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treatments-xml/data/03/EF/62/03EF6217BF7CFF8A0AC0FA2F995EAA6B.xml
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<document id="E2627A284E2C0D992AF48C9E230341C6" ID-CLB-Dataset="20762" ID-DOI="10.11646/zootaxa.3365.1.1" ID-GBIF-Dataset="1db9bc74-46dd-4263-9784-88a59bd05d59" ID-ISSN="1175­5334" ID-Zenodo-Dep="6038900" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629787146833" checkinUser="felipe" docAuthor="Garcia, Francisco Hita &amp; Fisher, Brian L." docDate="2012" docId="03EF6217BF7CFF8A0AC0FA2F995EAA6B" docLanguage="en" docName="zt03365p123.pdf" docOrigin="Zootaxa 3365" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Tetramorium mallenseana Garcia &amp; Fisher 2012, sp. n." docType="treatment" docVersion="8" lastPageNumber="73" masterDocId="FFD61A6FBF39FFC20A57FF889C5BAB77" masterDocTitle="The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups" masterLastPageNumber="123" masterPageNumber="1" pageNumber="70" updateTime="1699142484625" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="EA4851758EF8AFB9CD7C1680D196A59E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1C990C9357A2DA4B517B980C01AC7AE8">
<mods:title id="FD1D6AAE2D610D4D95A119E0D6AA60A5">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region — taxonomy of the T. bessonii, T. bonibony, T. dysalum, T. marginatum, T. tsingy, and T. weitzeckeri species groups</mods:title>
</mods:titleInfo>
<mods:name id="D66333D69E3A022E21FA152B5CA79A21" type="personal">
<mods:role id="1BB5C547DCC295F8EDE2E50172A4BCD2">
<mods:roleTerm id="5B619E73086BE184711B88CD5AE347DB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05AC7E3408B37010FEDB7D93A34188B8">Garcia, Francisco Hita</mods:namePart>
</mods:name>
<mods:name id="CBE84DF292E07CC39D05C5E59CD76CAB" type="personal">
<mods:role id="828E4EC98B07DFF0E4F7929D876362FB">
<mods:roleTerm id="4049BAD9729ADD2BFE9560D2EA1F4018">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB59535FFFB85388717DCB9A511B2632">Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="85C6224349ECE98E5B8BC287CB36D3E1">text</mods:typeOfResource>
<mods:relatedItem id="502E0C21322EAEB1971A4FF6124B5DD8" type="host">
<mods:titleInfo id="4A43AE1FCD44546B7EAA9C9B3C3C44D2">
<mods:title id="AF3668F9A0D3913F46DA67EA30D70D34">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CBA2691774CC78415AD2BF3BE5D7FA0B">
<mods:date id="F1BDF1A953FD8FE3C29C57CB8F6F5156">2012</mods:date>
<mods:detail id="5FD92F0D17E43B325E03920489249BDF" type="pubDate">
<mods:number id="A4815AFE852F98C1A9F0D6A8B7EAA705">2012-07-04</mods:number>
</mods:detail>
<mods:detail id="6ABA0B4E8F6552DDD960B0BB8BBDAB29" type="volume">
<mods:number id="A0331A31D3ADFCDA4671C15F9436DDC1">3365</mods:number>
</mods:detail>
<mods:extent id="9670B559BBC08189A31B6CEED0D1E22A" unit="page">
<mods:start id="1731C762035FD72EDC8FEB0A82DC00C2">1</mods:start>
<mods:end id="63AF1D568BEFA586575F35754C53C7EC">123</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BD6A6042830DCAA805D22BD7D416BEF8">journal article</mods:classification>
<mods:identifier id="6698392AE23AB3E1DD2624DBFD3D1F24" type="CLB-Dataset">20762</mods:identifier>
<mods:identifier id="FF82BBD536F14FCD59890F3DFC7F1B17" type="DOI">10.11646/zootaxa.3365.1.1</mods:identifier>
<mods:identifier id="C551833273CEF750140B37AA3BE519A1" type="GBIF-Dataset">1db9bc74-46dd-4263-9784-88a59bd05d59</mods:identifier>
<mods:identifier id="7E21C06244940B2EE8F0C29D0A4C6C9E" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="CD53DB9E002E2C68D003EB9589647935" type="Zenodo-Dep">6038900</mods:identifier>
</mods:mods>
<treatment id="03EF6217BF7CFF8A0AC0FA2F995EAA6B" ID-DOI="http://doi.org/10.5281/zenodo.5253664" ID-GBIF-Taxon="193281040" ID-Zenodo-Dep="5253664" LSID="urn:lsid:plazi:treatment:03EF6217BF7CFF8A0AC0FA2F995EAA6B" httpUri="http://treatment.plazi.org/id/03EF6217BF7CFF8A0AC0FA2F995EAA6B" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70">
<subSubSection id="C35C808ABF7CFF870AC0FA2F9E44AEB6" box="[151,543,1447,1473]" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="8BF9D301BF7CFF870AC0FA2F9E44AEB6" blockId="69.[151,543,1447,1506]" box="[151,543,1447,1473]" pageId="69" pageNumber="70">
<heading id="D0B1646DBF7CFF870AC0FA2F9E44AEB6" bold="true" box="[151,543,1447,1473]" fontSize="11" level="1" pageId="69" pageNumber="70" reason="1">
<emphasis id="B9320F13BF7CFF870AC0FA2F9E44AEB6" bold="true" box="[151,543,1447,1473]" pageId="69" pageNumber="70">
<taxonomicName id="4C46A882BF7CFF870AC0FA2F9D8DAEB6" authority="Garcia &amp; Fisher, 2012" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[151,470,1447,1473]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="mallenseana" status="sp. nov.">
<emphasis id="B9320F13BF7CFF870AC0FA2F9D8DAEB6" bold="true" box="[151,470,1447,1473]" italics="true" pageId="69" pageNumber="70">Tetramorium mallenseana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201B268BF7CFF870B8BFA2F9E44AEB6" box="[476,543,1447,1473]" pageId="69" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7CFF870AC0FA429DC5AE95" box="[151,414,1482,1506]" pageId="69" pageNumber="70" type="description">
<paragraph id="8BF9D301BF7CFF870AC0FA429DC5AE95" blockId="69.[151,543,1447,1506]" box="[151,414,1482,1506]" pageId="69" pageNumber="70">(figs 77, 104, 105, 106)</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7CFF870AC0F99A9E0FAC51" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="8BF9D301BF7CFF870AC0F99A9E0FAC51" blockId="69.[151,1436,1554,1830]" pageId="69" pageNumber="70">
<materialsCitation id="3B2ED95CBF7CFF870AC0F99A99C2AD05" ID-GBIF-Occurrence="3462705420" accessionNumber="BLF08400" collectingDate="2003-03-21" collectingDateMax="2003-03-31" collectingDateMin="2003-03-21" collectionCode="CASC" collectorName="B. L. Fisher &amp; C. Griswold" country="Madagascar" elevation="1100" latitude="-21.29" location="Vatoharanana River" longLatPrecision="555" longitude="47.43333" municipality="Parc National de Ranomafana" pageId="69" pageNumber="70" specimenCode="CASENT0039659" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="54FD6DA3BF7CFF870AC0F99A9CA4AD5D" box="[151,255,1554,1578]" pageId="69" pageNumber="70" type="holotype">Holotype</typeStatus>
worker,
<collectingCountry id="F3519391BF7CFF870B3CF99A9E6EAD5D" box="[363,565,1554,1578]" name="Madagascar" pageId="69" pageNumber="70">MADAGASCAR</collectingCountry>
,
<collectingRegion id="49821DE3BF7CFF870812F99A9E8CAD5D" box="[581,727,1554,1578]" country="Madagascar" name="Fianarantsoa" pageId="69" pageNumber="70">Fianarantsoa</collectingRegion>
,
<collectingMunicipality id="6B9D497BBF7CFF8708B0F99B981DAD5D" box="[743,1094,1554,1578]" pageId="69" pageNumber="70">Parc National de Ranomafana</collectingMunicipality>
,
<location id="8E9985DABF7CFF870E01F99A9961AD5D" LSID="urn:lsid:plazi:treatment:03EF6217BF7CFF8A0AC0FA2F995EAA6B:8E9985DABF7CFF870E01F99A9961AD5D" box="[1110,1338,1554,1578]" country="Madagascar" latitude="-21.29" longLatPrecision="555" longitude="47.43333" municipality="Parc National de Ranomafana" name="Vatoharanana River" pageId="69" pageNumber="70" stateProvince="Fianarantsoa">Vatoharanana River</location>
,
<locationDeviation id="9A4BCA23BF7CFF870F1DF99A9DC1AD39" location="Ranomafana" pageId="69" pageNumber="70">
<quantity id="4CBE7EE4BF7CFF870F1DF99A99C7AD5D" box="[1354,1436,1554,1578]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="69" pageNumber="70" unit="km" value="4.1">4.1 km</quantity>
231° SW Ranomafana
</locationDeviation>
,
<geoCoordinate id="EE72B5C6BF7CFF870BFFF9BE9E59AD39" box="[424,514,1590,1614]" degrees="21.29" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="555" value="-21.29">21.29 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF7CFF870846F9BE9ECCAD39" box="[529,663,1590,1614]" degrees="47.43333" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="1" value="47.43333">47.43333 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF7CFF8708F1F9BE9EA4AD39" box="[678,767,1590,1614]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="69" pageNumber="70" unit="m" value="1100.0">
<elevation id="006B3432BF7CFF8708F1F9BE9EA4AD39" box="[678,767,1590,1614]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="69" pageNumber="70" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF08400,
<date id="FFF8F5C1BF7CFF870BBBF9D29EC7AD05" box="[492,668,1626,1650]" pageId="69" pageNumber="70" value="2003-03-21" valueMax="2003-03-31" valueMin="2003-03-21">
<collectingDate id="EFBC0C29BF7CFF870BBBF9D29EC7AD05" box="[492,668,1626,1650]" pageId="69" pageNumber="70" value="2003-03-21" valueMax="2003-03-31" valueMin="2003-03-21">21.–31.III.2003</collectingDate>
</date>
(
<emphasis id="B9320F13BF7CFF8708E2F9D39863AD04" box="[693,1080,1627,1651]" italics="true" pageId="69" pageNumber="70">
<collectorName id="26B3B6D7BF7CFF8708E2F9D39F65AD04" box="[693,830,1627,1651]" pageId="69" pageNumber="70">B.L. Fisher</collectorName>
,
<collectorName id="26B3B6D7BF7CFF870904F9D39FBEAD04" box="[851,997,1627,1651]" pageId="69" pageNumber="70">C. Griswold</collectorName>
et al.
</emphasis>
) (
<collectionCode id="ED574BC4BF7CFF870E0DF9D298F2AD05" box="[1114,1193,1626,1650]" pageId="69" pageNumber="70">CASC</collectionCode>
:
<specimenCode id="DBE07B7ABF7CFF870EEAF9D299CAAD05" box="[1213,1425,1626,1650]" collectionCode="CASC" pageId="69" pageNumber="70">CASENT0039659</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B2ED95CBF7CFF870AC0F9F79857ADE1" ID-GBIF-Occurrence="3462706007" accessionNumber="BLF08400" box="[151,1036,1662,1687]" collectingDate="2003-03-21" collectingDateMax="2003-03-31" collectingDateMin="2003-03-21" collectionCode="CASC" collectorName="B. L. Fisher &amp; C. Griswold" country="Madagascar" elevation="1100" latitude="-21.29" location="Vatoharanana River" longLatPrecision="555" longitude="47.43333" municipality="Parc National de Ranomafana" pageId="69" pageNumber="70" specimenCode="CASENT039656" specimenCount="1" specimenCount-worker="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="54FD6DA3BF7CFF870AC0F9F79D5CADE0" box="[151,263,1663,1687]" pageId="69" pageNumber="70" type="paratype">Paratypes</typeStatus>
, one worker with same data as holotype (
<collectionCode id="ED574BC4BF7CFF8708B4F9F69F69ADE1" box="[739,818,1662,1686]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="69" pageNumber="70">CASC</collectionCode>
:
<specimenCode id="DBE07B7ABF7CFF870969F9F6985FADE1" box="[830,1028,1662,1686]" collectionCode="CASC" pageId="69" pageNumber="70">CASENT039656</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B2ED95CBF7CFF870E4EF9F69D36ADA9" ID-GBIF-Occurrence="3462705999" collectingDate="1999-02-23" collectingDateMax="1999-02-28" collectingDateMin="1999-02-23" collectingMethod="pitfall trap" collectionCode="MCZ" collectorName="W. E. Steiner" country="Madagascar" elevation="1000" location="montane rainforest" pageId="69" pageNumber="70" specimenCode="CASENT0247148" specimenCount="1" specimenCount-worker="1" stateProvince="Fianarantsoa" typeStatus="paratype">
five workers from
<collectingRegion id="49821DE3BF7CFF870EB8F9F699DAADE1" box="[1263,1409,1662,1686]" country="Madagascar" name="Fianarantsoa" pageId="69" pageNumber="70">Fianarantsoa</collectingRegion>
,
<locationDeviation id="9A4BCA23BF7CFF870FD9F9F79DECADCD" pageId="69" pageNumber="70">
<quantity id="4CBE7EE4BF7CFF870FD9F9F79CE0ADCD" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="69" pageNumber="70" unit="km" value="7.0">7 km</quantity>
W of Ranomafana
</locationDeviation>
,
<location id="8E9985DABF7CFF870B98F92B9EEAADCD" LSID="urn:lsid:plazi:treatment:03EF6217BF7CFF8A0AC0FA2F995EAA6B:8E9985DABF7CFF870B98F92B9EEAADCD" box="[463,689,1698,1722]" country="Madagascar" name="montane rainforest" pageId="69" pageNumber="70" stateProvince="Fianarantsoa">montane rainforest</location>
,
<collectingMethod id="5207AB16BF7CFF87089EF92A9F11ADCD" box="[713,842,1698,1722]" pageId="69" pageNumber="70">pitfall trap</collectingMethod>
<quantity id="4CBE7EE4BF7CFF870909F92A9F9AADCD" box="[862,961,1698,1722]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="69" pageNumber="70" unit="m" value="1000.0">
<elevation id="006B3432BF7CFF870909F92A9F9AADCD" box="[862,961,1698,1722]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="69" pageNumber="70" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFF8F5C1BF7CFF87098EF92A9824ADCD" box="[985,1151,1698,1722]" pageId="69" pageNumber="70" value="1999-02-23" valueMax="1999-02-28" valueMin="1999-02-23">
<collectingDate id="EFBC0C29BF7CFF87098EF92A9824ADCD" box="[985,1151,1698,1722]" pageId="69" pageNumber="70" value="1999-02-23" valueMax="1999-02-28" valueMin="1999-02-23">23.–28.II.1999</collectingDate>
</date>
(
<emphasis id="B9320F13BF7CFF870ECBF92B996AADCC" box="[1180,1329,1699,1723]" italics="true" pageId="69" pageNumber="70">
<collectorName id="26B3B6D7BF7CFF870ECBF92B996AADCC" box="[1180,1329,1699,1723]" pageId="69" pageNumber="70">W.E. Steiner</collectorName>
</emphasis>
) (
<collectionCode id="ED574BC4BF7CFF870F01F92A99C3ADCD" box="[1366,1432,1698,1722]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="69" pageNumber="70" type="Museum">MCZ</collectionCode>
:
<specimenCode id="DBE07B7ABF7CFF870AC0F94E9D31ADA9" box="[151,362,1734,1758]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="69" pageNumber="70" type="Museum">CASENT0247148</specimenCode>
;
</materialsCitation>
<materialsCitation id="3B2ED95CBF7CFF870B2EF94E9ED7ADA9" ID-GBIF-Occurrence="3462705554" box="[377,652,1734,1758]" collectingDate="1999-02-23" collectingDateMax="1999-02-28" collectingDateMin="1999-02-23" collectionCode="MCZ" collectorName="W. E. Steiner" country="Madagascar" elevation="1000" location="Ranomafana" pageId="69" pageNumber="70" specimenCode="CASENT0247148" specimenCount="1" specimenCount-worker="1" stateProvince="Fianarantsoa" typeStatus="paratype">CASENT0247157); and</materialsCitation>
<materialsCitation id="3B2ED95CBF7CFF8708C0F94E98EEAC75" ID-GBIF-Occurrence="3462705653" collectingDate="1999-03-08" collectingDateMax="1999-03-13" collectingDateMin="1999-03-08" collectingMethod="pitfall trap" collectionCode="MCZ" collectorName="W. E. Steiner" country="Madagascar" elevation="1000" location="Ranomafana" pageId="69" pageNumber="70" specimenCode="CASENT0247153" specimenCount="1" specimenCount-worker="1" stateProvince="Fianarantsoa" typeStatus="paratype">
six workers from
<collectingRegion id="49821DE3BF7CFF87093DF94E9FA7ADA9" box="[874,1020,1734,1758]" country="Madagascar" name="Fianarantsoa" pageId="69" pageNumber="70">Fianarantsoa</collectingRegion>
,
<quantity id="4CBE7EE4BF7CFF870E5CF94F9813ADA9" box="[1035,1096,1734,1758]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="69" pageNumber="70" unit="km" value="7.0">
<locationDeviation id="9A4BCA23BF7CFF870E5CF94F9813ADA9" box="[1035,1096,1734,1758]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="69" pageNumber="70" unit="km" value="7.0">7 km</locationDeviation>
</quantity>
W of
<location id="8E9985DABF7CFF870ECBF94E9977ADA9" LSID="urn:lsid:plazi:treatment:03EF6217BF7CFF8A0AC0FA2F995EAA6B:8E9985DABF7CFF870ECBF94E9977ADA9" box="[1180,1324,1734,1758]" country="Madagascar" name="Ranomafana" pageId="69" pageNumber="70" stateProvince="Fianarantsoa">Ranomafana</location>
, montane rainforest,
<collectingMethod id="5207AB16BF7CFF870B4EF9629DCCAC75" box="[281,407,1770,1794]" pageId="69" pageNumber="70">pitfall trap</collectingMethod>
<quantity id="4CBE7EE4BF7CFF870BF0F9629E5CAC75" box="[423,519,1770,1794]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="69" pageNumber="70" unit="m" value="1000.0">
<elevation id="006B3432BF7CFF870BF0F9629E5CAC75" box="[423,519,1770,1794]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="69" pageNumber="70" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFF8F5C1BF7CFF87084BF9629EE7AC75" box="[540,700,1770,1794]" pageId="69" pageNumber="70" value="1999-03-08" valueMax="1999-03-13" valueMin="1999-03-08">
<collectingDate id="EFBC0C29BF7CFF87084BF9629EE7AC75" box="[540,700,1770,1794]" pageId="69" pageNumber="70" value="1999-03-08" valueMax="1999-03-13" valueMin="1999-03-08">8.–13.III.1999</collectingDate>
</date>
(
<emphasis id="B9320F13BF7CFF870881F9639F33AC74" box="[726,872,1771,1795]" italics="true" pageId="69" pageNumber="70">
<collectorName id="26B3B6D7BF7CFF870881F9639F33AC74" box="[726,872,1771,1795]" pageId="69" pageNumber="70">W.E. Steiner</collectorName>
</emphasis>
) (
<collectionCode id="ED574BC4BF7CFF8709DEF9629F90AC75" box="[905,971,1770,1794]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="69" pageNumber="70" type="Museum">MCZ</collectionCode>
:
<specimenCode id="DBE07B7ABF7CFF870989F96298EAAC75" box="[990,1201,1770,1794]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="69" pageNumber="70" type="Museum">CASENT0247153</specimenCode>
;
</materialsCitation>
<materialsCitation id="3B2ED95CBF7CFF870E92F9629D36AC51" ID-GBIF-Occurrence="3462705476" collectingDate="1999-03-08" collectingDateMax="1999-03-13" collectingDateMin="1999-03-08" collectionCode="MCZ" collectorName="W. E. Steiner" country="Madagascar" elevation="1000" location="Ranomafana" pageId="69" pageNumber="70" specimenCode="CASENT0247153" specimenCount="1" specimenCount-worker="1" stateProvince="Fianarantsoa" typeStatus="paratype">CASENT0247154; CASENT0247155;</materialsCitation>
<materialsCitation id="3B2ED95CBF7CFF870B22F8869E0BAC51" ID-GBIF-Occurrence="3462705440" box="[373,592,1806,1830]" collectingDate="1999-03-08" collectingDateMax="1999-03-13" collectingDateMin="1999-03-08" collectionCode="MCZ" collectorName="W. E. Steiner" country="Madagascar" elevation="1000" location="Ranomafana" pageId="69" pageNumber="70" specimenCode="CASENT0247153" specimenCount="1" specimenCount-worker="1" stateProvince="Fianarantsoa" typeStatus="paratype">CASENT0247156)</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7CFF840AC0F8DE9FFFABA3" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="diagnosis">
<paragraph id="8BF9D301BF7CFF870AC0F8DE9D51AC18" blockId="69.[151,1436,1878,2010]" box="[151,266,1878,1903]" pageId="69" pageNumber="70">
<heading id="D0B1646DBF7CFF870AC0F8DE9D51AC18" bold="true" box="[151,266,1878,1903]" fontSize="10" level="2" pageId="69" pageNumber="70" reason="2">
<emphasis id="B9320F13BF7CFF870AC0F8DE9D51AC18" bold="true" box="[151,266,1878,1903]" pageId="69" pageNumber="70">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF7CFF840AC0F8F29FFFABA3" blockId="69.[151,1436,1878,2010]" lastBlockId="70.[151,1436,151,212]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
The following character combination distinguishes
<taxonomicName id="4C46A882BF7CFF8708A7F8F39FC5ACE4" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[752,926,1915,1939]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="mallenseana">
<emphasis id="B9320F13BF7CFF8708A7F8F39FC5ACE4" box="[752,926,1915,1939]" italics="true" pageId="69" pageNumber="70">T. mallenseana</emphasis>
</taxonomicName>
from the other species of the group: short antennal scapes (SI 72–75); long and curved propodeal spines (PSLI 38–43); very short propodeal lobes; petiolar node rounded high nodiform, in dorsal view only slightly wider than long (DPel 110–123), petiolar dorsum only weakly tapering backwards posteriorly; postpetiole in dorsal view 1.7 to 2 times wider than petiolar node (PPI 168–200); dorsum of mesosoma with longitudinally rugose sculpture.
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7FFF840AC0FE8B99D8AFD7" pageId="70" pageNumber="71" type="description">
<paragraph id="8BF9D301BF7FFF840AC0FE8B9D7AAA6B" blockId="70.[151,1436,259,1184]" box="[151,289,259,284]" pageId="70" pageNumber="71">
<heading id="D0B1646DBF7FFF840AC0FE8B9D7AAA6B" bold="true" box="[151,289,259,284]" fontSize="10" level="2" pageId="70" pageNumber="71" reason="2">
<emphasis id="B9320F13BF7FFF840AC0FE8B9D7AAA6B" bold="true" box="[151,289,259,284]" pageId="70" pageNumber="71">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF7FFF840AC0FEA09925AAB8" blockId="70.[151,1436,259,1184]" pageId="70" pageNumber="71">
HL 0.67–0.71 (0.70); HW 0.67–0.72 (0.70); SL 0.49–0.54 (0.51); EL 0.12–0.15 (0.14);
<collectingCountry id="F3519391BF7FFF840E2CFEA098C4AA48" box="[1147,1183,296,319]" name="Philippines" pageId="70" pageNumber="71">PH</collectingCountry>
0.37–0.40 (0.38); PW 0.50–0.54 (0.52); WL 0.84–0.95 (0.88); PSL 0.27–0.29 (0.28); PTL 0.15–0.17 (0.16); PTH 0.30–0.33 (0.31); PTW 0.17–0.20 (0.19); PPL 0.20–0.23 (0.22); PPH 0.29–0.32 (0.30); PPW 0.29–0.38 (0.34); CI 99–101 (100); SI 72–75 (73); OI 18–21 (20); DMI 57–61 (59); LMI 42–44 (43); PSLI 38–43 (40); PeNI 33–38 (36); LPeI 48–53 (51); DPeI 110–123 (117); PpNI 58–76 (72); LPpI 66–79 (72); DPpI 145–165 (156); PPI 168–200 (182) (seven measured).
</paragraph>
<paragraph id="8BF9D301BF7FFF840A90FE5499D8AFD7" blockId="70.[151,1436,259,1184]" pageId="70" pageNumber="71">Head as long as wide (CI 99–101). Anterior clypeal margin with distinct median impression. Frontal carinae strongly developed, usually ending at posterior head margin. Antennal scrobes narrow and faint. Antennal scapes short, not reaching posterior head margin (SI 72–75). Eyes comparatively small (OI 18–21). Mesosomal outline in profile comparatively flat, moderately weakly marginate from lateral to dorsal mesosoma, promesonotal suture and metanotal groove absent; mesosoma comparatively stout and compact (LMI 42–44). Propodeal spines relatively thin, very long, and back-curved (PSLI 38–43). Propodeal lobes weakly developed, very short and triangular. Petiolar node in profile rounded high nodiform, approximately two times higher than long (LPeI 48–53), anterior and posterior faces almost parallel, anterodorsal margin situated higher than posterodorsal margin, dorsum weakly tapering backwards posteriorly; node shape in dorsal view only weakly elliptical, almost rounded, only between 1.1 to 1.2 times wider than long (DPeI 110–123). Postpetiole in profile rounded, approximately 1.3 to 1.5 times higher than long (LPpI 66–79), in dorsal view 1.4 to 1.7 times wider than long (DPpI 145–165). Postpetiole in profile a bit more voluminous than petiolar node, in dorsal view much wider than petiolar node, between 1.7 to 2 times wider (PPI 168–200). Mandibles distinctly longitudinally rugose, in some specimens unsculptured and smooth or weakly sculptured; clypeus with strong median longitudinal ruga and one often weaker ruga at each side; cephalic dorsum between frontal carinae with six to eight longitudinal rugae running to posterior head margin, some rugae broken or with cross-meshes; lateral and ventral head with weak reticulate-rugose sculpture; head with faint reticulatepunctate to punctate ground sculpture. Mesosoma laterally and dorsally strongly longitudinally rugose, rugae moderately meandering but distinctly longitudinally arranged with few cross-meshes. Petiole and postpetiole almost unsculptured, with few weak rugulae. Gaster unsculptured, smooth and shining. All dorsal surfaces of body with long, erect or suberect pilosity. Head, mesosoma, waist segments dark brown, appendages of lighter brown.</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7FFF840AC0FB589F9CADA8" pageId="70" pageNumber="71" type="discussion">
<paragraph id="8BF9D301BF7FFF840AC0FB589C81AF9F" blockId="70.[151,1436,1232,1759]" box="[151,218,1232,1256]" pageId="70" pageNumber="71">
<heading id="D0B1646DBF7FFF840AC0FB589C81AF9F" bold="true" box="[151,218,1232,1256]" fontSize="10" level="2" pageId="70" pageNumber="71" reason="2">
<emphasis id="B9320F13BF7FFF840AC0FB589C81AF9F" bold="true" box="[151,218,1232,1256]" pageId="70" pageNumber="71">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF7FFF840AC0FB7D9F60AE58" blockId="70.[151,1436,1232,1759]" pageId="70" pageNumber="71">
<taxonomicName id="4C46A882BF7FFF840AC0FB7D9D9AAE7B" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[151,449,1268,1292]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="mallenseana">
<emphasis id="B9320F13BF7FFF840AC0FB7D9D9AAE7B" box="[151,449,1268,1292]" italics="true" pageId="70" pageNumber="71">Tetramorium mallenseana</emphasis>
</taxonomicName>
is only known from the Parc National de Ranomafana and its vicinity, which is a montane rainforest situated at an altitude of
<quantity id="4CBE7EE4BF7FFF8408D3FA9F9F6CAE58" box="[644,823,1303,1327]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="70" pageNumber="71" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000 to 1100 m</quantity>
.
</paragraph>
<paragraph id="8BF9D301BF7FFF840A90FAB49F9CADA8" blockId="70.[151,1436,1232,1759]" pageId="70" pageNumber="71">
Within the
<taxonomicName id="4C46A882BF7FFF840B13FAB59DE7AE23" authorityName="Bolton" authorityYear="1979" box="[324,444,1340,1364]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dysalum">
<emphasis id="B9320F13BF7FFF840B13FAB59DE7AE23" box="[324,444,1340,1364]" italics="true" pageId="70" pageNumber="71">T. dysalum</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C46A882BF7FFF840845FAB59EE0AE23" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[530,699,1340,1364]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="mallenseana">
<emphasis id="B9320F13BF7FFF840845FAB59EE0AE23" box="[530,699,1340,1364]" italics="true" pageId="70" pageNumber="71">T. mallenseana</emphasis>
</taxonomicName>
has an interesting character combination that makes it unlikely to be misidentified with another group member. The propodeal spines are very long and strongly backcurved. This shape is only comparable with the shape of the spines seen in
<taxonomicName id="4C46A882BF7FFF84090AFA0D9F8CAEEB" authorityName="Bolton" authorityYear="1979" box="[861,983,1412,1436]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dysalum">
<emphasis id="B9320F13BF7FFF84090AFA0D9F8CAEEB" box="[861,983,1412,1436]" italics="true" pageId="70" pageNumber="71">T. dysalum</emphasis>
</taxonomicName>
, although the species are unlikely to be confused since
<taxonomicName id="4C46A882BF7FFF840B14FA219DE0AEB7" authorityName="Bolton" authorityYear="1979" box="[323,443,1448,1472]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="dysalum">
<emphasis id="B9320F13BF7FFF840B14FA219DE0AEB7" box="[323,443,1448,1472]" italics="true" pageId="70" pageNumber="71">T. dysalum</emphasis>
</taxonomicName>
always has unsculptured, very smooth and shiny mandibles, whereas the mandibles of
<taxonomicName id="4C46A882BF7FFF840FD0FA219D7EAE93" authorityName="Garcia &amp; Fisher" authorityYear="2012" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="mallenseana">
<emphasis id="B9320F13BF7FFF840FD0FA219D7EAE93" italics="true" pageId="70" pageNumber="71">T. mallenseana</emphasis>
</taxonomicName>
are distinctly sculptured. The spines of
<taxonomicName id="4C46A882BF7FFF8408A7FA459F2FAE93" authorityName="Forel" authorityYear="1892" box="[752,884,1484,1508]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="steinheili">
<emphasis id="B9320F13BF7FFF8408A7FA459F2FAE93" box="[752,884,1484,1508]" italics="true" pageId="70" pageNumber="71">T. steinheili</emphasis>
</taxonomicName>
are usually straight and only rarely backcurved, and if they are, then the propodeal lobes are very well-developed, whereas the lobes of
<taxonomicName id="4C46A882BF7FFF840EDDFA79996EAD7F" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[1162,1333,1520,1544]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="mallenseana">
<emphasis id="B9320F13BF7FFF840EDDFA79996EAD7F" box="[1162,1333,1520,1544]" italics="true" pageId="70" pageNumber="71">T. mallenseana</emphasis>
</taxonomicName>
are very short and inconspicuous. The best character separating
<taxonomicName id="4C46A882BF7FFF84094AF99D9F92AD5B" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[797,969,1556,1580]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="mallenseana">
<emphasis id="B9320F13BF7FFF84094AF99D9F92AD5B" box="[797,969,1556,1580]" italics="true" pageId="70" pageNumber="71">T. mallenseana</emphasis>
</taxonomicName>
from the other species, however, is the shape of the petiolar node, especially the relationship between petiolar node width and postpetiole width. The petiolar node in dorsal view is only weakly wider than long (DPeI 110–123), and its shape is only weakly elliptical and almost rounded, which is rare in the group. Also, the postpetiole is always between 1.6 to 2 times wider than the comparatively narrow petiolar node (PPI 168–200), while most other species have a postpetiole that is usually distinctly less than 1.5 times wider than the petiolar node (PPI 119–152).
</paragraph>
</subSubSection>
<subSubSection id="C35C808ABF7FFF840AC0F8989ECAAC18" pageId="70" pageNumber="71" type="etymology">
<paragraph id="8BF9D301BF7FFF840AC0F8989D43AC5F" blockId="70.[151,1436,1808,1903]" box="[151,280,1808,1832]" pageId="70" pageNumber="71">
<heading id="D0B1646DBF7FFF840AC0F8989D43AC5F" bold="true" box="[151,280,1808,1832]" fontSize="10" level="2" pageId="70" pageNumber="71" reason="2">
<emphasis id="B9320F13BF7FFF840AC0F8989D43AC5F" bold="true" box="[151,280,1808,1832]" pageId="70" pageNumber="71">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF7FFF840AC0F8BC9ECAAC18" blockId="70.[151,1436,1808,1903]" pageId="70" pageNumber="71">The species epithet is an arbitrary combination of letters, and the new species is dedicated to Sean Allen for his support to discover and identify life on earth.</paragraph>
</subSubSection>
<caption id="DF398389BF7EFF850AC0F8349D87AC99" ID-DOI="http://doi.org/10.5281/zenodo.6039005" ID-Zenodo-Dep="6039005" httpUri="https://zenodo.org/record/6039005/files/figure.png" pageId="71" pageNumber="72" startId="71.[151,258,1980,2001]" targetBox="[186,1431,193,1973]" targetPageId="71">
<paragraph id="8BF9D301BF7EFF850AC0F8349D87AC99" blockId="71.[151,1436,1980,2030]" pageId="71" pageNumber="72">
<emphasis id="B9320F13BF7EFF850AC0F8349D39ACA6" bold="true" box="[151,354,1980,2001]" pageId="71" pageNumber="72">FIGURES 104–106.</emphasis>
<taxonomicName id="4C46A882BF7EFF850B3EF8349DA0ACA6" authorityName="Garcia &amp; Fisher" authorityYear="2012" box="[361,507,1980,2001]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="mallenseana">
<emphasis id="B9320F13BF7EFF850B3EF8349DA0ACA6" box="[361,507,1980,2001]" italics="true" pageId="71" pageNumber="72">T. mallenseana</emphasis>
</taxonomicName>
, holotype—CASENT0039659 (Estella Ortega 2011). 104. Body in profile. 105. Body in dorsal view. 106. Head in full-face view.
</paragraph>
</caption>
<subSubSection id="C35C808ABF71FF8A0AC0FF1F995EAA6B" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="8BF9D301BF71FF8A0AC0FF1F9D22ABC7" blockId="72.[151,1436,151,284]" box="[151,377,151,176]" pageId="72" pageNumber="73">
<heading id="D0B1646DBF71FF8A0AC0FF1F9D22ABC7" bold="true" box="[151,377,151,176]" fontSize="10" level="2" pageId="72" pageNumber="73" reason="2">
<emphasis id="B9320F13BF71FF8A0AC0FF1F9D22ABC7" bold="true" box="[151,377,151,176]" pageId="72" pageNumber="73">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9D301BF71FF8A0AC0FF33995EAA6B" blockId="72.[151,1436,151,284]" pageId="72" pageNumber="73">
<materialsCitation id="3B2ED95CBF71FF8A0AC0FF3399C7AB80" ID-GBIF-Occurrence="3462706106" pageId="72" pageNumber="73">
<emphasis id="B9320F13BF71FF8A0AC0FF339D3FABA3" bold="true" box="[151,356,187,212]" pageId="72" pageNumber="73">
<collectingCountry id="F3519391BF71FF8A0AC0FF339D3FABA3" box="[151,356,187,212]" name="Madagascar" pageId="72" pageNumber="73">MADAGASCAR</collectingCountry>
</emphasis>
:
<collectingRegion id="49821DE3BF71FF8A0B28FF349E4AABA3" box="[383,529,188,212]" country="Madagascar" name="Fianarantsoa" pageId="72" pageNumber="73">Fianarantsoa</collectingRegion>
, Parc National de Ranomafana, Vatoharanana River,
<quantity id="4CBE7EE4BF71FF8A0EEBFF34994DABA3" box="[1212,1302,188,212]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="72" pageNumber="73" unit="km" value="4.1">4.1 km</quantity>
231° SW Ranomafana,
<geoCoordinate id="EE72B5C6BF71FF8A0B64FF579DD7AB80" box="[307,396,223,247]" degrees="21.29" direction="south" orientation="latitude" pageId="72" pageNumber="73" precision="555" value="-21.29">21.29 S</geoCoordinate>
,
<geoCoordinate id="EE72B5C6BF71FF8A0BCDFF579E45AB80" box="[410,542,223,247]" degrees="47.43333" direction="east" orientation="longitude" pageId="72" pageNumber="73" precision="1" value="47.43333">47.43333 E</geoCoordinate>
,
<quantity id="4CBE7EE4BF71FF8A087CFF579ED9AB80" box="[555,642,223,247]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="72" pageNumber="73" unit="m" value="1100.0">1100 m</quantity>
, montane rainforest,
<date id="FFF8F5C1BF71FF8A0924FF579879AB80" box="[883,1058,223,247]" pageId="72" pageNumber="73" value="2003-03-21" valueMax="2003-03-31" valueMin="2003-03-21">21.–31.III.2003</date>
(
<emphasis id="B9320F13BF71FF8A0E64FF6999D0AB8F" box="[1075,1419,224,248]" italics="true" pageId="72" pageNumber="73">B.L. Fisher, C. Griswold et al.</emphasis>
);
</materialsCitation>
<materialsCitation id="3B2ED95CBF71FF8A0AC0FE8C98A4AA6B" ID-GBIF-Occurrence="3462705986" box="[151,1279,260,284]" pageId="72" pageNumber="73">
<collectingRegion id="49821DE3BF71FF8A0AC0FE8C9D72AA6B" box="[151,297,260,284]" country="Madagascar" name="Fianarantsoa" pageId="72" pageNumber="73">Fianarantsoa</collectingRegion>
,
<quantity id="4CBE7EE4BF71FF8A0B63FE8C9D37AA6B" box="[308,364,260,284]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="72" pageNumber="73" unit="km" value="7.0">7 km</quantity>
W of Ranomafana, montane rainforest,
<quantity id="4CBE7EE4BF71FF8A0967FE8C9FDDAA6C" box="[816,902,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FFF8F5C1BF71FF8A09C5FE8C9800AA6B" box="[914,1115,260,284]" pageId="72" pageNumber="73" value="1990-02-23" valueMax="1990-03-13" valueMin="1990-02-23">23.II.–13.III.1990</date>
(
<emphasis id="B9320F13BF71FF8A0E3CFE8D98AFAA6B" box="[1131,1268,260,284]" italics="true" pageId="72" pageNumber="73">W.E. Steiner</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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