159 lines
17 KiB
XML
159 lines
17 KiB
XML
<document id="5A24D16335D80886B03B15073164218B" ID-DOI="10.11646/zootaxa.3609.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5292698" ID-ZooBank="3C0A4EBB-B8C3-4D9B-9897-3F0300874621" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629839015983" checkinUser="felipe" docAuthor="Ramírez, Alonso & Gutiérrez-Fonseca, Pablo E." docDate="2013" docId="03EF87C74E09FFF4FF67ADB98B9FFC17" docLanguage="en" docName="zt03609p100.pdf" docOrigin="Zootaxa 3609 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heteragrion atrolineatum Donnelly 1992" docType="treatment" docVersion="4" lastPageNumber="99" masterDocId="FFD6FFBF4E0AFFF7FFF0AD2E8A59FF92" masterDocTitle="The larvae of Heteragrion majus Selys and H. atrolineatum Donnelly, with a key to known species from Costa Rica (Odonata: Megapodagrionidae)" masterLastPageNumber="100" masterPageNumber="96" pageNumber="99" updateTime="1699119446385" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="9803748DF76CDA924C12F3A41C59DD10" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="BB1444532E9626754F78077BDE27DB3F">
|
||
<mods:title id="5223417D15CB344727F10664DF6B05AE">The larvae of Heteragrion majus Selys and H. atrolineatum Donnelly, with a key to known species from Costa Rica (Odonata: Megapodagrionidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="20092787DFF75792C5B08F887C5173F7" type="personal">
|
||
<mods:role id="5A1C1695B330940EA5ADB2C69BE4247A">
|
||
<mods:roleTerm id="0A72A51690F71DA637D76D56F5139E2F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0F8514BD69AF4A60E9B591873EABA936">Ramírez, Alonso</mods:namePart>
|
||
<mods:nameIdentifier id="19F85CF0F2210CF338BEA9BE34C7901C" type="email">aramirez@ramirezlab.net</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="98903ACF13215722731E89197367E7FC" type="personal">
|
||
<mods:role id="B9AB7D74F4B3D32E38DB6651A328B205">
|
||
<mods:roleTerm id="88B45319B74E3BE762778FC35A66C4F4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B56A722BE70A3B42AD7E01C46225D19D">Gutiérrez-Fonseca, Pablo E.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="C206979395E81F7172A66A9BC39B018F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="2C74F9ABD36FE6DBB47A71C620402363" type="host">
|
||
<mods:titleInfo id="A64CFDAEE95F1B8208A46A6E5BA59450">
|
||
<mods:title id="9633B3985E2DB6AC394BF32E8B8C9B08">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="ECC35AC63F83CDDFFF3A678477F8B408">
|
||
<mods:date id="CE3D77CCA0041A7FE0A1719CB23563D0">2013</mods:date>
|
||
<mods:detail id="B191D244850085A713B17D4C6F0D7FAA" type="pubDate">
|
||
<mods:number id="5C8407A6B9E9C2073F6D476EDDA28206">2013-01-25</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="93F93E5520A6A5E3186C46ECA9AE6B8D" type="volume">
|
||
<mods:number id="6EBCEA781F1D6778225C3729B523E664">3609</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F8358450BB8E166552879DB0DA56DD4A" type="issue">
|
||
<mods:number id="37F7861CD299ACE1EAC1BF36B5437C8E">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="C20E05BC930CDBDCA7571A67079D9F07" unit="page">
|
||
<mods:start id="AB74C4AA14CD454A46002912489AFD81">96</mods:start>
|
||
<mods:end id="1D239EF6717C303D7EEDDA9B56C51356">100</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="C79DD8573EB1E295C461F4D70F438F17">journal article</mods:classification>
|
||
<mods:identifier id="27E5E485AADAB46080854AAEC34E5291" type="DOI">10.11646/zootaxa.3609.1.8</mods:identifier>
|
||
<mods:identifier id="E643EE72C72DA12A145A822E21DC7980" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="964A9008BA3393EE8DEEF5E330A89ABD" type="Zenodo-Dep">5292698</mods:identifier>
|
||
<mods:identifier id="C703E28CAFD4EA5BE75217A65D217BCA" type="ZooBank">3C0A4EBB-B8C3-4D9B-9897-3F0300874621</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03EF87C74E09FFF4FF67ADB98B9FFC17" ID-DOI="http://doi.org/10.5281/zenodo.5280151" ID-Zenodo-Dep="5280151" LSID="urn:lsid:plazi:treatment:03EF87C74E09FFF4FF67ADB98B9FFC17" httpUri="http://treatment.plazi.org/id/03EF87C74E09FFF4FF67ADB98B9FFC17" lastPageNumber="99" pageId="3" pageNumber="99">
|
||
<subSubSection id="C35C655A4E09FFF4FF67ADB9882EFF22" box="[151,631,151,176]" pageId="3" pageNumber="99" type="nomenclature">
|
||
<paragraph id="8BF936D14E09FFF4FF67ADB9882EFF22" blockId="3.[151,631,151,212]" box="[151,631,151,176]" pageId="3" pageNumber="99">
|
||
<heading id="D0B181BD4E09FFF4FF67ADB9882EFF22" bold="true" box="[151,631,151,176]" fontSize="10" level="2" pageId="3" pageNumber="99" reason="2">
|
||
<taxonomicName id="4C464D524E09FFF4FF67ADB9882EFF22" ID-CoL="3L73Q" authority="Donnelly, 1992" authorityName="Donnelly" authorityYear="1992" box="[151,631,151,176]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="99" phylum="Arthropoda" rank="species" species="atrolineatum">
|
||
<emphasis id="B932EAC34E09FFF4FF67ADB9882EFF22" bold="true" box="[151,631,151,176]" pageId="3" pageNumber="99">
|
||
<emphasis id="B932EAC34E09FFF4FF67ADB98B98FF22" bold="true" box="[151,449,151,176]" italics="true" pageId="3" pageNumber="99">Heteragrion atrolineatum</emphasis>
|
||
Donnelly, 1992
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C655A4E09FFF4FF67AD928B42FF46" box="[151,283,188,212]" pageId="3" pageNumber="99" type="description">
|
||
<paragraph id="8BF936D14E09FFF4FF67AD928B42FF46" blockId="3.[151,631,151,212]" box="[151,283,188,212]" pageId="3" pageNumber="99">
|
||
<figureCitation id="137D2A544E09FFF4FF67AD928B42FF46" box="[151,283,188,212]" captionStart="FIGURES 2–5" captionStartId="2.[151,264,1464,1487]" captionTargetBox="[178,1422,208,1408]" captionTargetId="figure-268@2.[147,1433,193,1443]" captionTargetPageId="2" captionText="FIGURES 2–5. Details of H. majus and H. atrolineatum. 2) Antennae of H. majus (a) and H. atrolineatum (b); 3) dorsal view of ligula of H. majus (a) and H. atrolineatum (b); 4) dorsal and ventral view of the abdomen and base of caudal gills for H. majus; 5) dorsal view of H. majus caudal gills." figureDoi="http://doi.org/10.5281/zenodo.5292704" httpUri="https://zenodo.org/record/5292704/files/figure.png" pageId="3" pageNumber="99">Figs. 2b, 3b</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C655A4E09FFF4FF67AC2D8976FEAD" pageId="3" pageNumber="99" type="materials_examined">
|
||
<paragraph id="8BF936D14E09FFF4FF67AC2D8976FEAD" blockId="3.[151,1436,259,901]" pageId="3" pageNumber="99">
|
||
<emphasis id="B932EAC34E09FFF4FF67AC2D8B5FFE8E" bold="true" box="[151,262,259,284]" inLineHeading="true" pageId="3" pageNumber="99" reason="1">Material.</emphasis>
|
||
<materialsCitation id="3B2E3C8C4E09FFF4FEFFAC2A8BB9FEAD" collectorName="Dos Novillos" country="Costa Rica" county="Guapiles" elevation="410" latitude="10.212777" location="Don Eladio" longLatPrecision="21" longitude="-83.59139" municipality="Dos Novillos" pageId="3" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Limon">
|
||
<specimenCount id="9D40FD584E09FFF4FEFFAC2A8B6BFE89" box="[271,306,260,284]" pageId="3" pageNumber="99" type="male">1♂</specimenCount>
|
||
exuvia (reared).
|
||
<collectingCountry id="F35176414E09FFF4FE06AC2A88F9FE8E" box="[502,672,260,284]" name="Costa Rica" pageId="3" pageNumber="99">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="4982F8334E09FFF4FD5CAC2A893DFE8E" box="[684,868,260,284]" country="Costa Rica" name="Limon" pageId="3" pageNumber="99">Limón province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62984E5D4E09FFF4FC81AC2A898EFE8E" box="[881,983,260,284]" pageId="3" pageNumber="99">Guápiles</collectingCounty>
|
||
,
|
||
<collectorName id="26B353074E09FFF4FC14AC2A8E23FE8E" box="[996,1146,260,284]" pageId="3" pageNumber="99">
|
||
<collectingMunicipality id="6B9DACAB4E09FFF4FC14AC2A8E23FE8E" box="[996,1146,260,284]" pageId="3" pageNumber="99">Dos Novillos</collectingMunicipality>
|
||
</collectorName>
|
||
stream,
|
||
<location id="8E99600A4E09FFF4FB2EAC2A8F39FE8E" LSID="urn:lsid:plazi:treatment:03EF87C74E09FFF4FF67ADB98B9FFC17:8E99600A4E09FFF4FB2EAC2A8F39FE8E" box="[1246,1376,260,284]" country="Costa Rica" county="Guapiles" latitude="10.212777" longLatPrecision="21" longitude="-83.59139" municipality="Dos Novillos" name="Don Eladio" pageId="3" pageNumber="99" stateProvince="Limon">Don Eladio</location>
|
||
(
|
||
<quantity id="4CBE9B344E09FFF4FA80AC2A8AE9FEAD" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="99" unit="m" value="410.0">
|
||
<elevation id="006BD1E24E09FFF4FA80AC2A8AE9FEAD" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="99" unit="m" value="410.0">410 m</elevation>
|
||
</quantity>
|
||
),
|
||
<geoCoordinate id="EE7250164E09FFF4FF34AC098B13FEAD" box="[196,330,295,319]" degrees="10" direction="north" minutes="12" orientation="latitude" pageId="3" pageNumber="99" precision="15" seconds="46" value="10.212777">10°12’46 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7250164E09FFF4FEA6AC098BB9FEAD" box="[342,480,295,319]" degrees="83" direction="west" minutes="35" orientation="longitude" pageId="3" pageNumber="99" precision="15" seconds="29" value="-83.59139">
|
||
83°35’
|
||
<specimenCount id="9D40FD584E09FFF4FE6FAC098BB9FEAD" box="[415,480,295,319]" pageId="3" pageNumber="99" type="generic">29 W</specimenCount>
|
||
</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
<date id="FFF810114E09FFF4FE1BAC098836FEAD" box="[491,623,295,319]" pageId="3" pageNumber="99" value="2006-04-16">16-IV-2006</date>
|
||
. P. Gutiérrez leg.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C655A4E09FFF4FF37AC658B67FC8B" pageId="3" pageNumber="99" type="description">
|
||
<paragraph id="8BF936D14E09FFF4FF37AC658839FE15" blockId="3.[151,1436,259,901]" pageId="3" pageNumber="99">
|
||
<emphasis id="B932EAC34E09FFF4FF37AC658B01FEF6" bold="true" box="[199,344,331,356]" inLineHeading="true" pageId="3" pageNumber="99" reason="1">Description.</emphasis>
|
||
Larvae uniformly light brown, body robust with cylindrical abdomen, integument densely covered with small setae, as in
|
||
<taxonomicName id="4C464D524E09FFF4FE04AC5F8800FE15" authorityName="Selys" authorityYear="1886" box="[500,601,368,392]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="99" phylum="Arthropoda" rank="species" species="majus">
|
||
<emphasis id="B932EAC34E09FFF4FE04AC5F8800FE15" box="[500,601,368,392]" italics="true" pageId="3" pageNumber="99">H. majus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF936D14E09FFF4FF37ACBA8B92FDD3" blockId="3.[151,1436,259,901]" pageId="3" pageNumber="99">
|
||
<emphasis id="B932EAC34E09FFF4FF37ACBA8B5DFE3E" box="[199,260,404,428]" italics="true" pageId="3" pageNumber="99">Head</emphasis>
|
||
. As in
|
||
<taxonomicName id="4C464D524E09FFF4FEA4ACBB8BE0FE3E" authorityName="Selys" authorityYear="1886" box="[340,441,405,428]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="99" phylum="Arthropoda" rank="species" species="majus">
|
||
<emphasis id="B932EAC34E09FFF4FEA4ACBB8BE0FE3E" box="[340,441,405,428]" italics="true" pageId="3" pageNumber="99">H. majus</emphasis>
|
||
</taxonomicName>
|
||
, but with following differences: Cephalic lobes with short setae. Antennae 7-segmented, 3
|
||
<superScript id="7C339B994E09FFF4FF55AC998AEBFE57" attach="left" box="[165,178,439,453]" fontSize="6" pageId="3" pageNumber="99">rd</superScript>
|
||
segment longest, relative length of antennomeres: 0.7, 0.7, 1, 0.8, 0.4, 0.3, 0.2. Antennomeres 3-7 without rings of fine setae (
|
||
<figureCitation id="137D2A544E09FFF4FEDFACCF8BDAFE6B" box="[303,387,481,505]" captionStart="FIGURES 2–5" captionStartId="2.[151,264,1464,1487]" captionTargetBox="[178,1422,208,1408]" captionTargetId="figure-268@2.[147,1433,193,1443]" captionTargetPageId="2" captionText="FIGURES 2–5. Details of H. majus and H. atrolineatum. 2) Antennae of H. majus (a) and H. atrolineatum (b); 3) dorsal view of ligula of H. majus (a) and H. atrolineatum (b); 4) dorsal and ventral view of the abdomen and base of caudal gills for H. majus; 5) dorsal view of H. majus caudal gills." figureDoi="http://doi.org/10.5281/zenodo.5292704" httpUri="https://zenodo.org/record/5292704/files/figure.png" pageId="3" pageNumber="99">Fig. 2b</figureCitation>
|
||
). Mandibles as in
|
||
<taxonomicName id="4C464D524E09FFF4FDBFACCC88EDFE6B" authorityName="Selys" authorityYear="1886" box="[591,692,482,505]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="99" phylum="Arthropoda" rank="species" species="majus">
|
||
<emphasis id="B932EAC34E09FFF4FDBFACCC88EDFE6B" box="[591,692,482,505]" italics="true" pageId="3" pageNumber="99">H. majus</emphasis>
|
||
</taxonomicName>
|
||
, but L 1 1' 2 3 4 0 a(m
|
||
<superScript id="7C339B994E09FFF4FC44ACF089B0FE7E" attach="left" box="[948,1001,478,492]" fontSize="6" pageId="3" pageNumber="99">1,2,3,4,5</superScript>
|
||
)b. Prementum longer than wide (2 x
|
||
<quantity id="4CBE9B344E09FFF4FA7DACCF8A9EFD8F" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="99" unit="mm" value="3.0">3 mm</quantity>
|
||
). Labial palp with few short setae on external margin; middle one of 3 apical teeth longest. Ligula with median incision U-shaped (
|
||
<figureCitation id="137D2A544E09FFF4FE80AF078BE7FDD3" box="[368,446,553,577]" captionStart="FIGURES 2–5" captionStartId="2.[151,264,1464,1487]" captionTargetBox="[178,1422,208,1408]" captionTargetId="figure-268@2.[147,1433,193,1443]" captionTargetPageId="2" captionText="FIGURES 2–5. Details of H. majus and H. atrolineatum. 2) Antennae of H. majus (a) and H. atrolineatum (b); 3) dorsal view of ligula of H. majus (a) and H. atrolineatum (b); 4) dorsal and ventral view of the abdomen and base of caudal gills for H. majus; 5) dorsal view of H. majus caudal gills." figureDoi="http://doi.org/10.5281/zenodo.5292704" httpUri="https://zenodo.org/record/5292704/files/figure.png" pageId="3" pageNumber="99">Fig 3b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF936D14E09FFF4FF37AF638BBAFD3F" blockId="3.[151,1436,259,901]" pageId="3" pageNumber="99">
|
||
<emphasis id="B932EAC34E09FFF4FF37AF638B68FDF7" box="[199,305,589,613]" italics="true" pageId="3" pageNumber="99">Abdomen</emphasis>
|
||
. As in
|
||
<taxonomicName id="4C464D524E09FFF4FE77AF608BB6FDF7" authorityName="Selys" authorityYear="1886" box="[391,495,590,613]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="99" phylum="Arthropoda" rank="species" species="majus">
|
||
<emphasis id="B932EAC34E09FFF4FE77AF608BB6FDF7" box="[391,495,590,613]" italics="true" pageId="3" pageNumber="99">H. majus</emphasis>
|
||
</taxonomicName>
|
||
, but posterior margin of abdominal S6–8 with row of strong setae in dorsal view; S9–10 with row of spines. Ventrally, S9–10 with incomplete row of spines in ventral view, more evident toward margins. Gills as in
|
||
<taxonomicName id="4C464D524E09FFF4FE87AFB88B85FD3F" authorityName="Selys" authorityYear="1886" box="[375,476,662,685]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="99" phylum="Arthropoda" rank="species" species="majus">
|
||
<emphasis id="B932EAC34E09FFF4FE87AFB88B85FD3F" box="[375,476,662,685]" italics="true" pageId="3" pageNumber="99">H. majus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF936D14E09FFF4FF37AF948B67FC8B" blockId="3.[151,1436,259,901]" pageId="3" pageNumber="99">
|
||
<emphasis id="B932EAC34E09FFF4FF37AF948B8CFD43" box="[199,469,697,721]" italics="true" pageId="3" pageNumber="99">Measurements (in mm)</emphasis>
|
||
: Total length without gills: 13 (larvae); posterior femur 4.0; posterior tibia 3.5; maximum width of head 3.0; lateral gills 5 without filament (7 with filament), middle gill 4 without filament (6 with filament).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C655A4E09FFF4FF37AE088B9FFC17" pageId="3" pageNumber="99" type="discussion">
|
||
<paragraph id="8BF936D14E09FFF4FF37AE088B9FFC17" blockId="3.[151,1436,259,901]" pageId="3" pageNumber="99">
|
||
<emphasis id="B932EAC34E09FFF4FF37AE088B62FCAF" box="[199,315,806,830]" inLineHeading="true" pageId="3" pageNumber="99" reason="1">
|
||
<emphasis id="B932EAC34E09FFF4FF37AE088B6DFCAC" bold="true" box="[199,308,806,830]" pageId="3" pageNumber="99">Remarks</emphasis>
|
||
.
|
||
</emphasis>
|
||
Larvae of both species were found among accumulations of detritus on the stream bottom, under rocks or where leaf litter accumulated. Streams were
|
||
<quantity id="4CBE9B344E09FFF4FD01AE678960FCF3" box="[753,825,841,865]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="99" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
|
||
wide and had abundant cobbles and continuous water flow, surrounded by forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |