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<document id="D83A8383A22279008AA7FCBB17E28396" ID-DOI="10.5852/ejt.2020.665" ID-GBIF-Dataset="06748f67-18d9-4c3e-a14c-77de1664d81b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3899047" ID-ZooBank="836C9171-0849-4F4D-BC8D-90C2D9E8B9D1" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1592413559169" checkinUser="valdenar" docAuthor="Fernandes, Maurício Romulo &amp; Pimenta, Alexandre Dias" docDate="2020" docId="03E58799FFB7AF79FE07FBF1FD6DFA0E" docLanguage="en" docName="ejt-665_ferrnandes_pimenta.pdf.imf" docOrigin="European Journal of Taxonomy 665 (665)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Isotriphora tricingulata Rolan &amp; Fernandez-Garces 2015" docType="treatment" docVersion="12" lastPageNumber="14" masterDocId="FFDCFFE1FFBCAF74FFC5FFE5FF9CFF92" masterDocTitle="Unraveling one of the ‘ Big Five’: update of the taxonomy of Triphoridae (Gastropoda, Triphoroidea) from Brazil" masterLastPageNumber="170" masterPageNumber="1" pageNumber="12" updateTime="1733402249213" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5A1580352C9E1F9A2B0AB3865418572F">Unraveling one of the ‘ Big Five’: update of the taxonomy of Triphoridae (Gastropoda, Triphoroidea) from Brazil</mods:title>
</mods:titleInfo>
<mods:name id="87477C721CF69946CF9E4001F5BD4D10" type="personal">
<mods:role id="E14B158D3CDCD262A62EB4DB2617102F">
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<mods:namePart id="422AD2056621AB7C7ED7EA848B14C276">Fernandes, Maurício Romulo</mods:namePart>
<mods:nameIdentifier id="1DE9364A2BB2CC132504F32E30224E6A" type="LSID">3B8B864F-3300-45B2-8D1F-61F282F83CDE</mods:nameIdentifier>
<mods:affiliation id="C29669283BFC6A89B0C7CB35B9A7D16B">Departamento de Zoologia, Instituto de Biociências, Universidade Federal do Estado do Rio de Janeiro, Av. Pasteur, 458, Urca, 22290 - 240, Rio de Janeiro, Brazil. &amp; Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20940 - 040, Rio de Janeiro, Brazil.</mods:affiliation>
<mods:nameIdentifier id="E90BF47D3FF1E67191D3322521D780BC" type="email">mauriciofernandes14@hotmail.com</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="E3EEE4BFD639353B73CF5BBD7B6970A4">Pimenta, Alexandre Dias</mods:namePart>
<mods:nameIdentifier id="C71B4BFF62C0C872581840C3C63F48FF" type="LSID">2A42AE1F-6AF8-4AB3-975C-D3CE4D678F22</mods:nameIdentifier>
<mods:affiliation id="0F9F3094108DC9E0929425891DEC6958">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20940 - 040, Rio de Janeiro, Brazil.</mods:affiliation>
<mods:nameIdentifier id="49D7D83F980681B60EF1996B1AC038C0" type="email">alexpim@mn.ufrj.br</mods:nameIdentifier>
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<mods:title id="78E423A28A65518CF5D5AED9F1AAFE0B">European Journal of Taxonomy</mods:title>
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<mods:part id="DA501ABFF048050EB728B6AF29392A9E">
<mods:date id="F68F1F623DA649179033C3130CFAA678">2020</mods:date>
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<mods:number id="A846E00A7C5BBE7CAA5D9BC861F6E0EF">2020-06-12</mods:number>
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<mods:number id="F79EE283FFECCF888B66D3C4511B7BAB">665</mods:number>
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<treatment id="03E58799FFB7AF79FE07FBF1FD6DFA0E" ID-GBIF-Taxon="164612994" LSID="urn:lsid:plazi:treatment:03E58799FFB7AF79FE07FBF1FD6DFA0E" httpUri="http://treatment.plazi.org/id/03E58799FFB7AF79FE07FBF1FD6DFA0E" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" scope_class="Gastropoda" scope_family="Triphoridae" scope_subFamily="Triphorinae" scope_superFamily="Triphoroidea">
<subSubSection id="C3566504FFB7AF7FFE07FBF1FBEEFBBD" box="[450,1138,1044,1071]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BF3368FFFB7AF7FFE07FBF1FBEEFBBD" blockId="11.[450,1138,1044,1106]" box="[450,1138,1044,1071]" pageId="11" pageNumber="12">
<heading id="D0BB81E3FFB7AF7FFE07FBF1FBEEFBBD" box="[450,1138,1044,1071]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<taxonomicName id="4C4C4D0CFFB7AF7FFE07FBF1FBEEFBBD" ID-CoL="6NBJM" authority="Rolan &amp; Fernandez-Garces, 2015" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="2015" box="[450,1138,1044,1071]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tricingulata">
<emphasis id="B938EA9DFFB7AF7FFE07FBF1FD7CFBBD" bold="true" box="[450,736,1044,1071]" italics="true" pageId="11" pageNumber="12">Isotriphora tricingulata</emphasis>
<bibRefCitation id="EFDD4B7EFFB7AF7FFD22FBF1FBEEFBBD" author="Rolan E. &amp; Fernandez-Garces R." box="[743,1138,1044,1071]" pageId="11" pageNumber="12" pagination="46 - 54" refId="ref45335" refString="Rolan E. &amp; Fernandez-Garces R. 2015. Triphorids of the Karubenthos Expedition to Guadeloupe Island. Gloria Maris 54: 46 - 54." type="journal article" year="2015">Rolán &amp; Fernández-Garcés, 2015</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3566504FFB7AF7FFD7AFBDDFCE8FBC0" box="[703,884,1079,1106]" pageId="11" pageNumber="12" type="description">
<paragraph id="8BF3368FFFB7AF7FFD7AFBDDFCE8FBC0" blockId="11.[450,1138,1044,1106]" box="[703,884,1079,1106]" pageId="11" pageNumber="12">
<figureCitation id="13772A0AFFB7AF7FFD7AFBDDFC95FBC0" box="[703,777,1080,1106]" captionStart="Fig" captionStartId="12.[189,232,1930,1956]" captionTargetBox="[189,1399,463,1886]" captionTargetId="figure@12.[189,1399,463,1886]" captionTargetPageId="12" captionText="Fig. 3. Isotriphora tricingulata Rolán &amp; Fernández-Garcés, 2015. A, C. MNRJ 32558*, 2.75 mm and 3.04 mm, respectively. B. MNRJ 17929*, 4.62 mm. D. MNRJ 32574*, 3.29 mm. E–I, K. Same shell as A. J. Same shell as D. Scale bars: A–E = 1 mm; F–H = 500 µm; I–K = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3899053" httpUri="https://zenodo.org/record/3899053/files/figure.png" pageId="11" pageNumber="12">Figs 3</figureCitation>
,
<figureCitation id="13772A0AFFB7AF7FFCD3FBDDFCD7FBC0" box="[790,843,1080,1106]" captionStart="Fig" captionStartId="49.[189,232,1717,1743]" captionTargetBox="[189,1399,354,1670]" captionTargetId="figure@49.[189,1399,354,1670]" captionTargetPageId="49" captionText="Fig. 23. Triphoridae from Brazil (2). A. “Inella” sp. 4. B. Iniforis carmelae Rolán &amp; Fernández- Garcés, 1993. C. Iniforis pseudothomae Rolán &amp; Fernández-Garcés, 1993. D. Isotriphora leo sp. nov. E. Isotriphora onca Fernandes, Pimenta &amp; Leal, 2013. F. Isotriphora tigrina Fernandes, Pimenta &amp; Leal, 2013. G. Isotriphora tricingulata Rolán &amp; Fernández-Garcés, 2015. H. Isotriphora uncia sp. nov. I. Isotriphora sp. 1. J. Latitriphora albida (A. Adams, 1854). K. Marshallora sp. L. Marshallora ostenta Rolán &amp; Fernández-Garcés, 2008. M. Marshallora sp. 1. N. Monophorus caracca (Dall, 1927) comb. nov. O. Monophorus olivaceus (Dall, 1889). P. Monophorus verecundus sp. nov. Q. Nanaphora leei Fernandes &amp; Pimenta, 2015. R. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989). S. Nanaphora sp. 1. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3899093" httpUri="https://zenodo.org/record/3899093/files/figure.png" pageId="11" pageNumber="12">23G</figureCitation>
,
<figureCitation id="13772A0AFFB7AF7FFC92FBD2FCE8FBC0" box="[855,884,1079,1106]" captionStart="Fig" captionStartId="81.[189,232,1615,1641]" captionTargetBox="[189,1399,265,1561]" captionTargetId="figure@81.[189,1399,265,1561]" captionTargetPageId="81" captionText="Fig. 50. Distribution map of Isotriphora tricingulata Rolán &amp; Fernández-Garcés, 2015 (red circle = type locality; black circles = additional localities) and Isotriphora onca Fernandes, Pimenta &amp; Leal, 2013 (brown circle = type locality; gray circle = additional locality)." figureDoi="http://doi.org/10.5281/zenodo.3899147" httpUri="https://zenodo.org/record/3899147/files/figure.png" pageId="11" pageNumber="12">50</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3566504FFB7AF7FFF78FB9CFD57FB47" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8BF3368FFFB7AF7FFF78FB9CFC79FB06" blockId="11.[189,997,1145,1172]" box="[189,997,1145,1172]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB5C11A1FFB7AF7FFF78FB9CFC79FB06" box="[189,997,1145,1172]" pageId="11" pageNumber="12">
<treatmentCitation id="0AED109EFFB7AF7FFF78FB9CFC0AFB06" author="Rolan E. &amp; Fernandez-Garces R." box="[189,918,1145,1172]" page="47" pageId="11" pageNumber="12" year="2015">
<taxonomicName id="4C4C4D0CFFB7AF7FFF78FB9CFC0AFB06" ID-CoL="6NBJM" authority="Rolan &amp; Fernandez-Garces, 2015: 47" authorityName="Rolan &amp; Fernandez-Garces" authorityPageNumber="47" authorityYear="2015" box="[189,918,1145,1172]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tricingulata">
<emphasis id="B938EA9DFFB7AF7FFF78FB9CFE49FB01" box="[189,469,1145,1171]" italics="true" pageId="11" pageNumber="12">Isotriphora tricingulata</emphasis>
<bibRefCitation id="EFDD4B7EFFB7AF7FFE19FB9CFC0AFB06" author="Rolan E. &amp; Fernandez-Garces R." box="[476,918,1145,1172]" pageId="11" pageNumber="12" pagination="46 - 54" refId="ref45335" refString="Rolan E. &amp; Fernandez-Garces R. 2015. Triphorids of the Karubenthos Expedition to Guadeloupe Island. Gloria Maris 54: 46 - 54." type="journal article" year="2015">Rolán &amp; Fernández-Garcés, 2015: 47</bibRefCitation>
</taxonomicName>
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, fig. 2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF3368FFFB7AF7FFF78FB5EFD57FB47" blockId="11.[189,715,1211,1238]" box="[189,715,1211,1238]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB5C11A1FFB7AF7FFF78FB5EFD57FB47" box="[189,715,1211,1238]" pageId="11" pageNumber="12">
<taxonomicName id="4C4C4D0CFFB7AF7FFF78FB5EFE19FB44" ID-CoL="7P5ZK" authorityName="Cotton &amp; Godfrey" authorityYear="1931" box="[189,389,1211,1238]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" isUncertain="true" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="undetermined">
“
<emphasis id="B938EA9DFFB7AF7FFF0FFB5EFECCFB47" box="[202,336,1211,1237]" italics="true" pageId="11" pageNumber="12">Isotriphora</emphasis>
” sp.
</taxonomicName>
–
<treatmentCitation id="0AED109EFFB7AF7FFE66FB5EFDC2FB47" author="Zhang D. Y." box="[419,606,1211,1237]" page="99" pageId="11" pageNumber="12" year="2011">
<bibRefCitation id="EFDD4B7EFFB7AF7FFE66FB5EFDC2FB47" author="Zhang D. Y." box="[419,606,1211,1237]" pageId="11" pageNumber="12" refId="ref46865" refString="Zhang D. Y. 2011. Antiguan Shallow-water Seashells - a Collection with 18 Years Study and Research of Shoreline Shells from Antigua, West Indies. MdM Publishing, Wellington." type="book" year="2011">Zhang 2011: 99</bibRefCitation>
</treatmentCitation>
, fig. 296.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3566504FFB7AF7FFF78FB18FD15F9CC" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BF3368FFFB7AF7FFF78FB18FE36FA85" blockId="11.[189,426,1277,1303]" box="[189,426,1277,1303]" pageId="11" pageNumber="12">
<heading id="D0BB81E3FFB7AF7FFF78FB18FE36FA85" bold="true" box="[189,426,1277,1303]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B938EA9DFFB7AF7FFF78FB18FE36FA85" bold="true" box="[189,426,1277,1303]" pageId="11" pageNumber="12">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFB7AF7FFF2FFAC9FEC0FAD4" blockId="11.[189,1298,1324,1386]" box="[234,348,1324,1350]" pageId="11" pageNumber="12">
<emphasis id="B938EA9DFFB7AF7FFF2FFAC9FEC0FAD4" bold="true" box="[234,348,1324,1350]" pageId="11" pageNumber="12">
<typeStatus id="54F7882DFFB7AF7FFF2FFAC9FEC0FAD4" box="[234,348,1324,1350]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF3368FFFB7AF7FFF78FAB5FA8EFAF8" blockId="11.[189,1298,1324,1386]" box="[189,1298,1359,1386]" pageId="11" pageNumber="12">
<materialsCitation id="3B243CD2FFB7AF7FFF78FAB5FA8EFAF8" ID-GBIF-Occurrence="2636393065" box="[189,1298,1359,1386]" collectionCode="MNHN" country="Guadeloupe" elevation="81" location="Port-Louis" municipality="Grand Cul-de-Sac Marin" pageId="11" pageNumber="12" specimenCode="IM-2000-30472" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F35B761FFFB7AF7FFF78FAB5FE1EFAF8" box="[189,386,1360,1386]" name="Guadeloupe" pageId="11" pageNumber="12">GUADELOUPE</collectingCountry>
•
<location id="8E936054FFB7AF7FFE5EFAAAFD80FAFB" LSID="urn:lsid:plazi:treatment:03E58799FFB7AF79FE07FBF1FD6DFA0E:8E936054FFB7AF7FFE5EFAAAFD80FAFB" box="[411,540,1359,1385]" country="Guadeloupe" municipality="Grand Cul-de-Sac Marin" name="Port-Louis" pageId="11" pageNumber="12">Port-Louis</location>
,
<collectingMunicipality id="6B97ACF5FFB7AF7FFDE2FAAAFCD2FAFB" box="[551,846,1359,1385]" pageId="11" pageNumber="12">Grand Cul-de-Sac Marin</collectingMunicipality>
; depth
<quantity id="4CB49B6AFFB7AF7FFC66FAB5FC7DFAF8" box="[931,993,1360,1386]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="11" pageNumber="12" unit="m" value="81.0">
<elevation id="0061D1BCFFB7AF7FFC66FAB5FC7DFAF8" box="[931,993,1360,1386]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="11" pageNumber="12" unit="m" value="81.0">81 m</elevation>
</quantity>
;
<collectionCode id="ED5DAE4AFFB7AF7FFC2BFAB5FBD4FAF8" box="[1006,1096,1360,1386]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="12">MNHN</collectionCode>
<specimenCode id="DBEA9EF4FFB7AF7FFB8AFAB5FA92FAF8" box="[1103,1294,1360,1386]" collectionCode="IM-" pageId="11" pageNumber="12">IM-2000-30472</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF3368FFFB7AF7FFF2FFA74FE3AFA39" blockId="11.[189,1399,1425,1630]" box="[234,422,1425,1451]" pageId="11" pageNumber="12">
<emphasis id="B938EA9DFFB7AF7FFF2FFA74FE3AFA39" bold="true" box="[234,422,1425,1451]" pageId="11" pageNumber="12">Other material</emphasis>
</paragraph>
<paragraph id="8BF3368FFFB7AF7FFF78FA50FD15F9CC" blockId="11.[189,1399,1425,1630]" pageId="11" pageNumber="12">
<materialsCitation id="3B243CD2FFB7AF7FFF78FA50FBCAFA42" ID-GBIF-Occurrence="2636393265" box="[189,1110,1459,1488]" collectionCode="MNRJ" country="Brazil" latitude="3.9786112" location="Amapa" longLatPrecision="21" longitude="-49.556667" pageId="11" pageNumber="12" specimenCode="MNRJ 32558" specimenCount="12" stateProvince="Amapa">
<collectingCountry id="F35B761FFFB7AF7FFF78FA50FEBAFA5D" box="[189,294,1461,1487]" name="Brazil" pageId="11" pageNumber="12">BRAZIL</collectingCountry>
–
<emphasis id="B938EA9DFFB7AF7FFE86FA51FE02FA5D" bold="true" box="[323,414,1460,1487]" pageId="11" pageNumber="12">
<collectingRegion id="4988F86DFFB7AF7FFE86FA51FE02FA5D" box="[323,414,1460,1487]" country="Brazil" name="Amapa" pageId="11" pageNumber="12">Amapá</collectingRegion>
</emphasis>
•
<specimenCount id="9D4AFD06FFB7AF7FFE73FA50FD81FA5D" box="[438,541,1461,1487]" pageId="11" pageNumber="12">12 specs</specimenCount>
;
<geoCoordinate id="EE785048FFB7AF7FFDECFA56FD20FA5D" box="[553,700,1459,1487]" degrees="03" direction="north" minutes="58" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="43" value="3.9786112">03º58′43″ N</geoCoordinate>
,
<geoCoordinate id="EE785048FFB7AF7FFD02FA56FCFCFA5D" box="[711,864,1459,1487]" degrees="49" direction="west" minutes="33" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="24" value="-49.556667">49º33′24″ W</geoCoordinate>
; 2001;
<specimenCode id="DBEA9EF4FFB7AF7FFC70FA50FBCAFA42" box="[949,1110,1461,1488]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ 32558</specimenCode>
</materialsCitation>
* •
<materialsCitation id="3B243CD2FFB7AF7FFBBEFA50FDD0FA61" ID-GBIF-Occurrence="2636392860" collectionCode="MNRJ" country="Brazil" latitude="3.9786112" longLatPrecision="21" longitude="-49.556667" pageId="11" pageNumber="12" specimenCode="MNRJ 32556" specimenCount="2" stateProvince="Amapa">
<specimenCount id="9D4AFD06FFB7AF7FFBBEFA50FB4FFA5D" box="[1147,1235,1461,1487]" pageId="11" pageNumber="12">2 specs</specimenCount>
;
<geoCoordinate id="EE785048FFB7AF7FFB1AFA56FAEEFA5D" box="[1247,1394,1459,1487]" degrees="03" direction="north" minutes="58" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="43" value="3.9786112">03º58′43″ N</geoCoordinate>
,
<geoCoordinate id="EE785048FFB7AF7FFF78FA32FEC9FA61" box="[189,341,1495,1523]" degrees="49" direction="west" minutes="33" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="24" value="-49.556667">49º33′24″ W</geoCoordinate>
; 2001;
<specimenCode id="DBEA9EF4FFB7AF7FFE6FFA3CFDD0FA61" box="[426,588,1496,1523]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ 32556</specimenCode>
</materialsCitation>
* •
<materialsCitation id="3B243CD2FFB7AF7FFDAAFA3DFB66FA61" ID-GBIF-Occurrence="2636392403" box="[623,1274,1495,1523]" collectionCode="MNRJ" country="Brazil" latitude="3.9786112" longLatPrecision="21" longitude="-49.556667" pageId="11" pageNumber="12" specimenCode="MNRJ 32563" specimenCount="1" stateProvince="Amapa">
<specimenCount id="9D4AFD06FFB7AF7FFDAAFA3DFD5FFA61" box="[623,707,1496,1523]" pageId="11" pageNumber="12" type="generic">1 spec.</specimenCount>
;
<geoCoordinate id="EE785048FFB7AF7FFD0BFA32FCFDFA61" box="[718,865,1495,1523]" degrees="03" direction="north" minutes="58" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="43" value="3.9786112">03º58′43″ N</geoCoordinate>
,
<geoCoordinate id="EE785048FFB7AF7FFCA9FA32FB98FA61" box="[876,1028,1495,1523]" degrees="49" direction="west" minutes="33" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="24" value="-49.556667">49º33′24″ W</geoCoordinate>
; 2001;
<specimenCode id="DBEA9EF4FFB7AF7FFB9CFA3CFB66FA61" box="[1113,1274,1496,1523]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ 32563</specimenCode>
</materialsCitation>
* •
<materialsCitation id="3B243CD2FFB7AF7FFAD8FA3DFD47F985" ID-GBIF-Occurrence="2636392563" collectionCode="MNRJ" country="Brazil" latitude="3.9786112" longLatPrecision="21" longitude="-49.556667" pageId="11" pageNumber="12" specimenCode="MNRJ 32569" specimenCount="1" stateProvince="Amapa">
<specimenCount id="9D4AFD06FFB7AF7FFAD8FA3DFAEEFA61" box="[1309,1394,1496,1523]" pageId="11" pageNumber="12" type="generic">1 spec.</specimenCount>
;
<geoCoordinate id="EE785048FFB7AF7FFF78FA1FFED1F984" box="[189,333,1530,1558]" degrees="03" direction="north" minutes="58" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="43" value="3.9786112">03º58′43″ N</geoCoordinate>
,
<geoCoordinate id="EE785048FFB7AF7FFE93FA1FFE70F984" box="[342,492,1530,1558]" degrees="49" direction="west" minutes="33" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="24" value="-49.556667">49º33′24″ W</geoCoordinate>
; 2001;
<specimenCode id="DBEA9EF4FFB7AF7FFDF9FA19FD47F985" box="[572,731,1532,1559]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ 32569</specimenCode>
</materialsCitation>
* •
<materialsCitation id="3B243CD2FFB7AF7FFD3CFA19FAFBF985" ID-GBIF-Occurrence="2636392341" box="[761,1383,1530,1559]" collectionCode="MNRJ" country="Brazil" latitude="3.9786112" longLatPrecision="21" longitude="-49.556667" pageId="11" pageNumber="12" specimenCode="MNRJ 32574" specimenCount="1" stateProvince="Amapa">
<specimenCount id="9D4AFD06FFB7AF7FFD3CFA19FCD7F984" box="[761,843,1532,1558]" pageId="11" pageNumber="12" type="generic">1 spec.</specimenCount>
;
<geoCoordinate id="EE785048FFB7AF7FFC91FA1FFC79F984" box="[852,997,1530,1558]" degrees="03" direction="north" minutes="58" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="43" value="3.9786112">03º58′43″ N</geoCoordinate>
,
<geoCoordinate id="EE785048FFB7AF7FFC28FA1FFB1CF984" box="[1005,1152,1530,1558]" degrees="49" direction="west" minutes="33" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="24" value="-49.556667">49º33′24″ W</geoCoordinate>
2001;
<specimenCode id="DBEA9EF4FFB7AF7FFB09FA19FAFBF985" box="[1228,1383,1532,1559]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ 32574</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B243CD2FFB7AF7FFF78F9C5FC21F9A8" ID-GBIF-Occurrence="2636392944" box="[189,957,1566,1594]" collectingDate="2008-11" collectionCode="MNRJ" country="Brazil" elevation="75" latitude="2.75" longLatPrecision="1307" longitude="-48.7" pageId="11" pageNumber="12" specimenCode="MNRJ 17929" specimenCount="1" stateProvince="Amapa">
<specimenCount id="9D4AFD06FFB7AF7FFF78F9C5FE8DF9A8" box="[189,273,1568,1594]" pageId="11" pageNumber="12" type="generic">1 spec.</specimenCount>
;
<geoCoordinate id="EE785048FFB7AF7FFEDEF9FBFE18F9A8" box="[283,388,1566,1594]" degrees="02" direction="north" minutes="45" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="2.75">02º45′ N</geoCoordinate>
,
<geoCoordinate id="EE785048FFB7AF7FFE4BF9FBFE60F9A8" box="[398,508,1566,1594]" degrees="48" direction="west" minutes="42" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="-48.7">48º42′ W</geoCoordinate>
; depth
<quantity id="4CB49B6AFFB7AF7FFD8BF9FAFD17F9A8" box="[590,651,1567,1594]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="12" unit="m" value="75.0">
<elevation id="0061D1BCFFB7AF7FFD8BF9FAFD17F9A8" box="[590,651,1567,1594]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="12" unit="m" value="75.0">75 m</elevation>
</quantity>
;
<date id="FFF2104FFFB7AF7FFD53F9C5FC8EF9A8" box="[662,786,1568,1594]" pageId="11" pageNumber="12" value="2008-11">
<collectingDate id="EFB6E9A7FFB7AF7FFD53F9C5FC8EF9A8" box="[662,786,1568,1594]" pageId="11" pageNumber="12" value="2008-11">Nov. 2008</collectingDate>
</date>
;
<specimenCode id="DBEA9EF4FFB7AF7FFCD8F9C5FC21F9A8" box="[797,957,1568,1594]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="12">MNRJ 17929</specimenCode>
</materialsCitation>
* •
<materialsCitation id="3B243CD2FFB7AF7FFC25F9C5FD19F9CC" ID-GBIF-Occurrence="2636393195" collectingDate="1997-03" collectionCode="MORG" country="Brazil" elevation="72" latitude="2.3533332" longLatPrecision="21" longitude="-48.498333" pageId="11" pageNumber="12" specimenCode="MORG 52263" specimenCount="73" specimenCount-male="72" stateProvince="Amapa">
<specimenCount id="9D4AFD06FFB7AF7FFC25F9C5FBAFF9A8" box="[992,1075,1568,1594]" pageId="11" pageNumber="12" type="generic">1 spec.</specimenCount>
;
<geoCoordinate id="EE785048FFB7AF7FFBFBF9FBFB4CF9A8" box="[1086,1232,1566,1594]" degrees="02" direction="north" minutes="21" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="12" value="2.3533332">02º21′12″ N</geoCoordinate>
,
<geoCoordinate id="EE785048FFB7AF7FFB1FF9FBFAEEF9A8" box="[1242,1394,1566,1594]" degrees="48" direction="west" minutes="29" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="54" value="-48.498333">48º29′54″ W</geoCoordinate>
; depth
<quantity id="4CB49B6AFFB7AF7FFEC0F9A6FED8F9CC" box="[261,324,1603,1630]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="11" pageNumber="12" unit="m" value="72.0">
<specimenCount id="9D4AFD06FFB7AF7FFEC0F9A6FED8F9CC" box="[261,324,1603,1630]" pageId="11" pageNumber="12" type="male">72 m</specimenCount>
</quantity>
;
<date id="FFF2104FFFB7AF7FFE95F9A1FE50F9CF" box="[336,460,1603,1630]" pageId="11" pageNumber="12" value="1997-03">
<collectingDate id="EFB6E9A7FFB7AF7FFE95F9A1FE50F9CF" box="[336,460,1603,1630]" pageId="11" pageNumber="12" value="1997-03">Mar. 1997</collectingDate>
</date>
;
<specimenCode id="DBEA9EF4FFB7AF7FFE1CF9A6FD19F9CC" box="[473,645,1603,1630]" collectionCode="MORG" pageId="11" pageNumber="12">MORG 52263</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3566504FFB7AF79FF78F960FD16FEDA" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="description">
<paragraph id="8BF3368FFFB7AF7FFF78F960FED2F90D" blockId="11.[189,334,1669,1695]" box="[189,334,1669,1695]" pageId="11" pageNumber="12">
<heading id="D0BB81E3FFB7AF7FFF78F960FED2F90D" bold="true" box="[189,334,1669,1695]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B938EA9DFFB7AF7FFF78F960FED2F90D" bold="true" box="[189,334,1669,1695]" pageId="11" pageNumber="12">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFB7AF78FF78F951FAEBFE1C" blockId="11.[189,1399,1716,2027]" lastBlockId="12.[189,1399,267,398]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Shell sinistral, small, cyrtoconoid, rectilinear profile, up to
<quantity id="4CB49B6AFFB7AF7FFCA1F951FC21F95C" box="[868,957,1716,1742]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="11" pageNumber="12" unit="mm" value="4.6">4.6 mm</quantity>
long,
<quantity id="4CB49B6AFFB7AF7FFBC6F951FBC7F95C" box="[1027,1115,1716,1742]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="12" unit="mm" value="1.4">1.4 mm</quantity>
wide, length/width ratio 2.8 to 3.1, up to 11 whorls. Protoconch paucispiral with truncated apex, without clear differentiation of the teleoconch; two strongly nodulose spiral cords starting from the slightly pointed and very narrow nucleus, situated at ~37% and ~73% of the last whorl height; median spiral cord usually emerging in the fifth whorl of the shell, sometimes as early as the end of fourth whorl or as late as the end of the sixth whorl, and readily reaching the same size of other cords; on the body whorl, distance between spiral cords is 0.9–1.2 × higher than width of cords; 17–20 orthocline to slightly opisthocline axial ribs at seventh whorl of shell; rounded to slightly elliptical nodules of medium size; distinct and welldeveloped suture, with small sutural cord; wavy subperipheral cord, mainly not developing nodules, two thin and nearly smooth basal cords, abapical one almost indistinct in some cases; no supranumerical cords; ovate aperture,
<quantity id="4CB49B6AFFB0AF78FE0DFECBFDE8FEDB" box="[456,628,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.2" metricValueMin="5.5" pageId="12" pageNumber="13" unit="mm" value="0.585" valueMax="0.62" valueMin="0.55">0.55–0.62 mm</quantity>
long,
<quantity id="4CB49B6AFFB0AF78FD07FECBFCF3FEDB" box="[706,879,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.699999999999999" pageId="12" pageNumber="13" unit="mm" value="0.485" valueMax="0.5" valueMin="0.47">0.47–0.50 mm</quantity>
wide, length/width ratio 1.2; anterior canal curved backwards/downwards, short, open or partly closed by projection of outer lip,
<quantity id="4CB49B6AFFB0AF78FB0FFEB4FAEBFEF9" box="[1226,1399,337,363]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" metricValueMax="3.0" metricValueMin="2.4" pageId="12" pageNumber="13" unit="mm" value="0.27" valueMax="0.3" valueMin="0.24">0.24–0.30 mm</quantity>
long,
<quantity id="4CB49B6AFFB0AF78FF3AFE91FE34FE1C" box="[255,424,372,398]" metricMagnitude="-4" metricUnit="m" metricValue="1.5500000000000003" metricValueMax="1.7000000000000002" metricValueMin="1.4000000000000001" pageId="12" pageNumber="13" unit="mm" value="0.155" valueMax="0.17" valueMin="0.14">0.14–0.17 mm</quantity>
wide, length/width ratio 1.7–1.8; posterior canal is small sinus,
<quantity id="4CB49B6AFFB0AF78FB48FE91FAA9FE1C" box="[1165,1333,372,398]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.8999999999999999" pageId="12" pageNumber="13" unit="mm" value="0.1" valueMax="0.11" valueMin="0.09">0.09–0.11 mm</quantity>
long,
</paragraph>
<caption id="DF336607FFB0AF78FF78F86FFBFFF879" ID-DOI="http://doi.org/10.5281/zenodo.3899053" ID-Zenodo-Dep="3899053" httpUri="https://zenodo.org/record/3899053/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1930,1956]" targetBox="[189,1399,463,1886]" targetPageId="12">
<paragraph id="8BF3368FFFB0AF78FF78F86FFBFFF879" blockId="12.[189,1398,1930,2027]" pageId="12" pageNumber="13">
<emphasis id="B938EA9DFFB0AF78FF78F86FFE97F836" bold="true" box="[189,267,1930,1956]" pageId="12" pageNumber="13">Fig. 3.</emphasis>
<taxonomicName id="4C4C4D0CFFB0AF78FED1F86FFC5AF837" authority="Rolan &amp; Fernandez-Garces, 2015" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="2015" box="[276,966,1930,1957]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="tricingulata">
<emphasis id="B938EA9DFFB0AF78FED1F86FFDB1F836" box="[276,557,1930,1956]" italics="true" pageId="12" pageNumber="13">Isotriphora tricingulata</emphasis>
<bibRefCitation id="EFDD4B7EFFB0AF78FDF0F86FFC5AF837" author="Rolan E. &amp; Fernandez-Garces R." box="[565,966,1930,1957]" pageId="12" pageNumber="13" pagination="46 - 54" refId="ref45335" refString="Rolan E. &amp; Fernandez-Garces R. 2015. Triphorids of the Karubenthos Expedition to Guadeloupe Island. Gloria Maris 54: 46 - 54." type="journal article" year="2015">Rolán &amp; Fernández-Garcés, 2015</bibRefCitation>
</taxonomicName>
.
<emphasis id="B938EA9DFFB0AF78FC17F86FFB90F836" bold="true" box="[978,1036,1930,1956]" pageId="12" pageNumber="13">A, C</emphasis>
. MNRJ 32558*, 2.75 mm and 3.04 mm, respectively.
<emphasis id="B938EA9DFFB0AF78FE08F848FE7DF855" bold="true" box="[461,481,1965,1991]" pageId="12" pageNumber="13">B</emphasis>
. MNRJ 17929*, 4.62 mm.
<emphasis id="B938EA9DFFB0AF78FCE4F848FCAAF855" bold="true" box="[801,822,1965,1991]" pageId="12" pageNumber="13">D</emphasis>
. MNRJ 32574*, 3.29 mm.
<emphasis id="B938EA9DFFB0AF78FBB3F848FB55F855" bold="true" box="[1142,1225,1965,1991]" pageId="12" pageNumber="13">E–I, K</emphasis>
. Same shell as A.
<emphasis id="B938EA9DFFB0AF78FF24F835FF6CF878" bold="true" box="[225,240,2000,2026]" pageId="12" pageNumber="13">J</emphasis>
. Same shell as D. Scale bars: A–E = 1 mm; F–H = 500 µm; I–K = 100 µm.
</paragraph>
</caption>
<paragraph id="8BF3368FFFB1AF79FF78FEE9FD16FEDA" blockId="13.[189,1399,267,328]" pageId="13" pageNumber="14">not detached from aperture. Two to three initial whorls white, remaining whorls homogeneously cream or light brown; worn shells dirty white.</paragraph>
</subSubSection>
<subSubSection id="C3566504FFB1AF79FF78FE8BFD6DFA0E" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BF3368FFFB1AF79FF78FE8BFEB3FE1A" blockId="13.[189,303,366,392]" box="[189,303,366,392]" pageId="13" pageNumber="14">
<heading id="D0BB81E3FFB1AF79FF78FE8BFEB3FE1A" bold="true" box="[189,303,366,392]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B938EA9DFFB1AF79FF78FE8BFEB3FE1A" bold="true" box="[189,303,366,392]" pageId="13" pageNumber="14">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFB1AF79FF78FE79FDC6FD16" blockId="13.[189,1399,412,644]" pageId="13" pageNumber="14">
As indicated in the original description, the shell of
<taxonomicName id="4C4C4D0CFFB1AF79FC95FE78FC65FE24" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="2015" box="[848,1017,412,438]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="tricingulata">
<emphasis id="B938EA9DFFB1AF79FC95FE78FCC6FE24" box="[848,858,413,438]" italics="true" pageId="13" pageNumber="14">I</emphasis>
.
<emphasis id="B938EA9DFFB1AF79FCABFE79FC65FE24" box="[878,1017,412,438]" italics="true" pageId="13" pageNumber="14">tricingulata</emphasis>
</taxonomicName>
superficially resembles that of
<taxonomicName id="4C4C4D0CFFB1AF79FF78FE5BFCDFFE4A" authority="(Moolenbeek &amp; Faber, 1989)" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[189,835,446,472]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="peetersae">
<emphasis id="B938EA9DFFB1AF79FF78FE5BFE5EFE4A" box="[189,450,446,472]" italics="true" pageId="13" pageNumber="14">Isotriphora peetersae</emphasis>
(
<bibRefCitation id="EFDD4B7EFFB1AF79FE1CFE5BFCA5FE4A" author="Moolenbeek R. G. &amp; Faber M. J." box="[473,825,446,472]" pageId="13" pageNumber="14" pagination="77 - 80" refId="ref43230" refString="Moolenbeek R. G. &amp; Faber M. J. 1989. Two new Triphora species from the West Indies (Gastropoda; Triphoridae). Basteria 53: 77 - 80." type="journal article" year="1989">Moolenbeek &amp; Faber, 1989</bibRefCitation>
)
</taxonomicName>
, a species widely distributed throughout the Caribbean (
<bibRefCitation id="EFDD4B7EFFB1AF79FE80FE05FD1BFE69" author="Moolenbeek R. G. &amp; Faber M. J." box="[325,647,480,507]" pageId="13" pageNumber="14" pagination="77 - 80" refId="ref43230" refString="Moolenbeek R. G. &amp; Faber M. J. 1989. Two new Triphora species from the West Indies (Gastropoda; Triphoridae). Basteria 53: 77 - 80." type="journal article" year="1989">Moolenbeek &amp; Faber 1989</bibRefCitation>
;
<bibRefCitation id="EFDD4B7EFFB1AF79FD51FE05FC33FE69" author="Rolan E. &amp; Espinosa J." box="[660,943,480,507]" pageId="13" pageNumber="14" pagination="63 - 68" refId="ref45029" refString="Rolan E. &amp; Espinosa J. 1994. The family Triphoridae (Mollusca, Gastropoda) in Cuba. 3. The genus Isotriphora, with description of a new species. Basteria 58: 63 - 68." type="journal article" year="1994">Rolán &amp; Espinosa 1994</bibRefCitation>
;
<bibRefCitation id="EFDD4B7EFFB1AF79FC7EFE05FBFDFE69" author="Redfern C." box="[955,1121,480,507]" pageId="13" pageNumber="14" refId="ref44555" refString="Redfern C. 2013. Bahamian Seashells: 1161 Species from Abaco, Bahamas. Bahamian Seashells Inc., Boca Raton." type="book" year="2013">Redfern 2013</bibRefCitation>
). However,
<taxonomicName id="4C4C4D0CFFB1AF79FB28FE04FAEAFE68" baseAuthorityName="Moolenbeek &amp; Faber" baseAuthorityYear="1989" box="[1261,1398,481,507]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="peetersae">
<emphasis id="B938EA9DFFB1AF79FB28FE04FB6BFE68" box="[1261,1271,481,506]" italics="true" pageId="13" pageNumber="14">I</emphasis>
.
<emphasis id="B938EA9DFFB1AF79FAC3FE04FAEAFE68" box="[1286,1398,481,506]" italics="true" pageId="13" pageNumber="14">peetersae</emphasis>
</taxonomicName>
is differentiated by a somewhat bottle-shaped shell, the presence of three spiral cords through most of the protoconch, a late median spiral cord of the teleoconch, the white coloration comprising the five initial whorls of the shell (but only two to three whorls in
<taxonomicName id="4C4C4D0CFFB1AF79FC9CFDADFC60FDF3" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="2015" box="[857,1020,583,610]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="tricingulata">
<emphasis id="B938EA9DFFB1AF79FC9CFDADFCFFFDF3" box="[857,867,584,609]" italics="true" pageId="13" pageNumber="14">I</emphasis>
.
<emphasis id="B938EA9DFFB1AF79FCB4FDA2FC60FDF3" box="[881,1020,583,609]" italics="true" pageId="13" pageNumber="14">tricingulata</emphasis>
</taxonomicName>
,
<figureCitation id="13772A0AFFB1AF79FBCCFDA2FBD7FDF0" box="[1033,1099,583,610]" captionStart="Fig" captionStartId="12.[189,232,1930,1956]" captionTargetBox="[189,1399,463,1886]" captionTargetId="figure@12.[189,1399,463,1886]" captionTargetPageId="12" captionText="Fig. 3. Isotriphora tricingulata Rolán &amp; Fernández-Garcés, 2015. A, C. MNRJ 32558*, 2.75 mm and 3.04 mm, respectively. B. MNRJ 17929*, 4.62 mm. D. MNRJ 32574*, 3.29 mm. E–I, K. Same shell as A. J. Same shell as D. Scale bars: A–E = 1 mm; F–H = 500 µm; I–K = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3899053" httpUri="https://zenodo.org/record/3899053/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
A–D) and a darker brown coloration in the remaining whorls.
</paragraph>
<paragraph id="8BF3368FFFB1AF79FF78FD4FFEADFCE2" blockId="13.[189,1399,682,880]" pageId="13" pageNumber="14">
<taxonomicName id="4C4C4D0CFFB1AF79FF78FD4FFE49FD56" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="2015" box="[189,469,682,708]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="tricingulata">
<emphasis id="B938EA9DFFB1AF79FF78FD4FFE49FD56" box="[189,469,682,708]" italics="true" pageId="13" pageNumber="14">Isotriphora tricingulata</emphasis>
</taxonomicName>
is also easily distinguished from
<taxonomicName id="4C4C4D0CFFB1AF79FC9AFD4FFF66FD75" authority="Fernandes, Pimenta &amp; Leal, 2013" authorityName="Fernandes, Pimenta &amp; Leal" authorityYear="2013" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="onca">
<emphasis id="B938EA9DFFB1AF79FC9AFD4FFBBAFD56" box="[863,1062,682,708]" italics="true" pageId="13" pageNumber="14">Isotriphora onca</emphasis>
<bibRefCitation id="EFDD4B7EFFB1AF79FBE8FD4FFF66FD75" author="Fernandes M. R. &amp; Pimenta A. D. &amp; Leal J. H." pageId="13" pageNumber="14" pagination="1 - 18" refId="ref40682" refString="Fernandes M. R., Pimenta A. D. &amp; Leal J. H. 2013. Taxonomic review of Triphorinae (Gastropoda: Triphoridae) from the Vitoria-Trindade Seamount Chain, southeastern Brazil. The Nautilus 127: 1 - 18." type="journal article" year="2013">Fernandes, Pimenta &amp; Leal, 2013</bibRefCitation>
</taxonomicName>
, from the Vitória-Trindade Seamount Chain (southeastern
<collectingCountry id="F35B761FFFB1AF79FC73FD28FB9EFD7A" box="[950,1026,717,744]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
), by the presence of an evident median spiral cord emerging in the fifth/sixth whorl (
<figureCitation id="13772A0AFFB1AF79FCFDFD0AFC0DFC9B" box="[824,913,751,777]" captionStart="Fig" captionStartId="12.[189,232,1930,1956]" captionTargetBox="[189,1399,463,1886]" captionTargetId="figure@12.[189,1399,463,1886]" captionTargetPageId="12" captionText="Fig. 3. Isotriphora tricingulata Rolán &amp; Fernández-Garcés, 2015. A, C. MNRJ 32558*, 2.75 mm and 3.04 mm, respectively. B. MNRJ 17929*, 4.62 mm. D. MNRJ 32574*, 3.29 mm. E–I, K. Same shell as A. J. Same shell as D. Scale bars: A–E = 1 mm; F–H = 500 µm; I–K = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3899053" httpUri="https://zenodo.org/record/3899053/files/figure.png" pageId="13" pageNumber="14">Fig. 3F</figureCitation>
), but only with a very reduced size after the 13
<superScript id="7C399BC7FFB1AF79FEC0FCF5FE8EFC8C" attach="left" box="[261,274,784,798]" fontSize="6" pageId="13" pageNumber="14">th</superScript>
whorl in
<taxonomicName id="4C4C4D0CFFB1AF79FE46FCF7FD73FCB9" authority="(Fernandes et al. 2013)" baseAuthorityName="Fernandes" baseAuthorityYear="2013" box="[387,751,785,812]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="onca">
<emphasis id="B938EA9DFFB1AF79FE46FCF7FE11FCB9" box="[387,397,786,811]" italics="true" pageId="13" pageNumber="14">I</emphasis>
.
<emphasis id="B938EA9DFFB1AF79FE5EFCF7FE48FCB9" box="[411,468,786,811]" italics="true" pageId="13" pageNumber="14">onca</emphasis>
(
<bibRefCitation id="EFDD4B7EFFB1AF79FE26FCF4FD7AFCB9" author="Fernandes M. R. &amp; Pimenta A. D. &amp; Leal J. H." box="[483,742,785,812]" pageId="13" pageNumber="14" pagination="1 - 18" refId="ref40682" refString="Fernandes M. R., Pimenta A. D. &amp; Leal J. H. 2013. Taxonomic review of Triphorinae (Gastropoda: Triphoridae) from the Vitoria-Trindade Seamount Chain, southeastern Brazil. The Nautilus 127: 1 - 18." type="journal article" year="2013">
Fernandes
<emphasis id="B938EA9DFFB1AF79FDA0FCF7FD04FCB9" box="[613,664,785,811]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. 2013
</bibRefCitation>
)
</taxonomicName>
. The shell of
<taxonomicName id="4C4C4D0CFFB1AF79FC4EFCF7FC40FCB9" box="[907,988,786,812]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="onca">
<emphasis id="B938EA9DFFB1AF79FC4EFCF7FC09FCB9" box="[907,917,786,811]" italics="true" pageId="13" pageNumber="14">I</emphasis>
.
<emphasis id="B938EA9DFFB1AF79FC66FCF7FC40FCB9" box="[931,988,786,811]" italics="true" pageId="13" pageNumber="14">onca</emphasis>
</taxonomicName>
is more elongated, with a long and almost closed anterior canal, instead of the short and open or partly closed anterior canal of
<taxonomicName id="4C4C4D0CFFB1AF79FB10FCD0FAEBFCDC" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="2015" box="[1237,1399,820,846]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="tricingulata">
<emphasis id="B938EA9DFFB1AF79FB10FCD0FB43FCDC" box="[1237,1247,821,846]" italics="true" pageId="13" pageNumber="14">I</emphasis>
.
<emphasis id="B938EA9DFFB1AF79FB29FCD1FAEBFCDC" box="[1260,1399,820,846]" italics="true" pageId="13" pageNumber="14">tricingulata</emphasis>
</taxonomicName>
(
<figureCitation id="13772A0AFFB1AF79FF03FCB3FEB8FCE2" box="[198,292,854,880]" captionStart="Fig" captionStartId="12.[189,232,1930,1956]" captionTargetBox="[189,1399,463,1886]" captionTargetId="figure@12.[189,1399,463,1886]" captionTargetPageId="12" captionText="Fig. 3. Isotriphora tricingulata Rolán &amp; Fernández-Garcés, 2015. A, C. MNRJ 32558*, 2.75 mm and 3.04 mm, respectively. B. MNRJ 17929*, 4.62 mm. D. MNRJ 32574*, 3.29 mm. E–I, K. Same shell as A. J. Same shell as D. Scale bars: A–E = 1 mm; F–H = 500 µm; I–K = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3899053" httpUri="https://zenodo.org/record/3899053/files/figure.png" pageId="13" pageNumber="14">Fig. 3H</figureCitation>
).
</paragraph>
<paragraph id="8BF3368FFFB1AF79FF78FC73FD6AFBA8" blockId="13.[189,1399,918,1082]" pageId="13" pageNumber="14">
The single difference between the
<typeStatus id="54F7882DFFB1AF79FDA9FC73FD4FFC22" box="[620,723,918,944]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4C4D0CFFB1AF79FCC0FC72FC32FC23" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="2015" box="[773,942,919,945]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="tricingulata">
<emphasis id="B938EA9DFFB1AF79FCC0FC72FC93FC22" box="[773,783,919,944]" italics="true" pageId="13" pageNumber="14">I</emphasis>
.
<emphasis id="B938EA9DFFB1AF79FCE6FC72FC32FC23" box="[803,942,919,945]" italics="true" pageId="13" pageNumber="14">tricingulata</emphasis>
</taxonomicName>
and shells from
<collectingCountry id="F35B761FFFB1AF79FB43FC73FB52FC22" box="[1158,1230,918,944]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
is the earlier emergence of the median spiral cord in the
<typeStatus id="54F7882DFFB1AF79FD78FC5CFCB8FC41" box="[701,804,953,979]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(fourth whorl vs usually fifth whorl in shells from
<collectingCountry id="F35B761FFFB1AF79FF78FC3EFE96FC64" box="[189,266,987,1014]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
). Despite the supposed intracapsular metamorphosis suggested by protoconch morphology (
<figureCitation id="13772A0AFFB1AF79FF03FC18FE85FB8A" box="[198,281,1021,1048]" captionStart="Fig" captionStartId="12.[189,232,1930,1956]" captionTargetBox="[189,1399,463,1886]" captionTargetId="figure@12.[189,1399,463,1886]" captionTargetPageId="12" captionText="Fig. 3. Isotriphora tricingulata Rolán &amp; Fernández-Garcés, 2015. A, C. MNRJ 32558*, 2.75 mm and 3.04 mm, respectively. B. MNRJ 17929*, 4.62 mm. D. MNRJ 32574*, 3.29 mm. E–I, K. Same shell as A. J. Same shell as D. Scale bars: A–E = 1 mm; F–H = 500 µm; I–K = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3899053" httpUri="https://zenodo.org/record/3899053/files/figure.png" pageId="13" pageNumber="14">Fig. 3I</figureCitation>
), different populations of this species may be connected by the stepping stones of the Lesser Antilles and the continental shelf of the
<collectingCountry id="F35B761FFFB1AF79FD56FBC5FD6EFBA8" box="[659,754,1056,1082]" name="Guyana" pageId="13" pageNumber="14">Guyana</collectingCountry>
.
</paragraph>
<paragraph id="8BF3368FFFB1AF79FF78FB84FEFCFB2D" blockId="13.[189,1399,1120,1215]" pageId="13" pageNumber="14">
One shell of
<taxonomicName id="4C4C4D0CFFB1AF79FE91FB84FE64FBE8" authorityName="Rolan &amp; Fernandez-Garces" authorityYear="2015" box="[340,504,1120,1147]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="tricingulata">
<emphasis id="B938EA9DFFB1AF79FE91FB84FEC2FBE8" box="[340,350,1121,1146]" italics="true" pageId="13" pageNumber="14">I</emphasis>
.
<emphasis id="B938EA9DFFB1AF79FEA8FB85FE64FBE8" box="[365,504,1120,1146]" italics="true" pageId="13" pageNumber="14">tricingulata</emphasis>
</taxonomicName>
from
<collectingCountry id="F35B761FFFB1AF79FDFAFB85FD1BFBE8" box="[575,647,1120,1146]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
has a late emergence of the adapical spiral cord, only at the end of the third whorl of the shell (
<figureCitation id="13772A0AFFB1AF79FDF7FB66FD1BFB0F" box="[562,647,1155,1181]" captionStart="Fig" captionStartId="12.[189,232,1930,1956]" captionTargetBox="[189,1399,463,1886]" captionTargetId="figure@12.[189,1399,463,1886]" captionTargetPageId="12" captionText="Fig. 3. Isotriphora tricingulata Rolán &amp; Fernández-Garcés, 2015. A, C. MNRJ 32558*, 2.75 mm and 3.04 mm, respectively. B. MNRJ 17929*, 4.62 mm. D. MNRJ 32574*, 3.29 mm. E–I, K. Same shell as A. J. Same shell as D. Scale bars: A–E = 1 mm; F–H = 500 µm; I–K = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3899053" httpUri="https://zenodo.org/record/3899053/files/figure.png" pageId="13" pageNumber="14">Fig. 3J</figureCitation>
). This might be a small abnormality in the development of the initial whorls.
</paragraph>
<paragraph id="8BF3368FFFB1AF79FF78FB00FE50FA92" blockId="13.[189,460,1253,1280]" box="[189,460,1253,1280]" pageId="13" pageNumber="14">
<heading id="D0BB81E3FFB1AF79FF78FB00FE50FA92" bold="true" box="[189,460,1253,1280]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B938EA9DFFB1AF79FF78FB00FE50FA92" bold="true" box="[189,460,1253,1280]" pageId="13" pageNumber="14">Geographical records</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFB1AF79FF78FAF6FBC6FABC" blockId="13.[189,1114,1299,1326]" box="[189,1114,1299,1326]" pageId="13" pageNumber="14">
<collectingCountry id="F35B761FFFB1AF79FF78FAF6FE80FABF" box="[189,284,1299,1325]" name="Antigua and Barbuda" pageId="13" pageNumber="14">Antigua</collectingCountry>
(
<bibRefCitation id="EFDD4B7EFFB1AF79FEEEFAF6FE21FABF" author="Zhang D. Y." box="[299,445,1299,1325]" pageId="13" pageNumber="14" refId="ref46865" refString="Zhang D. Y. 2011. Antiguan Shallow-water Seashells - a Collection with 18 Years Study and Research of Shoreline Shells from Antigua, West Indies. MdM Publishing, Wellington." type="book" year="2011">Zhang 2011</bibRefCitation>
); Guadeloupe (
<typeStatus id="54F7882DFFB1AF79FDAAFAF6FD3FFABF" box="[623,675,1299,1325]" pageId="13" pageNumber="14">type</typeStatus>
locality);
<collectingCountry id="F35B761FFFB1AF79FCD9FAF6FCF4FABF" box="[796,872,1299,1325]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
:
<collectingRegion id="4988F86DFFB1AF79FCB4FAF6FC5AFABF" box="[881,966,1299,1325]" country="Brazil" name="Amapa" pageId="13" pageNumber="14">Amapá</collectingRegion>
(this study).
</paragraph>
<paragraph id="8BF3368FFFB1AF79FF78FAB1FE69FAFC" blockId="13.[189,501,1364,1390]" box="[189,501,1364,1390]" pageId="13" pageNumber="14">
<heading id="D0BB81E3FFB1AF79FF78FAB1FE69FAFC" bold="true" box="[189,501,1364,1390]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B938EA9DFFB1AF79FF78FAB1FE69FAFC" bold="true" box="[189,501,1364,1390]" pageId="13" pageNumber="14">Bathymetric distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFB1AF79FF78FA64FD6DFA0E" blockId="13.[189,753,1409,1436]" box="[189,753,1409,1436]" pageId="13" pageNumber="14">
Depth:
<quantity id="4CB49B6AFFB1AF79FED1FA67FECCFA0E" box="[276,336,1410,1436]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="13" pageNumber="14" unit="m" value="72.0">72 m</quantity>
(this study) to
<quantity id="4CB49B6AFFB1AF79FDC4FA67FDA1FA0E" box="[513,573,1410,1436]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="14" unit="m" value="81.0">81 m</quantity>
(
<typeStatus id="54F7882DFFB1AF79FD89FA67FD1CFA0E" box="[588,640,1410,1436]" pageId="13" pageNumber="14">type</typeStatus>
locality).
</paragraph>
</subSubSection>
</treatment>
</document>
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