269 lines
38 KiB
XML
269 lines
38 KiB
XML
<document id="49D82FBAE6E941188F855C734556C1E5" ID-DOI="10.11646/zootaxa.5519.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="13915579" ID-ZooBank="C12023C8-FF3D-4058-A590-E1E2DC38B4FA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728584281405" checkinUser="plazi" docAuthor="Huang, Tian, Janák, Jiří & Zhou, Hong-Zhang" docDate="2024" docId="03E587A3FFF8FFD7B9C08FBAFDF5FFC5" docLanguage="en" docName="zootaxa.5519.1.5.pdf" docOrigin="Zootaxa 5519 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5519.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diochus papuanus Cameron 1939" docType="treatment" docVersion="1" lastPageNumber="117" masterDocId="FFDCFFDBFFF4FFD9B9578E0CFFB2FC0C" masterDocTitle="Taxonomy of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) of Papua New Guinea, Vanuatu and Solomon Islands" masterLastPageNumber="118" masterPageNumber="103" pageNumber="115" updateTime="1728585392433" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E2CBA92FDB1D53415F26898B2B09E22E">Taxonomy of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) of Papua New Guinea, Vanuatu and Solomon Islands</mods:title>
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<mods:namePart id="9E617A414441D31D03134E8F1417C0B5">Huang, Tian</mods:namePart>
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<mods:affiliation id="8AFE59FE97FC0E7D89E9501B4DBD7111">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
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</mods:role>
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<mods:namePart id="93CD7D7C6CC652D2E24C7792CB16BAA9">Janák, Jiří</mods:namePart>
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<mods:affiliation id="F89F8C357BE3832DC35FFEDFC15227FE">CZ- 41762 Rtyně nad Bílinou, č. 4, Czech Republic.</mods:affiliation>
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<mods:namePart id="24690C822069316F18C2408A78EBAAA9">Zhou, Hong-Zhang</mods:namePart>
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<mods:affiliation id="FDCDF99D87E6A626DDF60CDFD67D0662">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
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<mods:title id="484CDE2B94C869AEC93E0BB8BA60B8E5">Zootaxa</mods:title>
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<mods:date id="225B10A46058E502A316761598B9B3AF">2024</mods:date>
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<mods:number id="A25AD20452ADC68DF945ABB189449972">2024-10-07</mods:number>
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<treatment id="03E587A3FFF8FFD7B9C08FBAFDF5FFC5" LSID="urn:lsid:plazi:treatment:03E587A3FFF8FFD7B9C08FBAFDF5FFC5" httpUri="http://treatment.plazi.org/id/03E587A3FFF8FFD7B9C08FBAFDF5FFC5" lastPageId="14" lastPageNumber="117" pageId="12" pageNumber="115">
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<subSubSection id="C356653EFFF8FFD5B9C08FBAFDC2FDDC" box="[151,624,438,465]" pageId="12" pageNumber="115" type="nomenclature">
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<paragraph id="8BF336B5FFF8FFD5B9C08FBAFDC2FDDC" blockId="12.[151,624,438,501]" box="[151,624,438,465]" pageId="12" pageNumber="115">
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<heading id="D0BB81D9FFF8FFD5B9C08FBAFDC2FDDC" bold="true" box="[151,624,438,465]" fontSize="11" level="1" pageId="12" pageNumber="115" reason="1">
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<emphasis id="B938EAA7FFF8FFD5B9C08FBAFDC2FDDC" bold="true" box="[151,624,438,465]" pageId="12" pageNumber="115">
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4.
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<taxonomicName id="4C4C4D36FFF8FFD5B9908FBAFDC2FDDC" ID-CoL="369ZD" authority="Cameron, 1939" authorityName="Cameron" authorityYear="1939" box="[199,624,438,465]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="species" species="papuanus">
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<emphasis id="B938EAA7FFF8FFD5B9908FBAFE19FDDD" bold="true" box="[199,427,438,465]" italics="true" pageId="12" pageNumber="115">Diochus papuanus</emphasis>
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||
<bibRefCitation id="EFDD4B44FFF8FFD5B8E58FBAFDC2FDDC" author="Cameron, M." box="[434,624,438,464]" pageId="12" pageNumber="115" pagination="139 - 153" refId="ref8513" refString="Cameron, M. (1939) New Staphylinidae (Col.) from New Guinea. The Annals and Magazine of Natural History, Series 11, 3 (13), 139 - 153. https: // doi. org / 10.1080 / 03745481.1939.9723583" type="journal article" year="1939">Cameron, 1939</bibRefCitation>
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</taxonomicName>
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||
</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C356653EFFF8FFD5B9C08FD7FC00FE3B" pageId="12" pageNumber="115" type="description">
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<paragraph id="8BF336B5FFF8FFD5B9C08FD7FEF2FDF9" blockId="12.[151,624,438,501]" box="[151,320,474,501]" pageId="12" pageNumber="115">
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(
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||
<figureCitation id="13772A30FFF8FFD5B9C88FD7FF42FDF9" box="[159,240,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[154,1432,744,1846]" captionTargetId="figure-232@2.[151,1435,741,1850]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of four species from Papua New Guinea, Vanuatu and Solomon Islands in dorsal view:A. Diochus elongatus sp. nov.; B. D. chambrieri sp. nov.; C. D. papuanus Cameron, 1939, D. D. loebli sp. nov. (Scale bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915581" httpUri="https://zenodo.org/record/13915581/files/figure.png" pageId="12" pageNumber="115">Fig 1C</figureCitation>
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;
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<figureCitation id="13772A30FFF8FFD5B9AB8FD7FE85FDF9" box="[252,311,474,501]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="12" pageNumber="115">Fig 5</figureCitation>
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)
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||
</paragraph>
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<paragraph id="8BF336B5FFF8FFD5B9C08C13FC00FE3B" blockId="12.[151,946,543,567]" box="[151,946,543,567]" pageId="12" pageNumber="115">
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<materialsCitation id="3B243CE8FFF8FFD5B9C08C13FC1CFE3B" box="[151,942,543,567]" country="Guinea-Bissau" elevation="366" location="New" municipality="Kokoda" pageId="12" pageNumber="115" specimenCount="1">
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<bibRefCitation id="EFDD4B44FFF8FFD5B9C08C13FED5FE3B" author="Cameron, M." box="[151,359,543,567]" pageId="12" pageNumber="115" pagination="139 - 153" refId="ref8513" refString="Cameron, M. (1939) New Staphylinidae (Col.) from New Guinea. The Annals and Magazine of Natural History, Series 11, 3 (13), 139 - 153. https: // doi. org / 10.1080 / 03745481.1939.9723583" type="journal article" year="1939">Cameron, 1939: 145</bibRefCitation>
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(
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||
<typeStatus id="54F78817FFF8FFD5B8238C13FE1AFE3B" box="[372,424,543,567]" pageId="12" pageNumber="115">Type</typeStatus>
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locality:
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||
<location id="8E93606EFFF8FFD5BB5F8C2CFD8AFE3B" LSID="urn:lsid:plazi:treatment:03E587A3FFF8FFD7B9C08FBAFDF5FFC5:8E93606EFFF8FFD5BB5F8C2CFD8AFE3B" box="[520,568,544,567]" country="Guinea-Bissau" municipality="Kokoda" name="New" pageId="12" pageNumber="115">New</location>
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<collectingCountry id="F35B7625FFF8FFD5BB698C13FD39FE3B" box="[574,651,543,567]" name="Guinea-Bissau" pageId="12" pageNumber="115">Guinea</collectingCountry>
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||
:
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<collectingMunicipality id="6B97ACCFFFF8FFD5BBC38C13FD55FE3B" box="[660,743,543,567]" pageId="12" pageNumber="115">Kokoda</collectingMunicipality>
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||
, altitude
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||
<quantity id="4CB49B50FFF8FFD5BA148C13FC15FE3B" box="[835,935,543,567]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="12" pageNumber="115" unit="ft" value="1200.0">
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||
<elevation id="0061D186FFF8FFD5BA148C13FC15FE3B" box="[835,935,543,567]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="12" pageNumber="115" unit="ft" value="1200.0">1200 feet</elevation>
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||
</quantity>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C356653EFFF8FFD5B9C08C69FBFAFF5B" pageId="12" pageNumber="115" type="materials_examined">
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||
<paragraph id="8BF336B5FFF8FFD5B9C08C69FC7EFECB" blockId="12.[151,1437,613,2008]" pageId="12" pageNumber="115">
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||
<emphasis id="B938EAA7FFF8FFD5B9C08C69FE0DFE73" bold="true" box="[151,447,613,639]" pageId="12" pageNumber="115">Type material examined.</emphasis>
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<materialsCitation id="3B243CE8FFF8FFD5B8928C69FC7EFECB" collectingDate="1933-09" collectionCode="TYPE, NHMUK" collectorName="L. E. Cheesman." country="Papua New Guinea" elevation="366" location="Type" pageId="12" pageNumber="115" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="B938EAA7FFF8FFD5B8928C69FCD5FE73" bold="true" box="[453,871,613,639]" pageId="12" pageNumber="115">
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||
<collectingCountry id="F35B7625FFF8FFD5B8928C69FD57FE73" box="[453,741,613,639]" name="Papua New Guinea" pageId="12" pageNumber="115">PAPUA NEW GUINEA</collectingCountry>
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||
:
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||
<typeStatus id="54F78817FFF8FFD5BBA68C69FCD0FE73" box="[753,866,613,639]" pageId="12" pageNumber="115" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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male, “
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<location id="8E93606EFFF8FFD5BAEB8C69FC4AFE73" LSID="urn:lsid:plazi:treatment:03E587A3FFF8FFD7B9C08FBAFDF5FFC5:8E93606EFFF8FFD5BAEB8C69FC4AFE73" box="[956,1016,613,639]" country="Papua New Guinea" name="Type" pageId="12" pageNumber="115">Type</location>
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||
” (printed rounded white, red boarded label) // “PAPUA: Kokoda. /
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||
<quantity id="4CB49B50FFF8FFD5B88A8C85FD87FEAF" box="[477,565,649,675]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="12" pageNumber="115" unit="ft" value="1200.0">
|
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<elevation id="0061D186FFF8FFD5B88A8C85FD87FEAF" box="[477,565,649,675]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="12" pageNumber="115" unit="ft" value="1200.0">1,200ft.</elevation>
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</quantity>
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||
<date id="FFF21075FFF8FFD5BB6C8C85FD29FEAF" box="[571,667,649,675]" pageId="12" pageNumber="115" value="1933-09">
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||
<collectingDate id="EFB6E99DFFF8FFD5BB6C8C85FD29FEAF" box="[571,667,649,675]" pageId="12" pageNumber="115" value="1933-09">IX.1933</collectingDate>
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||
</date>
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||
. /
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||
<collectorName id="26B95363FFF8FFD5BBE38C85FCD2FEAF" box="[692,864,649,675]" pageId="12" pageNumber="115">L.E.Cheesman.</collectorName>
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||
/ B.M.1933-427.” (printed) // “
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||
<taxonomicName id="4C4C4D36FFF8FFD5BDEA8C85FAAFFEAF" box="[1213,1309,649,675]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="genus">Diochus</taxonomicName>
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||
/ papuanus /
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<collectionCode id="ED5DAE70FFF8FFD5B9F28CA2FF59FEC4" box="[165,235,686,712]" pageId="12" pageNumber="115">TYPE</collectionCode>
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||
Cam” (Cameronʼs handwriting, black and red inked) (
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||
<collectionCode id="ED5DAE70FFF8FFD5BA188CA1FC72FECB" box="[847,960,685,711]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="12" pageNumber="115" type="Museum">NHMUK</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFF8FFD5B9908CDDFBFAFF5B" blockId="12.[151,1437,613,2008]" pageId="12" pageNumber="115">
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||
<emphasis id="B938EAA7FFF8FFD5B9908CDDFEB4FEE7" bold="true" box="[199,262,721,747]" pageId="12" pageNumber="115">Note.</emphasis>
|
||
<materialsCitation id="3B243CE8FFF8FFD5B8458CDEFBF6FF5B" collectionCode="NHMUK" pageId="12" pageNumber="115" specimenCount="1" typeStatus="holotype">
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||
We examined the unique male type from the Cameronʼs collection (
|
||
<collectionCode id="ED5DAE70FFF8FFD5BD6B8CDDFB1FFEE7" box="[1084,1197,721,747]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="12" pageNumber="115" type="Museum">NHMUK</collectionCode>
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||
). The specimen was remounted, glued on a new card plate, aedeagus and last abdominal segments were embedded in Euparal on a plastic plate placed at the same pin as the specimens. As
|
||
<bibRefCitation id="EFDD4B44FFF8FFD5BB9B8D15FC37FF3F" author="Cameron, M." box="[716,901,793,819]" pageId="12" pageNumber="115" pagination="139 - 153" refId="ref8513" refString="Cameron, M. (1939) New Staphylinidae (Col.) from New Guinea. The Annals and Magazine of Natural History, Series 11, 3 (13), 139 - 153. https: // doi. org / 10.1080 / 03745481.1939.9723583" type="journal article" year="1939">Cameron (1939)</bibRefCitation>
|
||
mentioned in the description “Unique” and the specimen was labeled as “Type” there is no doubt that it is a
|
||
<typeStatus id="54F78817FFF8FFD5BA688D31FC13FF5B" box="[831,929,829,855]" pageId="12" pageNumber="115" type="holotype">holotype</typeStatus>
|
||
of the species
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||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356653EFFF8FFD7B9908D6DFE10FF51" lastPageId="14" lastPageNumber="117" pageId="12" pageNumber="115" type="description">
|
||
<paragraph id="8BF336B5FFF8FFD5B9908D6DFBA0FFAC" blockId="12.[151,1437,613,2008]" pageId="12" pageNumber="115">
|
||
<emphasis id="B938EAA7FFF8FFD5B9908D6DFECCFF77" bold="true" box="[199,382,865,891]" pageId="12" pageNumber="115">Measurements.</emphasis>
|
||
BL = 4.0 mm, FL =
|
||
<quantity id="4CB49B50FFF8FFD5BB338D6DFD0EFF70" box="[612,700,865,892]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="115" unit="mm" value="1.7">1.7 mm</quantity>
|
||
, HL =
|
||
<quantity id="4CB49B50FFF8FFD5BA5E8D6DFCDDFF70" box="[777,879,865,892]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="12" pageNumber="115" unit="mm" value="0.51">0.51 mm</quantity>
|
||
, HW =
|
||
<quantity id="4CB49B50FFF8FFD5BA928D6DFB99FF70" box="[965,1067,865,892]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="12" pageNumber="115" unit="mm" value="0.44">0.44 mm</quantity>
|
||
, EyL =
|
||
<quantity id="4CB49B50FFF8FFD5BDD48D6DFB5BFF70" box="[1155,1257,865,892]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="115" unit="mm" value="0.14">0.14 mm</quantity>
|
||
, TL =
|
||
<quantity id="4CB49B50FFF8FFD5BC658D6DFA2AFF70" box="[1330,1432,865,892]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="115" unit="mm" value="0.3">0.30 mm</quantity>
|
||
, PL =
|
||
<quantity id="4CB49B50FFF8FFD5B9828D89FE89FFAC" box="[213,315,901,928]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="12" pageNumber="115" unit="mm" value="0.71">0.71 mm</quantity>
|
||
, PW =
|
||
<quantity id="4CB49B50FFF8FFD5B8DA8D89FE41FFAC" box="[397,499,901,928]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="12" pageNumber="115" unit="mm" value="0.59">0.59 mm</quantity>
|
||
, EL =
|
||
<quantity id="4CB49B50FFF8FFD5BB698D89FD16FFAC" box="[574,676,901,928]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="12" pageNumber="115" unit="mm" value="0.62">0.62 mm</quantity>
|
||
, EW =
|
||
<quantity id="4CB49B50FFF8FFD5BBAF8D89FCECFFAC" box="[760,862,901,928]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="12" pageNumber="115" unit="mm" value="0.68">0.68 mm</quantity>
|
||
, SL =
|
||
<quantity id="4CB49B50FFF8FFD5BAF08D89FBBCFFAC" box="[935,1038,901,928]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="115" unit="mm" value="0.42">0.42 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFF8FFD5B9908DA5FCC2F823" blockId="12.[151,1437,613,2008]" pageId="12" pageNumber="115">
|
||
<emphasis id="B938EAA7FFF8FFD5B9908DA5FEC7FFCF" bold="true" box="[199,373,937,963]" pageId="12" pageNumber="115">Redescription.</emphasis>
|
||
Body (
|
||
<figureCitation id="13772A30FFF8FFD5B89B8DA5FDAFFFCF" box="[460,541,937,963]" captionStart="FIGURE 1" captionStartId="2.[151,250,1874,1898]" captionTargetBox="[154,1432,744,1846]" captionTargetId="figure-232@2.[151,1435,741,1850]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of four species from Papua New Guinea, Vanuatu and Solomon Islands in dorsal view:A. Diochus elongatus sp. nov.; B. D. chambrieri sp. nov.; C. D. papuanus Cameron, 1939, D. D. loebli sp. nov. (Scale bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915581" httpUri="https://zenodo.org/record/13915581/files/figure.png" pageId="12" pageNumber="115">Fig 1C</figureCitation>
|
||
) long, narrowed anteriorly and posteriorly, medium-sized. Body entirely dark brown to black, posterior margins of abdominal segments and abdominal segments VIII−X lighter, dark reddish brown. Legs yellowish brown, tarsi distinctly lighter. Antennomeres 1−4 light brown, 5−11 yellowish, apex of each antennomere lighter. Maxillary and labial palpi yellowish brown.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFF8FFD5B9908A35FF62F9B7" blockId="12.[151,1437,613,2008]" pageId="12" pageNumber="115">
|
||
<emphasis id="B938EAA7FFF8FFD5B9908A35FEB5F85F" bold="true" box="[199,263,1081,1107]" pageId="12" pageNumber="115">Head</emphasis>
|
||
(
|
||
<figureCitation id="13772A30FFF8FFD5B8408A35FEDFF858" box="[279,365,1081,1108]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="12" pageNumber="115">Fig 5A</figureCitation>
|
||
). Relativelly short and wide, 1.17 times as long as wide. Tempora roundly widened posteriad, posterior angles widely rounded. Dorsal surface shiny, bearing very fine transverse microstriae and mesh, also with coarse and very sparse punctures. Head with 5−7 irregular punctures scattered around each eye and numerous smaller punctures on deflexed portion of tempus and near posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, with temporal puncture and occipital puncture near basal margin; disc surface bearing two additional paired punctures in middle area. Frontal furrows and anteocular furrows absent. Eye large-sized, slightly protruding laterad, longitudinal diameter about half the length of tempora (eye: tempora = 0.45). Epistoma not flattened, abruptly oblique, anterior margin rounded. Ventral surface with scattered punctures, interspaces between them about 4 puncture diameters. Gular sutures deep, confluent at basal 1/4, then extended to base.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFF8FFD5B9908BC9FE1DFA47" blockId="12.[151,1437,613,2008]" pageId="12" pageNumber="115">
|
||
<emphasis id="B938EAA7FFF8FFD5B9908BC9FE88F9D3" bold="true" box="[199,314,1477,1503]" pageId="12" pageNumber="115">Antennae</emphasis>
|
||
(
|
||
<figureCitation id="13772A30FFF8FFD5B81D8BC9FE2EF9EC" box="[330,412,1477,1504]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="12" pageNumber="115">Fig 5B</figureCitation>
|
||
). Scape rod-shaped, slightly thickened apically, shorter than two subsequent antennomeres combined; antennomere 2 slightly elongate, antennomere 3 slightly longer than 2; antennomere 4 almost twice the width, antennomere 4 to 10 gradually shortened, antennomere 4 observably longer than 10; antennomere 11 distinctly longer than 10.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFF8FFD5B9908859FE7DFAB4" blockId="12.[151,1437,613,2008]" pageId="12" pageNumber="115">
|
||
<emphasis id="B938EAA7FFF8FFD5B9908859FEEFFA63" bold="true" box="[199,349,1621,1647]" pageId="12" pageNumber="115">Mouthparts.</emphasis>
|
||
Labrum oval-shaped, lateral margin rounded, widest at basal 1/2; width of anterior margin is about half the width at the widest point. Mandibles falciform, left one with three small teeth on inner margin, whereas right one with two.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFF8FFD5B99088CDFD82FB2F" blockId="12.[151,1437,613,2008]" pageId="12" pageNumber="115">
|
||
<emphasis id="B938EAA7FFF8FFD5B99088CDFEB9FAD7" bold="true" box="[199,267,1729,1755]" pageId="12" pageNumber="115">Neck.</emphasis>
|
||
Cylindrical, shiny, only bearing shallow microsculpture, shorter than 1/3 of head width. First half of dorsal surface depressed, with distinct groove. Ventral surface with gular sutures forming obvious triangular region, with deep transverse median furrow.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFF8FFD4B9908921FD9BFCD9" blockId="12.[151,1437,613,2008]" lastBlockId="13.[151,1436,151,501]" lastPageId="13" lastPageNumber="116" pageId="12" pageNumber="115">
|
||
<emphasis id="B938EAA7FFF8FFD5B9908921FEF3FB4B" bold="true" box="[199,321,1837,1863]" pageId="12" pageNumber="115">Prothorax</emphasis>
|
||
(
|
||
<figureCitation id="13772A30FFF8FFD5B8078921FE16FB44" box="[336,420,1837,1864]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="12" pageNumber="115">Fig 5A</figureCitation>
|
||
). Pronotum elongate (PL to PW ratio 1.21), distinctly longer and wider than head. Anterior region near anterior angles deflexed, lateral margins widened posteriad to basal 1/3 (widest at 1/3), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy, extensively with micropunctures, but without any microsculpture. Each side of longitudinal midline with one row of punctures composing of 3 large punctures and 2 additional punctures outside of puncture row, several punctures scatted near each margin (including anterior, lateral and posterior margins). Prosternum bearing observable transverse ridge between basisternum and furcasternum, angularly backwards.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFF9FFD4B9908ED3FB7EFD4D" blockId="13.[151,1436,151,501]" pageId="13" pageNumber="116">
|
||
<emphasis id="B938EAA7FFF9FFD4B9908ED3FEE5FCF5" bold="true" box="[199,343,223,249]" pageId="13" pageNumber="116">Pterothorax</emphasis>
|
||
(
|
||
<figureCitation id="13772A30FFF9FFD4B83D8ED3FE73FCF5" box="[362,449,222,249]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="13" pageNumber="116">Fig 5A</figureCitation>
|
||
). Mesoscutellum triangular and small, surface shiny, bearing distinct microstriae, but without any punctures. Transverse ridge on mesoventrite wave shaped; paired oblique furrows short but deep; carina limiting mesocoxal cavity slightly arcuate. Discrimen region on metaventrite normally elevated.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFF9FFD4B9908F47FDE8FDF9" blockId="13.[151,1436,151,501]" pageId="13" pageNumber="116">
|
||
<emphasis id="B938EAA7FFF9FFD4B9908F47FEA1FD69" bold="true" box="[199,275,331,357]" pageId="13" pageNumber="116">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="13772A30FFF9FFD4B8738F47FECBFD69" box="[292,377,330,357]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="13" pageNumber="116">Fig 5A</figureCitation>
|
||
). Slightly transverse (EL to EW ratio 0.95), only a sixth longer and but markedly wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, posterior margin of each elytron obliquely truncate. Dorsal surface shiny, flattened, without microsculpture, finely coriaceous; each elytron with row of 5 punctures along suture, row of 6 punctures in median, two isolated punctures between sutural and median row, also with 6−7 punctures on deflexed portion.
|
||
</paragraph>
|
||
<caption id="DF33663DFFF9FFD4B9C0895EFAD5FB83" ID-DOI="http://doi.org/10.5281/zenodo.13915598" ID-Zenodo-Dep="13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="13" pageNumber="116" startId="13.[151,250,1874,1898]" targetBox="[161,1426,535,1841]" targetPageId="13" targetType="figure">
|
||
<paragraph id="8BF336B5FFF9FFD4B9C0895EFAD5FB83" blockId="13.[151,1436,1874,1935]" pageId="13" pageNumber="116">
|
||
<emphasis id="B938EAA7FFF9FFD4B9C0895EFEA0FB67" bold="true" box="[151,274,1874,1899]" pageId="13" pageNumber="116">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C4C4D36FFF9FFD4B840895FFDC4FB67" authority="Cameron, 1939" authorityName="Cameron" authorityYear="1939" box="[279,630,1875,1899]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="116" phylum="Arthropoda" rank="species" species="papuanus">
|
||
<emphasis id="B938EAA7FFF9FFD4B840895FFE61FB66" box="[279,467,1875,1899]" italics="true" pageId="13" pageNumber="116">Diochus papuanus</emphasis>
|
||
<bibRefCitation id="EFDD4B44FFF9FFD4B88F895FFDC4FB67" author="Cameron, M." box="[472,630,1875,1899]" pageId="13" pageNumber="116" pagination="139 - 153" refId="ref8513" refString="Cameron, M. (1939) New Staphylinidae (Col.) from New Guinea. The Annals and Magazine of Natural History, Series 11, 3 (13), 139 - 153. https: // doi. org / 10.1080 / 03745481.1939.9723583" type="journal article" year="1939">Cameron, 1939</bibRefCitation>
|
||
</taxonomicName>
|
||
: A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336B5FFFAFFD7B9908E9BFA2EFCD9" blockId="14.[151,1437,151,969]" pageId="14" pageNumber="117">
|
||
<emphasis id="B938EAA7FFFAFFD7B9908E9BFF4DFCBD" bold="true" box="[199,255,151,177]" pageId="14" pageNumber="117">Legs</emphasis>
|
||
. First four segments of protarsi heart-shaped, moderately dilated, and those of meso- and metatarsi slender. Last segment of protarsi almost equal to 2−4 together, that of meso-, meta- slightly longer than length of each 2−3.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFFAFFD7B9908ED2FCACFDA1" blockId="14.[151,1437,151,969]" pageId="14" pageNumber="117">
|
||
<emphasis id="B938EAA7FFFAFFD7B9908ED2FE8EFCF4" bold="true" box="[199,316,222,248]" pageId="14" pageNumber="117">Abdomen</emphasis>
|
||
. Broadest at segment V. Tergites III–VII covered with very fine shallow transverse microstriae and mesh; each tergite with dense pubescence, also with row of darker and coarser setae at posterior margin of tergites III–VI. Tergites III–VII with basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; all abdominal tergites with fine and moderately dense puncturation, distance between punctures larger than diameter of puncture. Posterior margin of tergite VII with distinct palisade fringe. All abdominal sternites shiny, with microstriae and punctures as those on tergites.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFFAFFD7B9908FBBFD08FF35" blockId="14.[151,1437,151,969]" pageId="14" pageNumber="117">
|
||
<emphasis id="B938EAA7FFFAFFD7B9908FBBFEB6FDDD" bold="true" box="[199,260,439,465]" pageId="14" pageNumber="117">Male</emphasis>
|
||
(
|
||
<figureCitation id="13772A30FFFAFFD7B84D8FBBFEC6FDDD" box="[282,372,438,465]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="14" pageNumber="117">Fig 5A</figureCitation>
|
||
). Head with small shiny, slightly elongate, markedly protruding elevation between eyes. Posterior margin of sternite VIII markedly roundly emarginated medially, both sides apicolaterally obliquely truncate, apical margin with very short setae (
|
||
<figureCitation id="13772A30FFFAFFD7BBF18FF3FD49FE15" box="[678,763,510,537]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="14" pageNumber="117">Fig 5C</figureCitation>
|
||
). Posterior margin of tergite VIII arcuate (
|
||
<figureCitation id="13772A30FFFAFFD7BDB58FF3FA8AFE15" box="[1250,1336,510,537]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="14" pageNumber="117">Fig 5D</figureCitation>
|
||
). Tergite IX (
|
||
<figureCitation id="13772A30FFFAFFD7B9908C2FFEA4FE31" box="[199,278,546,573]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="14" pageNumber="117">Fig 5F</figureCitation>
|
||
) symmetrical, connected mediobasally, narrowed apically. Sternite IX (
|
||
<figureCitation id="13772A30FFFAFFD7BD058C2FFB11FE31" box="[1106,1187,546,573]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="14" pageNumber="117">Fig 5E</figureCitation>
|
||
) almost symmetrical, widest near basal 1/3; basal margin straight, apical margin normally emarginated. Tergite X (
|
||
<figureCitation id="13772A30FFFAFFD7BDF98C4BFB48FE6D" box="[1198,1274,582,609]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="14" pageNumber="117">Fig 5F</figureCitation>
|
||
) symmetrical, triangular. Aedeagus (
|
||
<figureCitation id="13772A30FFFAFFD7B8C38C67FE5BFE89" box="[404,489,618,645]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IX–X; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, B–G = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="14" pageNumber="117">Fig 5G</figureCitation>
|
||
) symmetrical, medium-sized, ca.
|
||
<quantity id="4CB49B50FFFAFFD7BA248C67FC69FE89" box="[883,987,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="14" pageNumber="117" unit="mm" value="0.49">0.49 mm</quantity>
|
||
long, normally sclerotized with apical margin triangularly emarginate. Parameres symmetrical, thick and relatively short, about 1/3 length of median lobe. Internal structures sclerotized, two paired strips shaped sclerites located on both sides, parallel, lateral one slender, in brown color, the other broader, in brown color; 3 sclerites in middle: paired rectangular sclerites in brown color, basally with one Y-shaped sclerite in brown color; proximally with paired of brush like structures consisting of long fibers. Sperm pump coiled, thinned toward apex.
|
||
</paragraph>
|
||
<paragraph id="8BF336B5FFFAFFD7B9908D4FFE10FF51" blockId="14.[151,1437,151,969]" box="[199,418,835,861]" pageId="14" pageNumber="117">
|
||
<emphasis id="B938EAA7FFFAFFD7B9908D4FFEACFF51" bold="true" box="[199,286,835,861]" pageId="14" pageNumber="117">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356653EFFFAFFD7B9908D6AFC15FFA9" pageId="14" pageNumber="117" type="diagnosis">
|
||
<paragraph id="8BF336B5FFFAFFD7B9908D6AFC15FFA9" blockId="14.[151,1437,151,969]" pageId="14" pageNumber="117">
|
||
<emphasis id="B938EAA7FFFAFFD7B9908D6AFEF0FF8C" bold="true" box="[199,322,870,896]" pageId="14" pageNumber="117">Diagnosis.</emphasis>
|
||
The species can be easily separated from other species by its short elytra, which are a sixth shorter than the thorax and by deeply roundly emarginated male sternite VIII.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356653EFFFAFFD7B9908DA2FDF5FFC5" box="[199,583,942,969]" pageId="14" pageNumber="117" type="distribution">
|
||
<paragraph id="8BF336B5FFFAFFD7B9908DA2FDF5FFC5" blockId="14.[151,1437,151,969]" box="[199,583,942,969]" pageId="14" pageNumber="117">
|
||
<emphasis id="B938EAA7FFFAFFD7B9908DA2FED3FFC4" bold="true" box="[199,353,942,968]" pageId="14" pageNumber="117">Distribution.</emphasis>
|
||
<collectingCountry id="F35B7625FFFAFFD7B8308DA3FDF6FFC5" box="[359,580,943,969]" name="Papua New Guinea" pageId="14" pageNumber="117">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |