253 lines
36 KiB
XML
253 lines
36 KiB
XML
<document id="D45111DBBE9A777FAF316C1A4E369196" ID-CLB-Dataset="295175" ID-DOI="10.15298/rusentj.29.1.06" ID-GBIF-Dataset="89f2997f-194a-42d6-9061-afc4f703dc9b" ID-ISSN="0132-8069" ID-Zenodo-Dep="10978415" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1713265834206" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2020" docId="D6009B3AFFF53B47FF137DDD0DCEFE0D" docLanguage="en" docName="RussEntomolJ.29.1.38-52.pdf" docOrigin="Russian Entomological Journal 29 (1)" docSource="http://dx.doi.org/10.15298/rusentj.29.1.06" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.2:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="2" docTitle="(Tausternus) Fedorenko 2020, subgen.n." docType="treatment" docVersion="4" lastPageNumber="40" masterDocId="2A39E342FFF43B45FFE17C710E41FFFC" masterDocTitle="New taxa of Pterostichini (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber="52" masterPageNumber="38" pageNumber="39" updateTime="1714489442679" updateUser="ExternalLinkService">
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<mods:title id="36704503DE58D97956152D19818C3C11">New taxa of Pterostichini (Coleoptera: Carabidae) from Vietnam</mods:title>
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<mods:namePart id="190901D41028C940294CB4E2F5B15228">Fedorenko, D. N.</mods:namePart>
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<mods:title id="C079AA1D72E6D1C015AF0865AB0497BC">Russian Entomological Journal</mods:title>
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<mods:part id="25DBD6B945F2C3739D2151D10AA3BCDB">
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<mods:date id="87026FFF6C78EA4B953ABD651566A8E8">2020</mods:date>
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<treatment id="D6009B3AFFF53B47FF137DDD0DCEFE0D" ID-DOI="http://doi.org/10.5281/zenodo.11093713" ID-Zenodo-Dep="11093713" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D" httpUri="http://treatment.plazi.org/id/D6009B3AFFF53B47FF137DDD0DCEFE0D" lastPageId="2" lastPageNumber="40" pageId="1" pageNumber="39">
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<subSubSection id="16B379A7FFF53B44FF137DDD0C93FE39" box="[242,722,428,453]" pageId="1" pageNumber="39" type="nomenclature">
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<paragraph id="5E162A2CFFF53B44FF137DDD0C93FE39" blockId="1.[189,776,428,2063]" box="[242,722,428,453]" pageId="1" pageNumber="39">
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<heading id="055E9D40FFF53B44FF137DDD0C93FE39" box="[242,722,428,453]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="39" reason="3">
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Subgenus
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<taxonomicName id="99A951AFFFF53B44FE807DDC0C1BFE38" ID-CoL="8MCQ7" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[353,602,428,452]" kingdom="Animalia" pageId="1" pageNumber="39" rank="subGenus" status="subgen. nov." subGenus="Tausternus">
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<emphasis id="6CDDF63EFFF53B44FE807DDC0F9CFE38" bold="true" box="[353,477,429,452]" italics="true" pageId="1" pageNumber="39">Tausternus</emphasis>
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Fedorenko
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</taxonomicName>
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,
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<emphasis id="6CDDF63EFFF53B44FD847DDC0C93FE39" bold="true" box="[613,722,429,453]" pageId="1" pageNumber="39">
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<taxonomicNameLabel id="77EE4B45FFF53B44FD847DDC0C93FE39" box="[613,722,429,453]" pageId="1" pageNumber="39" rank="subGenus">subgen.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="16B379A7FFF53B44FE487DBB0C5AFE1E" box="[425,539,458,482]" pageId="1" pageNumber="39" type="description">
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<paragraph id="5E162A2CFFF53B44FE487DBB0C5AFE1E" blockId="1.[189,776,428,2063]" box="[425,539,458,482]" pageId="1" pageNumber="39">
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<figureCitation id="C69236A9FFF53B44FE487DBB0C56FE1E" box="[425,535,458,482]" captionStart-0="Figs 1–9" captionStart-1="Figs 10–13" captionStartId-0="2.[226,263,1921,1940]" captionStartId-1="3.[226,263,1873,1892]" captionTargetBox-0="[207,1387,521,1894]" captionTargetBox-1="[192,1396,1226,1849]" captionTargetId-0="figure-254@2.[189,1399,517,1899]" captionTargetId-1="figure-689@3.[189,1399,1221,1853]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs 1–9. Pterostichus (Tausternus) hoii sp.n.: 1 — habitus of paratype ♀; 2, 3 — head and pronotum; 4 — variant form of pronotum; 5 — body apex; 6–9 — left metatibia of male (6, 8) and female (7, 9); 1, 4 — dorsal aspect; 2 — ventral aspect; 3, 5, 8–9 — left lateral aspect; 6–7 — caudal aspect. Рис. 1–9. Pterostichus (Tausternus) hoii sp.n.: 1 — габитус дорЗально, паратип ♀; 2, 3 — голова и переднеспинка; 4 — вариант формы переднеспинки; 5 — верШина тела; 6–9 — ЗаднЯЯ леваЯ голень самца (6, 8) и самки (7, 9); 1, 4 — дорЗально; 2 — вентрально; 3, 5, 8–9 — слева; 6–7 — каудально." captionText-1="Figs 10–13. Pterostichus (Tausternus) hoii sp.n.: 10–11 — median lobe of aedeagus; 12 — right paramere; 13 — female genitalia and reproductive tract (spermathecal gland and major part of spermatheca removed), ventral aspect; 10 — left lateral aspect; 11 — dorsal aspect; bc — bursa copulatrix; gsc — gonosubcoxite; LTIX — laterotergite IX; ov — common oviduct; sbs — spermathecal basal sclerite; sc — seminal canal. Scale bars: 1 mm. Рис. 10–13. Pterostichus (Tausternus) hoii sp.n.: 10–11 — среднЯЯ долЯ Эдеагуса; 12 — праваЯ парамера; 13 — половой аппарат самки (больШаЯ часть сперматеки и желеЗа сперматеки удалены), вентрально; 10 — слева; 11 — дорЗально; bc — копулЯтивнаЯ сумка; gsc — гоносубкоксит; LTIX — латеротергит IX; ov — обЩий Яйцевод; sbs — баЗальный склерит сперматеки; sc — семенной канал. МасШтаб: 1 мм." figureDoi-0="http://doi.org/10.5281/zenodo.10978419" figureDoi-1="http://doi.org/10.5281/zenodo.10978421" httpUri-0="https://zenodo.org/record/10978419/files/figure.png" httpUri-1="https://zenodo.org/record/10978421/files/figure.png" pageId="1" pageNumber="39">Figs 1–13</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="16B379A7FFF53B44FF037D830CD1FDF8" box="[226,656,497,517]" pageId="1" pageNumber="39" type="type_taxon">
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<paragraph id="5E162A2CFFF53B44FF037D830CD1FDF8" blockId="1.[189,776,428,2063]" box="[226,656,497,517]" pageId="1" pageNumber="39">
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<typeStatus id="8112948EFFF53B44FF037D830F4FFDF9" box="[226,270,498,517]" pageId="1" pageNumber="39">Type</typeStatus>
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species:
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<taxonomicName id="99A951AFFFF53B44FE807D800FB7FDF8" authorityName="Fedorenko" authorityYear="2020" box="[353,502,497,516]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="39" phylum="Arthropoda" rank="species" species="hoii">
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<emphasis id="6CDDF63EFFF53B44FE807D800FB7FDF8" box="[353,502,497,516]" italics="true" pageId="1" pageNumber="39">Pterostichus hoii</emphasis>
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</taxonomicName>
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Fedorenko,
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<emphasis id="6CDDF63EFFF53B44FD847D800CD1FDF8" bold="true" box="[613,656,497,516]" pageId="1" pageNumber="39">
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<taxonomicNameLabel id="77EE4B45FFF53B44FD847D800CD1FDF8" box="[613,656,497,516]" pageId="1" pageNumber="39" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="16B379A7FFF53B44FF037E780CE2FA38" pageId="1" pageNumber="39" type="diagnosis">
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<paragraph id="5E162A2CFFF53B44FF037E780CE8FAF6" blockId="1.[189,776,428,2063]" pageId="1" pageNumber="39">
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<materialsCitation id="EEC12071FFF53B44FF037E780CE4FAF6" county="Other" location="Head" municipality="Body" pageId="1" pageNumber="39" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chin">
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DIAGNOSIS. A subgenus of
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<taxonomicName id="99A951AFFFF53B44FE1A7E780C30FDE2" authorityName="Bonelli" authorityYear="1810" box="[507,625,521,542]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="39" phylum="Arthropoda" rank="genus">
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<emphasis id="6CDDF63EFFF53B44FE1A7E780C30FDE2" box="[507,625,521,542]" italics="true" pageId="1" pageNumber="39">Pterostichus</emphasis>
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</taxonomicName>
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recognizable in having prosternum with a very deep T-shaped sulcus, combined with metatibiae and metatarsi incrassate in male.
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<collectingCounty id="B77752A0FFF53B44FD307E4F0D46FDA8" box="[721,775,574,596]" pageId="1" pageNumber="39">Other</collectingCounty>
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features are as follows:
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<collectingMunicipality id="BE72B056FFF53B44FE437E280F94FD93" box="[418,469,601,623]" pageId="1" pageNumber="39">Body</collectingMunicipality>
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apterous, medium-sized for the genus, shiny, metallic, violaceous blue.
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<location id="5B767CF7FFF53B44FDA17E050C33FD76" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FDA17E050C33FD76" box="[576,626,628,650]" county="Other" municipality="Body" name="Head" pageId="1" pageNumber="39" stateProvince="Chin">Head</location>
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medium-sized, microphthalmic, with long genae; frontal foveae impunctate, in form of long and very deep round brackets; terminal palpomeres subfusiform.
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<location id="5B767CF7FFF53B44FE7D7EB50C41FD26" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FE7D7EB50C41FD26" box="[412,512,708,730]" county="Other" municipality="Body" name="Mandibles" pageId="1" pageNumber="39" stateProvince="Chin">Mandibles</location>
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moderately long, slender, nearly straight, with ventral scrobal ridge sinuate in dorsal view.
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<collectingRegion id="9C6DE4CEFFF53B44FF177E880F07FCF2" box="[246,326,761,782]" country="Myanmar" name="Chin" pageId="1" pageNumber="39">Mentum</collectingRegion>
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tooth triangular and blunt.
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<location id="5B767CF7FFF53B44FDA97E880C93FCF2" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FDA97E880C93FCF2" box="[584,722,761,782]" county="Other" municipality="Body" name="Antennomeres" pageId="1" pageNumber="39" stateProvince="Chin">Antennomeres</location>
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4–11 pubescent, scape and pedicel unisetose, segment 3 without setae other than verticellate ones.
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<location id="5B767CF7FFF53B44FDE57F5E0C23FCB8" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FDE57F5E0C23FCB8" box="[516,610,815,836]" county="Other" municipality="Body" name="Pronotum" pageId="1" pageNumber="39" stateProvince="Chin">Pronotum</location>
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subcordate, with two basal sulci on each side, inner long, outer vestigial; disc convex up to lateral bead, without explanate lateral margin.
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<location id="5B767CF7FFF53B44FF5C7F0F0EB6FC68" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FF5C7F0F0EB6FC68" box="[189,247,894,916]" county="Other" municipality="Body" name="Elytra" pageId="1" pageNumber="39" stateProvince="Chin">Elytra</location>
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connate, oblong, with apices blunt and narrowly separated along suture by an oblong fissure; humeri distinct, humeral angle obtuse, basal ridge outside stria 4.
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<location id="5B767CF7FFF53B44FD647FC50C84FC36" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FD647FC50C84FC36" box="[645,709,948,970]" county="Other" municipality="Body" name="Elytral" pageId="1" pageNumber="39" stateProvince="Chin">Elytral</location>
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parascutellar striole missing.
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<location id="5B767CF7FFF53B44FE787FBF0FB8FC18" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FE787FBF0FB8FC18" box="[409,505,974,996]" county="Other" municipality="Body" name="Prosternal" pageId="1" pageNumber="39" stateProvince="Chin">Prosternal</location>
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process conspicuously beaded.
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<location id="5B767CF7FFF53B44FF007F980F0DFC03" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FF007F980F0DFC03" box="[225,332,1001,1023]" county="Other" municipality="Body" name="Abdominal" pageId="1" pageNumber="39" stateProvince="Chin">Abdominal</location>
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sternite VII with a long horn-like process in male.
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<location id="5B767CF7FFF53B44FF1A78750F68FBE5" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FF1A78750F68FBE5" box="[251,297,1028,1049]" county="Other" municipality="Body" name="Legs" pageId="1" pageNumber="39" stateProvince="Chin">Legs</location>
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slender; tarsomeres neither sulcate nor carinate laterally.
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<location id="5B767CF7FFF53B44FEF7786F0F0BFBC8" LSID="urn:lsid:plazi:treatment:D6009B3AFFF53B47FF137DDD0DCEFE0D:5B767CF7FFF53B44FEF7786F0F0BFBC8" box="[278,330,1054,1076]" county="Other" municipality="Body" name="Body" pageId="1" pageNumber="39" stateProvince="Chin">Body</location>
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on each side with fixed setae as follows: head: 2 supra-ocular, 1 clypeal; submentum — 2; mentum — 1; penultimate labial palpomere — 2; maxillary stipes — 2; pronotum — 2; elytra: parascutellar seta in stria 2, discal setae missing, apical setae —
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<quantity id="995187C9FFF53B44FE4878F80F8FFB63" box="[425,462,1161,1183]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="39" unit="in" value="2.0">2 in</quantity>
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stria 7; abdominal sternites IV– VII — 1, sternite VII — 1 (
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<specimenCount id="48AFE1A5FFF53B44FE2578D00F9BFB45" box="[452,474,1185,1209]" count="1" pageId="1" pageNumber="39" type="male">♂</specimenCount>
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) or 2 (
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<specimenCount id="48AFE1A5FFF53B44FDFD78D00C68FB45" box="[540,553,1185,1209]" count="1" pageId="1" pageNumber="39" type="female">♀</specimenCount>
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); legs: trochanter — 1; metacoxa — 3; profemur — 3, posterior; mesofemur — 2, anteroventral; metafemur — 1, anteroventral basal; metatibia without outer setae; tarsomere 5 glabrous ventrally
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="5E162A2CFFF53B44FF03797F0CE2FA38" blockId="1.[189,776,428,2063]" pageId="1" pageNumber="39">
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Aedeagus with right paramere small, apical orifice of median lobe left dorsolateral. Female reproductive tract similar to that of
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<taxonomicName id="99A951AFFFF53B44FED679350EAEFA8F" authority="Tscitscherine, 1898" authorityName="Tscitscherine" authorityYear="1898" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Morphohaptoderus">
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<emphasis id="6CDDF63EFFF53B44FED679350FEEFAA5" box="[311,431,1348,1369]" italics="true" pageId="1" pageNumber="39">Pterostichus</emphasis>
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(
|
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<emphasis id="6CDDF63EFFF53B44FE5A79350C34FAA5" box="[443,629,1348,1369]" italics="true" pageId="1" pageNumber="39">Morphohaptoderus</emphasis>
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) Tscitschérine, 1898
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</taxonomicName>
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, except that spermathecal duct is extremely long and medially balled (receptacle subcylindric, gradually broadened apicad, meeting spermathecal duct at acute angle, with spermathecal gland duct entering just the corner).
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</paragraph>
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</subSubSection>
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<subSubSection id="16B379A7FFF53B44FF0379B80A50F8A7" pageId="1" pageNumber="39" type="description">
|
||
<paragraph id="5E162A2CFFF53B44FF0379B80F5CFA06" blockId="1.[189,776,428,2063]" pageId="1" pageNumber="39">
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DESCRIPTION. Body (
|
||
<figureCitation id="C69236A9FFF53B44FE3079B80C4AFA23" box="[465,523,1481,1503]" captionStart="Figs 1–9" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[207,1387,521,1894]" captionTargetId="figure-254@2.[189,1399,517,1899]" captionTargetPageId="2" captionText="Figs 1–9. Pterostichus (Tausternus) hoii sp.n.: 1 — habitus of paratype ♀; 2, 3 — head and pronotum; 4 — variant form of pronotum; 5 — body apex; 6–9 — left metatibia of male (6, 8) and female (7, 9); 1, 4 — dorsal aspect; 2 — ventral aspect; 3, 5, 8–9 — left lateral aspect; 6–7 — caudal aspect. Рис. 1–9. Pterostichus (Tausternus) hoii sp.n.: 1 — габитус дорЗально, паратип ♀; 2, 3 — голова и переднеспинка; 4 — вариант формы переднеспинки; 5 — верШина тела; 6–9 — ЗаднЯЯ леваЯ голень самца (6, 8) и самки (7, 9); 1, 4 — дорЗально; 2 — вентрально; 3, 5, 8–9 — слева; 6–7 — каудально." figureDoi="http://doi.org/10.5281/zenodo.10978419" httpUri="https://zenodo.org/record/10978419/files/figure.png" pageId="1" pageNumber="39">Fig. 1</figureCitation>
|
||
) apterous, oblong, medium-sized.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FF03798F0D46F948" blockId="1.[189,776,428,2063]" pageId="1" pageNumber="39">Head medium-sized, without neck constriction, each side with a deep supra-ocular groove slightly surpassing eye; eyes small yet moderately convex, dorsolateral; genae long; posterior supra-ocular seta midway between posterior margin of eye and the angle. Frontal foveae C-shaped, long and deep. Antennae nearly filiform, length ratio of antennomeres 1–4 (1.13: 0.6: 1: 0.8), scape with dorsal seta, pedicel with ventral seta.</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FF037AC80FACF8C6" blockId="1.[189,776,428,2063]" pageId="1" pageNumber="39">
|
||
Mandibular scrobe medially with a deep longitudinal sulcus reaching its apex.
|
||
<collectingRegion id="9C6DE4CEFFF53B44FE4B7AA50FBDF915" box="[426,508,1748,1769]" country="Myanmar" name="Chin" pageId="1" pageNumber="39">Mentum</collectingRegion>
|
||
with two close setae at base of a simple and blunt median tooth. Genae ventrally just behind submentum deeply transversely grooved. Terminal labial palpomeres subfusiform.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FF037B4E0A99FE90" blockId="1.[189,776,428,2063]" lastBlockId="1.[812,1399,315,2070]" pageId="1" pageNumber="39">Pronotum rather long, base and apex truncate, basal angles slightly obtuse and blunt, apical angles acute and porrect; basal bead missing, apical bead obliterate in middle third, lateral bead reaching posterolateral seta; basal and apical transverse impressions rather shallow. Inner basal sulci disappearing a third from base, diverging on each side of basal transverse impression and almost reaching base; outer sulcus half as long, running close to and anteriorly joining lateral bead, reaching and rather deep at basal margin; basal fovea impunctate. Posterolateral seta distinctly in front of base.</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FCB37D000D92FD6C" blockId="1.[812,1399,315,2070]" pageId="1" pageNumber="39">
|
||
Elytron with a slight preapical sinuation and narrowly rounded apex, preapical plica well-developed yet internal and invisible in lateral view. Base truncate and narrow, without humeral tooth; humeri distinct yet apically rounded; sides diverging and straight in basal 1/2–2/5. Reflexed lateral margin finely beaded up to apex, neither costate nor carinate, about as wide as interval 9. Striae deep impunctate or indistinctly crenulate. Intervals subequally wide, 9
|
||
<superScript id="A9DC8764FFF53B44FB127E5A0ABFFDCB" attach="left" box="[1267,1278,555,567]" fontSize="5" pageId="1" pageNumber="39">th</superScript>
|
||
with 15–18
|
||
<collectingCountry id="26BE6ABCFFF53B44FCCD7E370D0AFDA7" box="[812,843,582,603]" name="United States of America" pageId="1" pageNumber="39">US</collectingCountry>
|
||
(mostly 16) arranged into anterior group in basal 1/3–1/4 (6–7
|
||
<collectingCountry id="26BE6ABCFFF53B44FCBF7E100D3EFD8A" box="[862,895,609,630]" name="United States of America" pageId="1" pageNumber="39">US</collectingCountry>
|
||
, mostly six), posterior group behind middle, and one intermediate
|
||
<collectingCountry id="26BE6ABCFFF53B44FC4C7E0A0D8EFD6C" box="[941,975,635,656]" name="United States of America" pageId="1" pageNumber="39">US</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FCB37EE70A6EFC9B" blockId="1.[812,1399,315,2070]" pageId="1" pageNumber="39">Underside nearly smooth. Prosternal T-shaped sulcus very deep and almost reaching apex of prosternal process; this process apically rounded or subtruncate and conspicuously beaded; inclination posteriorly flat, wide dorsally, less so ventrally, constricted in between. Metepisternum ca. 0.83 times as long as wide. Abdomen entirely beaded laterally, sternite III with a wide and rather shallow median groove broadened slightly apicad.</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FCB37F1D0A25FB3D" blockId="1.[812,1399,315,2070]" pageId="1" pageNumber="39">
|
||
Legs fairly slender and lightly armed with spiniform setae. Protibia slightly dilated apicad, apically with one posterior and two posterolateral spinules. Metatrochanter rounded apically, less than half as long as metafemur. Mesofemur with two subequally spaced anteroventral setae and two anterodorsal setae in apical two fifths; mesotibia with three outer setae, distal seta inserted closer to anterior ridge. Tarsomeres 1–4 each with one pair of latero-apical setae, tarsomeres 1–2 each with one pair of dorso-apical setae; meso- and metatarsomeres 1–4 with ventral setae rather sparse, being totally reduced in inner (posterior) row, except for 1–3 short setae survived on mesotarsomeres 1–4 and/or on metatarsomere
|
||
<quantity id="995187C9FFF53B44FC7378DA0DF9FB3D" box="[914,952,1195,1217]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="39" unit="in" value="1.0">1 in</quantity>
|
||
some specimens.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FCB378B70B21FB27" blockId="1.[812,1399,315,2070]" box="[850,1376,1222,1244]" pageId="1" pageNumber="39">
|
||
Aedeagus (
|
||
<figureCitation id="C69236A9FFF53B44FC5E78B70A6EFB20" box="[959,1071,1222,1244]" captionStart="Figs 10–13" captionStartId="3.[226,263,1873,1892]" captionTargetBox="[192,1396,1226,1849]" captionTargetId="figure-689@3.[189,1399,1221,1853]" captionTargetPageId="3" captionText="Figs 10–13. Pterostichus (Tausternus) hoii sp.n.: 10–11 — median lobe of aedeagus; 12 — right paramere; 13 — female genitalia and reproductive tract (spermathecal gland and major part of spermatheca removed), ventral aspect; 10 — left lateral aspect; 11 — dorsal aspect; bc — bursa copulatrix; gsc — gonosubcoxite; LTIX — laterotergite IX; ov — common oviduct; sbs — spermathecal basal sclerite; sc — seminal canal. Scale bars: 1 mm. Рис. 10–13. Pterostichus (Tausternus) hoii sp.n.: 10–11 — среднЯЯ долЯ Эдеагуса; 12 — праваЯ парамера; 13 — половой аппарат самки (больШаЯ часть сперматеки и желеЗа сперматеки удалены), вентрально; 10 — слева; 11 — дорЗально; bc — копулЯтивнаЯ сумка; gsc — гоносубкоксит; LTIX — латеротергит IX; ov — обЩий Яйцевод; sbs — баЗальный склерит сперматеки; sc — семенной канал. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978421" httpUri="https://zenodo.org/record/10978421/files/figure.png" pageId="1" pageNumber="39">Figs 10–13</figureCitation>
|
||
) characteristic of
|
||
<taxonomicName id="99A951AFFFF53B44FB0078B70B1BFB27" authorityName="Bonelli" authorityYear="1810" box="[1249,1370,1222,1243]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6CDDF63EFFF53B44FB0078B70B1BFB27" box="[1249,1370,1222,1243]" italics="true" pageId="1" pageNumber="39">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FCB378900B35F9CB" blockId="1.[812,1399,315,2070]" pageId="1" pageNumber="39">
|
||
Female genitalia and reproductive tract. Laterotergite with dense marginal setae; gonosubcoxite setulose along inner margin, with a few (four) strong setae along latero-apical margin. Gonocoxite falcate, with a double nematiform seta near apex and three ensiform setae, two ventral (outer) and one dorsal (inner), and 1–2 smaller additional setae at inner margin.
|
||
<emphasis id="6CDDF63EFFF53B44FC5C79F00A25FA6A" box="[957,1124,1409,1430]" italics="true" pageId="1" pageNumber="39">Bursa copulatrix</emphasis>
|
||
with three large yet fairly weak, contiguous, subtriangular sclerites, and a nodule, sphincter, near base of seminal canal. Spermatheca geniculate, seminal canal sclerotized at base, extremely long, serpentine, medially rolled into an incompact ball, meeting receptacle at acute angle; duct of spermathecal gland enters the corner; receptacle straight, subclavate, gradually broadened apicad.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FCB37A4A0B36F8F0" blockId="1.[812,1399,315,2070]" pageId="1" pageNumber="39">
|
||
Secondary sexual differences (of male): protarsomeres 1– 3 dilated and biseriately squamose on ventral side as usual in the genus; abdominal sternite VII (
|
||
<figureCitation id="C69236A9FFF53B44FB9C7A000AF6F97A" box="[1149,1207,1648,1671]" captionStart="Figs 1–9" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[207,1387,521,1894]" captionTargetId="figure-254@2.[189,1399,517,1899]" captionTargetPageId="2" captionText="Figs 1–9. Pterostichus (Tausternus) hoii sp.n.: 1 — habitus of paratype ♀; 2, 3 — head and pronotum; 4 — variant form of pronotum; 5 — body apex; 6–9 — left metatibia of male (6, 8) and female (7, 9); 1, 4 — dorsal aspect; 2 — ventral aspect; 3, 5, 8–9 — left lateral aspect; 6–7 — caudal aspect. Рис. 1–9. Pterostichus (Tausternus) hoii sp.n.: 1 — габитус дорЗально, паратип ♀; 2, 3 — голова и переднеспинка; 4 — вариант формы переднеспинки; 5 — верШина тела; 6–9 — ЗаднЯЯ леваЯ голень самца (6, 8) и самки (7, 9); 1, 4 — дорЗально; 2 — вентрально; 3, 5, 8–9 — слева; 6–7 — каудально." figureDoi="http://doi.org/10.5281/zenodo.10978419" httpUri="https://zenodo.org/record/10978419/files/figure.png" pageId="1" pageNumber="39">Fig. 5</figureCitation>
|
||
) strongly modified, with a shallow round impression at middle and a fine longitudinal carina at its bottom that gives rise to a large, slightly flattened, horn-like process; meso- and metatibia (
|
||
<figureCitation id="C69236A9FFF53B44FAF97AB00B31F92A" box="[1304,1392,1728,1751]" captionStart="Figs 1–9" captionStartId="2.[226,263,1921,1940]" captionTargetBox="[207,1387,521,1894]" captionTargetId="figure-254@2.[189,1399,517,1899]" captionTargetPageId="2" captionText="Figs 1–9. Pterostichus (Tausternus) hoii sp.n.: 1 — habitus of paratype ♀; 2, 3 — head and pronotum; 4 — variant form of pronotum; 5 — body apex; 6–9 — left metatibia of male (6, 8) and female (7, 9); 1, 4 — dorsal aspect; 2 — ventral aspect; 3, 5, 8–9 — left lateral aspect; 6–7 — caudal aspect. Рис. 1–9. Pterostichus (Tausternus) hoii sp.n.: 1 — габитус дорЗально, паратип ♀; 2, 3 — голова и переднеспинка; 4 — вариант формы переднеспинки; 5 — верШина тела; 6–9 — ЗаднЯЯ леваЯ голень самца (6, 8) и самки (7, 9); 1, 4 — дорЗально; 2 — вентрально; 3, 5, 8–9 — слева; 6–7 — каудально." figureDoi="http://doi.org/10.5281/zenodo.10978419" httpUri="https://zenodo.org/record/10978419/files/figure.png" pageId="1" pageNumber="39">Figs 6–9</figureCitation>
|
||
) plurituberculate along both inner ridges, anterior and posterior; metatibia and metatarsus incrassate, tarsomeres cordiform.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FCB37B600A50F8A7" blockId="1.[812,1399,315,2070]" pageId="1" pageNumber="39">
|
||
<collectionCode id="38B8B2E9FFF53B44FCB37B600DDDF8DA" box="[850,924,1809,1830]" pageId="1" pageNumber="39">NAME</collectionCode>
|
||
. Masculine, combination of Greek letter ‘τ’ [tau] and ‘
|
||
<emphasis id="6CDDF63EFFF53B44FCBE7B5D0DECF8BD" box="[863,941,1836,1857]" italics="true" pageId="1" pageNumber="39">sternum</emphasis>
|
||
’, referring to the very deep T-shaped impression of the prosternum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFF53B44FCB37B100D2CF7EA" pageId="1" pageNumber="39" type="distribution">
|
||
<paragraph id="5E162A2CFFF53B44FCB37B100B10F86D" blockId="1.[812,1399,315,2070]" pageId="1" pageNumber="39">
|
||
DISTRIBUTION. The only species of the subgenus is known to occur in the Ngoc Linh Mt., Central
|
||
<collectingCountry id="26BE6ABCFFF53B44FB167B0A0B0CF86D" box="[1271,1357,1915,1937]" name="Vietnam" pageId="1" pageNumber="39">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF53B44FCB37BE70D2CF7EA" blockId="1.[812,1399,315,2070]" pageId="1" pageNumber="39">
|
||
HABITS AND HABITATS. All specimens of
|
||
<taxonomicName id="99A951AFFFF53B44FACE7BE60DD3F83A" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="39" phylum="Arthropoda" rank="species" species="hoii" status="sp. nov.">
|
||
<emphasis id="6CDDF63EFFF53B44FACE7BE60DD3F83A" italics="true" pageId="1" pageNumber="39">Pterostichus hoii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF53B44FC797BC00D88F83A" bold="true" box="[920,969,1969,1990]" pageId="1" pageNumber="39">
|
||
<taxonomicNameLabel id="77EE4B45FFF53B44FC797BC00D88F83A" box="[920,969,1969,1990]" pageId="1" pageNumber="39" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were collected by pitfall traps in a monsoon forest at
|
||
<quantity id="995187C9FFF53B44FC627BBA0A48F81D" box="[899,1033,1995,2017]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="2.15" metricValueMin="1.6" pageId="1" pageNumber="39" unit="m" value="1875.0" valueMax="2150.0" valueMin="1600.0">1600–2150 m</quantity>
|
||
altitudes, together with some species of the subgenera
|
||
<taxonomicName id="99A951AFFFF53B44FC3F7B960A22F800" box="[990,1123,2023,2044]" pageId="1" pageNumber="38" rank="subGenus" subGenus="Vietosteropus">
|
||
<emphasis id="6CDDF63EFFF53B44FC3F7B960A22F800" box="[990,1123,2023,2044]" italics="true" pageId="1" pageNumber="39">Vietosteropus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="99A951AFFFF53B44FB7B7B970B47F807" authorityName="Andrewes" authorityYear="1937" box="[1178,1286,2022,2043]" pageId="1" pageNumber="52" rank="subGenus" subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF53B44FB7B7B970B47F807" box="[1178,1286,2022,2043]" italics="true" pageId="1" pageNumber="39">Steropanus</emphasis>
|
||
</taxonomicName>
|
||
of
|
||
<taxonomicName id="99A951AFFFF53B44FACE7B960D27F7EA" authorityName="Bonelli" authorityYear="1810" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6CDDF63EFFF53B44FACE7B960D27F7EA" italics="true" pageId="1" pageNumber="39">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFF63B47FF037D4A0DCEFE0D" pageId="2" pageNumber="40" type="discussion">
|
||
<paragraph id="5E162A2CFFF63B47FF037D4A0DCEFE0D" blockId="2.[189,775,315,497]" lastBlockId="2.[812,1399,315,497]" pageId="2" pageNumber="40">
|
||
COMMENTS. This subgenus is monobasic. It is very distinctive in body appearance, including colouration; deeply transversely sulcate prosternum; frontal sulci unusual in shape, deep and wide; mandibles long and sinuate laterally; elytra apically separated and lacking discal setae; hind two leg pairs sexually dimorphic, with tarsi scarcely setose ventrally; abdominal sternite VII strongly modified in male; etc. On the other hand, there is only a little difference between this and some other subgenera of
|
||
<taxonomicName id="99A951AFFFF63B47FBFC7D270AD2FE97" authorityName="Bonelli" authorityYear="1810" box="[1053,1171,342,363]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6CDDF63EFFF63B47FBFC7D270AD2FE97" box="[1053,1171,342,363]" italics="true" pageId="2" pageNumber="40">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
from Southeast Asia in body setation and female genital apparatus. Among them,
|
||
<emphasis id="6CDDF63EFFF63B47FCCD7DFA0DA3FE5C" box="[812,994,395,416]" italics="true" pageId="2" pageNumber="40">Morphohaptoderus</emphasis>
|
||
and probably also
|
||
<taxonomicName id="99A951AFFFF63B47FB457DFA0D1DFE47" authority="Berg, 1898" authorityName="Berg" authorityYear="1898" class="Insecta" family="Carabidae" genus="Tschitscherinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6CDDF63EFFF63B47FB457DFA0B7BFE5C" box="[1188,1338,395,416]" italics="true" pageId="2" pageNumber="40">Tschitscherinea</emphasis>
|
||
Berg, 1898
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="99A951AFFFF63B47FC6B7DD70A2DFE47" authority="Sciaky, 1997" authorityName="Sciaky" authorityYear="1997" box="[906,1132,422,444]" class="Insecta" family="Carabidae" genus="Jedlickaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6CDDF63EFFF63B47FC6B7DD70DAAFE47" box="[906,1003,422,443]" italics="true" pageId="2" pageNumber="40">Jedlickaia</emphasis>
|
||
Sciaky, 1997
|
||
</taxonomicName>
|
||
seem be closest to
|
||
<taxonomicName id="99A951AFFFF63B47FAC77DD60D0CFE2A" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Tausternus">
|
||
<emphasis id="6CDDF63EFFF63B47FAC77DD60D0CFE2A" italics="true" pageId="2" pageNumber="40">Tausternus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF63B47FCB57DB00DF5FE2A" bold="true" box="[852,948,449,470]" pageId="2" pageNumber="40">
|
||
<taxonomicNameLabel id="77EE4B45FFF63B47FCB57DB00DF5FE2A" box="[852,948,449,470]" pageId="2" pageNumber="40" rank="subGenus">subgen.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, all having the metacoxa trisetose (
|
||
<emphasis id="6CDDF63EFFF63B47FAEB7DB00B5FFE2A" box="[1290,1310,449,470]" italics="true" pageId="2" pageNumber="40">vs</emphasis>
|
||
. bisetose in others).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |