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<document id="344657E810A62FEA9901A7EC7F41F7A7" ID-DOI="10.11646/zootaxa.3635.3.5" ID-GBIF-Dataset="aa1e5dd3-98d2-4982-a52e-1bcec0907bb5" ID-ISSN="1175-5326" ID-Zenodo-Dep="222217" ID-ZooBank="E36F75A7-73F4-4A4E-869A-427C2EE788B1" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460586202895" checkinUser="plazi" docAuthor="Melnitsky, Stanislav I. &amp; Ivanov, Vladimir D." docDate="2013" docId="03FC5411FFBCFFFFFF2998C42165F8D5" docLanguage="en" docName="zt03635p268.pdf" docOrigin="Zootaxa 3635 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archaeotinodes petropolitana Melnitsky &amp; Ivanov, new species" docType="treatment" docVersion="8" lastPageNumber="263" masterDocId="FFC52C69FFBDFFFDFFBE996A230EFF8F" masterDocTitle="Three new caddisflies species of the fossil genus Archaeotinodes (Insecta: Trichoptera: Ecnomidae) from the Baltic Amber" masterLastPageNumber="268" masterPageNumber="261" pageNumber="262" updateTime="1698325239223" updateUser="plazi">
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<mods:title id="90598FBA022DDA812D98C6044003BB4C">Three new caddisflies species of the fossil genus Archaeotinodes (Insecta: Trichoptera: Ecnomidae) from the Baltic Amber</mods:title>
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<mods:name id="135B95DC13AC2F92C5AB8CC5B5C4679F" type="personal">
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<mods:namePart id="EACA26147450C2AF25787D86034DD9D2">Melnitsky, Stanislav I.</mods:namePart>
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<mods:namePart id="8D7FBE9E217772FF3FEEABE466FB0405">Ivanov, Vladimir D.</mods:namePart>
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<mods:title id="777D35D72C04C5D8E661B5E447A45591">Zootaxa</mods:title>
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<mods:date id="C626583AF729EE45A07137D2FD91D974">2013</mods:date>
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<mods:identifier id="5C453C63B5D75279541473BB6663CD51" type="DOI">10.11646/zootaxa.3635.3.5</mods:identifier>
<mods:identifier id="90AFAE32BCF68CD315F3944E2A44AF65" type="GBIF-Dataset">aa1e5dd3-98d2-4982-a52e-1bcec0907bb5</mods:identifier>
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<treatment id="03FC5411FFBCFFFFFF2998C42165F8D5" ID-DOI="http://doi.org/10.5281/zenodo.6164362" ID-GBIF-Taxon="119565018" ID-Zenodo-Dep="6164362" LSID="urn:lsid:plazi:treatment:03FC5411FFBCFFFFFF2998C42165F8D5" httpUri="http://treatment.plazi.org/id/03FC5411FFBCFFFFFF2998C42165F8D5" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262">
<subSubSection id="C34FB68CFFBCFFFCFF2998C423F1FE66" pageId="1" pageNumber="262" type="nomenclature">
<paragraph id="8BEAE507FFBCFFFCFF2998C42094FE47" blockId="1.[151,922,430,489]" box="[151,922,430,456]" pageId="1" pageNumber="262">
<heading id="D0A2526BFFBCFFFCFF2998C42094FE47" bold="true" box="[151,922,430,456]" fontSize="11" level="1" pageId="1" pageNumber="262" reason="1">
<emphasis id="B9213915FFBCFFFCFF2998C42094FE47" bold="true" box="[151,922,430,456]" pageId="1" pageNumber="262">
<taxonomicName id="4C559E84FFBCFFFCFF2998C421F0FE47" authority="Melnitsky &amp; Ivanov" authorityName="Melnitsky &amp; Ivanov" box="[151,766,430,456]" class="Insecta" family="Ecnomidae" genus="Archaeotinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="petropolitana" status="sp. nov.">
<emphasis id="B9213915FFBCFFFCFF2998C422F2FE47" bold="true" box="[151,508,430,456]" italics="true" pageId="1" pageNumber="262">Archaeotinodes petropolitana</emphasis>
Melnitsky &amp; Ivanov
</taxonomicName>
,
<taxonomicNameLabel id="A212846EFFBCFFFCFCB498C42094FE47" box="[778,922,430,456]" pageId="1" pageNumber="262" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEAE507FFBCFFFCFF2998BB23F1FE66" blockId="1.[151,922,430,489]" box="[151,255,465,489]" pageId="1" pageNumber="262">
<figureCitation id="136EF982FFBCFFFCFF2998BB23F1FE66" box="[151,255,465,489]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1869,1892]" captionStartId-1="2.[151,250,1406,1429]" captionTargetBox-0="[178,1409,905,1845]" captionTargetBox-1="[253,1368,410,1341]" captionTargetId-0="figure@1.[178,1410,905,1846]" captionTargetId-1="figure@2.[188,1398,373,1385]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Archaeotinodes petropolitana sp. nov., holotype PIN, № 364 / 789." captionText-1="FIGURE 2. Archaeotinodes petropolitana sp. nov., holotype PIN, № 364 / 789, male genitalia, lateral. Scale bar = 0.1 mm. Abbreviations: AE—aedeagus, DL—dorsal lobe of lower appendage, PA—preanal appendage, VL—ventral lobe of lower appendage." httpUri-0="https://zenodo.org/record/222218/files/figure.png" httpUri-1="https://zenodo.org/record/222219/files/figure.png" pageId="1" pageNumber="262">Figs. 1–2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34FB68CFFBCFFFFFF299B7222E8F945" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262" type="description">
<paragraph id="8BEAE507FFBCFFFCFF299B72263DFCDE" blockId="1.[151,1436,536,849]" pageId="1" pageNumber="262">
<emphasis id="B9213915FFBCFFFCFF299B72222CFDBE" bold="true" box="[151,290,536,561]" pageId="1" pageNumber="262">Description</emphasis>
(
<figureCitation id="136EF982FFBCFFFCFE8E9B732295FDBE" box="[304,411,537,561]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1869,1892]" captionStartId-1="2.[151,250,1406,1429]" captionTargetBox-0="[178,1409,905,1845]" captionTargetBox-1="[253,1368,410,1341]" captionTargetId-0="figure@1.[178,1410,905,1846]" captionTargetId-1="figure@2.[188,1398,373,1385]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Archaeotinodes petropolitana sp. nov., holotype PIN, № 364 / 789." captionText-1="FIGURE 2. Archaeotinodes petropolitana sp. nov., holotype PIN, № 364 / 789, male genitalia, lateral. Scale bar = 0.1 mm. Abbreviations: AE—aedeagus, DL—dorsal lobe of lower appendage, PA—preanal appendage, VL—ventral lobe of lower appendage." httpUri-0="https://zenodo.org/record/222218/files/figure.png" httpUri-1="https://zenodo.org/record/222219/files/figure.png" pageId="1" pageNumber="262">Figs. 1–2</figureCitation>
). Abdomen and thorax light brown. Head, antennae and palpi slightly lighter. Wings brown, heavily covered with fine hairs. Anterior branch of CuA in each forewing with additional fork. Third segment of each maxillary palp slightly longer and thicker than second and fourth. Forewings each with five forks. Antennae reaching twothirds of forewing length. Medial hair warts on pronotum large, swollen and united. Scutal hair warts of mesonotum with oval shape and closely approximate. Occipital hair warts of head large, each with rounded posterior margin. Ocellar hair warts narrow and elongated. Anterior margin of abdominal sternite V with distinct, clearly marked cuticular plate, darker than yellow sternites. Small bulge apparently associated with pheromone gland situated postero-medially to this structure. Cuticle of abdominal sternite V covered by small network pattern making rough surface. Spur formula 3.4.4. Measurements: body length
<quantity id="4CAD48E2FFBCFFFCFC069A53271CFCDE" box="[952,1042,825,849]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="262" unit="mm" value="4.7">4.7 mm</quantity>
; forewing length
<quantity id="4CAD48E2FFBCFFFCFB689A53263EFCDE" box="[1238,1328,825,849]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="262" unit="mm" value="5.3">5.3 mm</quantity>
.
</paragraph>
<caption id="DF2AB58FFFBCFFFCFF299E2720B9F8EB" box="[151,951,1869,1892]" httpUri="https://zenodo.org/record/222218/files/figure.png" pageId="1" pageNumber="262" targetBox="[178,1409,905,1845]" targetPageId="1">
<paragraph id="8BEAE507FFBCFFFCFF299E2720B9F8EB" blockId="1.[151,951,1869,1892]" box="[151,951,1869,1892]" pageId="1" pageNumber="262">
<emphasis id="B9213915FFBCFFFCFF299E27221DF8EC" bold="true" box="[151,275,1869,1892]" pageId="1" pageNumber="262">FIGURE 1.</emphasis>
<taxonomicName id="4C559E84FFBCFFFCFEA79E242148F8EB" box="[281,582,1870,1892]" class="Insecta" family="Ecnomidae" genus="Archaeotinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="petropolitana" status="sp. nov.">
<emphasis id="B9213915FFBCFFFCFEA79E242148F8EB" box="[281,582,1870,1892]" italics="true" pageId="1" pageNumber="262">Archaeotinodes petropolitana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212846EFFBCFFFCFDF29E242192F8EC" box="[588,668,1870,1891]" pageId="1" pageNumber="262" rank="species">
<emphasis id="B9213915FFBCFFFCFDF29E242192F8EC" bold="true" box="[588,668,1870,1891]" pageId="1" pageNumber="262">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype PIN, № 364/789.
</paragraph>
</caption>
<paragraph id="8BEAE507FFBCFFFFFF799EFA2608FEB0" blockId="1.[151,1436,1936,2032]" lastBlockId="2.[151,1436,151,320]" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262">
Male genitalia (
<figureCitation id="136EF982FFBCFFFCFE3C9EFA22E6F827" box="[386,488,1936,1960]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1869,1892]" captionStartId-1="2.[151,250,1406,1429]" captionTargetBox-0="[178,1409,905,1845]" captionTargetBox-1="[253,1368,410,1341]" captionTargetId-0="figure@1.[178,1410,905,1846]" captionTargetId-1="figure@2.[188,1398,373,1385]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Archaeotinodes petropolitana sp. nov., holotype PIN, № 364 / 789." captionText-1="FIGURE 2. Archaeotinodes petropolitana sp. nov., holotype PIN, № 364 / 789, male genitalia, lateral. Scale bar = 0.1 mm. Abbreviations: AE—aedeagus, DL—dorsal lobe of lower appendage, PA—preanal appendage, VL—ventral lobe of lower appendage." httpUri-0="https://zenodo.org/record/222218/files/figure.png" httpUri-1="https://zenodo.org/record/222219/files/figure.png" pageId="1" pageNumber="262">Fig. 1–2</figureCitation>
). Lower appendages (gonopods) small, subdivided into dorsal and ventral lobes (branches), widely separated. Dorsal lobes oblong and rounded at apex, outer surfaces covered with numerous setae. Ventral lobe of each gonopod long and thin, with apical portion extended into long sickle-shaped and pointed process directed posteroventrad. Aedeagus strong and well sclerotized. Apical part of aedeagus reaching ventral lobes of lower appendages. Sternite IX broad, projecting backwards, with two incisions on ventral margin. Preanal appendages (cerci) very long and curved, enlarged apically; their apical parts without spine. Numerous hairs covering subapical part of outer surface of preanal appendages. The lateral margins of preanal appendages heavily sclerotized. Segment X membranous. Ventral branches of segment X very long, s-shaped and curved.
</paragraph>
<caption id="DF2AB58FFFBFFFFFFF299C142204FA5D" httpUri="https://zenodo.org/record/222219/files/figure.png" pageId="2" pageNumber="263" targetBox="[253,1368,410,1341]" targetPageId="2">
<paragraph id="8BEAE507FFBFFFFFFF299C142204FA5D" blockId="2.[151,1437,1406,1490]" pageId="2" pageNumber="263">
<emphasis id="B9213915FFBFFFFFFF299C14221BFA1B" bold="true" box="[151,277,1406,1429]" pageId="2" pageNumber="263">FIGURE 2.</emphasis>
<taxonomicName id="4C559E84FFBFFFFFFEA09C152140FA1A" box="[286,590,1407,1429]" class="Insecta" family="Ecnomidae" genus="Archaeotinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="petropolitana" status="sp. nov.">
<emphasis id="B9213915FFBFFFFFFEA09C152140FA1A" box="[286,590,1407,1429]" italics="true" pageId="2" pageNumber="263">Archaeotinodes petropolitana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212846EFFBFFFFFFDE99C1521A4FA1B" box="[599,682,1407,1428]" pageId="2" pageNumber="263" rank="species">
<emphasis id="B9213915FFBFFFFFFDE99C1521A4FA1B" bold="true" box="[599,682,1407,1428]" pageId="2" pageNumber="263">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype PIN, № 364/789, male genitalia, lateral. Scale bar = 0.1 mm. Abbreviations: AE—aedeagus, DL—dorsal lobe of lower appendage, PA—preanal appendage, VL—ventral lobe of lower appendage.
</paragraph>
</caption>
<paragraph id="8BEAE507FFBFFFFFFF799F4822E8F945" blockId="2.[151,1436,1570,1882]" pageId="2" pageNumber="263">
<emphasis id="B9213915FFBFFFFFFF799F48226CF9B4" bold="true" box="[199,354,1570,1595]" pageId="2" pageNumber="263">Comparison.</emphasis>
The new species is similar to
<taxonomicName id="4C559E84FFBFFFFFFD7E9F492786F9B5" authority="Ulmer, 1912" authorityName="Ulmer" authorityYear="1912" box="[704,1160,1570,1595]" class="Insecta" family="Ecnomidae" genus="Archaeotinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="lanceolata">
<emphasis id="B9213915FFBFFFFFFD7E9F4920E0F9B4" box="[704,1006,1571,1595]" italics="true" pageId="2" pageNumber="263">Archaeotinodes lanceolata</emphasis>
Ulmer, 1912
</taxonomicName>
,
<taxonomicName id="4C559E84FFBFFFFFFB2B9F492226F9D0" authority="Ulmer, 1912" authorityName="Ulmer" authorityYear="1912" class="Insecta" family="Ecnomidae" genus="Archaeotinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="B9213915FFBFFFFFFB2B9F492695F9B4" box="[1173,1435,1571,1595]" italics="true" pageId="2" pageNumber="263">Archaeotinodes pauper</emphasis>
Ulmer, 1912
</taxonomicName>
, and
<taxonomicName id="4C559E84FFBFFFFFFED99F2D2050F9D0" authority="Melnitsky, 2009" authorityName="Melnitsky" authorityYear="2009" box="[359,862,1607,1631]" class="Insecta" family="Ecnomidae" genus="Archaeotinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="igneusaper">
<emphasis id="B9213915FFBFFFFFFED99F2D2195F9D0" box="[359,667,1607,1631]" italics="true" pageId="2" pageNumber="263">Archaeotinodes igneusaper</emphasis>
Melnitsky, 2009
</taxonomicName>
, from Baltic amber.
<taxonomicName id="4C559E84FFBFFFFFFBF29F2D2692F9D0" box="[1100,1436,1607,1631]" class="Insecta" family="Ecnomidae" genus="Archaeotinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="petropolitana" status="sp. nov.">
<emphasis id="B9213915FFBFFFFFFBF29F2D2692F9D0" box="[1100,1436,1607,1631]" italics="true" pageId="2" pageNumber="263">Archaeotinodes petropolitana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212846EFFBFFFFFFF299F0123FCF90C" box="[151,242,1643,1667]" pageId="2" pageNumber="263" rank="species">
<emphasis id="B9213915FFBFFFFFFF299F0123FCF90C" bold="true" box="[151,242,1643,1667]" pageId="2" pageNumber="263">sp. nov.</emphasis>
</taxonomicNameLabel>
differs by its longer and more strongly curved apically ventral lobe of the lower appendage, by the more massive aedeagus and the absence of a long black inner spine in the apical part of each preanal appendage (the ventral branch of segment X).
</paragraph>
</subSubSection>
<subSubSection id="C34FB68CFFBFFFFFFF799FBD204EF960" box="[199,832,1751,1775]" pageId="2" pageNumber="263" type="materials_examined">
<paragraph id="8BEAE507FFBFFFFFFF799FBD204EF960" blockId="2.[151,1436,1570,1882]" box="[199,832,1751,1775]" pageId="2" pageNumber="263">
<emphasis id="B9213915FFBFFFFFFF799FBD2275F960" bold="true" box="[199,379,1751,1775]" pageId="2" pageNumber="263">
<typeStatus id="54EE5BA5FFBFFFFFFF799FBD223AF960" box="[199,308,1751,1775]" pageId="2" pageNumber="263" type="holotype">Holotype</typeStatus>
male.
</emphasis>
PIN, № 364/789, Baltic amber, Eocene.
</paragraph>
</subSubSection>
<subSubSection id="C34FB68CFFBFFFFFFF799F912165F8D5" pageId="2" pageNumber="263" type="etymology">
<paragraph id="8BEAE507FFBFFFFFFF799F912165F8D5" blockId="2.[151,1436,1570,1882]" pageId="2" pageNumber="263">
<emphasis id="B9213915FFBFFFFFFF799F912240F89C" bold="true" box="[199,334,1787,1811]" pageId="2" pageNumber="263">Etymology.</emphasis>
From the Latin Petropolitana—St. Petersburg. The gender of
<taxonomicName id="4C559E84FFBFFFFFFBB19F9127B0F89C" box="[1039,1214,1787,1811]" class="Insecta" family="Ecnomidae" genus="Archaeotinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B9213915FFBFFFFFFBB19F9127B0F89C" box="[1039,1214,1787,1811]" italics="true" pageId="2" pageNumber="263">Archaeotinodes</emphasis>
</taxonomicName>
is feminine. Ulmer (1912) selected
<taxonomicName id="4C559E84FFBFFFFFFEF79E752106F8B8" authority="Pictet" authorityName="Pictet" box="[329,520,1823,1847]" class="Insecta" family="Ecnomidae" genus="Archaeotinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="prisca">
<emphasis id="B9213915FFBFFFFFFEF79E752254F8B8" box="[329,346,1823,1847]" italics="true" pageId="2" pageNumber="263">A</emphasis>
.
<emphasis id="B9213915FFBFFFFFFED69E4A22A0F8B8" box="[360,430,1824,1847]" italics="true" pageId="2" pageNumber="263">prisca</emphasis>
(Pictet)
</taxonomicName>
as the
<typeStatus id="54EE5BA5FFBFFFFFFDE89E4A2188F8B7" box="[598,646,1824,1848]" pageId="2" pageNumber="263">type</typeStatus>
of the genus, and thus, designated the gender of the genus as feminine (See ICZN article 30.1.4.4.) intentionally.
</paragraph>
</subSubSection>
</treatment>
</document>
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