229 lines
30 KiB
XML
229 lines
30 KiB
XML
<document id="4604111BB4A4CCA1BFFA70978BC0820A" ID-CLB-Dataset="295175" ID-DOI="10.15298/rusentj.29.1.06" ID-GBIF-Dataset="89f2997f-194a-42d6-9061-afc4f703dc9b" ID-ISSN="0132-8069" ID-Zenodo-Dep="10978415" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1713265834206" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2020" docId="D6009B3AFFF33B4DFCCF7E010B21FD8B" docLanguage="en" docName="RussEntomolJ.29.1.38-52.pdf" docOrigin="Russian Entomological Journal 29 (1)" docSource="http://dx.doi.org/10.15298/rusentj.29.1.06" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.2:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="2" docTitle="Pterostichus (Steropanus) fossifrons Fedorenko 2020, sp.n." docType="treatment" docVersion="6" lastPageNumber="46" masterDocId="2A39E342FFF43B45FFE17C710E41FFFC" masterDocTitle="New taxa of Pterostichini (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber="52" masterPageNumber="38" pageNumber="45" updateTime="1715862488959" updateUser="guilherme">
|
||
<mods:mods id="251B96CD33515453FA8C5ECE88466FF6" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="F48B86299A2C4770359EB4A93C43A8F9">
|
||
<mods:title id="4C44E97B3413D985C95DC32205D08FA2">New taxa of Pterostichini (Coleoptera: Carabidae) from Vietnam</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="68CFD35A0A3ECD9FF24DDA6BB03D20EC" type="personal">
|
||
<mods:role id="AA377DEDC87E6F943053BA992C0659BA">
|
||
<mods:roleTerm id="643ED72458D30EF08B7A2BE21A90F3F1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CC1B583D8B265F0FB67CEFF0A75336FC">Fedorenko, D. N.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="4175BF2D116032654C4339F10F198DAF">text</mods:typeOfResource>
|
||
<mods:relatedItem id="8965C0B9334320F612CC1F5AC57C11D9" type="host">
|
||
<mods:titleInfo id="D399124AE0B8CC7A434AFF8F5FDA43FC">
|
||
<mods:title id="8AFD3AE5346C9EC05459B4DD4C4BB3F7">Russian Entomological Journal</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="047C72BC0BF1113AA89C7D83E2667599">
|
||
<mods:date id="17AE793F15D3C6CBD12DEFAB8AE8DA66">2020</mods:date>
|
||
<mods:detail id="8E9CB1B799320344C8A3F79E926E1011" type="pubDate">
|
||
<mods:number id="22E906BE45F1562AA8A5D383BAECF16A">2020-03-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="6FF50D288F78A0ABE13C3C81C6908DBF" type="volume">
|
||
<mods:number id="DAE69F266FA1F8DEE66F71130627A8FA">29</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F2A1B0C5F24F4EBE9DD4A0DE6C53BCF6" type="issue">
|
||
<mods:number id="330F6F0E95BA216C93134E60C612D2D4">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="6855D35C5E8205B31F9F9744805EE14B" unit="page">
|
||
<mods:start id="27C4B4BB7A0D38DB3F10F833D2023B3D">38</mods:start>
|
||
<mods:end id="F5AF09666CB9FD37BEE30D6C37D80008">52</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="B8727AAE14EB0814D5950585F1066D62">
|
||
<mods:url id="0A51A5E13F11DADEF255F80C900668BB">http://dx.doi.org/10.15298/rusentj.29.1.06</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="D4FC6505220B7BAAC3C515F9C4789461">journal article</mods:classification>
|
||
<mods:identifier id="62DC426EDEA2A37D0989B6293918CAF6" type="CLB-Dataset">295175</mods:identifier>
|
||
<mods:identifier id="BB7DCA9607F322FFC194944BC04CE9EF" type="DOI">10.15298/rusentj.29.1.06</mods:identifier>
|
||
<mods:identifier id="B1B5C7AF4E01F065A77B34F5112D6A24" type="GBIF-Dataset">89f2997f-194a-42d6-9061-afc4f703dc9b</mods:identifier>
|
||
<mods:identifier id="835992D69ECE0BD5C93845D4060463AA" type="ISSN">0132-8069</mods:identifier>
|
||
<mods:identifier id="4BA38EED75AD29802A28FD29A488408B" type="Zenodo-Dep">10978415</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="D6009B3AFFF33B4DFCCF7E010B21FD8B" ID-DOI="http://doi.org/10.5281/zenodo.11093719" ID-GBIF-Taxon="228853514" ID-Zenodo-Dep="11093719" LSID="urn:lsid:plazi:treatment:D6009B3AFFF33B4DFCCF7E010B21FD8B" httpUri="http://treatment.plazi.org/id/D6009B3AFFF33B4DFCCF7E010B21FD8B" lastPageId="8" lastPageNumber="46" pageId="7" pageNumber="45">
|
||
<subSubSection id="16B379A7FFF33B42FCCF7E010B35FD74" box="[814,1396,624,648]" pageId="7" pageNumber="45" type="nomenclature">
|
||
<paragraph id="5E162A2CFFF33B42FCCF7E010B35FD74" blockId="7.[812,1399,624,2064]" box="[814,1396,624,648]" pageId="7" pageNumber="45">
|
||
<heading id="055E9D40FFF33B42FCCF7E010B35FD74" box="[814,1396,624,648]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="45" reason="4">
|
||
<taxonomicName id="99A951AFFFF33B42FCCF7E010B72FD74" ID-CoL="8M739" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[814,1331,624,648]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="fossifrons" subGenus="Steropanus">
|
||
<emphasis id="6CDDF63EFFF33B42FCCF7E010DF5FD74" box="[814,948,624,648]" italics="true" pageId="7" pageNumber="45">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="6CDDF63EFFF33B42FC247E010A7CFD7B" box="[965,1085,624,647]" italics="true" pageId="7" pageNumber="45">Steropanus</emphasis>
|
||
)
|
||
<emphasis id="6CDDF63EFFF33B42FBAF7E010AF7FD74" box="[1102,1206,624,648]" italics="true" pageId="7" pageNumber="45">fossifrons</emphasis>
|
||
Fedorenko
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="6CDDF63EFFF33B42FADE7E010B35FD74" bold="true" box="[1343,1396,624,648]" pageId="7" pageNumber="45">
|
||
<taxonomicNameLabel id="77EE4B45FFF33B42FADE7E010B35FD74" box="[1343,1396,624,648]" pageId="7" pageNumber="45" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFF33B42FC087EFC0AFBFD59" box="[1001,1210,653,677]" pageId="7" pageNumber="45" type="description">
|
||
<paragraph id="5E162A2CFFF33B42FC087EFC0AFBFD59" blockId="7.[812,1399,624,2064]" box="[1001,1210,653,677]" pageId="7" pageNumber="45">
|
||
<figureCitation id="C69236A9FFF33B42FC087EFC0A22FD59" box="[1001,1123,653,677]" captionStart-0="Figs 14–16" captionStart-1="Figs 17–22" captionStartId-0="4.[226,263,1969,1988]" captionStartId-1="5.[226,262,1921,1940]" captionTargetBox-0="[193,1367,1150,1943]" captionTargetBox-1="[194,1391,758,1882]" captionTargetId-0="figure-586@4.[189,1381,1138,1949]" captionTargetId-1="figure-432@5.[189,1399,755,1886]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figs 14–16. Dorsal habitus, holotypes: 14 — Pterostichus (Steropanus) securipenis sp.n; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n. Рис. 14–16. Габитус дорЗально, голотипы: 14 — Pterostichus (Steropanus) securipenis sp.n.; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n." captionText-1="Figs 17–22. Aedeagus of Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 17–20 — median lobe; 17, 19, 21 — left lateral aspect; 18, 20, 22 — right lateral aspect. Scale bars: 1 mm. Рис. 17–22. Эдеагус Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 17–20 — среднЯЯ долЯ; 17, 19, 21 — слева; 18, 20, 22 — справа. МасШтаб: 1 мм." figureDoi-0="http://doi.org/10.5281/zenodo.10978425" figureDoi-1="http://doi.org/10.5281/zenodo.10978427" httpUri-0="https://zenodo.org/record/10978425/files/figure.png" httpUri-1="https://zenodo.org/record/10978427/files/figure.png" pageId="7" pageNumber="45">Figs 16–18</figureCitation>
|
||
,
|
||
<figureCitation id="C69236A9FFF33B42FB8E7EFC0AF7FD59" box="[1135,1206,653,677]" captionStart="Figs 23–29" captionStartId="6.[226,263,1921,1940]" captionTargetBox="[192,1364,1003,1906]" captionTargetId="figure-533@6.[189,1369,999,1909]" captionTargetPageId="6" captionText="Figs 23–29. Aedeagus of Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (apical margin slightly damaged); 25, 28 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — median lobe, ventral aspect; 26–28 — inside of right paramere. Scale bars: 1 mm. Рис. 23–29. Эдеагус Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (апикальный край слегка обломан); 25, 28 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — среднЯЯ долЯ? вентрально; 26–28 — праваЯ парамера, иЗнутри. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978429" httpUri="https://zenodo.org/record/10978429/files/figure.png" pageId="7" pageNumber="45">23, 26</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFF33B42FCB37EC30B2CFD09" pageId="7" pageNumber="45" type="materials_examined">
|
||
<paragraph id="5E162A2CFFF33B42FCB37EC30B2CFD09" blockId="7.[812,1399,624,2064]" pageId="7" pageNumber="45">
|
||
<materialsCitation id="EEC12071FFF33B42FCB37EC30B2CFD09" ID-GBIF-Occurrence="4850364317" collectingDate="2018-05-22" collectingDateMax="2018-06-06" collectingDateMin="2018-05-22" collectionCode="ZMMU" collectorName="Phia Oac Mt., E & A. Abramov'" elevation="1700" latitude="22.6075" location="Phia Oac Mt." longLatPrecision="21" longitude="105.86667" pageId="7" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Cao Bang" typeStatus="holotype">
|
||
MATERIAL.
|
||
<typeStatus id="8112948EFFF33B42FC2F7EC30A5FFD39" box="[974,1054,690,709]" pageId="7" pageNumber="45" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="48AFE1A5FFF33B42FBC77EDE0A73FD39" box="[1062,1074,687,709]" count="1" pageId="7" pageNumber="45" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="38B8B2E9FFF33B42FBA17EC30AC6FD39" box="[1088,1159,690,709]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="7" pageNumber="45" type="Museum">ZMMU</collectionCode>
|
||
), labelled: ‘N-Vietnam,
|
||
<quantity id="995187C9FFF33B42FA807EC30D06FD21" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="45" unit="km" value="40.0">40 km</quantity>
|
||
W of
|
||
<collectingRegion id="9C6DE4CEFFF33B42FC647EBB0D9FFD21" box="[901,990,714,733]" country="Vietnam" name="Cao Bang" pageId="7" pageNumber="45">Cao Bang</collectingRegion>
|
||
,
|
||
<collectorName id="F35C4FFAFFF33B42FC0B7EBB0A3AFD21" box="[1002,1147,714,733]" pageId="7" pageNumber="45">
|
||
<location id="5B767CF7FFF33B42FC0B7EBB0A22FD21" LSID="urn:lsid:plazi:treatment:D6009B3AFFF33B4DFCCF7E010B21FD8B:5B767CF7FFF33B42FC0B7EBB0A22FD21" box="[1002,1123,714,733]" latitude="22.6075" longLatPrecision="21" longitude="105.86667" name="Phia Oac Mt." pageId="7" pageNumber="45" stateProvince="Cao Bang">Phia Oac Mt.</location>
|
||
, E
|
||
</collectorName>
|
||
[astern]-slope, h=
|
||
<quantity id="995187C9FFF33B42FAF47EBB0D7EFD09" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="7" pageNumber="45" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
|
||
<elevation id="D584CD1FFFF33B42FAF47EBB0D7EFD09" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="7" pageNumber="45" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">1600–1800 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="3B9D4CEBFFF33B42FCAB7E930DF4FD09" box="[842,949,738,757]" degrees="22" direction="north" minutes="36" orientation="latitude" pageId="7" pageNumber="45" precision="15" seconds="27" value="22.6075">22°36´27´´N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="3B9D4CEBFFF33B42FC227E930A6FFD09" box="[963,1070,738,757]" degrees="105" direction="east" minutes="52" orientation="longitude" pageId="7" pageNumber="45" precision="15" seconds="0" value="105.86667">105°52´0´´E</geoCoordinate>
|
||
,
|
||
<date id="2A170CECFFF33B42FBD87E930A86FD09" box="[1081,1223,738,757]" pageId="7" pageNumber="45" value="2018-05-22" valueMax="2018-06-06" valueMin="2018-05-22">
|
||
<collectingDate id="3A53F504FFF33B42FBD87E930A86FD09" box="[1081,1223,738,757]" pageId="7" pageNumber="45" value="2018-05-22" valueMax="2018-06-06" valueMin="2018-05-22">22.V–6.VI.2018</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="F35C4FFAFFF33B42FB187E930B2BFD09" box="[1273,1386,738,757]" pageId="7" pageNumber="45">A.Abramov’</collectorName>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFF33B42FCB37E8B0D92FBE6" pageId="7" pageNumber="45" type="diagnosis">
|
||
<paragraph id="5E162A2CFFF33B42FCB37E8B0D92FBE6" blockId="7.[812,1399,624,2064]" pageId="7" pageNumber="45">
|
||
DIAGNOSIS. This species is similar to
|
||
<taxonomicName id="99A951AFFFF33B42FB017E8A0DF9FCD7" authority="(Andrewes, 1937)" baseAuthorityName="Andrewes" baseAuthorityYear="1937" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="aequus">
|
||
<emphasis id="6CDDF63EFFF33B42FB017E8A0B00FCEC" box="[1248,1345,763,784]" italics="true" pageId="7" pageNumber="45">P. aequus</emphasis>
|
||
(
|
||
<bibRefCitation id="3A3857DDFFF33B42FAAE7E8B0DF0FCD7" author="Andrewes H. E." pageId="7" pageNumber="45" pagination="489 - 495" refId="ref13093" refString="Andrewes H. E. 1937. Papers on Oriental Carabidae. - XXXI // Ann. Mag. Nat. Hist. Ser. 9. Vol. 19. P. 489 - 495." type="journal article" year="1937">Andrewes, 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in appearance. According to the description,
|
||
<taxonomicName id="99A951AFFFF33B42FCCD7F410DD3FCB9" baseAuthorityName="Andrewes" baseAuthorityYear="1937" box="[812,914,816,837]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="aequus">
|
||
<emphasis id="6CDDF63EFFF33B42FCCD7F410DD3FCB9" box="[812,914,816,837]" italics="true" pageId="7" pageNumber="45">P. aequus</emphasis>
|
||
</taxonomicName>
|
||
has pronotal sides indistinctly sinuate, frontal sulci less deep, pronotal basal angles obtuse and more rounded, and impressed line (= outer basal sulcus extended into a remnant of basal bead) less deep than in
|
||
<taxonomicName id="99A951AFFFF33B42FAFE7FF10D24FC4C" baseAuthorityName="Andrewes" baseAuthorityYear="1937" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="glymmiger">
|
||
<emphasis id="6CDDF63EFFF33B42FAFE7FF10D24FC4C" italics="true" pageId="7" pageNumber="45">P. glymmiger</emphasis>
|
||
</taxonomicName>
|
||
. In the new species, pronotal sides are distinctly sinuate just in front of basal angles, frontal sulci very deep, much deeper than those of
|
||
<taxonomicName id="99A951AFFFF33B42FBD47FA10AF8FC18" baseAuthorityName="Andrewes" baseAuthorityYear="1937" box="[1077,1209,975,997]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="glymmiger">
|
||
<emphasis id="6CDDF63EFFF33B42FBD47FA10AF8FC18" box="[1077,1209,975,997]" italics="true" pageId="7" pageNumber="45">P. glymmiger</emphasis>
|
||
</taxonomicName>
|
||
, pronotal angles in form of a blunt obtuse tooth, and the impressed line just as in
|
||
<taxonomicName id="99A951AFFFF33B42FCA678740D8CFBE6" baseAuthorityName="Andrewes" baseAuthorityYear="1937" box="[839,973,1029,1050]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="glymmiger">
|
||
<emphasis id="6CDDF63EFFF33B42FCA678740D8CFBE6" box="[839,973,1029,1050]" italics="true" pageId="7" pageNumber="45">P. glymmiger</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFF33B4DFCB3786E0A17FDF7" lastPageId="8" lastPageNumber="46" pageId="7" pageNumber="45" type="description">
|
||
<paragraph id="5E162A2CFFF33B42FCB3786E0B20FB0C" blockId="7.[812,1399,624,2064]" pageId="7" pageNumber="45">
|
||
DESCRIPTION. BL
|
||
<quantity id="995187C9FFF33B42FBCD786E0ACDFBC9" box="[1068,1164,1055,1077]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="7" pageNumber="45" unit="mm" value="13.9">13.9 mm</quantity>
|
||
. Body (
|
||
<figureCitation id="C69236A9FFF33B42FB03786E0B6FFBC9" box="[1250,1326,1055,1077]" captionStart="Figs 14–16" captionStartId="4.[226,263,1969,1988]" captionTargetBox="[193,1367,1150,1943]" captionTargetId="figure-586@4.[189,1381,1138,1949]" captionTargetPageId="4" captionText="Figs 14–16. Dorsal habitus, holotypes: 14 — Pterostichus (Steropanus) securipenis sp.n; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n. Рис. 14–16. Габитус дорЗально, голотипы: 14 — Pterostichus (Steropanus) securipenis sp.n.; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n." figureDoi="http://doi.org/10.5281/zenodo.10978425" httpUri="https://zenodo.org/record/10978425/files/figure.png" pageId="7" pageNumber="45">Fig. 16</figureCitation>
|
||
) shiny black. Antennae, palps, femoral apices, tibiae and tarsi rather dark reddish brown. Dorsal microsculpture fine-meshed: isodiametric and almost imperceptible on head; superficial yet distinct over pronotum and elytra, consisting of barely transverse or narrow transverse meshes, respectively. Head rather densely micropunctate, pronotal punctation much sparser and much finer in part; elytral micropunctation indistinct.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF33B42FCB378840A93F9E9" blockId="7.[812,1399,624,2064]" pageId="7" pageNumber="45">Head: Eyes as in previous species. Frontal sulci indistinctly crenulate at bottom, very deep S-shaped, running parallel to each other just behind clypeus, strongly diverging on frons, slightly incurved and then abruptly disappearing just in front of the level of anterior supra-ocular seta; finest posterior extensions of frontal sulci diverging toward, running just inside, and disappearing just behind the level of postero-ocular setae. Supra-ocular groove deep, slightly incurved anteriorly. Labrum subsinuate apically, sexsetose, with inner four setae slightly separate from lateral ones. Antennae not quite reaching pronotal base.</paragraph>
|
||
<paragraph id="5E162A2CFFF33B42FCB37A6B0D9BF97C" blockId="7.[812,1399,624,2064]" pageId="7" pageNumber="45">
|
||
Penultimate labial palpomere as in
|
||
<taxonomicName id="99A951AFFFF33B42FB537A6A0B7CF9CC" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[1202,1341,1563,1584]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="securipenis" status="sp.n.">
|
||
<emphasis id="6CDDF63EFFF33B42FB537A6A0B7CF9CC" box="[1202,1341,1563,1584]" italics="true" pageId="7" pageNumber="45">P. securipenis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFF33B42FAA77A6B0B37F9D3" bold="true" box="[1350,1398,1562,1583]" pageId="7" pageNumber="45">
|
||
<taxonomicNameLabel id="77EE4B45FFF33B42FAA77A6B0B37F9D3" box="[1350,1398,1562,1583]" pageId="7" pageNumber="45" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Terminal labial palpomere two thirds as wide at apex as long at inner margin, somewhat pentagonal because of very convex outer margin.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFF33B4DFCB37AF40C7CFE7B" blockId="7.[812,1399,624,2064]" lastBlockId="8.[189,775,315,1057]" lastPageId="8" lastPageNumber="46" pageId="7" pageNumber="45">
|
||
Pronotum cordate,
|
||
<collectionCode id="38B8B2E9FFF33B42FBEC7AF40A70F966" box="[1037,1073,1669,1690]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="45">PW</collectionCode>
|
||
/
|
||
<collectionCode id="38B8B2E9FFF33B42FBD97AF40A15F966" box="[1080,1108,1669,1690]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="45" type="Museum">PL</collectionCode>
|
||
1.18,
|
||
<collectionCode id="38B8B2E9FFF33B42FB727AF40AF6F966" box="[1171,1207,1669,1690]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="45">PW</collectionCode>
|
||
/HW 1.65, broadest two fifths from apex, PLw/
|
||
<collectionCode id="38B8B2E9FFF33B42FBDE7AEE0A1AF948" box="[1087,1115,1695,1716]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="45" type="Museum">PL</collectionCode>
|
||
0.42; sides evenly rounded, sinuate just in front of basal angles. Base slightly sinuate medially, oblique towards basal angles, as wide as apex, PB/ PA 1.02; basal angles as fairly small, obtuse and blunt tooth. Apex gently sinuate, with a slight convexity towards subrectangular and blunt apical angles. Apical bead vestigial, extremely fine and hardly traceable, obliterate in middle third; lateral bead entire, rather thin; lateral groove crenulate at bottom, deep and narrow in basal half, barely wider and slightly more shallow in front. Inner basal sulci vestigial, almost indistinct yet long, running parallel to each other on basal two fifths, obliterate basally; basal sulcus, combined with a remnant of basal bead, as a rectangular impressed line separated by a fold from basal angle, this fold being as wide as lateral bead. Disc smooth, except for sparse, ill-defined transverse rugosities. Median line, transverse impressions and lateral setae as in previous species.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFFC3B4DFF037DFA0C4CFCB0" blockId="8.[189,775,315,1057]" pageId="8" pageNumber="46">
|
||
Elytra connate, elliptic, broadest medially, EW/EL 1.55, EW/
|
||
<collectionCode id="38B8B2E9FFFC3B4DFF097DD70F4DFE47" box="[232,268,422,443]" country="Thailand" name="Paleontological Collections" pageId="8" pageNumber="46">PW</collectionCode>
|
||
1.20, apices rounded combined, preapical sinuation imperceptible, plica internal. Base slightly oblique, humeri rather widely rounded, humeral tooth just traceable as a minute tubercle between lateral bead and basal ridge; this latter reaching stria 1. Striae deep, minutely punctate before apex, otherwise nearly smooth, 10
|
||
<superScript id="A9DC8764FFFC3B4DFE797E5A0FE2FDCB" attach="left" box="[408,419,555,567]" fontSize="5" pageId="8" pageNumber="46">th</superScript>
|
||
finely yet distinctly punctate in basal three fifths; striae 1–6 adjoining basal ridge; parascutellar striole missing. Intervals convex, slightly more so laterally and apically, 7
|
||
<superScript id="A9DC8764FFFC3B4DFEFF7E0A0F68FD7B" attach="none" box="[286,297,635,647]" fontSize="5" pageId="8" pageNumber="46">th</superScript>
|
||
, 5
|
||
<superScript id="A9DC8764FFFC3B4DFEA17E0A0F0AFD7B" attach="left" box="[320,331,635,647]" fontSize="5" pageId="8" pageNumber="46">th</superScript>
|
||
and 3
|
||
<superScript id="A9DC8764FFFC3B4DFE627E0A0FCFFD7B" attach="left" box="[387,398,635,647]" fontSize="5" pageId="8" pageNumber="46">rd</superScript>
|
||
confluent apicad in succession and then joining 2
|
||
<superScript id="A9DC8764FFFC3B4DFEF37EE40F61FD5D" attach="left" box="[274,288,661,673]" fontSize="5" pageId="8" pageNumber="46">nd</superScript>
|
||
and 8
|
||
<superScript id="A9DC8764FFFC3B4DFEBB7EE40F24FD5D" attach="none" box="[346,357,661,673]" fontSize="5" pageId="8" pageNumber="46">th</superScript>
|
||
; 1
|
||
<superScript id="A9DC8764FFFC3B4DFE9F7EE40FC6FD5D" attach="left" box="[382,391,661,673]" fontSize="5" pageId="8" pageNumber="46">st</superScript>
|
||
merging in lateral bead at sutural angle; 8
|
||
<superScript id="A9DC8764FFFC3B4DFF297EC10E92FD40" attach="left" box="[200,211,688,700]" fontSize="5" pageId="8" pageNumber="46">th</superScript>
|
||
costate and very narrow, subcarinate before preapical plica, as wide as 9
|
||
<superScript id="A9DC8764FFFC3B4DFECE7EBA0F7BFD2B" attach="left" box="[303,314,715,727]" fontSize="5" pageId="8" pageNumber="46">th</superScript>
|
||
and less than half as wide as 7
|
||
<superScript id="A9DC8764FFFC3B4DFDBA7EBA0C27FD2B" attach="none" box="[603,614,715,727]" fontSize="5" pageId="8" pageNumber="46">th</superScript>
|
||
. Reflexed lateral margin narrow, slightly broadened apicad and barely convex in apical third. Parascutellar seta near base of stria
|
||
<quantity id="995187C9FFFC3B4DFD9E7F700CE2FCEB" box="[639,675,769,791]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="46" unit="in" value="2.0">2 in</quantity>
|
||
interval 2, discal setae missing, stria 7 with one preapical seta, anterior, distant far from apex; USS: 23–25.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFFC3B4DFF037F200CF5FBDD" blockId="8.[189,775,315,1057]" pageId="8" pageNumber="46">Underside impunctate, except for mesepisternum moderately and densely punctate across middle. Prosternum with a shallow median impression in front of procoxae. Prosternal process in ventral view truncate, with rounded angles; in lateral view rectangular, rounded apically; inclination wide, barely concave between just traceable lateral beads, slightly constricted at middle; vertical and straight in lateral view. Abdominal lateral bead entire; sternite VII similar in both sexes.</paragraph>
|
||
<paragraph id="5E162A2CFFFC3B4DFCB37D4D0B34FE5D" blockId="8.[812,1399,315,631]" pageId="8" pageNumber="46">
|
||
Legs as in
|
||
<taxonomicName id="99A951AFFFFC3B4DFC5B7D4D0AC0FEAC" authority="Fedorenko, 2020" authorityName="Fedorenko" authorityYear="2020" box="[954,1153,315,337]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="pseudoglymmiger" status="sp.n.">
|
||
<emphasis id="6CDDF63EFFFC3B4DFC5B7D4D0AC0FEAC" box="[954,1153,315,337]" italics="true" pageId="8" pageNumber="46">P. pseudoglymmiger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6CDDF63EFFFC3B4DFB667D4A0AF9FEAC" bold="true" box="[1159,1208,315,336]" pageId="8" pageNumber="46">
|
||
<taxonomicNameLabel id="77EE4B45FFFC3B4DFB667D4A0AF9FEAC" box="[1159,1208,315,336]" pageId="8" pageNumber="46" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, except for protibia with 5–6 spinules at apex of posterolateral edge; tarsomere 5 glabrous ventrally; apical setal brush of mesotibia triangular, more than one-row; and metatibia glabrous at outer margin.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFFC3B4DFCB37DD70A17FDF7" blockId="8.[812,1399,315,631]" pageId="8" pageNumber="46">
|
||
Aedeagus (
|
||
<figureCitation id="C69236A9FFFC3B4DFC5E7DD70A6EFE47" box="[959,1071,422,444]" captionStart="Figs 17–22" captionStartId="5.[226,262,1921,1940]" captionTargetBox="[194,1391,758,1882]" captionTargetId="figure-432@5.[189,1399,755,1886]" captionTargetPageId="5" captionText="Figs 17–22. Aedeagus of Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 17–20 — median lobe; 17, 19, 21 — left lateral aspect; 18, 20, 22 — right lateral aspect. Scale bars: 1 mm. Рис. 17–22. Эдеагус Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 17–20 — среднЯЯ долЯ; 17, 19, 21 — слева; 18, 20, 22 — справа. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978427" httpUri="https://zenodo.org/record/10978427/files/figure.png" pageId="8" pageNumber="46">Figs 17–18</figureCitation>
|
||
,
|
||
<figureCitation id="C69236A9FFFC3B4DFBD87DD70A3BFE40" box="[1081,1146,422,444]" captionStart="Figs 23–29" captionStartId="6.[226,263,1921,1940]" captionTargetBox="[192,1364,1003,1906]" captionTargetId="figure-533@6.[189,1369,999,1909]" captionTargetPageId="6" captionText="Figs 23–29. Aedeagus of Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (apical margin slightly damaged); 25, 28 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — median lobe, ventral aspect; 26–28 — inside of right paramere. Scale bars: 1 mm. Рис. 23–29. Эдеагус Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (апикальный край слегка обломан); 25, 28 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — среднЯЯ долЯ? вентрально; 26–28 — праваЯ парамера, иЗнутри. МасШтаб: 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10978429" httpUri="https://zenodo.org/record/10978429/files/figure.png" pageId="8" pageNumber="46">23, 26</figureCitation>
|
||
): Median lobe geniculate; apex in dorsal view large, with sides barely diverging apicad, widely rounded apically. Right paramere moderately long, crescent, more curved at apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFFC3B4DFCB37E600AA6FDBD" pageId="8" pageNumber="46" type="distribution">
|
||
<paragraph id="5E162A2CFFFC3B4DFCB37E600B20FDDB" blockId="8.[812,1399,315,631]" box="[850,1377,529,551]" pageId="8" pageNumber="46">
|
||
DISTRIBUTION. Known from the
|
||
<typeStatus id="8112948EFFFC3B4DFB517E600A9BFDDA" box="[1200,1242,529,550]" pageId="8" pageNumber="46">type</typeStatus>
|
||
locality only.
|
||
</paragraph>
|
||
<paragraph id="5E162A2CFFFC3B4DFCB37E5A0AA6FDBD" blockId="8.[812,1399,315,631]" box="[850,1255,555,577]" pageId="8" pageNumber="46">
|
||
<collectionCode id="38B8B2E9FFFC3B4DFCB37E5A0DDDFDBC" box="[850,924,555,576]" pageId="8" pageNumber="46">NAME</collectionCode>
|
||
. Refers to very deep frontal sulci.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16B379A7FFFC3B4DFCB37E370B21FD8B" pageId="8" pageNumber="46" type="description">
|
||
<paragraph id="5E162A2CFFFC3B4DFCB37E370B21FD8B" blockId="8.[812,1399,315,631]" pageId="8" pageNumber="46">
|
||
HABITATS AND HABITS. The
|
||
<typeStatus id="8112948EFFFC3B4DFB777E370AAAFDA0" box="[1174,1259,582,604]" pageId="8" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
specimen was collected by pitfall trap in a montane broad-leaved forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |