283 lines
34 KiB
XML
283 lines
34 KiB
XML
<document id="984B44CF091E12DC4293297DF126B9F6" ID-DOI="10.5281/zenodo.189200" ID-GBIF-Dataset="89ec9a06-f83a-416b-8f1d-ce502cf9b0c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="189200" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461100149847" checkinUser="plazi" docAuthor="Ya, Igor & Vikhrev, Nikita E." docDate="2009" docId="03F83511D000FFE5FF4CF9B2FB4A8BD9" docLanguage="en" docName="zt02170p052.pdf" docOrigin="Zootaxa 2170" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Medetera varvara Grichanov & Vikhrev, sp. nov." docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="FFC14D69D002FFE0FFDBFFD7FFB58A3E" masterDocTitle="Mediterranean species of the Medetera plumbella species group with description of a new peculiar species from Morocco (Diptera: Dolichopodidae)" masterLastPageNumber="52" masterPageNumber="46" pageNumber="48" updateTime="1698590920703" updateUser="plazi">
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<mods:mods id="8A90781AF9BF7EEDA701ADA90ACC2BCB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="330F0AF186E6947FDF2596F3033E69D7">
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<mods:title id="5986EF68DA0CDB440DB7594391A3EFC5">Mediterranean species of the Medetera plumbella species group with description of a new peculiar species from Morocco (Diptera: Dolichopodidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="9F2EA0C50F4055D8652495168838D54E" type="personal">
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<mods:roleTerm id="367E716CAE01B0BE9E981F087B3992E7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BBB9B429F529C36157ACF9C00EC56880">Ya, Igor</mods:namePart>
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</mods:name>
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<mods:name id="F50E87BB4471591D56F39C5131170AC0" type="personal">
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<mods:role id="02C0F66546B9EE46C89EA97D900CE848">
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<mods:roleTerm id="CA2FC6C54CE1FCA46DA0FD0EAD74BD41">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="67AAB4A0B654779A3925C8D4D0BD6317">Vikhrev, Nikita E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="87759B4BEF535FECF1CD793EC63ADB87">text</mods:typeOfResource>
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<mods:titleInfo id="D2AE2710D43B6231FC5F043552632AAA">
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<mods:title id="5ECE24FF6650478D5762C0023AA67CEE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BFC9270B624D90813D26C539E1196AB6">
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<mods:date id="160C469828E1F694C03C6ACCB95B9A19">2009</mods:date>
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<mods:detail id="F7B8158854683178703F263AB628BA6C" type="volume">
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<mods:number id="8C2F778F84E8768CE6F97CD25A986456">2170</mods:number>
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</mods:detail>
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<mods:extent id="913B4926D48BF28230A549B108C6B0CD" unit="page">
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<mods:start id="98ACA9D976B830AB22DDA62AF57BFF7B">46</mods:start>
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<mods:end id="00AB11B688E49F92DD01F8AAC1AF2563">52</mods:end>
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</mods:extent>
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<mods:classification id="8419B41409B0E6D67C216C583C03C4E3">journal article</mods:classification>
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<mods:identifier id="C3B014073FF48E7583A859E6358DE04D" type="DOI">10.5281/zenodo.189200</mods:identifier>
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<mods:identifier id="D3C7192598CB8C099B6BF869A688D0C3" type="GBIF-Dataset">89ec9a06-f83a-416b-8f1d-ce502cf9b0c9</mods:identifier>
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<mods:identifier id="ADEE658C28C3A68F0AC8032898C6AB15" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="60C3283189BC45C3C7FADFB16F09C4BD" type="Zenodo-Dep">189200</mods:identifier>
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</mods:mods>
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<treatment id="03F83511D000FFE5FF4CF9B2FB4A8BD9" ID-DOI="http://doi.org/10.5281/zenodo.6214683" ID-GBIF-Taxon="119621444" ID-Zenodo-Dep="6214683" LSID="urn:lsid:plazi:treatment:03F83511D000FFE5FF4CF9B2FB4A8BD9" httpUri="http://treatment.plazi.org/id/03F83511D000FFE5FF4CF9B2FB4A8BD9" lastPageId="5" lastPageNumber="51" pageId="2" pageNumber="48">
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<subSubSection id="C34BD78CD000FFE2FF4CF9B2FEA28C9C" pageId="2" pageNumber="48" type="nomenclature">
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<paragraph id="8BEE8407D000FFE2FF4CF9B2FD5F8C41" blockId="2.[151,746,1637,1698]" box="[151,746,1637,1663]" pageId="2" pageNumber="48">
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||
<heading id="D0A6336BD000FFE2FF4CF9B2FD5F8C41" bold="true" box="[151,746,1637,1663]" fontSize="11" level="1" pageId="2" pageNumber="48" reason="1">
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||
<emphasis id="B9255815D000FFE2FF4CF9B2FD5F8C41" bold="true" box="[151,746,1637,1663]" pageId="2" pageNumber="48">
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<taxonomicName id="4C51FF84D000FFE2FF4CF9B2FD338C41" ID-CoL="9HXQJ" authority="Grichanov & Vikhrev" authorityName="Grichanov & Vikhrev" box="[151,646,1637,1663]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="varvara" status="sp. nov.">
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<emphasis id="B9255815D000FFE2FF4CF9B2FED88C41" bold="true" box="[151,365,1637,1663]" italics="true" pageId="2" pageNumber="48">Medetera varvara</emphasis>
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Grichanov & Vikhrev
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</taxonomicName>
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<taxonomicNameLabel id="A216E56ED000FFE2FD56F9B2FD5F8C41" box="[653,746,1637,1663]" pageId="2" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEE8407D000FFE2FF4CF95FFEA28C9C" blockId="2.[151,746,1637,1698]" box="[151,279,1672,1698]" pageId="2" pageNumber="48">
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(
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<figureCitation id="136A9882D000FFE2FF44F95FFEBB8C9C" box="[159,270,1672,1698]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1712,1736]" captionTargetBox="[165,1421,301,1670]" captionTargetId="figure@3.[158,858,966,1153]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Medetera varvara Grichanov et Vikhrev sp. nov. 1. male holotype, habitus. 2. female paratype, habitus. 3. male fore tarsus. 4. wing. 5. hypopygium, left lateral aspect. 6. apex of hypopygium, dorsal aspect. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/189201/files/figure.png" pageId="2" pageNumber="48">Figs. 1–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34BD78CD000FFE2FF4CF902FB088D28" pageId="2" pageNumber="48" type="materials_examined">
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<paragraph id="8BEE8407D000FFE2FF4CF902FB088D28" blockId="2.[151,1438,1748,2007]" pageId="2" pageNumber="48">
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<emphasis id="B9255815D000FFE2FF4CF902FEE58CD1" bold="true" box="[151,336,1749,1775]" pageId="2" pageNumber="48">
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<typeStatus id="54EA3AA5D000FFE2FF4CF902FF628CD1" box="[151,215,1749,1775]" pageId="2" pageNumber="48">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="54EA3AA5D000FFE2FEBBF902FE668CD1" box="[352,467,1749,1775]" pageId="2" pageNumber="48" type="holotype">Holotype</typeStatus>
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3:
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<emphasis id="B9255815D000FFE2FDDFF902FDC28CD1" bold="true" box="[516,631,1749,1775]" pageId="2" pageNumber="48">
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<collectingCountry id="F346C497D000FFE2FDDFF902FDC28CD1" box="[516,631,1749,1775]" name="Morocco" pageId="2" pageNumber="48">Morocco</collectingCountry>
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</emphasis>
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, near
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<collectingCountry id="F346C497D000FFE2FD1CF902FCF68CD1" box="[711,835,1749,1775]" name="Morocco" pageId="2" pageNumber="48">Essaouira</collectingCountry>
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,
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<geoCoordinate id="EE65E2C0D000FFE2FC8BF902FC708CD1" box="[848,965,1749,1775]" direction="north" orientation="latitude" pageId="2" pageNumber="48" precision="55" value="31.563">31.563ºN</geoCoordinate>
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<geoCoordinate id="EE65E2C0D000FFE2FC15F902FB8F8CD1" box="[974,1082,1749,1775]" direction="west" orientation="longitude" pageId="2" pageNumber="48" precision="55" value="-9.714">9.714ºW</geoCoordinate>
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, sand dune,
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<date id="FFEFA2C7D000FFE2FB0DF902FA2D8CD1" box="[1238,1432,1749,1775]" pageId="2" pageNumber="48" value="2009-03-29">29 March 2009</date>
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, N.Vikhrev (ZMU).
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<typeStatus id="54EA3AA5D000FFE2FEA7F92BFE478D28" box="[380,498,1788,1814]" pageId="2" pageNumber="48" type="paratype">Paratypes</typeStatus>
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. 2 ƤƤ with same data as
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<typeStatus id="54EA3AA5D000FFE2FCC6F92BFC338D28" box="[797,902,1788,1814]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
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,
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<date id="FFEFA2C7D000FFE2FC49F92BFBEC8D28" box="[914,1113,1788,1814]" pageId="2" pageNumber="48">28 and 29 March</date>
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(ZMU).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34BD78CD000FFE3FF1EF8F5FC108AC1" lastPageId="3" lastPageNumber="49" pageId="2" pageNumber="48" type="diagnosis">
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<paragraph id="8BEE8407D000FFE3FF1EF8F5FC108AC1" blockId="2.[151,1438,1748,2007]" lastBlockId="3.[151,1436,152,255]" lastPageId="3" lastPageNumber="49" pageId="2" pageNumber="48">
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<emphasis id="B9255815D000FFE2FF1EF8F5FE8B8D02" bold="true" box="[197,318,1826,1852]" pageId="2" pageNumber="48">Diagnosis</emphasis>
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.
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<taxonomicName id="4C51FF84D000FFE2FE96F8F4FD968D02" box="[333,547,1827,1852]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="varvara">
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<emphasis id="B9255815D000FFE2FE96F8F4FD968D02" box="[333,547,1827,1852]" italics="true" pageId="2" pageNumber="48">Medetera varvara</emphasis>
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</taxonomicName>
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is related to Egyptian
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<taxonomicName id="4C51FF84D000FFE2FCEAF8F4FB878D02" authority="Parent" authorityName="Parent" box="[817,1074,1826,1852]" class="Insecta" family="Dolichopodidae" genus="Oligochaetus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="albescens">
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<emphasis id="B9255815D000FFE2FCEAF8F4FCFC8D02" box="[817,841,1827,1852]" italics="true" pageId="2" pageNumber="48">M</emphasis>
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.
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<emphasis id="B9255815D000FFE2FC82F8F4FC798D02" box="[857,972,1827,1852]" italics="true" pageId="2" pageNumber="48">albescens</emphasis>
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(Parent)
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</taxonomicName>
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which differs from the former by silvery-white frons and face, hind coxa bearing three outer setae and fore tarsus unmodified. Ornamentation of fore legs is unknown for other species of Palearctic
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<taxonomicName id="4C51FF84D000FFE2FC13F8A7FB8D8DB7" box="[968,1080,1904,1929]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="genus">
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<emphasis id="B9255815D000FFE2FC13F8A7FB8D8DB7" box="[968,1080,1904,1929]" italics="true" pageId="2" pageNumber="48">Medetera</emphasis>
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</taxonomicName>
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including
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<taxonomicName id="4C51FF84D000FFE2FB6EF8A7FAFB8DB7" box="[1205,1358,1904,1930]" class="Insecta" family="Dolichopodidae" genus="Oligochaetus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="albescens">
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<emphasis id="B9255815D000FFE2FB6EF8A7FB788DB7" box="[1205,1229,1904,1929]" italics="true" pageId="2" pageNumber="48">M</emphasis>
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.
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<emphasis id="B9255815D000FFE2FB07F8A7FAFB8DB7" box="[1244,1358,1904,1929]" italics="true" pageId="2" pageNumber="48">albescens</emphasis>
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</taxonomicName>
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which was published with a rather detailed description. In the Afrotropics, only
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<taxonomicName id="4C51FF84D000FFE2FC27F840FAE68D8E" authority="Parent, 1936" authorityName="Parent" authorityYear="1936" box="[1020,1363,1942,1968]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="luteoscutata">
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<emphasis id="B9255815D000FFE2FC27F840FB068D8E" box="[1020,1203,1943,1968]" italics="true" pageId="2" pageNumber="48">M. luteoscutata</emphasis>
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Parent, 1936
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</taxonomicName>
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, has a small apicoventral process on tarsomere 1 and 3 of fore leg. Mainly Pantropical
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<taxonomicName id="4C51FF84D000FFE2FB9FF86AFA228DE9" authority="Becker, 1914" authorityName="Becker" authorityYear="1914" box="[1092,1431,1981,2007]" class="Insecta" family="Dolichopodidae" genus="Saccopheronta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="48" phylum="Arthropoda" rank="genus">
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<emphasis id="B9255815D000FFE2FB9FF86AFB468DE8" box="[1092,1267,1981,2006]" italics="true" pageId="2" pageNumber="48">Saccopheronta</emphasis>
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Becker, 1914
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</taxonomicName>
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, a sister genus (
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<bibRefCitation id="EFC0F9F6D001FFE3FE95FF4FFD928A8C" author="Grichanov" box="[334,551,152,178]" pageId="3" pageNumber="51" refString="Grichanov, I. Ya. (1997 b) Four new species of Saccopheronta Becker (Diptera: Dolichopodidae) from Tropical Africa with notes on the world fauna of the genus. International Journal of Dipterological Research, 8 (3), 123 - 134." type="journal article" year="1997" yearSuffix="b">Grichanov 1997b</bibRefCitation>
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) or “
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<taxonomicName id="4C51FF84D001FFE3FDBCFF4FFD608A8F" box="[615,725,152,177]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="aberrans">
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<emphasis id="B9255815D001FFE3FDBCFF4FFD608A8F" box="[615,725,152,177]" italics="true" pageId="3" pageNumber="49">aberrans</emphasis>
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</taxonomicName>
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’ group of
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<taxonomicName id="4C51FF84D001FFE3FC81FF4FFC7A8A8F" box="[858,975,152,177]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="genus">
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<emphasis id="B9255815D001FFE3FC81FF4FFC7A8A8F" box="[858,975,152,177]" italics="true" pageId="3" pageNumber="49">Medetera</emphasis>
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</taxonomicName>
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species (
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<bibRefCitation id="EFC0F9F6D001FFE3FBE4FF4FFB6F8A8C" author="Bickel" box="[1087,1242,152,178]" pageId="3" pageNumber="51" refString="Bickel, D. J. (1985) A revision of the Nearctic Medetera (Diptera: Dolichopodidae). United States Department of Agriculture Technical Bulletin, 1692, 1 - 109." type="journal article" year="1985">Bickel 1985</bibRefCitation>
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,
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<bibRefCitation id="EFC0F9F6D001FFE3FB3CFF4FFA908A8C" author="Bickel" box="[1255,1317,152,178]" pageId="3" pageNumber="51" refString="Bickel, D. J. (1987) A Revision of the Oriental and Australasian Medetera (Diptera: Dolichopodidae). Records of the Australian Museum, 39, 131 - 182." type="journal article" year="1987">1987</bibRefCitation>
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) includes species with thickened or widened tarsomeres 2 and 3 of male fore leg. The absolute majority of medeterine species of the world fauna have no remarkable fore leg decoration.
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</paragraph>
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</subSubSection>
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||
<caption id="DF2ED48FD001FFE3FF4CF967FA298CD4" httpUri="https://zenodo.org/record/189201/files/figure.png" pageId="3" pageNumber="49" targetBox="[165,1421,301,1670]" targetPageId="3">
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<paragraph id="8BEE8407D001FFE3FF4CF967FA298CD4" blockId="3.[151,1436,1712,1771]" pageId="3" pageNumber="49">
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<emphasis id="B9255815D001FFE3FF4CF967FEF78CF6" bold="true" box="[151,322,1712,1736]" pageId="3" pageNumber="49">FIGURES 1–6.</emphasis>
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<taxonomicName id="4C51FF84D001FFE3FE92F967FD438CF9" authority="Grichanov et Vikhrev" authorityName="Grichanov et Vikhrev" box="[329,758,1712,1735]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="varvara" status="sp. nov.">
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<emphasis id="B9255815D001FFE3FE92F967FDBC8CF9" box="[329,521,1712,1735]" italics="true" pageId="3" pageNumber="49">Medetera varvara</emphasis>
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Grichanov et Vikhrev
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</taxonomicName>
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<taxonomicNameLabel id="A216E56ED001FFE3FD27F967FCE48CF9" box="[764,849,1712,1736]" pageId="3" pageNumber="49" rank="species">
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<emphasis id="B9255815D001FFE3FD27F967FCFE8CF6" bold="true" box="[764,843,1712,1736]" pageId="3" pageNumber="49">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
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||
<emphasis id="B9255815D001FFE3FC8CF967FCDE8CF6" bold="true" box="[855,875,1712,1736]" pageId="3" pageNumber="49">1.</emphasis>
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male holotype, habitus.
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<emphasis id="B9255815D001FFE3FBABF967FB318CF6" bold="true" box="[1136,1156,1712,1736]" pageId="3" pageNumber="49">2.</emphasis>
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female paratype, habitus.
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<emphasis id="B9255815D001FFE3FF4CF904FF1F8CD5" bold="true" box="[151,170,1747,1771]" pageId="3" pageNumber="49">3.</emphasis>
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male fore tarsus.
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<emphasis id="B9255815D001FFE3FEBCF904FEC18CD5" bold="true" box="[359,372,1747,1771]" pageId="3" pageNumber="49">4</emphasis>
|
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. wing.
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<emphasis id="B9255815D001FFE3FE18F904FE658CD5" bold="true" box="[451,464,1747,1771]" pageId="3" pageNumber="49">5</emphasis>
|
||
. hypopygium, left lateral aspect.
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<emphasis id="B9255815D001FFE3FCE9F904FC8A8CD5" bold="true" box="[818,831,1747,1771]" pageId="3" pageNumber="49">6</emphasis>
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. apex of hypopygium, dorsal aspect. Scale bars: 0.2 mm.
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</paragraph>
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</caption>
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||
<subSubSection id="C34BD78CD001FFE4FF1EF8C1FBB58DB7" lastPageId="4" lastPageNumber="50" pageId="3" pageNumber="49" type="description">
|
||
<paragraph id="8BEE8407D001FFE3FF1EF8C1FA848D0F" blockId="3.[151,1437,1814,2034]" box="[197,1329,1814,1841]" pageId="3" pageNumber="49">
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||
<emphasis id="B9255815D001FFE3FF1EF8C1FE118D0E" bold="true" box="[197,420,1814,1840]" pageId="3" pageNumber="49">Description. Male</emphasis>
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(
|
||
<figureCitation id="136A9882D001FFE3FE6FF8C0FE4D8D0F" box="[436,504,1815,1841]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1712,1736]" captionTargetBox="[165,1421,301,1670]" captionTargetId="figure@3.[158,858,966,1153]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Medetera varvara Grichanov et Vikhrev sp. nov. 1. male holotype, habitus. 2. female paratype, habitus. 3. male fore tarsus. 4. wing. 5. hypopygium, left lateral aspect. 6. apex of hypopygium, dorsal aspect. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/189201/files/figure.png" pageId="3" pageNumber="49">Fig. 1</figureCitation>
|
||
): Length (mm): body 2.0, wing 2.0/0.7, antenna 0.7, hypopygium 0.8.
|
||
</paragraph>
|
||
<paragraph id="8BEE8407D001FFE4FF1EF8E9FC0A8AC1" blockId="3.[151,1437,1814,2034]" lastBlockId="4.[151,1437,152,2007]" lastPageId="4" lastPageNumber="50" pageId="3" pageNumber="49">
|
||
<emphasis id="B9255815D001FFE3FF1EF8E9FEB18D69" box="[197,260,1854,1879]" italics="true" pageId="3" pageNumber="49">Head</emphasis>
|
||
: Frons, face, clypeus, palpus and postcranium greenish, evenly and densely dusted greyish-white, so frontoclypeal suture between face and clypeus not distinct. Postocular setae white, somewhat thickened, strongly increasing in length downward. Ventral postcranium shining greenish, with row of long white thickened setae. Frons with pair of strong vertical setae and pair of ocellar setae slightly stronger than verticals. Postverticals absent. Face relatively wide; ratio of height of face to its maximal width to height of clypeus to height of palpus,
|
||
<date id="FFEFA2C7D006FFE4FE28FF4FFDF68A8C" box="[499,579,152,178]" pageId="4" pageNumber="50" value="1915-05-08">15/8/5</date>
|
||
/5. Antennal segments short, with short white hairs; scape and pedicel orange-brownish, grey dusted at apex; postpedicel black, rounded. Stylus subapical, bare, about 2 times as long as ocellar seta. Proboscis short, black, shining, with white hairs.
|
||
</paragraph>
|
||
<paragraph id="8BEE8407D006FFE4FF1EFEDBFD978BD9" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="50">
|
||
<emphasis id="B9255815D006FFE4FF1EFEDBFEA98B1B" box="[197,284,268,293]" italics="true" pageId="4" pageNumber="50">Thorax</emphasis>
|
||
: Dark, densely grey dusted; mesonotum with 3 narrow bronze stripes distinct in anterior view. Three pairs of strong black dorsocentral setae, slightly decreasing in size anteriorly. Notopleural setae 2, sutural 1, supraalar 1, all black; 1 white humeral seta. Acrostichals very short, biseriate, white, extending to mesonotal flattening. Several white setulae in front of first dorsocentral and sutural setae. Propleuron with 5 white thickened setae, lower one almost 2 times longer than others. Scutellum with pair of strong black median setae, lateral setae absent.
|
||
</paragraph>
|
||
<paragraph id="8BEE8407D006FFE4FF1EFE23FB1A8EF7" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="50">
|
||
<emphasis id="B9255815D006FFE4FF1EFE23FF498833" box="[197,252,500,525]" italics="true" pageId="4" pageNumber="50">Legs</emphasis>
|
||
: Yellow, except fore legs whitish-yellow, hind tibia darkened at apex, tarsomeres 4 and 5 and apical part of 2 and 3 of mid and hind legs brownish; mid and hind coxae mostly dark, grey dusted, yellow at apex. Fore coxa with dense brush of long white flattened setae on anterior surface. Fore femur and tibia without setae. Fore tarsomeres 1 to 5 each with pair of small brown apical postero- and anteroventral setulae; tarsomeres 2 to 5 each with pair of very small brown ventral setulae; tarsomeres 2 to 4 slightly flattened laterally; apical 1/3 of tarsomere 1 and tarsomeres 2 to 4 with comb of white thickened cilia on dorsal surface (
|
||
<figureCitation id="136A9882D006FFE4FF7BFD0BFF5F88C8" box="[160,234,732,758]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1712,1736]" captionTargetBox="[165,1421,301,1670]" captionTargetId="figure@3.[158,858,966,1153]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Medetera varvara Grichanov et Vikhrev sp. nov. 1. male holotype, habitus. 2. female paratype, habitus. 3. male fore tarsus. 4. wing. 5. hypopygium, left lateral aspect. 6. apex of hypopygium, dorsal aspect. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/189201/files/figure.png" pageId="4" pageNumber="50">Fig. 3</figureCitation>
|
||
). Mid coxa with dense brush of long thickened white setae on anterior surface; mid trochanter with single white seta on anterior position; mid femur without setae. Mid tibia with pair of antero- and posterodorsals at 1/4 and long apicoventral seta, all white. Mid tarsus with four brown apical setae on each segment; basitarsus in apical half with 4–5 short brown setulae, somewhat irregularly placed, but in either antero- or posteroventral positions, tarsomere 2 with 3 such setulae, 3rd with 2–3, 4th and 5th with 2 ones each; all these setulae gradually decreasing in size apically.
|
||
<collectingCountry id="F346C497D006FFE4FCCAFC77FCF98984" box="[785,844,928,954]" name="India" pageId="4" pageNumber="50">Hind</collectingCountry>
|
||
coxa with single white seta on outer surface; hind femur with row of white dorsal setulae in basal half; hind tibia slightly thickened at apex, with posterodorsal setula at 1/4 and ventral apical one, both white; with very short black posterodorsal apical spur; hind tibia on apical 2/3, basitarsus and tarsomere 2 on basal 2/3 with row of short dense white posteroventral cilia; hind basitarsus short, with 2 apicals: brown anteroventral and white ventral, with small white posteroventral basal tooth and shallow basal excavation; tarsomeres 2 to 5 each with 1–3 brown anteroventral and 2–3 brown apical setulae, these setulae gradually decreasing in size apically. Fore leg length ratio (from coxa to tarsomere 5): 24/40/
|
||
<date id="FFEFA2C7D006FFE4FED5FB78FEC38EF7" box="[270,374,1199,1225]" pageId="4" pageNumber="50" value="1935-10-20">35/20/10</date>
|
||
/8/5/5, mid leg: 16/42/
|
||
<date id="FFEFA2C7D006FFE4FDA1FB78FD518EF7" box="[634,740,1199,1225]" pageId="4" pageNumber="50" value="1945-10-22">45/22/10</date>
|
||
/8/6/5, hind leg: 14/45/
|
||
<date id="FFEFA2C7D006FFE4FC35FB78FBEC8EF7" box="[1006,1113,1199,1225]" pageId="4" pageNumber="50" value="1951-12-24">51/12/24</date>
|
||
/
|
||
<date id="FFEFA2C7D006FFE4FBB9FB78FB1A8EF7" box="[1122,1199,1199,1225]" pageId="4" pageNumber="50" value="1914-06-08">14/8/6.</date>
|
||
</paragraph>
|
||
<paragraph id="8BEE8407D006FFE4FF1EFB01FDEB8FAD" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="50">
|
||
<emphasis id="B9255815D006FFE4FF1EFB01FEBE8ED1" box="[197,267,1238,1263]" italics="true" pageId="4" pageNumber="50">Wings</emphasis>
|
||
: Hyaline, veins yellow in anterobasal quarter of wing, brown in other parts (
|
||
<figureCitation id="136A9882D006FFE4FB43FB02FB548ED1" box="[1176,1249,1237,1263]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1712,1736]" captionTargetBox="[165,1421,301,1670]" captionTargetId="figure@3.[158,858,966,1153]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Medetera varvara Grichanov et Vikhrev sp. nov. 1. male holotype, habitus. 2. female paratype, habitus. 3. male fore tarsus. 4. wing. 5. hypopygium, left lateral aspect. 6. apex of hypopygium, dorsal aspect. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/189201/files/figure.png" pageId="4" pageNumber="50">Fig. 4</figureCitation>
|
||
). Costa without long hairs. R1 short, extending to basal third of wing, R4+5 and M1+2 distinctly convergent at apex. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1+2, 25/5. Ratio of apical to basal part of M1+2, 17/18. Ratio of cross-vein
|
||
<emphasis id="B9255815D006FFE4FEA6FA87FE0D8F57" box="[381,440,1360,1385]" italics="true" pageId="4" pageNumber="50">m-cu</emphasis>
|
||
to maximal distance between R4+5 and M1+2 to distal part of CuA1,
|
||
<date id="FFEFA2C7D006FFE4FB1AFA98FA858F57" box="[1217,1328,1359,1385]" pageId="4" pageNumber="50" value="1910-12-11">10/11/12.</date>
|
||
Calypter yellow, with white cilia. Halter yellow.
|
||
</paragraph>
|
||
<paragraph id="8BEE8407D006FFE4FF1EFA77FF6D8C9F" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="50">
|
||
<emphasis id="B9255815D006FFE4FF1EFA77FE818F87" box="[197,308,1440,1465]" italics="true" pageId="4" pageNumber="50">Abdomen</emphasis>
|
||
: Covered with short white setulae, olive-grey dusted, with fore margin of tergites 3 to 5 bronze dorsally; posterior margin of tergite 1 with 5–6 white flattened setae on each side. Tergite 6 slightly longer than tergite 5; segment 7 longer than preceding, with short hairs; segment 8 large, left basolateral, shorthaired. Epandrium (
|
||
<figureCitation id="136A9882D006FFE4FE58F9C4FE7C8C13" box="[387,457,1555,1581]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1712,1736]" captionTargetBox="[165,1421,301,1670]" captionTargetId="figure@3.[158,858,966,1153]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Medetera varvara Grichanov et Vikhrev sp. nov. 1. male holotype, habitus. 2. female paratype, habitus. 3. male fore tarsus. 4. wing. 5. hypopygium, left lateral aspect. 6. apex of hypopygium, dorsal aspect. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/189201/files/figure.png" pageId="4" pageNumber="50">Fig. 5</figureCitation>
|
||
) black, elongate-triangular; hypandrium basoventral, slightly swollen at base, then thinned, pointed at apex; phallus simple, pointed; epandrial lobe small, hardly divided, bearing pair of long simple setae; surstylus and cercus (
|
||
<figureCitation id="136A9882D006FFE4FDEFF9B6FDCF8C45" box="[564,634,1633,1659]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1712,1736]" captionTargetBox="[165,1421,301,1670]" captionTargetId="figure@3.[158,858,966,1153]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Medetera varvara Grichanov et Vikhrev sp. nov. 1. male holotype, habitus. 2. female paratype, habitus. 3. male fore tarsus. 4. wing. 5. hypopygium, left lateral aspect. 6. apex of hypopygium, dorsal aspect. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/189201/files/figure.png" pageId="4" pageNumber="50">Fig. 6</figureCitation>
|
||
) dark-brown; cercus fused almost to apex, covered with short white hairs.
|
||
</paragraph>
|
||
<paragraph id="8BEE8407D006FFE4FF1EF979FBB58DB7" blockId="4.[151,1437,152,2007]" pageId="4" pageNumber="50">
|
||
<emphasis id="B9255815D006FFE4FF1EF979FE958CF6" bold="true" box="[197,288,1710,1736]" pageId="4" pageNumber="50">Female</emphasis>
|
||
(
|
||
<figureCitation id="136A9882D006FFE4FEEBF979FECC8CF6" box="[304,377,1710,1736]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,269,1712,1736]" captionTargetBox="[165,1421,301,1670]" captionTargetId="figure@3.[158,858,966,1153]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Medetera varvara Grichanov et Vikhrev sp. nov. 1. male holotype, habitus. 2. female paratype, habitus. 3. male fore tarsus. 4. wing. 5. hypopygium, left lateral aspect. 6. apex of hypopygium, dorsal aspect. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/189201/files/figure.png" pageId="4" pageNumber="50">Fig. 2</figureCitation>
|
||
): Length (mm): body 2.2, wing 2.2; similar to male except lacking male secondary sexual characters. Each hemitergite bearing 1 acanthophorite and 1 simple seta; acanthophorites thin, much longer than cercus; cercus small, with short hairs. Dense brush of thickened white setae on anterior surface of fore and hind coxae, but setae about two times shorter and more equal in length than those of male. Fore tarsi unmodified; tarsomeres 4 and 5 and apical part of 2 and 3 of fore legs brownish, as on mid and hind tarsi.
|
||
<collectingCountry id="F346C497D006FFE4FF4CF8B8FF678DB7" box="[151,210,1903,1929]" name="India" pageId="4" pageNumber="50">Hind</collectingCountry>
|
||
tibia without apical spur; hind basitarsus simple, without basal tooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34BD78CD006FFE4FF1EF841FE6A8D8E" box="[197,479,1942,1968]" pageId="4" pageNumber="50" type="distribution">
|
||
<paragraph id="8BEE8407D006FFE4FF1EF841FE6A8D8E" blockId="4.[151,1437,152,2007]" box="[197,479,1942,1968]" pageId="4" pageNumber="50">
|
||
<emphasis id="B9255815D006FFE4FF1EF841FEEB8D8E" bold="true" box="[197,350,1942,1968]" pageId="4" pageNumber="50">Distribution</emphasis>
|
||
:
|
||
<collectingCountry id="F346C497D006FFE4FEB6F841FE6E8D8E" box="[365,475,1942,1968]" name="Morocco" pageId="4" pageNumber="50">Morocco</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34BD78CD006FFE5FF1EF86BFB4A8BD9" lastPageId="5" lastPageNumber="51" pageId="4" pageNumber="50" type="etymology">
|
||
<paragraph id="8BEE8407D006FFE4FF1EF86BFAC38DE9" blockId="4.[151,1437,152,2007]" box="[197,1398,1980,2007]" pageId="4" pageNumber="50">
|
||
<emphasis id="B9255815D006FFE4FF1EF86BFEFF8DE8" bold="true" box="[197,330,1980,2006]" pageId="4" pageNumber="50">Etymology</emphasis>
|
||
. The species is named for
|
||
<taxonomicName id="4C51FF84D006FFE4FD58F86AFD6B8DE9" box="[643,734,1981,2007]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="varvara">Varvara</taxonomicName>
|
||
Vikhreva who kindly helped to collect flies in
|
||
<collectingCountry id="F346C497D006FFE4FADEF86AFAC78DE9" box="[1285,1394,1981,2007]" name="Morocco" pageId="4" pageNumber="50">Morocco</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEE8407D007FFE5FF1EFF40FB4A8BD9" blockId="5.[151,1437,151,487]" pageId="5" pageNumber="51">
|
||
<emphasis id="B9255815D007FFE5FF1EFF40FE908A8F" bold="true" box="[197,293,151,177]" pageId="5" pageNumber="51">Habitat</emphasis>
|
||
: All three specimens of the
|
||
<typeStatus id="54EA3AA5D007FFE5FDB7FF4FFD2B8A8C" box="[620,670,152,178]" pageId="5" pageNumber="51">type</typeStatus>
|
||
series were collected from sandy substrates. This habitat is rather unusual for mainly dendrophilous, sometimes petrophilous species of
|
||
<taxonomicName id="4C51FF84D007FFE5FC12FF68FB8C8AE6" box="[969,1081,191,216]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9255815D007FFE5FC12FF68FB8C8AE6" box="[969,1081,191,216]" italics="true" pageId="5" pageNumber="51">Medetera</emphasis>
|
||
</taxonomicName>
|
||
, although many species of the Nearctic
|
||
<taxonomicName id="4C51FF84D007FFE5FED8FF32FE378AC0" box="[259,386,229,254]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="petulca">
|
||
<emphasis id="B9255815D007FFE5FED8FF32FE378AC0" box="[259,386,229,254]" italics="true" pageId="5" pageNumber="51">M. petulca</emphasis>
|
||
</taxonomicName>
|
||
group occur in such habitats (Bickel pers. comm. 2009). Trees on sand dunes were also examined, but all specimens of tree trunk
|
||
<taxonomicName id="4C51FF84D007FFE5FD51FEDBFD4E8B1B" box="[650,763,268,293]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9255815D007FFE5FD51FEDBFD4E8B1B" box="[650,763,268,293]" italics="true" pageId="5" pageNumber="51">Medetera</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C51FF84D007FFE5FCD6FEDBFBF58B18" authority="Meigen, 1824" authorityName="Meigen" authorityYear="1824" box="[781,1088,268,294]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="flavipes">
|
||
<emphasis id="B9255815D007FFE5FCD6FEDBFC248B1B" box="[781,913,268,293]" italics="true" pageId="5" pageNumber="51">M. flavipes</emphasis>
|
||
<bibRefCitation id="EFC0F9F6D007FFE5FC42FEDBFBF58B18" author="Meigen" box="[921,1088,268,294]" pageId="5" pageNumber="51" refString="Meigen, J. W. (1824) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Vierter Theil. Schulz-Wundermann, Hamm, xii + 428 pp." type="book" year="1824">Meigen, 1824</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C51FF84D007FFE5FBA4FEDBFF528B72" authority="Zetterstedt, 1843" authorityName="Zetterstedt" authorityYear="1843" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="pallipes">
|
||
<emphasis id="B9255815D007FFE5FBA4FEDBFAB18B1B" box="[1151,1284,268,293]" italics="true" pageId="5" pageNumber="51">M. pallipes</emphasis>
|
||
(Zetterstedt, 1843))
|
||
</taxonomicName>
|
||
collected on the
|
||
<collectingCountry id="F346C497D007FFE5FE63FEE5FD988B72" box="[440,557,306,332]" name="Morocco" pageId="5" pageNumber="51">Essaouira</collectingCountry>
|
||
dunes (between
|
||
<date id="FFEFA2C7D007FFE5FD20FEE5FC7E8B72" box="[763,971,306,332]" pageId="5" pageNumber="51">24 and 29 March</date>
|
||
) belong to other species groups of the genus. The senior author observed
|
||
<taxonomicName id="4C51FF84D007FFE5FDE7FE8EFD788B4C" box="[572,717,345,370]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="pallidior">
|
||
<emphasis id="B9255815D007FFE5FDE7FE8EFD788B4C" box="[572,717,345,370]" italics="true" pageId="5" pageNumber="51">M. pallidior</emphasis>
|
||
</taxonomicName>
|
||
imagos in Southern
|
||
<collectingCountry id="F346C497D007FFE5FC1CFE8EFB898B4D" box="[967,1084,345,371]" name="Tajikistan" pageId="5" pageNumber="51">Tajikistan</collectingCountry>
|
||
(the Tigrovaya Balka Nature Reserve) in
|
||
<date id="FFEFA2C7D007FFE5FEF5FE57FE1F8BA4" box="[302,426,384,410]" pageId="5" pageNumber="51" value="1978-07">July 1978</date>
|
||
, where they populated rodent holes (to
|
||
<quantity id="4CA929E2D007FFE5FC43FE57FC608BA4" box="[920,981,384,410]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="51" unit="cm" value="5.0">5 cm</quantity>
|
||
in diameter) on a rather dry and flat semidesert plot not far from the border with a large area of riparian marshes. Males and females of the species concentrated around holes, and frightened or disturbed flies dropped immediately into the holes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |