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<document id="1C4AC88A37DEB4F3EA5F6C1DB6D6EB5A" ID-DOI="10.5852/ejt.2020.665" ID-GBIF-Dataset="06748f67-18d9-4c3e-a14c-77de1664d81b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3899047" ID-ZooBank="836C9171-0849-4F4D-BC8D-90C2D9E8B9D1" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1592413559169" checkinUser="valdenar" docAuthor="Fernandes, Maurício Romulo &amp; Pimenta, Alexandre Dias" docDate="2020" docId="03E58799FFB3AF65FD4AFA40FEE8F9E4" docLanguage="en" docName="ejt-665_ferrnandes_pimenta.pdf.imf" docOrigin="European Journal of Taxonomy 665 (665)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Isotriphora leo Fernandes &amp; Pimenta 2020, sp. nov." docType="treatment" docUuid="D7BFF5C6-9436-4BA3-BE0F-8551DCB43F8A" docUuidSource="ZooBank" docVersion="12" lastPageNumber="18" masterDocId="FFDCFFE1FFBCAF74FFC5FFE5FF9CFF92" masterDocTitle="Unraveling one of the ‘ Big Five’: update of the taxonomy of Triphoridae (Gastropoda, Triphoroidea) from Brazil" masterLastPageNumber="170" masterPageNumber="1" pageNumber="16" updateTime="1733402249213" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F45312C77926981A7B553A1001E2F629" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0EA12C0416417C198312028A4E911633">
<mods:title id="1FA2E5BBA55F3DDBD158773CF87D0B0F">Unraveling one of the ‘ Big Five’: update of the taxonomy of Triphoridae (Gastropoda, Triphoroidea) from Brazil</mods:title>
</mods:titleInfo>
<mods:name id="B8DEC8DA41047E1BABD968B40A1C62FA" type="personal">
<mods:role id="9A7FCF441AA9B5CEAAE0CD6B6A017C90">
<mods:roleTerm id="356DD93CF42AC5D3455A494E4C6D5368">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6A5296406E5D5A3E5310A348232E6314">Fernandes, Maurício Romulo</mods:namePart>
<mods:nameIdentifier id="11024720CC6471D9FF59E8BCCB052834" type="LSID">3B8B864F-3300-45B2-8D1F-61F282F83CDE</mods:nameIdentifier>
<mods:affiliation id="FB26F54664A51AA00195590BA30D4B82">Departamento de Zoologia, Instituto de Biociências, Universidade Federal do Estado do Rio de Janeiro, Av. Pasteur, 458, Urca, 22290 - 240, Rio de Janeiro, Brazil. &amp; Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20940 - 040, Rio de Janeiro, Brazil.</mods:affiliation>
<mods:nameIdentifier id="F41D7976796FFC50742F95968FD84F1B" type="email">mauriciofernandes14@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B76AF2909CB09A624DFEFCC7A945AA6B" type="personal">
<mods:role id="F6401B0113554EF3553C8919773F9D6D">
<mods:roleTerm id="4BDE4204E630E0805E7D072F2EF0A83A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6FE08BD56F743C71CD42F87057038013">Pimenta, Alexandre Dias</mods:namePart>
<mods:nameIdentifier id="0CBD826B3D3350DA972701B397188F5A" type="LSID">2A42AE1F-6AF8-4AB3-975C-D3CE4D678F22</mods:nameIdentifier>
<mods:affiliation id="6002EE7B126037FB7DC157ED3B7B8A27">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20940 - 040, Rio de Janeiro, Brazil.</mods:affiliation>
<mods:nameIdentifier id="311666BCA94DE84CCEA4370523BA6573" type="email">alexpim@mn.ufrj.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="BA8E01B59B9A642DBC3A111B67BE0B06">text</mods:typeOfResource>
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<mods:titleInfo id="345B37EBC8145222F4732CBC05E794ED">
<mods:title id="162180EFB304C0DB1BC9F70A5365E998">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="61E9BD35689509E454A1B1A8B4739D11">
<mods:date id="641BF3AC6485F04D6614F798768C7C15">2020</mods:date>
<mods:detail id="3DD25D10E8E3350A2894B98C5DAA9E47" type="pubDate">
<mods:number id="5240DBAB98827F12F8015871D88535F3">2020-06-12</mods:number>
</mods:detail>
<mods:detail id="6BEE78B344CB84AF88CC7C9906246726" type="volume">
<mods:number id="472B2BBAE630A356182E740E5FC69687">665</mods:number>
</mods:detail>
<mods:detail id="CE2B7F0987FBAB994288D5067F5D7CAF" type="issue">
<mods:number id="9AAC79C6A92D06F646671B8088096EE6">665</mods:number>
</mods:detail>
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<mods:start id="7892792BC3F712F7A2862BAD2364A43C">1</mods:start>
<mods:end id="8E326F5B352145C7E63C7D9C3E82997C">170</mods:end>
</mods:extent>
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<mods:classification id="41A1E9252DBE45BE00C134F6C7F06515">journal article</mods:classification>
<mods:identifier id="FDBC5E752007C275DC3AAEC72C3CC610" type="DOI">10.5852/ejt.2020.665</mods:identifier>
<mods:identifier id="5FC3486FC879CD14F06B964237049AC0" type="GBIF-Dataset">06748f67-18d9-4c3e-a14c-77de1664d81b</mods:identifier>
<mods:identifier id="FA854DF10F8D3B78729215A82C06EEDD" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="C266DEFA3C60DCE33A2EAC694B079F9E" type="Zenodo-Dep">3899047</mods:identifier>
<mods:identifier id="BF331851456F30287E5D95FF0356F4FC" type="ZooBank">836C9171-0849-4F4D-BC8D-90C2D9E8B9D1</mods:identifier>
</mods:mods>
<treatment id="03E58799FFB3AF65FD4AFA40FEE8F9E4" ID-GBIF-Taxon="164612992" LSID="urn:lsid:zoobank.org:act:D7BFF5C6-9436-4BA3-BE0F-8551DCB43F8A" httpUri="http://treatment.plazi.org/id/03E58799FFB3AF65FD4AFA40FEE8F9E4" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" scope_class="Gastropoda" scope_family="Triphoridae" scope_subFamily="Triphorinae" scope_superFamily="Triphoroidea">
<subSubSection id="C3566504FFB3AF7BFD4AFA40FB50FA75" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BF3368FFFB3AF7BFD4AFA40FC39FA52" blockId="15.[361,1228,1445,1550]" box="[655,933,1445,1472]" pageId="15" pageNumber="16">
<heading id="D0BB81E3FFB3AF7BFD4AFA40FC39FA52" box="[655,933,1445,1472]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<taxonomicName id="4C4C4D0CFFB3AF7BFD4AFA40FCDFFA52" ID-CoL="7RRHR" authority="Fernandes &amp; Pimenta, 2020" authorityName="Fernandes &amp; Pimenta" authorityYear="2020" box="[655,835,1445,1472]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="leo" status="sp. nov.">
<emphasis id="B938EA9DFFB3AF7BFD4AFA40FCDFFA52" bold="true" box="[655,835,1445,1472]" italics="true" pageId="15" pageNumber="16">Isotriphora leo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57E6FFB3AF7BFC8FFA43FC39FA52" box="[842,933,1446,1472]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BF3368FFFB3AF7BFEACFA29FB50FA75" blockId="15.[361,1228,1445,1550]" box="[361,1228,1484,1511]" pageId="15" pageNumber="16">
<uri id="FFDD3A8DFFB3AF7BFEACFA29FB50FA75" box="[361,1228,1484,1511]" pageId="15" pageNumber="16">
urn:lsid:zoobank.org:act:
<uuid id="FFEA0C5AFFB3AF7BFD4FFA29FB50FA75" box="[650,1228,1484,1511]" pageId="15" pageNumber="16">D7BFF5C6-9436-4BA3-BE0F-8551DCB43F8A</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3566504FFB3AF7BFD7AFA16FCE8F99C" box="[703,884,1523,1550]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BF3368FFFB3AF7BFD7AFA16FCE8F99C" blockId="15.[361,1228,1445,1550]" box="[703,884,1523,1550]" pageId="15" pageNumber="16">
<figureCitation id="13772A0AFFB3AF7BFD7AFA16FC95F99C" box="[703,777,1523,1550]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[189,1399,465,1888]" captionTargetId="figure@16.[189,1399,465,1888]" captionTargetPageId="16" captionText="Fig. 5. Isotriphora leo sp. nov. A. MNRJ 29392*, holotype, 2.39 mm. B. MORG 25615*, 2.00 mm. C–D. MORG 25645*, 1.76 mm and 1.79 mm, respectively. E–I. HolotypE. Scale bars: A–E = 1 mm; F–G = 100 µm; H–I = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3899057" httpUri="https://zenodo.org/record/3899057/files/figure.png" pageId="15" pageNumber="16">Figs 5</figureCitation>
,
<figureCitation id="13772A0AFFB3AF7BFCD3FA11FCD0F99C" box="[790,844,1524,1550]" captionStart="Fig" captionStartId="49.[189,232,1717,1743]" captionTargetBox="[189,1399,354,1670]" captionTargetId="figure@49.[189,1399,354,1670]" captionTargetPageId="49" captionText="Fig. 23. Triphoridae from Brazil (2). A. “Inella” sp. 4. B. Iniforis carmelae Rolán &amp; Fernández- Garcés, 1993. C. Iniforis pseudothomae Rolán &amp; Fernández-Garcés, 1993. D. Isotriphora leo sp. nov. E. Isotriphora onca Fernandes, Pimenta &amp; Leal, 2013. F. Isotriphora tigrina Fernandes, Pimenta &amp; Leal, 2013. G. Isotriphora tricingulata Rolán &amp; Fernández-Garcés, 2015. H. Isotriphora uncia sp. nov. I. Isotriphora sp. 1. J. Latitriphora albida (A. Adams, 1854). K. Marshallora sp. L. Marshallora ostenta Rolán &amp; Fernández-Garcés, 2008. M. Marshallora sp. 1. N. Monophorus caracca (Dall, 1927) comb. nov. O. Monophorus olivaceus (Dall, 1889). P. Monophorus verecundus sp. nov. Q. Nanaphora leei Fernandes &amp; Pimenta, 2015. R. Nanaphora verbernei (Moolenbeek &amp; Faber, 1989). S. Nanaphora sp. 1. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3899093" httpUri="https://zenodo.org/record/3899093/files/figure.png" pageId="15" pageNumber="16">23D</figureCitation>
,
<figureCitation id="13772A0AFFB3AF7BFC92FA16FCE8F99C" box="[855,884,1523,1550]" captionStart="Fig" captionStartId="83.[189,232,1624,1650]" captionTargetBox="[189,1399,265,1561]" captionTargetId="figure@83.[189,1399,265,1561]" captionTargetPageId="83" captionText="Fig. 52. Distribution map of Isotriphora sp. 1 (gray circle) and Isotriphora leo sp. nov. (red circle)." figureDoi="http://doi.org/10.5281/zenodo.3899151" httpUri="https://zenodo.org/record/3899151/files/figure.png" pageId="15" pageNumber="16">52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3566504FFB3AF7BFF78F9DCFB0EF93E" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8BF3368FFFB3AF7BFF78F9DCFEAAF9C1" blockId="15.[189,310,1593,1619]" box="[189,310,1593,1619]" pageId="15" pageNumber="16">
<heading id="D0BB81E3FFB3AF7BFF78F9DCFEAAF9C1" bold="true" box="[189,310,1593,1619]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B938EA9DFFB3AF7BFF78F9DCFEAAF9C1" bold="true" box="[189,310,1593,1619]" pageId="15" pageNumber="16">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFB3AF7BFF78F98EFB0EF93E" blockId="15.[189,1398,1643,1708]" pageId="15" pageNumber="16">Small shell with a truncated apex; late adapical spiral cord, emerging in the third whorl of shell; white color, but light brown in the adapical spiral, subperipheral and adapical basal cords.</paragraph>
</subSubSection>
<subSubSection id="C3566504FFB3AF7BFF78F93DFD51F8DE" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BF3368FFFB3AF7BFF78F93DFED8F960" blockId="15.[189,324,1752,1778]" box="[189,324,1752,1778]" pageId="15" pageNumber="16">
<heading id="D0BB81E3FFB3AF7BFF78F93DFED8F960" bold="true" box="[189,324,1752,1778]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B938EA9DFFB3AF7BFF78F93DFED8F960" bold="true" box="[189,324,1752,1778]" pageId="15" pageNumber="16">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFB3AF7BFF78F8EFFD51F8DE" blockId="15.[189,1399,1802,1868]" pageId="15" pageNumber="16">
The specific name alludes to the lion (
<taxonomicName id="4C4C4D0CFFB3AF7BFD46F8EFFC64F8B7" authority="(Linnaeus, 1758))" authorityName="" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[643,1016,1802,1829]" class="Mammalia" family="Felidae" genus="Panthera" kingdom="Animalia" order="Carnivora" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="leo">
<emphasis id="B938EA9DFFB3AF7BFD46F8EFFC81F8B6" box="[643,797,1802,1828]" italics="true" pageId="15" pageNumber="16">Panthera leo</emphasis>
(Linnaeus, 1758))
</taxonomicName>
owing to the habit of authors to name species of
<taxonomicName id="4C4C4D0CFFB3AF7BFE45F8D7FD9AF8DE" authorityName="Cotton &amp; Godfrey" authorityYear="1931" box="[384,518,1842,1868]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus">
<emphasis id="B938EA9DFFB3AF7BFE45F8D7FD9AF8DE" box="[384,518,1842,1868]" italics="true" pageId="15" pageNumber="16">Isotriphora</emphasis>
</taxonomicName>
after big felines.
</paragraph>
</subSubSection>
<subSubSection id="C3566504FFB3AF64FF78F892FEE0FE1D" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BF3368FFFB3AF7BFF78F892FE36F803" blockId="15.[189,426,1911,1937]" box="[189,426,1911,1937]" pageId="15" pageNumber="16">
<heading id="D0BB81E3FFB3AF7BFF78F892FE36F803" bold="true" box="[189,426,1911,1937]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B938EA9DFFB3AF7BFF78F892FE36F803" bold="true" box="[189,426,1911,1937]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFB3AF7BFF2FF84FFEC0F856" blockId="15.[189,1340,1962,2027]" box="[234,348,1962,1988]" pageId="15" pageNumber="16">
<emphasis id="B938EA9DFFB3AF7BFF2FF84FFEC0F856" bold="true" box="[234,348,1962,1988]" pageId="15" pageNumber="16">
<typeStatus id="54F7882DFFB3AF7BFF2FF84FFEC0F856" box="[234,348,1962,1988]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF3368FFFB3AF7BFF78F834FAA0F879" blockId="15.[189,1340,1962,2027]" box="[189,1340,2000,2027]" pageId="15" pageNumber="16">
<materialsCitation id="3B243CD2FFB3AF7BFF78F834FAA0F879" ID-GBIF-Occurrence="2636392984" box="[189,1340,2000,2027]" collectionCode="MNRJ" collectorName="Barcellos &amp; Laurino" country="Brazil" elevation="5" location="Trindade Island" municipality="Beberibe" pageId="15" pageNumber="16" specimenCode="MNRJ 29392" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F35B761FFFB3AF7BFF78F834FEBAF879" box="[189,294,2001,2027]" name="Brazil" pageId="15" pageNumber="16">BRAZIL</collectingCountry>
–
<emphasis id="B938EA9DFFB3AF7BFE86F835FD90F879" bold="true" box="[323,524,2000,2027]" pageId="15" pageNumber="16">
<location id="8E936054FFB3AF7BFE86F835FD90F879" LSID="urn:lsid:plazi:treatment:03E58799FFB3AF65FD4AFA40FEE8F9E4:8E936054FFB3AF7BFE86F835FD90F879" box="[323,524,2000,2027]" country="Brazil" municipality="Beberibe" name="Trindade Island" pageId="15" pageNumber="16">Trindade Island</location>
</emphasis>
•
<collectingMunicipality id="6B97ACF5FFB3AF7BFDE0F834FD0FF879" box="[549,659,2001,2027]" pageId="15" pageNumber="16">Beberibe</collectingMunicipality>
; depth
<quantity id="4CB49B6AFFB3AF7BFD22F834FCA8F879" box="[743,820,2001,2027]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="15" pageNumber="16" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">
<elevation id="0061D1BCFFB3AF7BFD22F834FCA8F879" box="[743,820,2001,2027]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="15" pageNumber="16" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4–6 m</elevation>
</quantity>
;
<collectorName id="26B95359FFB3AF7BFC85F834FC33F879" box="[832,943,2001,2027]" pageId="15" pageNumber="16">Barcellos</collectorName>
and
<collectorName id="26B95359FFB3AF7BFC2DF834FBD9F879" box="[1000,1093,2001,2027]" pageId="15" pageNumber="16">Laurino</collectorName>
leg.;
<specimenCode id="DBEA9EF4FFB3AF7BFB42F834FAB1F879" box="[1159,1325,2001,2027]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="15" pageNumber="16">MNRJ 29392</specimenCode>
*.
</materialsCitation>
</paragraph>
<paragraph id="8BF3368FFFACAF64FF2FFEEEFE3AFEB7" blockId="16.[189,1399,267,399]" box="[234,422,267,293]" pageId="16" pageNumber="17">
<emphasis id="B938EA9DFFACAF64FF2FFEEEFE3AFEB7" bold="true" box="[234,422,267,293]" pageId="16" pageNumber="17">Other material</emphasis>
</paragraph>
<paragraph id="8BF3368FFFACAF64FF78FECBFEE0FE1D" blockId="16.[189,1399,267,399]" pageId="16" pageNumber="17">
<materialsCitation id="3B243CD2FFACAF64FF78FECBFE3BFEFE" ID-GBIF-Occurrence="2636392399" collectionCode="MORG" collectorName="Barcellos &amp; Laurino" country="Brazil" elevation="5" location="Trindade Island" municipality="Beberibe" pageId="16" pageNumber="17" specimenCode="MORG 25615" specimenCount="1" stateProvince="Trindade Island">
<collectingCountry id="F35B761FFFACAF64FF78FECBFEBAFEDA" box="[189,294,302,328]" name="Brazil" pageId="16" pageNumber="17">BRAZIL</collectingCountry>
–
<emphasis id="B938EA9DFFACAF64FE8FFECBFD8AFEDA" bold="true" box="[330,534,302,328]" pageId="16" pageNumber="17">
<collectingRegion id="4988F86DFFACAF64FE8FFECBFD8AFEDA" box="[330,534,302,328]" pageId="16" pageNumber="17">Trindade Island</collectingRegion>
</emphasis>
•
<specimenCount id="9D4AFD06FFACAF64FDF2FECBFD13FEDB" box="[567,655,302,329]" pageId="16" pageNumber="17" type="generic">1 spec.</specimenCount>
; same collection data as for holotype;
<collectorName id="26B95359FFACAF64FBAEFECBFB46FEDA" box="[1131,1242,302,328]" pageId="16" pageNumber="17">Barcellos</collectorName>
and
<collectorName id="26B95359FFACAF64FADFFECBFAEBFEDA" box="[1306,1399,302,328]" pageId="16" pageNumber="17">Laurino</collectorName>
leg.;
<specimenCode id="DBEA9EF4FFACAF64FF3FFEB4FE3BFEFE" box="[250,423,337,364]" collectionCode="MORG" pageId="16" pageNumber="17">MORG 25615</specimenCode>
</materialsCitation>
* •
<materialsCitation id="3B243CD2FFACAF64FE14FEB4FEF0FE1D" ID-GBIF-Occurrence="2636392602" collectingDate="1987-09-15" collectionCode="MORG" collectorName="Barcellos &amp; Laurino" country="Brazil" elevation="15" location="Fundeadouro" pageId="16" pageNumber="17" specimenCode="MORG 25645" specimenCount="2" stateProvince="Trindade Island">
<specimenCount id="9D4AFD06FFACAF64FE14FEB4FDB0FEFE" box="[465,556,337,364]" pageId="16" pageNumber="17">2 specs</specimenCount>
;
<location id="8E936054FFACAF64FDFFFEB4FD45FEF9" LSID="urn:lsid:plazi:treatment:03E58799FFB3AF65FD4AFA40FEE8F9E4:8E936054FFACAF64FDFFFEB4FD45FEF9" box="[570,729,337,363]" country="Brazil" name="Fundeadouro" pageId="16" pageNumber="17" stateProvince="Trindade Island">Fundeadouro</location>
; depth
<quantity id="4CB49B6AFFACAF64FCF4FEB4FCEEFEFE" box="[817,882,337,364]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="m" value="15.0">
<elevation id="0061D1BCFFACAF64FCF4FEB4FCEEFEFE" box="[817,882,337,364]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="m" value="15.0">15 m</elevation>
</quantity>
;
<date id="FFF2104FFFACAF64FC44FEB4FBBEFEF9" box="[897,1058,337,364]" pageId="16" pageNumber="17" value="1987-09-15">
<collectingDate id="EFB6E9A7FFACAF64FC44FEB4FBBEFEF9" box="[897,1058,337,364]" pageId="16" pageNumber="17" value="1987-09-15">15 Sep. 1987</collectingDate>
</date>
;
<collectorName id="26B95359FFACAF64FBF4FEB4FB3CFEF9" box="[1073,1184,337,363]" pageId="16" pageNumber="17">Barcellos</collectorName>
and
<collectorName id="26B95359FFACAF64FB18FEB4FAA6FEF9" box="[1245,1338,337,363]" pageId="16" pageNumber="17">Laurino</collectorName>
leg.;
<specimenCode id="DBEA9EF4FFACAF64FF78FE90FEF0FE1D" box="[189,364,372,399]" collectionCode="MORG" pageId="16" pageNumber="17">MORG 25645</specimenCode>
</materialsCitation>
*.
</paragraph>
</subSubSection>
<caption id="DF336607FFACAF64FF78F86DFDE1F87B" ID-DOI="http://doi.org/10.5281/zenodo.3899057" ID-Zenodo-Dep="3899057" httpUri="https://zenodo.org/record/3899057/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1928,1954]" targetBox="[189,1399,465,1888]" targetPageId="16">
<paragraph id="8BF3368FFFACAF64FF78F86DFDE1F87B" blockId="16.[189,1399,1928,2026]" pageId="16" pageNumber="17">
<emphasis id="B938EA9DFFACAF64FF78F86DFE8AF831" bold="true" box="[189,278,1928,1955]" pageId="16" pageNumber="17">Fig. 5.</emphasis>
<taxonomicName id="4C4C4D0CFFACAF64FEEDF86DFE78F830" authority="Fernandes &amp; Pimenta, 2020" authorityName="Fernandes &amp; Pimenta" authorityYear="2020" box="[296,484,1928,1954]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="leo" status="sp. nov.">
<emphasis id="B938EA9DFFACAF64FEEDF86DFE78F830" box="[296,484,1928,1954]" italics="true" pageId="16" pageNumber="17">Isotriphora leo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57E6FFACAF64FE32F86CFDC0F831" box="[503,604,1929,1955]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B938EA9DFFACAF64FDAAF86DFD18F830" bold="true" box="[623,644,1928,1954]" pageId="16" pageNumber="17">A</emphasis>
. MNRJ 29392*, holotype, 2.39 mm.
<emphasis id="B938EA9DFFACAF64FBBAF86CFB0FF831" bold="true" box="[1151,1171,1929,1955]" pageId="16" pageNumber="17">B</emphasis>
. MORG 25615*, 2.00 mm.
<emphasis id="B938EA9DFFACAF64FEF7F84EFEF7F857" bold="true" box="[306,363,1963,1989]" pageId="16" pageNumber="17">C–D</emphasis>
. MORG 25645*, 1.76 mm and 1.79 mm, respectively.
<emphasis id="B938EA9DFFACAF64FC2EF849FB85F854" bold="true" box="[1003,1049,1964,1990]" pageId="16" pageNumber="17">E–I</emphasis>
. HolotypE. Scale bars: A–E = 1 mm; F–G = 100 µm; H–I = 500 µm.
</paragraph>
</caption>
<subSubSection id="C3566504FFADAF65FF78FEEEFD45FC17" pageId="17" pageNumber="18" type="description">
<paragraph id="8BF3368FFFADAF65FF78FEEEFED2FEB7" blockId="17.[189,334,267,293]" box="[189,334,267,293]" pageId="17" pageNumber="18">
<heading id="D0BB81E3FFADAF65FF78FEEEFED2FEB7" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B938EA9DFFADAF65FF78FEEEFED2FEB7" bold="true" box="[189,334,267,293]" pageId="17" pageNumber="18">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFADAF65FF78FEDBFD45FC17" blockId="17.[189,1399,318,901]" pageId="17" pageNumber="18">
Shell sinistral, small, cyrtoconoid to ovoid, slightly convex profile, up to
<quantity id="4CB49B6AFFADAF65FBECFEDBFB19FECB" box="[1065,1157,318,345]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="18" unit="mm" value="2.4">2.4 mm</quantity>
long,
<quantity id="4CB49B6AFFADAF65FB17FEDBFAB1FECB" box="[1234,1325,318,345]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="18" unit="mm" value="0.9">0.9 mm</quantity>
wide, length/width ratio 2.4 to 2.5, up to 7.5 whorls. Protoconch paucispiral with truncated apex, without clear differentiation from teleoconch; two strongly nodulose spiral cords (median and abapical) of equal size starting from slightly pointed and very narrow nucleus, respectively situated at ~34% and ~71% of whorl height; third (adapical) spiral cord evident since third whorl of shell, bordering median cord and reaching same size of other cords after 1.5 to two whorls, being usually slightly more prominent than others on body whorl; on body whorl, spiral cords are 1.0–1.2 × as wide as distance between them; 17– 20 axial ribs at sixth whorl of shell, with varying arrangement, often orthocline, but slightly prosocline or slightly opisthocline in some whorls of certain shells; rounded nodules of medium to moderately large size; distinct and well-developed suture, with small sutural cord; smooth subperipheral and basal cords, abapical one very reduced, even indistinct in some shells; supranumerical cords absent; ovate to elliptical aperture,
<quantity id="4CB49B6AFFADAF65FE59FD11FDD4FC9C" box="[412,584,756,783]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="5.699999999999999" metricValueMin="3.4" pageId="17" pageNumber="18" unit="mm" value="0.45499999999999996" valueMax="0.57" valueMin="0.34">0.34–0.57 mm</quantity>
long,
<quantity id="4CB49B6AFFADAF65FD51FD11FCDCFC9C" box="[660,832,756,782]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="4.3" metricValueMin="3.1" pageId="17" pageNumber="18" unit="mm" value="0.37" valueMax="0.43" valueMin="0.31">0.31–0.43 mm</quantity>
wide, length/width ratio 1.1–1.4; anterior canal very short, open,
<quantity id="4CB49B6AFFADAF65FE43FCF9FDACFCA4" box="[390,560,796,822]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="17" pageNumber="18" unit="mm" value="0.11" valueMax="0.14" valueMin="0.08">0.08–0.14 mm</quantity>
long,
<quantity id="4CB49B6AFFADAF65FDBDFCF9FCBDFCA4" box="[632,801,796,822]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="17" pageNumber="18" unit="mm" value="0.135" valueMax="0.14" valueMin="0.13">0.13–0.14 mm</quantity>
wide, length/width ratio 0.6–1.0; posterior canal as acute sinus,
<quantity id="4CB49B6AFFADAF65FE8EFCA1FE69FCCC" box="[331,501,836,862]" metricMagnitude="-5" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7" pageId="17" pageNumber="18" unit="mm" value="0.095" valueMax="0.12" valueMin="0.07">0.07–0.12 mm</quantity>
long, not detached from aperture. Shell white, but light brown in the adapical spiral, subperipheral and adapical basal cords.
</paragraph>
</subSubSection>
<subSubSection id="C3566504FFADAF65FF78FC57FEE8F9E4" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BF3368FFFADAF65FF78FC57FEB3FC5E" blockId="17.[189,303,946,972]" box="[189,303,946,972]" pageId="17" pageNumber="18">
<heading id="D0BB81E3FFADAF65FF78FC57FEB3FC5E" bold="true" box="[189,303,946,972]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B938EA9DFFADAF65FF78FC57FEB3FC5E" bold="true" box="[189,303,946,972]" pageId="17" pageNumber="18">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFADAF65FF78FC01FAAFFBE4" blockId="17.[189,1399,996,1143]" pageId="17" pageNumber="18">
This species is unique in
<taxonomicName id="4C4C4D0CFFADAF65FE26FC00FDF5FC6D" authorityName="Cotton &amp; Godfrey" authorityYear="1931" box="[483,617,997,1023]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="B938EA9DFFADAF65FE26FC00FDF5FC6D" box="[483,617,997,1023]" italics="true" pageId="17" pageNumber="18">Isotriphora</emphasis>
</taxonomicName>
by the late emergence of the adapical spiral cord of the teleoconch (
<figureCitation id="13772A0AFFADAF65FF03FBE9FE95FBB5" box="[198,265,1036,1063]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[189,1399,465,1888]" captionTargetId="figure@16.[189,1399,465,1888]" captionTargetPageId="16" captionText="Fig. 5. Isotriphora leo sp. nov. A. MNRJ 29392*, holotype, 2.39 mm. B. MORG 25615*, 2.00 mm. C–D. MORG 25645*, 1.76 mm and 1.79 mm, respectively. E–I. HolotypE. Scale bars: A–E = 1 mm; F–G = 100 µm; H–I = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3899057" httpUri="https://zenodo.org/record/3899057/files/figure.png" pageId="17" pageNumber="18">Fig. 5</figureCitation>
E–F), thus raising doubts about the generic allocation. In fact, the shell of
<taxonomicName id="4C4C4D0CFFADAF65FBA3FBE9FA8BFBB4" authority="Fernandes &amp; Pimenta, 2020" authorityName="Fernandes &amp; Pimenta" authorityYear="2020" box="[1126,1303,1036,1062]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="leo" status="sp. nov.">
<emphasis id="B938EA9DFFADAF65FBA3FBE9FA8BFBB4" box="[1126,1303,1036,1062]" italics="true" pageId="17" pageNumber="18">Isotriphora leo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57E6FFADAF65FAD8FBE8FAEBFBB5" box="[1309,1399,1037,1063]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to that of some species of the family
<taxonomicName id="4C4C4D0CFFADAF65FCF8FBD1FC74FBDC" authorityName="Adams &amp; Adams" authorityYear="1853" box="[829,1000,1076,1102]" class="Gastropoda" family="Cerithiopsidae" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="family">Cerithiopsidae</taxonomicName>
in shape, small size and the late development of the adapical spiral cord, constituting an example of convergence in
<taxonomicName id="4C4C4D0CFFADAF65FB56FBB9FAB3FBE4" authorityName="Gray" authorityYear="1847" box="[1171,1327,1116,1142]" order="Null" pageId="17" pageNumber="137" rank="superFamily" superFamily="Triphoroidea">Triphoroidea</taxonomicName>
.
</paragraph>
<paragraph id="8BF3368FFFADAF65FF78FB47FDAEFA11" blockId="17.[189,1398,1186,1412]" pageId="17" pageNumber="18">
The late emergence of the adapical spiral cord distinguishes
<taxonomicName id="4C4C4D0CFFADAF65FC67FB46FC7EFB2E" authority="Fernandes &amp; Pimenta, 2020" authorityName="Fernandes &amp; Pimenta" authorityYear="2020" box="[930,994,1186,1213]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="leo" status="sp. nov.">
<emphasis id="B938EA9DFFADAF65FC67FB46FC30FB2E" box="[930,940,1187,1212]" italics="true" pageId="17" pageNumber="18">I</emphasis>
.
<emphasis id="B938EA9DFFADAF65FC7BFB47FC7EFB2E" box="[958,994,1186,1212]" italics="true" pageId="17" pageNumber="18">leo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57E6FFADAF65FC29FB46FBD7FB2F" box="[1004,1099,1187,1213]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
from the also small and slightly ovoid shell of
<taxonomicName id="4C4C4D0CFFADAF65FE07FB2EFD87FB76" authority="Fernandes &amp; Pimenta, 2020" authorityName="Fernandes &amp; Pimenta" authorityYear="2020" box="[450,539,1226,1252]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="uncia" status="sp. nov.">
<emphasis id="B938EA9DFFADAF65FE07FB2EFE50FB76" box="[450,460,1227,1252]" italics="true" pageId="17" pageNumber="18">I</emphasis>
.
<emphasis id="B938EA9DFFADAF65FE1FFB2EFD87FB76" box="[474,539,1227,1252]" italics="true" pageId="17" pageNumber="18">uncia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57E6FFADAF65FDE4FB2FFDE7FB76" box="[545,635,1226,1252]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
from Fernando de Noronha and Atol das Rocas, and the similarly colored shell of
<taxonomicName id="4C4C4D0CFFADAF65FE54FB17FBB7FA9E" authority="Fernandes, Pimenta &amp; Leal, 2013" authorityName="Fernandes, Pimenta &amp; Leal" authorityYear="2013" box="[401,1067,1266,1292]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="tigrina">
<emphasis id="B938EA9DFFADAF65FE54FB17FDE9FA9E" box="[401,629,1266,1292]" italics="true" pageId="17" pageNumber="18">Isotriphora tigrina</emphasis>
<bibRefCitation id="EFDD4B7EFFADAF65FD46FB17FBB7FA9E" author="Fernandes M. R. &amp; Pimenta A. D. &amp; Leal J. H." box="[643,1067,1266,1292]" pageId="17" pageNumber="18" pagination="1 - 18" refId="ref40682" refString="Fernandes M. R., Pimenta A. D. &amp; Leal J. H. 2013. Taxonomic review of Triphorinae (Gastropoda: Triphoridae) from the Vitoria-Trindade Seamount Chain, southeastern Brazil. The Nautilus 127: 1 - 18." type="journal article" year="2013">Fernandes, Pimenta &amp; Leal, 2013</bibRefCitation>
</taxonomicName>
from the Vitória-Trindade Chain.
<taxonomicName id="4C4C4D0CFFADAF65FED0FAFCFE55FAA1" authority="Fernandes &amp; Pimenta, 2020" authorityName="Fernandes &amp; Pimenta" authorityYear="2020" box="[277,457,1305,1331]" class="Gastropoda" family="Triphoridae" genus="Isotriphora" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="leo" status="sp. nov.">
<emphasis id="B938EA9DFFADAF65FED0FAFCFE55FAA1" box="[277,457,1305,1331]" italics="true" pageId="17" pageNumber="18">Isotriphora leo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57E6FFADAF65FE11FAFFFDAEFAA6" box="[468,562,1306,1332]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
probably is restricted to the shallow waters of Trindade Island, as it was not sampled in deeper waters of the Vitória-Trindade Chain by expeditions MD55 and REVIZEE-Central (
<bibRefCitation id="EFDD4B7EFFADAF65FEE7FA8CFDBAFA11" author="Fernandes M. R. &amp; Pimenta A. D. &amp; Leal J. H." box="[290,550,1385,1412]" pageId="17" pageNumber="18" pagination="1 - 18" refId="ref40682" refString="Fernandes M. R., Pimenta A. D. &amp; Leal J. H. 2013. Taxonomic review of Triphorinae (Gastropoda: Triphoridae) from the Vitoria-Trindade Seamount Chain, southeastern Brazil. The Nautilus 127: 1 - 18." type="journal article" year="2013">
Fernandes
<emphasis id="B938EA9DFFADAF65FE61FA8FFE44FA11" box="[420,472,1385,1411]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. 2013
</bibRefCitation>
).
</paragraph>
<paragraph id="8BF3368FFFADAF65FF78FA4AFE50FA5B" blockId="17.[189,460,1455,1481]" box="[189,460,1455,1481]" pageId="17" pageNumber="18">
<heading id="D0BB81E3FFADAF65FF78FA4AFE50FA5B" bold="true" box="[189,460,1455,1481]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B938EA9DFFADAF65FF78FA4AFE50FA5B" bold="true" box="[189,460,1455,1481]" pageId="17" pageNumber="18">Geographical records</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFADAF65FF78FA07FE4FFA6E" blockId="17.[189,467,1506,1532]" box="[189,467,1506,1532]" pageId="17" pageNumber="18">
<collectingCountry id="F35B761FFFADAF65FF78FA07FE95FA6E" box="[189,265,1506,1532]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
: Trindade Island.
</paragraph>
<paragraph id="8BF3368FFFADAF65FF78F9CDFE69F9D0" blockId="17.[189,501,1576,1602]" box="[189,501,1576,1602]" pageId="17" pageNumber="18">
<heading id="D0BB81E3FFADAF65FF78F9CDFE69F9D0" bold="true" box="[189,501,1576,1602]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B938EA9DFFADAF65FF78F9CDFE69F9D0" bold="true" box="[189,501,1576,1602]" pageId="17" pageNumber="18">Bathymetric distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3368FFFADAF65FF78F9BEFEE8F9E4" blockId="17.[189,372,1627,1654]" box="[189,372,1627,1654]" pageId="17" pageNumber="18">
Depth:
<quantity id="4CB49B6AFFADAF65FED1F9BEFEECF9E4" box="[276,368,1627,1654]" metricMagnitude="0" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="17" pageNumber="18" unit="m" value="9.5" valueMax="15.0" valueMin="4.0">
4–
<specimenCount id="9D4AFD06FFADAF65FEF1F9BEFEECF9E4" box="[308,368,1627,1654]" pageId="17" pageNumber="18" type="generic">15 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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