp; Murphy" authorityYear="1984" box="[151,735,151,177]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF33B1FFF448EAFFD63CF8F" bold="true" box="[151,735,151,177]" pageId="5" pageNumber="48">
<emphasis id="B952ED18FFF33B1FFF448EAFFE24CF8F" bold="true" box="[151,408,151,177]" italics="true" pageId="5" pageNumber="48">Trachyzelotes stubbsi</emphasis>
<bibRefCitation id="EFB74CFBFFF33B1FFE4C8EAFFD63CF8F" author="Platnick" box="[415,735,151,177]" pageId="5" pageNumber="48" refString="Platnick, N. I. &amp; Murphy, J. A. (1984) A revision of the spider genera Trachyzelotes and Urozelotes (Araneae, Gnaphosidae). American Museum Novitates, 2792, 1 - 30." type="journal article" year="1984">Platnick &amp; Murphy, 1984</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B99310AFFF33B1FFF448E82FEB7CFEA" blockId="5.[151,735,151,212]" box="[151,267,186,212]" pageId="5" pageNumber="48">
<figureCitation id="131D2D8FFFF33B1FFF448E82FEB7CFEA" box="[151,267,186,212]" captionStart-0="FIGURES 6 9" captionStart-1="FIGURES 10 13" captionStartId-0="5.[151,271,1971,1995]" captionStartId-1="6.[151,274,1979,2003]" captionTargetBox-0="[318,1275,482,1935]" captionTargetBox-1="[187,1391,697,1978]" captionTargetId-0="figure@5.[308,1278,482,1950]" captionTargetId-1="figure@6.[175,1411,658,2074]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 6 9. Trachyzelotes stubbsi male. Specimen from Cyprus (6, 8), specimen from Antikythira (7, 9). Ventral view (6 7). Retrolateral view (8 9). rp = retrolateral process." captionText-1="FIGURES 10 13. Trachyzelotes stubbsi female, specimen from Cyprus (10 11), specimen from Antikythira (12). Trachyzelotes adriaticus female, specimen from Crete (13). Epigyne (10, 12, 13). Vulva (11)." httpUri-0="https://zenodo.org/record/197220/files/figure.png" httpUri-1="https://zenodo.org/record/197221/files/figure.png" pageId="5" pageNumber="48">Figs 612</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33C6281FFF33B1FFF448F3AFD5FCE87" pageId="5" pageNumber="48" type="reference_group">
<paragraph id="8B99310AFFF33B1FFF448F3AFDE1CE07" blockId="5.[151,1436,258,441]" pageId="5" pageNumber="48">
<treatmentCitationGroup id="AB361624FFF33B1FFF448F3AFDE1CE07" pageId="5" pageNumber="48">
<taxonomicName id="4C264A89FFF33B1FFF448F3AFD8DCE27" ID-CoL="7CMJK" authority="Platnick &amp; Murphy, 1984: 9" authorityName="Platnick &amp; Murphy" authorityPageNumber="9" authorityYear="1984" box="[151,561,258,281]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF33B1FFF448F3AFF41CE27" box="[151,253,258,281]" italics="true" pageId="5" pageNumber="48">T. stubbsi</emphasis>
<treatmentCitation id="0A87171BFFF33B1FFED78F3AFD8DCE27" author="Platnick" box="[260,561,258,281]" page="9" pageId="5" pageNumber="48" year="1984">
<bibRefCitation id="EFB74CFBFFF33B1FFED78F3AFDA4CE27" author="Platnick" box="[260,536,258,281]" pageId="5" pageNumber="48" refString="Platnick, N. I. &amp; Murphy, J. A. (1984) A revision of the spider genera Trachyzelotes and Urozelotes (Araneae, Gnaphosidae). American Museum Novitates, 2792, 1 - 30." type="journal article" year="1984">Platnick &amp; Murphy, 1984</bibRefCitation>
: 9
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="131D2D8FFFF33B1FFDEE8F3AFD0ACE27" box="[573,694,258,281]" captionStart="FIGURES 14 19" captionStartId="10.[151,270,1927,1951]" captionTargetBox="[193,1391,205,1903]" captionTargetId="figure@10.[191,1396,197,1910]" captionTargetPageId="10" captionText="FIGURES 14 19. Zelotes helicoides n. sp. male (14, 16, 18). Zelotes cingarus male (15, 17, 19). Ventral view (14 15); retrolateral view (16 17); dorsal view (18 19). p = prolateral process, TA = terminal apophysis, EB = embolar basis, EM = embolus, M = median apophysis, X = dorsal extension. (From Chatzaki et al. 2003)." httpUri="https://zenodo.org/record/197222/files/figure.png" pageId="5" pageNumber="48">Figs 1516</figureCitation>
, (Dm)
<typeStatus id="549D8FA8FFF33B1FFCD18F3AFCE3CE27" box="[770,863,258,281]" pageId="5" pageNumber="48" type="holotype">holotype</typeStatus>
from
<collectingCountry id="F331719AFFF33B1FFC728F3AFC4DCE27" box="[929,1009,258,281]" name="Cyprus" pageId="5" pageNumber="48">Cyprus</collectingCountry>
, Cape Kiti, near Larnaka airport (contra
<bibRefCitation id="EFB74CFBFFF33B1FFF148F1AFE1ACE07" author="Chatzaki" box="[199,422,290,313]" pageId="5" pageNumber="48" refString="Chatzaki, M., Thaler, K. &amp; Mylonas, M. (2003) Ground spiders (Gnaphosidae; Araneae) from Crete and adjacent areas of Greece. Taxonomy and distribution III. Zelotes and allied genera. Revue Suisse de Zoology, 110 (1), 45 - 89." type="journal article" year="2003">
Chatzaki
<emphasis id="B952ED18FFF33B1FFEFF8F1AFEDECE07" box="[300,354,290,313]" italics="true" pageId="5" pageNumber="48">et al.</emphasis>
, 2003
</bibRefCitation>
) (not examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B99310AFFF33B1FFF448F7AFB8FCE67" blockId="5.[151,1436,258,441]" box="[151,1075,322,345]" pageId="5" pageNumber="48">
<treatmentCitationGroup id="AB361624FFF33B1FFF448F7AFB8FCE67" box="[151,1075,322,345]" pageId="5" pageNumber="48">
<taxonomicName id="4C264A89FFF33B1FFF448F7AFE06CE67" ID-CoL="7CMJK" authority="Levy, 1998: 106" authorityName="Levy" authorityPageNumber="106" authorityYear="1998" box="[151,442,322,345]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF33B1FFF448F7AFF40CE67" box="[151,252,322,345]" italics="true" pageId="5" pageNumber="48">T. stubbsi</emphasis>
:
<treatmentCitation id="0A87171BFFF33B1FFED88F7AFE06CE67" author="Levy" box="[267,442,322,345]" page="106" pageId="5" pageNumber="48" year="1998">
<bibRefCitation id="EFB74CFBFFF33B1FFED88F7AFE3ACE67" author="Levy" box="[267,390,322,345]" pageId="5" pageNumber="48" refString="Levy, G. (1998) The ground-spider genera Setaphis, Trachyzelotes, Zelotes and Drassyllus (Araneae: Gnaphosidae) in Israel. Israel Journal of Zoology, 44, 93 - 158." type="journal article" year="1998">Levy, 1998</bibRefCitation>
: 106
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="131D2D8FFFF33B1FFE158F7AFD81CE67" box="[454,573,322,345]" captionStart-0="FIGURES 20 23" captionStart-1="FIGURE 24" captionStartId-0="11.[151,270,1656,1680]" captionStartId-1="12.[151,255,1358,1382]" captionTargetBox-0="[256,1333,192,1634]" captionTargetBox-1="[151,1436,194,1336]" captionTargetId-0="figure@11.[251,1336,178,1634]" captionTargetId-1="figure@12.[151,1436,194,1336]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 20 23. Zelotes helicoides n. sp. female (20 21). Zelotes cingarus female (22 23). Epigyne (20, 22). Vulva (21, 23). (From Chatzaki et al. 2003)." captionText-1="FIGURE 24. Zelotes helicoides n. sp. SEM photos of ♂ palpal organ: ventral view (A); dorsal view (B); ventral to retrolateral view (C); prolateral view (D); embolar base detail (E); retrolateral view (F). p = prolateral process, TA = terminal apophysis, EB = embolar basis, EM = embolus, M = median apophysis, X = dorsal extension." httpUri-0="https://zenodo.org/record/197223/files/figure.png" httpUri-1="https://zenodo.org/record/197224/files/figure.png" pageId="5" pageNumber="48">Figs 2324</figureCitation>
(m), from
<collectingCountry id="F331719AFFF33B1FFD618F7AFD51CE67" box="[690,749,322,345]" name="Israel" pageId="5" pageNumber="48">Israel</collectingCountry>
(contra
<bibRefCitation id="EFB74CFBFFF33B1FFC958F7AFB9BCE67" author="Chatzaki" box="[838,1063,322,345]" pageId="5" pageNumber="48" refString="Chatzaki, M., Thaler, K. &amp; Mylonas, M. (2003) Ground spiders (Gnaphosidae; Araneae) from Crete and adjacent areas of Greece. Taxonomy and distribution III. Zelotes and allied genera. Revue Suisse de Zoology, 110 (1), 45 - 89." type="journal article" year="2003">
Chatzaki
<emphasis id="B952ED18FFF33B1FFC788F7AFC5DCE67" box="[939,993,322,345]" italics="true" pageId="5" pageNumber="48">et al.</emphasis>
, 2003
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B99310AFFF33B1FFF448F5AFE29CEA7" blockId="5.[151,1436,258,441]" pageId="5" pageNumber="48">
<treatmentCitationGroup id="AB361624FFF33B1FFF448F5AFE29CEA7" pageId="5" pageNumber="48">
<taxonomicName id="4C264A89FFF33B1FFF448F5AFDA8CE47" ID-CoL="7CMHV" authority="Caporiacco, 1953" authorityName="Caporiacco" authorityYear="1953" box="[151,532,354,377]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="B952ED18FFF33B1FFF448F5AFE9BCE47" box="[151,295,354,377]" italics="true" pageId="5" pageNumber="48">T. adriaticus</emphasis>
(Caporiacco, 1953)
</taxonomicName>
:
<treatmentCitation id="0A87171BFFF33B1FFDFB8F5AFCE6CE47" author="Chatzaki" box="[552,858,354,377]" page="57" pageId="5" pageNumber="48" year="2003">
<bibRefCitation id="EFB74CFBFFF33B1FFDFB8F5AFC94CE47" author="Chatzaki" box="[552,808,354,377]" pageId="5" pageNumber="48" refString="Chatzaki, M., Thaler, K. &amp; Mylonas, M. (2003) Ground spiders (Gnaphosidae; Araneae) from Crete and adjacent areas of Greece. Taxonomy and distribution III. Zelotes and allied genera. Revue Suisse de Zoology, 110 (1), 45 - 89." type="journal article" year="2003">
Chatzaki
<emphasis id="B952ED18FFF33B1FFD4B8F5AFD64CE47" box="[664,728,354,377]" italics="true" pageId="5" pageNumber="48">et al.</emphasis>
, 2003
</bibRefCitation>
: 57
</treatmentCitation>
, Figs 3637 (mf), from
<collectingCountry id="F331719AFFF33B1FFB5E8F5AFB67CE47" box="[1165,1243,354,377]" name="Greece" pageId="5" pageNumber="48">Greece</collectingCountry>
, Antikythira isl. (misidentification).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B99310AFFF33B1FFF448F9AFD5FCE87" blockId="5.[151,1436,258,441]" box="[151,739,418,441]" pageId="5" pageNumber="48">
<treatmentCitationGroup id="AB361624FFF33B1FFF448F9AFD5FCE87" box="[151,739,418,441]" pageId="5" pageNumber="48">
<taxonomicName id="4C264A89FFF33B1FFF448F9AFE15CE87" ID-CoL="7CMJK" authority="Levy 2009: 26" authorityName="Levy" authorityPageNumber="26" authorityYear="2009" box="[151,425,418,441]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF33B1FFF448F9AFF40CE87" box="[151,252,418,441]" italics="true" pageId="5" pageNumber="48">T. stubbsi</emphasis>
:
<treatmentCitation id="0A87171BFFF33B1FFED88F9AFE15CE87" ID-DOI="http://doi.org/10.5281/zenodo.3508421" ID-Zenodo-Dep="3508421" author="Levy" box="[267,425,418,441]" httpUri="http://treatment.plazi.org/id/0E2987C979280C373DF6F9EBFD53FADC" page="26" pageId="5" pageNumber="48" year="2009">
<bibRefCitation id="EFB74CFBFFF33B1FFED88F9AFE3DCE87" author="Levy" box="[267,385,418,441]" pageId="5" pageNumber="48" refString="Levy, G. (2009) New ground-spider genera and species with annexed checklist of the Gnaphosidae (Araneae) of Israel. Zootaxa, 2066, 1 - 49." type="journal article" year="2009">Levy 2009</bibRefCitation>
: 26
</treatmentCitation>
</taxonomicName>
, Figs 5758 (Df) from
<collectingCountry id="F331719AFFF33B1FFD738F9AFD63CE87" box="[672,735,418,441]" name="Israel" pageId="5" pageNumber="48">Israel</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF596182FFF33B1FFF44898BFCA1C8D4" httpUri="https://zenodo.org/record/197220/files/figure.png" pageId="5" pageNumber="48" targetBox="[318,1275,482,1935]" targetPageId="5">
<paragraph id="8B99310AFFF33B1FFF44898BFCA1C8D4" blockId="5.[151,1437,1971,2026]" pageId="5" pageNumber="48">
<emphasis id="B952ED18FFF33B1FFF44898BFEFAC8F5" bold="true" box="[151,326,1971,1995]" pageId="5" pageNumber="48">FIGURES 69.</emphasis>
<taxonomicName id="4C264A89FFF33B1FFE9D898BFD84C8F4" box="[334,568,1971,1994]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF33B1FFE9D898BFD84C8F4" box="[334,568,1971,1994]" italics="true" pageId="5" pageNumber="48">Trachyzelotes stubbsi</emphasis>
</taxonomicName>
male. Specimen from Cyprus (6, 8), specimen from Antikythira (7, 9). Ventral view (67). Retrolateral view (89). rp= retrolateral process.
</paragraph>
</caption>
<subSubSection id="C33C6281FFF03B1CFF448EAFFE9ECE52" pageId="6" pageNumber="49" type="diagnosis">
<paragraph id="8B99310AFFF03B1CFF448EAFFA80CE1C" blockId="6.[151,1436,151,626]" pageId="6" pageNumber="49">
<emphasis id="B952ED18FFF03B1CFF448EAFFEA5CF8F" bold="true" box="[151,281,151,177]" pageId="6" pageNumber="49">Diagnosis:</emphasis>
Males of
<taxonomicName id="4C264A89FFF03B1CFE5E8EA0FE40CF8F" box="[397,508,152,177]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF03B1CFE5E8EA0FE40CF8F" box="[397,508,152,177]" italics="true" pageId="6" pageNumber="49">T. stubbsi</emphasis>
</taxonomicName>
are distinguished by other congeners by the bifid and turned tibial apophysis of the palpal organ as well as the characteristic hammer-like shape of the prolateral extension of the terminal apophysis. Females are distinguished by the small spermathecae and the long M-shaped median ridge of the epigyne. Females of
<taxonomicName id="4C264A89FFF03B1CFE588F30FDA3CE1F" box="[395,543,264,289]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="B952ED18FFF03B1CFE588F30FDA3CE1F" box="[395,543,264,289]" italics="true" pageId="6" pageNumber="49">T. adriaticus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C264A89FFF03B1CFD848F30FD7BCE1F" box="[599,711,264,289]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF03B1CFD848F30FD7BCE1F" box="[599,711,264,289]" italics="true" pageId="6" pageNumber="49">T. stubbsi</emphasis>
</taxonomicName>
are inseparable (see details in taxonomic comments).
</paragraph>
<paragraph id="8B99310AFFF03B1CFF158F15FE9ECE52" blockId="6.[151,1436,151,626]" pageId="6" pageNumber="49">
<emphasis id="B952ED18FFF03B1CFF158F15FEC5CE79" bold="true" box="[198,377,301,327]" pageId="6" pageNumber="49">Identification:</emphasis>
<bibRefCitation id="EFB74CFBFFF03B1CFE518F15FD0BCE79" author="Platnick" box="[386,695,301,327]" pageId="6" pageNumber="49" refString="Platnick, N. I. &amp; Murphy, J. A. (1984) A revision of the spider genera Trachyzelotes and Urozelotes (Araneae, Gnaphosidae). American Museum Novitates, 2792, 1 - 30." type="journal article" year="1984">Platnick &amp; Murphy, 1984</bibRefCitation>
: 9,
<figureCitation id="131D2D8FFFF03B1CFD308F15FCD7CE79" box="[739,875,301,327]" captionStart="FIGURES 14 19" captionStartId="10.[151,270,1927,1951]" captionTargetBox="[193,1391,205,1903]" captionTargetId="figure@10.[191,1396,197,1910]" captionTargetPageId="10" captionText="FIGURES 14 19. Zelotes helicoides n. sp. male (14, 16, 18). Zelotes cingarus male (15, 17, 19). Ventral view (14 15); retrolateral view (16 17); dorsal view (18 19). p = prolateral process, TA = terminal apophysis, EB = embolar basis, EM = embolus, M = median apophysis, X = dorsal extension. (From Chatzaki et al. 2003)." httpUri="https://zenodo.org/record/197222/files/figure.png" pageId="6" pageNumber="49">Figs 1516</figureCitation>
,
<bibRefCitation id="EFB74CFBFFF03B1CFCAA8F15FBBECE79" author="Levy" box="[889,1026,301,327]" pageId="6" pageNumber="49" refString="Levy, G. (1998) The ground-spider genera Setaphis, Trachyzelotes, Zelotes and Drassyllus (Araneae: Gnaphosidae) in Israel. Israel Journal of Zoology, 44, 93 - 158." type="journal article" year="1998">Levy, 1998</bibRefCitation>
: 106,
<figureCitation id="131D2D8FFFF03B1CFB988F15FB6FCE79" box="[1099,1235,301,327]" captionStart-0="FIGURES 20 23" captionStart-1="FIGURE 24" captionStartId-0="11.[151,270,1656,1680]" captionStartId-1="12.[151,255,1358,1382]" captionTargetBox-0="[256,1333,192,1634]" captionTargetBox-1="[151,1436,194,1336]" captionTargetId-0="figure@11.[251,1336,178,1634]" captionTargetId-1="figure@12.[151,1436,194,1336]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 20 23. Zelotes helicoides n. sp. female (20 21). Zelotes cingarus female (22 23). Epigyne (20, 22). Vulva (21, 23). (From Chatzaki et al. 2003)." captionText-1="FIGURE 24. Zelotes helicoides n. sp. SEM photos of ♂ palpal organ: ventral view (A); dorsal view (B); ventral to retrolateral view (C); prolateral view (D); embolar base detail (E); retrolateral view (F). p = prolateral process, TA = terminal apophysis, EB = embolar basis, EM = embolus, M = median apophysis, X = dorsal extension." httpUri-0="https://zenodo.org/record/197223/files/figure.png" httpUri-1="https://zenodo.org/record/197224/files/figure.png" pageId="6" pageNumber="49">Figs 2324</figureCitation>
,
<bibRefCitation id="EFB74CFBFFF03B1CFB338F15FAD6CE79" author="Levy" box="[1248,1386,301,327]" pageId="6" pageNumber="49" refString="Levy, G. (2009) New ground-spider genera and species with annexed checklist of the Gnaphosidae (Araneae) of Israel. Zootaxa, 2066, 1 - 49." type="journal article" year="2009">Levy, 2009</bibRefCitation>
: 26, Figs 5758.
</paragraph>
</subSubSection>
<subSubSection id="C33C6281FFF03B1DFF158F4FFD96CBD2" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49" type="materials_examined">
<paragraph id="8B99310AFFF03B1CFF158F4FFDBACD3C" blockId="6.[151,1436,151,626]" pageId="6" pageNumber="49">
<emphasis id="B952ED18FFF03B1CFF158F4FFE81CEAF" bold="true" box="[198,317,375,401]" pageId="6" pageNumber="49">Material:</emphasis>
<collectingCountry id="F331719AFFF03B1CFE958F40FE01CEAC" box="[326,445,376,402]" name="Greece" pageId="6" pageNumber="49">GREECE</collectingCountry>
: Antikythira: Potamos village,
<quantity id="4CDE9CEFFFF03B1CFCE28F40FCC2CEAC" box="[817,894,376,402]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="49" unit="m" value="700.0">700 m</quantity>
west: sparse phrygana on sandy soil close to the village: 10 ɗɗ 3 ΨΨ (
<date id="FF9817CAFFF03B1CFE128FA5FD94CE89" box="[449,552,413,439]" pageId="6" pageNumber="49" value="2001-03-27">27.III.01</date>
to 05.VIII.01) (leg. M. Chatzaki);
<collectingCountry id="F331719AFFF03B1CFC138FA5FB8BCE89" box="[960,1079,413,439]" name="Cyprus" pageId="6" pageNumber="49">CYPRUS</collectingCountry>
: Lemesos, Kourio: 6 ɗɗ 4 ΨΨ (
<date id="FF9817CAFFF03B1CFF4C8FFAFF47CEE2" box="[159,251,450,476]" pageId="6" pageNumber="49" value="2007-05-27">27.V.07</date>
to 01.VIII.07); 9 ɗɗ 4 ΨΨ (
<date id="FF9817CAFFF03B1CFD9B8FFAFD0CCEE2" box="[584,688,450,476]" pageId="6" pageNumber="49" value="2008-03-15">15.III.08</date>
to 17.V.08); 1 ɗ 1 Ψ (
<date id="FF9817CAFFF03B1CFC658FFAFBA1CEE2" box="[950,1053,450,476]" pageId="6" pageNumber="49" value="2007-03-15">15.III.07</date>
to 27.V.07); 1 ɗ 1 Ψ (
<date id="FF9817CAFFF03B1CFAF18FFAFAC1CEE2" box="[1314,1405,450,476]" pageId="6" pageNumber="49" value="2006-05-13">13.V.06</date>
to 09.VII.06) [all leg. D. Kaltsas].
</paragraph>
<paragraph id="8B99310AFFF03B1CFF158C35FE8CCD4C" blockId="6.[151,1436,151,626]" pageId="6" pageNumber="49">
<emphasis id="B952ED18FFF03B1CFF158C35FE5DCD19" bold="true" box="[198,481,525,551]" pageId="6" pageNumber="49">Taxonomic comments.</emphasis>
This species was first described by
<bibRefCitation id="EFB74CFBFFF03B1CFC718C35FB47CD19" author="Platnick" box="[930,1275,525,551]" pageId="6" pageNumber="49" refString="Platnick, N. I. &amp; Murphy, J. A. (1984) A revision of the spider genera Trachyzelotes and Urozelotes (Araneae, Gnaphosidae). American Museum Novitates, 2792, 1 - 30." type="journal article" year="1984">Platnick and Murphy (1984)</bibRefCitation>
from a male collected in
<collectingCountry id="F331719AFFF03B1CFEF88C0AFE39CD72" box="[299,389,562,588]" name="Cyprus" pageId="6" pageNumber="49">Cyprus</collectingCountry>
.
<bibRefCitation id="EFB74CFBFFF03B1CFE428C0AFD9ACD72" author="Levy" box="[401,550,562,588]" pageId="6" pageNumber="49" refString="Levy, G. (1998) The ground-spider genera Setaphis, Trachyzelotes, Zelotes and Drassyllus (Araneae: Gnaphosidae) in Israel. Israel Journal of Zoology, 44, 93 - 158." type="journal article" year="1998">Levy (1998)</bibRefCitation>
confirmed its presence in
<collectingCountry id="F331719AFFF03B1CFCB58C0AFC15CD72" box="[870,937,562,588]" name="Israel" pageId="6" pageNumber="49">Israel</collectingCountry>
and later found and described the female (
<bibRefCitation id="EFB74CFBFFF03B1CFF4C8C60FE9FCD4C" author="Levy" box="[159,291,600,626]" pageId="6" pageNumber="49" refString="Levy, G. (2009) New ground-spider genera and species with annexed checklist of the Gnaphosidae (Araneae) of Israel. Zootaxa, 2066, 1 - 49." type="journal article" year="2009">Levy 2009</bibRefCitation>
).
</paragraph>
<caption id="DF596182FFF03B1CFF448983FBC6C8CD" httpUri="https://zenodo.org/record/197221/files/figure.png" pageId="6" pageNumber="49" targetBox="[187,1391,697,1978]" targetPageId="6">
<paragraph id="8B99310AFFF03B1CFF448983FBC6C8CD" blockId="6.[151,1437,1979,2035]" pageId="6" pageNumber="49">
<emphasis id="B952ED18FFF03B1CFF448983FEDAC8ED" bold="true" box="[151,358,1979,2003]" pageId="6" pageNumber="49">FIGURES 1013.</emphasis>
<taxonomicName id="4C264A89FFF03B1CFEBE8984FDE3C8ED" box="[365,607,1980,2003]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF03B1CFEBE8984FDE3C8ED" box="[365,607,1980,2003]" italics="true" pageId="6" pageNumber="49">Trachyzelotes stubbsi</emphasis>
</taxonomicName>
female, specimen from Cyprus (1011), specimen from Antikythira (12).
<taxonomicName id="4C264A89FFF03B1CFF4489E4FE20C8CD" box="[151,412,2012,2035]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="B952ED18FFF03B1CFF4489E4FE20C8CD" box="[151,412,2012,2035]" italics="true" pageId="6" pageNumber="49">Trachyzelotes adriaticus</emphasis>
</taxonomicName>
female, specimen from Crete (13). Epigyne (10, 12, 13). Vulva (11).
</paragraph>
</caption>
<paragraph id="8B99310AFFF13B1DFF168EA0FA28CEAC" blockId="7.[151,1436,152,1298]" pageId="7" pageNumber="50">
Males and females from the Greek island Antikythira fit the descriptions recorded above. Distinctive characters of the species are also present in the newly collected material from
<collectingCountry id="F331719AFFF13B1DFBF58E85FBC3CFE9" box="[1062,1151,189,215]" name="Cyprus" pageId="7" pageNumber="50">Cyprus</collectingCountry>
: the prolateral extension of the terminal apophysis of the palpal organ has a characteristic hammer-like shape (see
<figureCitation id="131D2D8FFFF13B1DFB028EDAFA80CFC2" box="[1233,1340,226,252]" captionStart="FIGURES 6 9" captionStartId="5.[151,271,1971,1995]" captionTargetBox="[318,1275,482,1935]" captionTargetId="figure@5.[308,1278,482,1950]" captionTargetPageId="5" captionText="FIGURES 6 9. Trachyzelotes stubbsi male. Specimen from Cyprus (6, 8), specimen from Antikythira (7, 9). Ventral view (6 7). Retrolateral view (8 9). rp = retrolateral process." httpUri="https://zenodo.org/record/197220/files/figure.png" pageId="7" pageNumber="50">Figs 67</figureCitation>
) and an extra retrolateral process (rp) of the tegular sclerite exists in all populations examined (see also
<bibRefCitation id="EFB74CFBFFF13B1DFAC88F30FEFECE79" author="Platnick" pageId="7" pageNumber="50" refString="Platnick, N. I. &amp; Murphy, J. A. (1984) A revision of the spider genera Trachyzelotes and Urozelotes (Araneae, Gnaphosidae). American Museum Novitates, 2792, 1 - 30." type="journal article" year="1984">Platnick &amp; Murphy, 1984</bibRefCitation>
, p.10,
<figureCitation id="131D2D8FFFF13B1DFE408F15FE58CE79" box="[403,484,301,327]" captionStart="FIGURES 14 19" captionStartId="10.[151,270,1927,1951]" captionTargetBox="[193,1391,205,1903]" captionTargetId="figure@10.[191,1396,197,1910]" captionTargetPageId="10" captionText="FIGURES 14 19. Zelotes helicoides n. sp. male (14, 16, 18). Zelotes cingarus male (15, 17, 19). Ventral view (14 15); retrolateral view (16 17); dorsal view (18 19). p = prolateral process, TA = terminal apophysis, EB = embolar basis, EM = embolus, M = median apophysis, X = dorsal extension. (From Chatzaki et al. 2003)." httpUri="https://zenodo.org/record/197222/files/figure.png" pageId="7" pageNumber="50">Fig.15</figureCitation>
). Intraspecific variation also occurs in males of this species: in specimens from Antikythira (
<figureCitation id="131D2D8FFFF13B1DFEFD8F6AFEC8CE52" box="[302,372,338,364]" captionStart="FIGURES 6 9" captionStartId="5.[151,271,1971,1995]" captionTargetBox="[318,1275,482,1935]" captionTargetId="figure@5.[308,1278,482,1950]" captionTargetPageId="5" captionText="FIGURES 6 9. Trachyzelotes stubbsi male. Specimen from Cyprus (6, 8), specimen from Antikythira (7, 9). Ventral view (6 7). Retrolateral view (8 9). rp = retrolateral process." httpUri="https://zenodo.org/record/197220/files/figure.png" pageId="7" pageNumber="50">Fig. 7</figureCitation>
) the hammer-like extension is less stout compared to those from
<collectingCountry id="F331719AFFF13B1DFBA08F6AFB74CE52" box="[1139,1224,338,364]" name="Cyprus" pageId="7" pageNumber="50">Cyprus</collectingCountry>
(
<figureCitation id="131D2D8FFFF13B1DFB0B8F6AFAA2CE52" box="[1240,1310,338,364]" captionStart="FIGURES 6 9" captionStartId="5.[151,271,1971,1995]" captionTargetBox="[318,1275,482,1935]" captionTargetId="figure@5.[308,1278,482,1950]" captionTargetPageId="5" captionText="FIGURES 6 9. Trachyzelotes stubbsi male. Specimen from Cyprus (6, 8), specimen from Antikythira (7, 9). Ventral view (6 7). Retrolateral view (8 9). rp = retrolateral process." httpUri="https://zenodo.org/record/197220/files/figure.png" pageId="7" pageNumber="50">Fig. 6</figureCitation>
) and has a more slender base; also specimens from Antikythira are smaller in size than those from
<collectingCountry id="F331719AFFF13B1DFB498F40FB53CEAC" box="[1178,1263,376,402]" name="Cyprus" pageId="7" pageNumber="50">Cyprus</collectingCountry>
(see
<tableCitation id="C6A404B1FFF13B1DFAFF8F40FA3ECEAC" box="[1324,1410,376,402]" captionStart="TABLE 1" captionStartId="4.[151,239,935,959]" captionTargetBox="[151,1436,1061,1809]" captionTargetPageId="4" captionText="TABLE 1. Comparison of somatic and genitalic characters of T. adriaticus and T. stubbsi (abbreviations: CL = carapace" httpUri="http://table.plazi.org/id/DF596182FFF23B1EFF448D9FFA20CC81" pageId="7" pageNumber="50" tableUuid="DF596182FFF23B1EFF448D9FFA20CC81">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8B99310AFFF13B1DFF168FA5FD05CD82" blockId="7.[151,1436,152,1298]" pageId="7" pageNumber="50">
Its epigyne however cannot be distinguished from that of the closely related
<taxonomicName id="4C264A89FFF13B1DFB8F8FA5FB4FCE88" box="[1116,1267,413,438]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="B952ED18FFF13B1DFB8F8FA5FB4FCE88" box="[1116,1267,413,438]" italics="true" pageId="7" pageNumber="50">T. adriaticus</emphasis>
</taxonomicName>
(
<figureCitation id="131D2D8FFFF13B1DFAD78FA5FA33CE89" box="[1284,1423,413,439]" captionStart="FIGURES 10 13" captionStartId="6.[151,274,1979,2003]" captionTargetBox="[187,1391,697,1978]" captionTargetId="figure@6.[175,1411,658,2074]" captionTargetPageId="6" captionText="FIGURES 10 13. Trachyzelotes stubbsi female, specimen from Cyprus (10 11), specimen from Antikythira (12). Trachyzelotes adriaticus female, specimen from Crete (13). Epigyne (10, 12, 13). Vulva (11)." httpUri="https://zenodo.org/record/197221/files/figure.png" pageId="7" pageNumber="50">Figs 1013</figureCitation>
).
<bibRefCitation id="EFB74CFBFFF13B1DFF448FFAFE97CEE2" author="Levy" box="[151,299,450,476]" pageId="7" pageNumber="50" refString="Levy, G. (2009) New ground-spider genera and species with annexed checklist of the Gnaphosidae (Araneae) of Israel. Zootaxa, 2066, 1 - 49." type="journal article" year="2009">Levy (2009)</bibRefCitation>
argued that the M-shaped median ridge (
<emphasis id="B952ED18FFF13B1DFCCD8FFAFCE6CEE5" box="[798,858,450,475]" italics="true" pageId="7" pageNumber="50">keels</emphasis>
according to Levy) of the epigyne takes up two thirds of the epigynal space (
<emphasis id="B952ED18FFF13B1DFE358FD0FD3DCD3F" box="[486,641,488,513]" italics="true" pageId="7" pageNumber="50">sunken space</emphasis>
) as delimited by the anterior margin and that “…
<emphasis id="B952ED18FFF13B1DFB6F8FD0FD68CD18" italics="true" pageId="7" pageNumber="50">the spatial position of the median keels and their lateral expansions</emphasis>
” is the distinctive character of the species as far as females are concerned. Unfortunately he did not give any comparative detail of
<taxonomicName id="4C264A89FFF13B1DFC238C0AFB3ACD75" box="[1008,1158,562,587]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="B952ED18FFF13B1DFC238C0AFB3ACD75" box="[1008,1158,562,587]" italics="true" pageId="7" pageNumber="50">T. adriaticus</emphasis>
</taxonomicName>
which seems to follow the same configuration (compare
<figureCitation id="131D2D8FFFF13B1DFDE68C60FCA2CD4C" box="[565,798,600,626]" captionStart="FIGURES 10 13" captionStartId="6.[151,274,1979,2003]" captionTargetBox="[187,1391,697,1978]" captionTargetId="figure@6.[175,1411,658,2074]" captionTargetPageId="6" captionText="FIGURES 10 13. Trachyzelotes stubbsi female, specimen from Cyprus (10 11), specimen from Antikythira (12). Trachyzelotes adriaticus female, specimen from Crete (13). Epigyne (10, 12, 13). Vulva (11)." httpUri="https://zenodo.org/record/197221/files/figure.png" pageId="7" pageNumber="50">Figs 10, 12 and 13</figureCitation>
); no consistent differences can be seen between the epigynes of both species in specimens from
<collectingCountry id="F331719AFFF13B1DFD778C45FD40CDA9" box="[676,764,637,663]" name="Cyprus" pageId="7" pageNumber="50">Cyprus</collectingCountry>
, Antikythira, Crete and Gavdos-Gavdopoula or from the illustrations given by Levy (p.26, Figs 5758).
</paragraph>
<paragraph id="8B99310AFFF13B1DFF168CF0FC23CCFC" blockId="7.[151,1436,152,1298]" pageId="7" pageNumber="50">
Levy also commented on the distribution of the two species on the Greek territory arguing that no females of
<taxonomicName id="4C264A89FFF13B1DFF648CD5FE9BCC38" box="[183,295,749,774]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF13B1DFF648CD5FE9BCC38" box="[183,295,749,774]" italics="true" pageId="7" pageNumber="50">T. stubbsi</emphasis>
</taxonomicName>
could be identified on the Greek islands as deduced by the illustrations of
<bibRefCitation id="EFB74CFBFFF13B1DFB488CD5FA20CC39" box="[1179,1436,749,775]" pageId="7" pageNumber="50" refString="Chatzaki, M., Thaler, K. &amp; Mylonas, M. (2003) Ground spiders (Gnaphosidae; Araneae) from Crete and adjacent areas of Greece. Taxonomy and distribution III. Zelotes and allied genera. Revue Suisse de Zoology, 110 (1), 45 - 89." type="journal article">
Chatzaki
<emphasis id="B952ED18FFF13B1DFAD88CD5FA82CC38" box="[1291,1342,749,774]" italics="true" pageId="7" pageNumber="50">et al</emphasis>
. (2003)
</bibRefCitation>
but instead only females of
<taxonomicName id="4C264A89FFF13B1DFE358D2AFDC3CC15" box="[486,639,786,811]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="B952ED18FFF13B1DFE358D2AFDC3CC15" box="[486,639,786,811]" italics="true" pageId="7" pageNumber="50">T. adriaticus</emphasis>
</taxonomicName>
were present. However, since Antikythira is a very small island it seems improbable that both species do occur on it sympatrically (males and females were collected at the same traps), especially taking into consideration that the two species are not known to occur on Crete, which is a much larger island. Male and female specimens from
<collectingCountry id="F331719AFFF13B1DFC9D8DBAFC18CCA2" box="[846,932,898,924]" name="Cyprus" pageId="7" pageNumber="50">Cyprus</collectingCountry>
were also collected sympatrically and the females appear no different from the females of the Greek islands.
</paragraph>
<paragraph id="8B99310AFFF13B1DFF168DF5FDD2CB0C" blockId="7.[151,1436,152,1298]" pageId="7" pageNumber="50">Therefore the females of the two species are inseparable in terms of genitalic comparison. Somatic characters are also variable within species so only the presence of males or the evidence from distributional patterns may help to their identification.</paragraph>
<paragraph id="8B99310AFFF13B1DFF168A05FD96CBD2" blockId="7.[151,1436,152,1298]" pageId="7" pageNumber="50">
In conclusion,
<taxonomicName id="4C264A89FFF13B1DFEA68A05FDB7CB68" box="[373,523,1085,1110]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="B952ED18FFF13B1DFEA68A05FDB7CB68" box="[373,523,1085,1110]" italics="true" pageId="7" pageNumber="50">T. adriaticus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C264A89FFF13B1DFD958A05FD04CB68" box="[582,696,1085,1110]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF13B1DFD958A05FD04CB68" box="[582,696,1085,1110]" italics="true" pageId="7" pageNumber="50">T. stubbsi</emphasis>
</taxonomicName>
are quite similar and intraspecific variation occurs in both for the same characters of males (details of the prolateral extension of the terminal apophysis). However, because the two species also differ in the habitus (size, color) and in the general pattern of the prolateral extension of the terminal apophysis of males (hammer-like in
<taxonomicName id="4C264A89FFF13B1DFCD98A95FC39CBF8" box="[778,901,1197,1222]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B952ED18FFF13B1DFCD98A95FC39CBF8" box="[778,901,1197,1222]" italics="true" pageId="7" pageNumber="50">T. stubbsi</emphasis>
</taxonomicName>
and pointed in
<taxonomicName id="4C264A89FFF13B1DFB9E8A95FB51CBF8" box="[1101,1261,1197,1222]" class="Arachnida" family="Gnaphosidae" genus="Trachyzelotes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="adriaticus">
<emphasis id="B952ED18FFF13B1DFB9E8A95FB51CBF8" box="[1101,1261,1197,1222]" italics="true" pageId="7" pageNumber="50">T. adriaticus</emphasis>
</taxonomicName>
) they may be considered as two distinct species.
</paragraph>
</subSubSection>
<subSubSection id="C33C6281FFF13B1DFF158ACFFC8ACA2C" box="[198,822,1271,1298]" pageId="7" pageNumber="50" type="distribution">
<paragraph id="8B99310AFFF13B1DFF158ACFFC8ACA2C" blockId="7.[151,1436,152,1298]" box="[198,822,1271,1298]" pageId="7" pageNumber="50">
<emphasis id="B952ED18FFF13B1DFF158ACFFEDBCA2F" bold="true" box="[198,359,1271,1297]" pageId="7" pageNumber="50">Distribution.</emphasis>
<collectingCountry id="F331719AFFF13B1DFEBD8AC0FE0FCA2C" box="[366,435,1272,1298]" name="Israel" pageId="7" pageNumber="50">Israel</collectingCountry>
;
<collectingCountry id="F331719AFFF13B1DFE6D8AC0FDAACA2C" box="[446,534,1272,1298]" name="Cyprus" pageId="7" pageNumber="50">Cyprus</collectingCountry>
;
<collectingCountry id="F331719AFFF13B1DFDF08AC0FDC7CA2C" box="[547,635,1272,1298]" name="Greece" pageId="7" pageNumber="50">Greece</collectingCountry>
: Antikythira isl.
</paragraph>
</subSubSection>
</treatment>
</document>