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<mods:title id="B254139CC11FD6F98A5139EA008FB3B5">Systematics and palaeobiology of kangaroos of the late Cenozoic genus Protemnodon (Marsupialia, Macropodidae)</mods:title>
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<mods:namePart id="4E45015477D09B014767BE3907F6717A">Kerr, Isaac A. R.</mods:namePart>
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<mods:namePart id="9BB116F6ACFE824DABA2DA5ED7E14617">Van Zoelen, Jacob D.</mods:namePart>
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<mods:date id="D070FCB23F93157A6690B5568F1C4D49">2024</mods:date>
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<treatment id="03E587FDFFE4D53CFCA2724CFAF6FA09" LSID="urn:lsid:plazi:treatment:03E587FDFFE4D53CFCA2724CFAF6FA09" httpUri="http://treatment.plazi.org/id/03E587FDFFE4D53CFCA2724CFAF6FA09" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="C3566560FFE4D53DFCA2724CFB2FF9C9" box="[810,1279,1546,1573]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BF336EBFFE4D53DFCA2724CFB2FF9C9" blockId="8.[810,1279,1546,1573]" box="[810,1279,1546,1573]" pageId="8" pageNumber="9">
<heading id="D0BB8187FFE4D53DFCA2724CFB2FF9C9" bold="true" box="[810,1279,1546,1573]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="1">
<emphasis id="B938EAF9FFE4D53DFCA2724CFB2FF9C9" bold="true" box="[810,1279,1546,1573]" pageId="8" pageNumber="9">
Genus
<taxonomicName id="4C4C4D68FFE4D53DFC09724DFB2FF9C9" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" box="[897,1279,1546,1573]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE4D53DFC09724DFBB5F9C9" bold="true" box="[897,1125,1547,1573]" italics="true" pageId="8" pageNumber="9">PROTEMNODON</emphasis>
<bibRefCitation id="EFDD4B1AFFE4D53DFBE4724CFB2FF9C9" author="Owen, R." box="[1132,1279,1546,1573]" pageId="8" pageNumber="9" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="8BF336EBFFE4D53DFCA27208FC7EF968" blockId="8.[810,1452,1614,1732]" pageId="8" pageNumber="9">
<taxonomicName id="4C4C4D68FFE4D53DFCA27208FBF7F988" authority="Owen, 1873" authorityName="Owen" authorityYear="1873" box="[810,1063,1614,1636]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE4D53DFCA27208FC7CF98F" box="[810,940,1614,1635]" italics="true" pageId="8" pageNumber="9">Protemnodon</emphasis>
Owen, 1873
</taxonomicName>
;
<emphasis id="B938EAF9FFE4D53DFBB87209FB26F98F" box="[1072,1270,1614,1636]" italics="true" pageId="8" pageNumber="9">Proc. Roy. Soc. Lond</emphasis>
. 21, p. 128 (
<emphasis id="B938EAF9FFE4D53DFAE37209FC70F96F" italics="true" pageId="8" pageNumber="9">nomen nudum</emphasis>
).
</paragraph>
<paragraph id="8BF336EBFFE4D53DFCA272C8FA7CF948" blockId="8.[810,1452,1614,1732]" box="[810,1452,1678,1700]" pageId="8" pageNumber="9">
<taxonomicName id="4C4C4D68FFE4D53DFCA272C8FBFBF948" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" box="[810,1067,1678,1700]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE4D53DFCA272C8FC7CF94F" box="[810,940,1678,1699]" italics="true" pageId="8" pageNumber="9">Protemnodon</emphasis>
<bibRefCitation id="EFDD4B1AFFE4D53DFC3972C9FBFBF948" author="Owen, R." box="[945,1067,1679,1700]" pageId="8" pageNumber="9" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
</taxonomicName>
:
<emphasis id="B938EAF9FFE4D53DFBBB72C8FAD4F948" box="[1075,1284,1678,1700]" italics="true" pageId="8" pageNumber="9">Phil. Trans. Roy. Soc.</emphasis>
164, p. 274275.
</paragraph>
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<subSubSection id="C3566560FFE4D53DFCA272E9FAE2F928" box="[810,1330,1710,1732]" pageId="8" pageNumber="9" type="type_taxon">
<paragraph id="8BF336EBFFE4D53DFCA272E9FAE2F928" blockId="8.[810,1452,1614,1732]" box="[810,1330,1710,1732]" pageId="8" pageNumber="9">
<typeStatus id="54F78849FFE4D53DFCA272E9FC8AF928" box="[810,858,1711,1732]" pageId="8" pageNumber="9">Type</typeStatus>
species:
<taxonomicName id="4C4C4D68FFE4D53DFC3972E8FAFEF928" authority="Owen, 1874" authorityName=", Owen" authorityYear="1874" box="[945,1326,1710,1732]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="anak">
<emphasis id="B938EAF9FFE4D53DFC3972E8FBB8F92F" box="[945,1128,1710,1731]" italics="true" pageId="8" pageNumber="9">Protemnodon anak</emphasis>
<bibRefCitation id="EFDD4B1AFFE4D53DFBE672E9FB35F928" author="Owen, R." box="[1134,1253,1711,1732]" pageId="8" pageNumber="9" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
, p. 278
</taxonomicName>
.
</paragraph>
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<paragraph id="8BF336EBFFE4D53DFCA272ACFBC0F8E8" blockId="8.[810,1452,1770,2085]" box="[810,1040,1770,1796]" pageId="8" pageNumber="9">
<emphasis id="B938EAF9FFE4D53DFCA272ACFBC0F8E8" bold="true" box="[810,1040,1770,1796]" pageId="8" pageNumber="9">Revised diagnosis:</emphasis>
</paragraph>
<paragraph id="8BF336EBFFE4D53DFCA2734DFAEBF7C9" blockId="8.[810,1452,1770,2085]" pageId="8" pageNumber="9">
Within
<taxonomicName id="4C4C4D68FFE4D53DFCF7734DFBF4F8C9" ID-CoL="628L6" authorityName="Gray" authorityYear="1821" box="[895,1060,1803,1829]" class="Mammalia" family="Macropodidae" kingdom="Animalia" order="Diprotodontia" pageId="8" pageNumber="9" phylum="Chordata" rank="subFamily" subFamily="Macropodinae">Macropodinae</taxonomicName>
, the species of
<taxonomicName id="4C4C4D68FFE4D53DFB5D734DFABCF8C9" authorityName="Owen" authorityYear="1874" box="[1237,1388,1803,1829]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE4D53DFB5D734DFABCF8C9" box="[1237,1388,1803,1829]" italics="true" pageId="8" pageNumber="9">Protemnodon</emphasis>
</taxonomicName>
most closely resemble those of
<taxonomicName id="4C4C4D68FFE4D53DFBCF736DFAADF8A9" authority="McNamara, 1994" authorityName="McNamara" authorityYear="1994" box="[1095,1405,1835,1861]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE4D53DFBCF736DFB65F8A9" box="[1095,1205,1835,1861]" italics="true" pageId="8" pageNumber="9">Congruus</emphasis>
<bibRefCitation id="EFDD4B1AFFE4D53DFB33736DFAADF8A9" author="McNamara, J. A." box="[1211,1405,1835,1861]" pageId="8" pageNumber="9" pagination="111 - 115" refId="ref152008" refString="McNamara, J. A. (1994) A new fossil wallaby (Marsupialia; Macropodidae) from the south east of South Australia. Records of the South Australian Museum, 27 (2), 111 - 115." type="journal article" year="1994">McNamara, 1994</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C4C4D68FFE4D53DFCA2730DFC5EF889" ID-CoL="87G7" authorityName="Trouessart" authorityYear="1904" box="[810,910,1867,1893]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE4D53DFCA2730DFC5EF889" box="[810,910,1867,1893]" italics="true" pageId="8" pageNumber="9">Wallabia</emphasis>
</taxonomicName>
in craniodental features. Species of
<taxonomicName id="4C4C4D68FFE4D53DFA9C730DFA7BF889" authorityName="Owen" authorityYear="1874" box="[1300,1451,1867,1893]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE4D53DFA9C730DFA7BF889" box="[1300,1451,1867,1893]" italics="true" pageId="8" pageNumber="9">Protemnodon</emphasis>
</taxonomicName>
differ from all known macropodines in having: an ilium with a highly reduced cranial iliac spine, a craniocaudally deeper caudal iliac spine and an associated deeply concave gluteal fossa; and a middle pedal phalanx IV that is short, broad, and highly dorsoplantarly compressed, with its shaft only slightly narrower than the distal end.
</paragraph>
</subSubSection>
<subSubSection id="C3566560FFE5D53CFF0074EDFAE6FC89" pageId="9" pageNumber="10" type="multiple">
<paragraph id="8BF336EBFFE5D53CFF0074EDFE06FF28" blockId="9.[136,779,170,1669]" box="[136,470,170,197]" pageId="9" pageNumber="10">
<emphasis id="B938EAF9FFE5D53CFF0074EDFE06FF28" bold="true" box="[136,470,170,197]" pageId="9" pageNumber="10">
<taxonomicName id="4C4C4D68FFE5D53CFF0074EDFEE3FF29" authorityName="Owen" authorityYear="1874" box="[136,307,171,197]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">Protemnodon</taxonomicName>
<emphasis id="B938EAF9FFE5D53CFEB274EDFE82FF29" bold="true" box="[314,338,171,197]" italics="true" pageId="9" pageNumber="10">vs</emphasis>
<taxonomicName id="4C4C4D68FFE5D53CFED274ECFE06FF28" box="[346,470,170,196]" class="Mammalia" family="Macropodidae" genus="Congruus" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="undetermined">Congruus</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BF336EBFFE5D53CFF00748DFE48FC49" blockId="9.[136,779,170,1669]" pageId="9" pageNumber="10">
Species of
<taxonomicName id="4C4C4D68FFE5D53CFE8C748DFE4BFF09" authorityName="Owen" authorityYear="1874" box="[260,411,203,229]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFE8C748DFE4BFF09" box="[260,411,203,229]" italics="true" pageId="9" pageNumber="10">Protemnodon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D68FFE5D53CFE5B748DFD91FF09" authorityName="McNamara" authorityYear="1994" box="[467,577,203,229]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFE5B748DFD91FF09" box="[467,577,203,229]" italics="true" pageId="9" pageNumber="10">Congruus</emphasis>
</taxonomicName>
share 14 thoracic and five lumbar vertebrae to the exclusion of all other macropodines, which have 13 and six, respectively. Dentally, the species of
<taxonomicName id="4C4C4D68FFE5D53CFE2D756DFDECFEA9" authorityName="Owen" authorityYear="1874" box="[421,572,299,325]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFE2D756DFDECFEA9" box="[421,572,299,325]" italics="true" pageId="9" pageNumber="10">Protemnodon</emphasis>
</taxonomicName>
are distinguished fromthoseof
<taxonomicName id="4C4C4D68FFE5D53CFE91750DFE57FE89" authorityName="McNamara" authorityYear="1994" box="[281,391,331,357]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFE91750DFE57FE89" box="[281,391,331,357]" italics="true" pageId="9" pageNumber="10">Congruus</emphasis>
</taxonomicName>
byhavinglongerpremolarsrelative to molars and a more dorsally inclined lower incisor. Species of
<taxonomicName id="4C4C4D68FFE5D53CFE8B75CDFE4AFE49" authorityName="Owen" authorityYear="1874" box="[259,410,395,421]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFE8B75CDFE4AFE49" box="[259,410,395,421]" italics="true" pageId="9" pageNumber="10">Protemnodon</emphasis>
</taxonomicName>
differ from species of
<taxonomicName id="4C4C4D68FFE5D53CFD1375CDFCD9FE49" authorityName="McNamara" authorityYear="1994" box="[667,777,395,421]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFD1375CDFCD9FE49" box="[667,777,395,421]" italics="true" pageId="9" pageNumber="10">Congruus</emphasis>
</taxonomicName>
in the following cranial attributes: cranium larger and more robust; premaxilla lacks an anterior sulcus within the nasal cavity, and is taller and more robust anteriorly; nasal less domed and less anteriorly projected; larger masseteric process; thicker, less laterally projected orbital lip on jugal; taller zygomatic arch; foramen ovale with a single anteroposterior ridge abutting the medial margin, rather than being abutted at the anterior by a shallow but distinct anteroposterior groove; occipital condyles that project posteriorly well beyond the posterior margin of the nuchal crest and occiput; and a broader foramen magnum relative to occipital height and width. The dentary of the species of
<taxonomicName id="4C4C4D68FFE5D53CFE8B776DFE4AFCA9" authorityName="Owen" authorityYear="1874" box="[259,410,811,837]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFE8B776DFE4AFCA9" box="[259,410,811,837]" italics="true" pageId="9" pageNumber="10">Protemnodon</emphasis>
</taxonomicName>
differs from those of
<taxonomicName id="4C4C4D68FFE5D53CFD13776DFCD9FCA9" authorityName="McNamara" authorityYear="1994" box="[667,777,811,837]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFD13776DFCD9FCA9" box="[667,777,811,837]" italics="true" pageId="9" pageNumber="10">Congruus</emphasis>
</taxonomicName>
in its more inflected diastema, more posteriorly situated mental foramen, and deeper medial pterygoid fossa with higher posterior margin.
</paragraph>
<paragraph id="8BF336EBFFE5D53CFF3077EDFE28F969" blockId="9.[136,779,170,1669]" pageId="9" pageNumber="10">
The forelimb differs by having: humerus more robust, with more dorsally projected greater and lesser tubercles and a deeper proximal shaft that deepens proximally; proximal ulna more transversely compressed; more elongate metacarpals; and manual phalanges considerably more dorsopalmarly compressed, particularly the distal phalanges. Within the hindlimb, species of
<taxonomicName id="4C4C4D68FFE5D53CFDFA702DFCD9FB69" authorityName="Owen" authorityYear="1874" box="[626,777,1131,1157]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFDFA702DFCD9FB69" box="[626,777,1131,1157]" italics="true" pageId="9" pageNumber="10">Protemnodon</emphasis>
</taxonomicName>
are distinguished from those of
<taxonomicName id="4C4C4D68FFE5D53CFE6A70CDFD80FB49" authorityName="McNamara" authorityYear="1994" box="[482,592,1163,1189]" class="Mammalia" family="Macropodidae" genus="Congruus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFE6A70CDFD80FB49" box="[482,592,1163,1189]" italics="true" pageId="9" pageNumber="10">Congruus</emphasis>
</taxonomicName>
by: a pelvis with a broader ilium and a smaller, blunter iliopubic eminence; straighter femur in dorsal view with a larger, more dorsally deflected head, more distally extensive lesser trochanteric ridge, less medially displaced quadratus tubercle, and a more proximally situated lateral gastrocnemial fossa; tibia with a deeper, flatter and more distinct distal fibular facet; talus with a more caudoplantarly extensive navicular facet, and a cuboid facet on the navicular head; a taller, narrower navicular; a more robust metatarsal IV; metatarsal V with a proximal plantar groove present, larger lateral plantar tuberosity and medial plantar tubercle, and lacking a slight kink of the arch of the shaft immediately proximal to midpoint in lateral view; and dorsoplantarly shorter distal phalanges with a more rounded dorsal peak and a less plantarly curved shaft.
</paragraph>
<paragraph id="8BF336EBFFE5D53CFF0072EDFE1AF928" blockId="9.[136,778,1706,2085]" box="[136,458,1706,1733]" pageId="9" pageNumber="10">
<emphasis id="B938EAF9FFE5D53CFF0072EDFE1AF928" bold="true" box="[136,458,1706,1733]" pageId="9" pageNumber="10">
<taxonomicName id="4C4C4D68FFE5D53CFF0072EDFEE3F929" authorityName="Owen" authorityYear="1874" box="[136,307,1707,1733]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">Protemnodon</taxonomicName>
<emphasis id="B938EAF9FFE5D53CFEB272EDFE82F929" bold="true" box="[314,338,1707,1733]" italics="true" pageId="9" pageNumber="10">vs</emphasis>
<taxonomicName id="4C4C4D68FFE5D53CFED272ECFE1AF928" authorityName="Trouessart" authorityYear="1904" box="[346,458,1706,1732]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">Wallabia</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BF336EBFFE5D53CFF00728DFAB4FE69" blockId="9.[136,778,1706,2085]" lastBlockId="9.[810,1452,171,869]" pageId="9" pageNumber="10">
Besides greater size, species of
<taxonomicName id="4C4C4D68FFE5D53CFDAA728DFD69F909" authorityName="Owen" authorityYear="1874" box="[546,697,1739,1765]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B938EAF9FFE5D53CFDAA728DFD69F909" box="[546,697,1739,1765]" italics="true" pageId="9" pageNumber="10">Protemnodon</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4C4C4D68FFE5D53CFEBC72ADFE25F8E9" baseAuthorityName="Desmarest" baseAuthorityYear="1804" box="[308,501,1771,1797]" class="Mammalia" family="Macropodidae" genus="Wallabia" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B938EAF9FFE5D53CFEBC72ADFE25F8E9" box="[308,501,1771,1797]" italics="true" pageId="9" pageNumber="10">Wallabia bicolor</emphasis>
</taxonomicName>
in the following ways. Their dentitions are distinguished by: P3 with larger posterolingual basin relative to length of P3; and upper molars with postparacrista and premetacrista that curve or deflect lingually toward the centre of the interloph valley, rather than continuing straight from the cones along the buccal margin of the lophs to merge above the buccal edge of interloph valley. The cranium differs in: a more ventrally projected anterior part of the premaxilla; nasal less convex and less anteriorly projected; masseteric process anteroposteriorly shorter and more ventrally projected; occipital condyles that project posteriorly well beyond the posterior margins of the nuchal crest and occiput; foramen magnum broader relative to the posterior cranium height; a more raised anteroposterior ridge meeting the medial margin of the foramen ovale; and a more laterally projected postglenoid process.
</paragraph>
<paragraph id="8BF336EBFFE5D53CFCD275CDFAE6FC89" blockId="9.[810,1452,171,869]" pageId="9" pageNumber="10">The forelimb differs in its humerus with a more medially projected humeral head, lower (less raised) and less elongate deltoid tuberosity, straighter pectoral crest, and a broader lateral supracondylar ridge; and ulna with olecranon lacking cranial deflection. The manus differs in its deeper, squarer capitatum. The hindlimb differs in its pelvis with broader ilium and more rounded, less projected iliopubic eminence; and femur with larger, more dorsally deflected femoral head and less dorsoventrally compressed distal shaft. The pes is differentiated by: calcaneus with a broader medial talar facet; talus with a more rounded posterior plantar process in medial view; metatarsal IV more robust; metatarsal V with larger proximolateral process relative to metatarsal V length; and broader pedal phalanges relative to length.</paragraph>
</subSubSection>
<subSubSection id="C3566560FFE5D53CFCA277CDFB25FBE9" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BF336EBFFE5D53CFCA277CDFC6BFC49" blockId="9.[810,1451,907,1029]" box="[810,955,907,933]" pageId="9" pageNumber="10">
<emphasis id="B938EAF9FFE5D53CFCA277CDFC6BFC49" bold="true" box="[810,955,907,933]" pageId="9" pageNumber="10">Etymology:</emphasis>
</paragraph>
<paragraph id="8BF336EBFFE5D53CFCA277EDFB25FBE9" blockId="9.[810,1451,907,1029]" pageId="9" pageNumber="10">
Greek, neutral gender;
<emphasis id="B938EAF9FFE5D53CFBA377EAFB81FC29" box="[1067,1105,940,965]" italics="true" pageId="9" pageNumber="10">pro</emphasis>
(before),
<emphasis id="B938EAF9FFE5D53CFB3777EAFAD3FC29" box="[1215,1283,940,965]" italics="true" pageId="9" pageNumber="10">temno</emphasis>
(to cut),
<emphasis id="B938EAF9FFE5D53CFAE077EDFA7BFC29" box="[1384,1451,939,965]" italics="true" pageId="9" pageNumber="10">odous</emphasis>
(tooth). …in reference to the sectorial form of the anterior molar or premolar (
<bibRefCitation id="EFDD4B1AFFE5D53CFB8477ADFB43FBE9" author="Owen, R." box="[1036,1171,1003,1029]" pageId="9" pageNumber="10" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen 1874</bibRefCitation>
, p. 274).
</paragraph>
</subSubSection>
<subSubSection id="C3566560FFE5D53CFCA2706DFAF6FA09" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BF336EBFFE5D53CFCA2706DFC75FBA9" blockId="9.[810,1452,1067,1509]" box="[810,933,1067,1093]" pageId="9" pageNumber="10">
<emphasis id="B938EAF9FFE5D53CFCA2706DFC75FBA9" bold="true" box="[810,933,1067,1093]" pageId="9" pageNumber="10">Remarks:</emphasis>
</paragraph>
<paragraph id="8BF336EBFFE5D53CFCA2700DFAF6FA09" blockId="9.[810,1452,1067,1509]" pageId="9" pageNumber="10">
We contend that
<taxonomicName id="4C4C4D68FFE5D53CFC67700DFB17FB89" authorityName=", Owen" authorityYear="1874" box="[1007,1223,1099,1125]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="anak">
<emphasis id="B938EAF9FFE5D53CFC67700DFB17FB89" box="[1007,1223,1099,1125]" italics="true" pageId="9" pageNumber="10">Protemnodon anak</emphasis>
</taxonomicName>
was designated the
<typeStatus id="54F78849FFE5D53CFCA2702DFC8AFB69" box="[810,858,1131,1157]" pageId="9" pageNumber="10">type</typeStatus>
species for the genus by
<bibRefCitation id="EFDD4B1AFFE5D53CFB15702DFAE9FB69" author="Owen, R." box="[1181,1337,1131,1157]" pageId="9" pageNumber="10" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen (1874)</bibRefCitation>
when he described
<taxonomicName id="4C4C4D68FFE5D53CFC1670CDFACCFB49" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" box="[926,1308,1163,1189]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mimas">
<emphasis id="B938EAF9FFE5D53CFC1670CDFB58FB49" box="[926,1160,1163,1189]" italics="true" pageId="9" pageNumber="10">Protemnodon mimas</emphasis>
<bibRefCitation id="EFDD4B1AFFE5D53CFB1970CDFACCFB49" author="Owen, R." box="[1169,1308,1163,1189]" pageId="9" pageNumber="10" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen, 1874</bibRefCitation>
</taxonomicName>
on the page following the description of
<taxonomicName id="4C4C4D68FFE5D53CFB0270EAFB34FB29" authorityName=", Owen" authorityYear="1874" box="[1162,1252,1195,1221]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="anak">
<emphasis id="B938EAF9FFE5D53CFB0270EAFB34FB29" box="[1162,1252,1195,1221]" italics="true" pageId="9" pageNumber="10">P. anak</emphasis>
</taxonomicName>
: “The postbasal ridge though narrow is definite; the parabasal ridge is proportionately as well developed as in
<taxonomicName id="4C4C4D68FFE5D53CFA9C70ADFCB4FAC9" authorityName=", Owen" authorityYear="1874" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="anak">
<emphasis id="B938EAF9FFE5D53CFA9C70ADFCB4FAC9" italics="true" pageId="9" pageNumber="10">Protemnodon Anak</emphasis>
</taxonomicName>
; its link also, and that of the two chief lobes, are more distinct than in the
<typeStatus id="54F78849FFE5D53CFBDB716DFB53FAA9" box="[1107,1155,1323,1349]" pageId="9" pageNumber="10">type</typeStatus>
species” (
<bibRefCitation id="EFDD4B1AFFE5D53CFB76716DFA58FAA9" author="Owen, R." box="[1278,1416,1323,1349]" pageId="9" pageNumber="10" pagination="245 - 288" refId="ref152824" refString="Owen, R. (1874) On the fossil mammals of Australia, Part VIII. Family Macropodidae: genera Macropus, Osphranter, Phascolagus, Sthenurus and Protemnodon. Philosophical Transactions of the Royal Society, 164, 245 - 288. https: // doi. org / 10.1098 / rstl. 1874.0008" type="journal article" year="1874">Owen 1874</bibRefCitation>
, p. 278). Here, Owen clearly referred to
<taxonomicName id="4C4C4D68FFE5D53CFB5C710AFAF7FA89" authorityName=", Owen" authorityYear="1874" box="[1236,1319,1355,1381]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="anak">
<emphasis id="B938EAF9FFE5D53CFB5C710AFAF7FA89" box="[1236,1319,1355,1381]" italics="true" pageId="9" pageNumber="10">P. anak</emphasis>
</taxonomicName>
as the
<typeStatus id="54F78849FFE5D53CFAF3710DFA7BFA89" box="[1403,1451,1355,1381]" pageId="9" pageNumber="10">type</typeStatus>
species, demonstrating his intent, not otherwise stated, that
<taxonomicName id="4C4C4D68FFE5D53CFCD371CAFC7CFA49" authorityName=", Owen" authorityYear="1874" box="[859,940,1419,1445]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="anak">
<emphasis id="B938EAF9FFE5D53CFCD371CAFC7CFA49" box="[859,940,1419,1445]" italics="true" pageId="9" pageNumber="10">P. anak</emphasis>
</taxonomicName>
be given this status,
<emphasis id="B938EAF9FFE5D53CFB1071CAFB31FA49" box="[1176,1249,1420,1445]" italics="true" pageId="9" pageNumber="10">contra</emphasis>
<bibRefCitation id="EFDD4B1AFFE5D53CFB6171CDFCA6FA2A" author="Mahoney, J. A. &amp; Ride, W. D. L." pageId="9" pageNumber="10" pagination="1 - 250" refId="ref151155" refString="Mahoney, J. A. &amp; Ride, W. D. L. (1975) Index to the genera and species of fossil Mammalia described from Australia and New Guinea between 1838 and 1968 (including citations of type species and primary type specimens). Special Publication of the Western Australian Museum, 6, 1 - 250." type="journal article" year="1975">Mahoney &amp; Ride (1975)</bibRefCitation>
, who stated that
<taxonomicName id="4C4C4D68FFE5D53CFBCB71EAFB67FA29" authority="is" authorityName="IS" box="[1091,1207,1451,1477]" class="Mammalia" family="Macropodidae" genus="Protemnodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="9" pageNumber="240" phylum="Chordata" rank="species" species="anak">
<emphasis id="B938EAF9FFE5D53CFBCB71EAFB49FA29" box="[1091,1177,1451,1477]" italics="true" pageId="9" pageNumber="10">P. anak</emphasis>
is
</taxonomicName>
the
<typeStatus id="54F78849FFE5D53CFB6771EDFACFFA29" box="[1263,1311,1451,1477]" pageId="9" pageNumber="10">type</typeStatus>
species by subsequent designation... by
<bibRefCitation id="EFDD4B1AFFE5D53CFBE5718DFACDFA09" author="Simpson, G. G." box="[1133,1309,1483,1509]" pageId="9" pageNumber="10" refId="ref155106" refString="Simpson, G. G. (1930) Post-Mesozoic Marsupialia. Fossilium Catalogus. 1: Animalia Part 47. W. Junk, Berlin, 87 pp." type="book" year="1930">Simpson, 1930</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>