555 lines
89 KiB
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555 lines
89 KiB
XML
<document id="3A370427A2B7BF560426B694372555E2" ID-CLB-Dataset="21532" ID-DOI="10.11646/zootaxa.4809.3.6" ID-GBIF-Dataset="4bd957a1-4346-4339-bb01-293e2f3c31f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3936887" ID-ZooBank="46DA13F2-1B01-4E07-B134-BCBEFAB44A73" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594291990048" checkinUser="plazi" docAuthor="Mitra, Santanu, Trivedi, Jigneshkumar N. & Mendoza, Jose Christopher E." docDate="2020" docId="03E58796FFCDFFF130E1F8A2FBC792A9" docLanguage="en" docName="zootaxa.4809.3.6.pdf" docOrigin="Zootaxa 4809 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dotilla fraternalis Mitra & Trivedi & Mendoza 2020, n. sp." docType="treatment" docVersion="7" lastPageNumber="532" masterDocId="FFDCFFEEFFC8FFFA3076FF81FFA29005" masterDocTitle="On the two ‘ forms’ of Dotilla intermedia De Man, 1888 (Crustacea: Brachyura Dotillidae) from the Bay of Bengal, India, with description of a new species" masterLastPageNumber="534" masterPageNumber="521" pageNumber="526" updateTime="1701999157486" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3E21327AF7669FBE744EA4A923EAF909">On the two ‘ forms’ of Dotilla intermedia De Man, 1888 (Crustacea: Brachyura Dotillidae) from the Bay of Bengal, India, with description of a new species</mods:title>
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<mods:name id="0F11B2D00F36FEB4138F5F4C6D672D06" type="personal">
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<mods:roleTerm id="2F9DF73B1E3E087119FE3B82E77BFA3A">Author</mods:roleTerm>
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<mods:namePart id="36FE189A41F98AF5E80E2979F5A7C3DC">Mitra, Santanu</mods:namePart>
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<mods:affiliation id="154EF0D3EA26ED3824C0B9F352FE87E8">Zoological Survey of India, F. P. S. Building, 27 J. L. Nehru Road, Kolkata- 700016, West Bengal, India.</mods:affiliation>
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<mods:namePart id="C6D1A685EBA3053D66703E9177335D57">Trivedi, Jigneshkumar N.</mods:namePart>
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<mods:affiliation id="3DC6731588A482243461CAAFA083F08F">Department of Life Sciences, Hemchandracharya North Gujarat University, Patan, Gujarat, India.</mods:affiliation>
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<mods:namePart id="38CBDEA81108D6D25B33BB6A64E179D3">Mendoza, Jose Christopher E.</mods:namePart>
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<mods:affiliation id="0B102206252CC5B9041D4F755E03022B">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, 117377 Singapore. jcmendoza @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 9729 - 5703</mods:affiliation>
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<mods:title id="9C086FFAFF55502B411E19E8CF3C6AD2">Zootaxa</mods:title>
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<mods:part id="AD8863FAE3744B4BCCE4AA5E819B6C37">
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<mods:date id="D459F3A7E6918FB8438F36252AA9D530">2020</mods:date>
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<mods:number id="FB1A5E3D4A133DF2AA7DD7BA4DD9B25D">2020-07-08</mods:number>
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<mods:number id="AA80B83191D720A2F9BB965680A11CEB">4809</mods:number>
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<treatment id="03E58796FFCDFFF130E1F8A2FBC792A9" ID-DOI="http://doi.org/10.5281/zenodo.4324275" ID-GBIF-Taxon="164986704" ID-Zenodo-Dep="4324275" LSID="urn:lsid:plazi:treatment:03E58796FFCDFFF130E1F8A2FBC792A9" httpUri="http://treatment.plazi.org/id/03E58796FFCDFFF130E1F8A2FBC792A9" lastPageId="11" lastPageNumber="532" pageId="5" pageNumber="526">
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<subSubSection id="C356650BFFCDFFFF30E1F8A2FE19973B" box="[151,443,1827,1854]" pageId="5" pageNumber="526" type="nomenclature">
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<paragraph id="8BF33680FFCDFFFF30E1F8A2FE19973B" blockId="5.[151,443,1827,1890]" box="[151,443,1827,1854]" pageId="5" pageNumber="526">
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<heading id="D0BB81ECFFCDFFFF30E1F8A2FE19973B" bold="true" box="[151,443,1827,1854]" fontSize="11" level="1" pageId="5" pageNumber="526" reason="1">
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<emphasis id="B938EA92FFCDFFFF30E1F8A2FE19973B" bold="true" box="[151,443,1827,1854]" pageId="5" pageNumber="526">
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<taxonomicName id="4C4C4D03FFCDFFFF30E1F8A2FED3973B" ID-CoL="84GT6" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[151,369,1827,1854]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
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<emphasis id="B938EA92FFCDFFFF30E1F8A2FED3973B" bold="true" box="[151,369,1827,1854]" italics="true" pageId="5" pageNumber="526">Dotilla fraternalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B57E9FFCDFFFF310EF8A5FE19973B" box="[376,443,1828,1854]" pageId="5" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C356650BFFCDFFFF30E1F8C9FE119767" box="[151,435,1864,1890]" pageId="5" pageNumber="526" type="description">
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<paragraph id="8BF33680FFCDFFFF30E1F8C9FE119767" blockId="5.[151,443,1827,1890]" box="[151,435,1864,1890]" pageId="5" pageNumber="526">
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(
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<figureCitation id="13772A05FFCDFFFF30E9F8C9FF4D9767" box="[159,239,1864,1890]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="5" pageNumber="526">Figs. 6</figureCitation>
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,
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<figureCitation id="13772A05FFCDFFFF308DF8C9FEED9767" box="[251,335,1864,1890]" captionStart="FIGURE 7" captionStartId="9.[151,250,1809,1834]" captionTargetBox="[162,1423,219,1699]" captionTargetId="figure@9.[151,1436,211,1708]" captionTargetPageId="9" captionText="FIGURE 7. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal.A, pleon; D, spoon tipped seta of inner margin of merus of second maxilliped. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, pleon; E, spoon tipped setae of inner margin of merus of second maxilliped. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. C, pleon." figureDoi="http://doi.org/10.5281/zenodo.3936901" httpUri="https://zenodo.org/record/3936901/files/figure.png" pageId="5" pageNumber="526">7A, 7D</figureCitation>
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,
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<figureCitation id="13772A05FFCDFFFF312DF8C9FE099767" box="[347,427,1864,1890]" captionStart="FIGURE 8" captionStartId="10.[151,250,1806,1831]" captionTargetBox="[161,1428,388,1776]" captionTargetId="figure@10.[151,1436,381,1782]" captionTargetPageId="10" captionText="FIGURE 8. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2). A, left G1; E, left G2. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, left G1; F, left G2. Dotilla intermedia De Man, 1888 paralectotype, male (CW 4.2 mm, CL 3.5 mm) (ZSI-C8236/6), Sullivan I., Mergui Archipelago. C, left G1. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. D, left G1; G, left G2." figureDoi="http://doi.org/10.5281/zenodo.3936903" httpUri="https://zenodo.org/record/3936903/files/figure.png" pageId="5" pageNumber="526">8A, 8E</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C356650BFFCDFFFF30E1F80DFC0997E4" pageId="5" pageNumber="526" type="reference_group">
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<paragraph id="8BF33680FFCDFFFF30E1F80DFEC297C6" blockId="5.[151,1436,1931,2018]" pageId="5" pageNumber="526">
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<treatmentCitationGroup id="AB5C11AEFFCDFFFF30E1F80DFEC297C6" pageId="5" pageNumber="526">
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<taxonomicName id="4C4C4D03FFCDFFFF30E1F80DFEF097A1" ID-CoL="37HDQ" authorityName="De Man" authorityYear="1888" box="[151,338,1932,1956]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="intermedia">
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<emphasis id="B938EA92FFCDFFFF30E1F80DFEF097A1" box="[151,338,1932,1956]" italics="true" pageId="5" pageNumber="526">Dotilla intermedia</emphasis>
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</taxonomicName>
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(“low form”),
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<treatmentCitation id="0AED1091FFCDFFFF3191F80DFD2D97A1" author="Kemp, S." box="[487,655,1932,1956]" page="331" pageId="5" pageNumber="526" year="1919">
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<bibRefCitation id="EFDD4B71FFCDFFFF3191F80DFD2D97A1" author="Kemp, S." box="[487,655,1932,1956]" pageId="5" pageNumber="526" pagination="305 - 348" refId="ref7192" refString="Kemp, S. (1919) Notes on Crustacea Decapoda in the Indian Museum. XII. Scopimerinae. Records of the Indian Museum, 16, 305 - 348. https: // doi. org / 10.5962 / bhl. part. 25926" type="journal article" year="1919">Kemp 1919: 331</bibRefCitation>
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||
</treatmentCitation>
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||
(in part), figs. 10b, c, f;
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||
<treatmentCitation id="0AED1091FFCDFFFF3309F80DFB0C97A1" author="Dev Roy, M. K. & Bhadra, S." box="[895,1198,1931,1956]" page="512" pageId="5" pageNumber="526" year="2005">
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||
<bibRefCitation id="EFDD4B71FFCDFFFF3309F80DFB0C97A1" author="Dev Roy, M. K. & Bhadra, S." box="[895,1198,1931,1956]" pageId="5" pageNumber="526" pagination="357 - 535" refId="ref6627" refString="Dev Roy, M. K. & Bhadra, S. (2005) Marine and Estuarine crabs. In: Director ZSI, Kolkata (Ed.), Fauna of Andhra Pardesh, State Fauna Series. Part 5. Zoological Survey of India, Kolkata, pp. 357 - 535." type="book chapter" year="2005">Dev Roy & Bhadra 2005: 512</bibRefCitation>
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||
</treatmentCitation>
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. Not
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||
<taxonomicName id="4C4C4D03FFCDFFFF3497F80DFEFE97C6" ID-CoL="37HDQ" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="intermedia">
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<emphasis id="B938EA92FFCDFFFF3497F80DFA3E97A1" box="[1249,1436,1932,1956]" italics="true" pageId="5" pageNumber="526">Dotilla intermedia</emphasis>
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<bibRefCitation id="EFDD4B71FFCDFFFF30B1F82AFEFE97C6" author="De Man, J. G." box="[199,348,1963,1987]" pageId="5" pageNumber="526" pagination="1 - 312" refId="ref6453" refString="De Man, J. G. (1888) Report of the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, FRS, Superintendent of the Museum, Journal of the Linnaean Society, London, Zoology, 22, 1 - 312. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1888">De Man, 1888</bibRefCitation>
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</taxonomicName>
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||
.
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</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8BF33680FFCDFFFF30E1F848FC0997E4" blockId="5.[151,1436,1931,2018]" box="[151,939,1993,2018]" pageId="5" pageNumber="526">
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<treatmentCitationGroup id="AB5C11AEFFCDFFFF30E1F848FC0997E4" box="[151,939,1993,2018]" pageId="5" pageNumber="526">
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||
<treatmentCitation id="0AED1091FFCDFFFF30E1F848FDB897E4" author="Bairagi, N." box="[151,538,1993,2018]" page="278" pageId="5" pageNumber="526" year="1995">
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||
<taxonomicName id="4C4C4D03FFCDFFFF30E1F848FDB897E4" ID-CoL="37HDQ" authority=", Bairagi 1995: 278" authorityName="Bairagi" authorityPageNumber="278" authorityYear="1995" box="[151,538,1993,2018]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="intermedia">
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<emphasis id="B938EA92FFCDFFFF30E1F848FEF197E4" box="[151,339,1993,2017]" italics="true" pageId="5" pageNumber="526">Dotilla intermedia</emphasis>
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,
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<bibRefCitation id="EFDD4B71FFCDFFFF3116F848FDB897E4" author="Bairagi, N." box="[352,538,1993,2018]" pageId="5" pageNumber="526" pagination="263 - 288" refId="ref6028" refString="Bairagi, N. (1995) Ocypodidae: Decapoda: Crustacea. In: Director ZSI, Kolkata (Ed.), Hugli Matla Estuary, West Bengal. Esturaine Ecosystem Series. Part 2. Zoological Survey Of India, Kolkata, pp. 263 - 288." type="book chapter" year="1995">Bairagi 1995: 278</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
. Not
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||
<taxonomicName id="4C4C4D03FFCDFFFF3226F848FC0597E4" ID-CoL="37HDQ" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" box="[592,935,1993,2017]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="intermedia">
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<emphasis id="B938EA92FFCDFFFF3226F848FCAE97E4" box="[592,780,1993,2017]" italics="true" pageId="5" pageNumber="526">Dotilla intermedia</emphasis>
|
||
<bibRefCitation id="EFDD4B71FFCDFFFF3364F84BFC0597E4" author="De Man, J. G." box="[786,935,1993,2017]" pageId="5" pageNumber="526" pagination="1 - 312" refId="ref6453" refString="De Man, J. G. (1888) Report of the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, FRS, Superintendent of the Museum, Journal of the Linnaean Society, London, Zoology, 22, 1 - 312. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1888">De Man, 1888</bibRefCitation>
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</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
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||
<caption id="DF336608FFCEFFFC30E1FB0AFD2494E9" ID-DOI="http://doi.org/10.5281/zenodo.3936897" ID-Zenodo-Dep="3936897" httpUri="https://zenodo.org/record/3936897/files/figure.png" pageId="6" pageNumber="527" startId="6.[151,250,1163,1188]" targetBox="[154,1433,181,1138]" targetPageId="6">
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||
<paragraph id="8BF33680FFCEFFFC30E1FB0AFD2494E9" blockId="6.[151,1437,1163,1260]" pageId="6" pageNumber="527">
|
||
<emphasis id="B938EA92FFCEFFFC30E1FB0AFEB094A1" bold="true" box="[151,274,1163,1188]" pageId="6" pageNumber="527">FIGURE 5.</emphasis>
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Type specimens with accompanying museum labels. A,
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<taxonomicName id="4C4C4D03FFCEFFFC333CFB0DFB3994A1" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" box="[842,1179,1164,1188]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="intermedia">
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||
<emphasis id="B938EA92FFCEFFFC333CFB0DFBA794A1" box="[842,1029,1164,1188]" italics="true" pageId="6" pageNumber="527">Dotilla intermedia</emphasis>
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||
<bibRefCitation id="EFDD4B71FFCEFFFC347DFB0DFB3994A1" author="De Man, J. G." box="[1035,1179,1164,1188]" pageId="6" pageNumber="527" pagination="1 - 312" refId="ref6453" refString="De Man, J. G. (1888) Report of the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, FRS, Superintendent of the Museum, Journal of the Linnaean Society, London, Zoology, 22, 1 - 312. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1888">De Man, 1888</bibRefCitation>
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||
</taxonomicName>
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paralectotype, male (CW 4.2 mm, CL 3.5 mm) (ZSI-C8236/6) Sullivan I., Mergui Archipelago. B,
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||
<taxonomicName id="4C4C4D03FFCEFFFC33E4FB31FA8F94CD" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[914,1325,1199,1224]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="clepsydrodactylus">
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<emphasis id="B938EA92FFCEFFFC33E4FB31FB3A94CD" box="[914,1176,1200,1224]" italics="true" pageId="6" pageNumber="527">Dotilla clepsydrodactylus</emphasis>
|
||
<bibRefCitation id="EFDD4B71FFCEFFFC34D6FB2EFA8F94CD" author="Alcock, A." box="[1184,1325,1199,1224]" pageId="6" pageNumber="527" pagination="279 - 456" refId="ref5901" refString="Alcock, A. (1900) Materials for carcinological fauna of India, No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, 69, 279 - 456. https: // doi. org / 10.5962 / bhl. title. 15344" type="journal article" year="1900">Alcock, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C356650BFFCEFFFC30E1FA98FD2195A5" pageId="6" pageNumber="527" type="materials_examined">
|
||
<paragraph id="8BF33680FFCEFFFC30E1FA98FE73955D" blockId="6.[151,1437,1305,2017]" pageId="6" pageNumber="527">
|
||
<emphasis id="B938EA92FFCEFFFC30E1FA98FE239531" bold="true" box="[151,385,1305,1332]" pageId="6" pageNumber="527">Material examined.</emphasis>
|
||
<materialsCitation id="3B243CDDFFCEFFFC31FFFA9BFE73955D" ID-GBIF-Occurrence="2806771304" collectingDate="2017-07-17" collectionCode="ZSI-C" collectorName="S. Mitra" country="India" location="West Bengal" pageId="6" pageNumber="527" specimenCode="ZSI-C7125" specimenCount="1" stateProvince="West Bengal" typeStatus="holotype">
|
||
<typeStatus id="54F78822FFCEFFFC31FFFA9BFE569531" box="[393,500,1306,1332]" pageId="6" pageNumber="527" type="holotype">Holotype</typeStatus>
|
||
, male, CW
|
||
<quantity id="4CB49B65FFCEFFFC320CFA9BFD719530" box="[634,723,1306,1333]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="6" pageNumber="527" unit="mm" value="7.9">7.9 mm</quantity>
|
||
, CL
|
||
<quantity id="4CB49B65FFCEFFFC337CFA98FCC29530" box="[778,864,1305,1333]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="6" pageNumber="527" unit="mm" value="5.9">5.9 mm</quantity>
|
||
(
|
||
<specimenCode id="DBEA9EFBFFCEFFFC3306FA98FC519531" box="[880,1011,1305,1332]" collectionCode="ZSI-C" pageId="6" pageNumber="527">ZSI-C7125</specimenCode>
|
||
/2); Digha Beach,
|
||
<collectingRegion id="4988F862FFCEFFFC34C8FA9AFAF29531" box="[1214,1360,1306,1333]" country="India" name="West Bengal" pageId="6" pageNumber="527">West Bengal</collectingRegion>
|
||
,
|
||
<collectingCountry id="F35B7610FFCEFFFC352AFA9BFA3A9531" box="[1372,1432,1306,1332]" name="India" pageId="6" pageNumber="527">India</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26B95356FFCEFFFC30B9FABFFE8C955D" box="[207,302,1342,1368]" pageId="6" pageNumber="527">S. Mitra</collectorName>
|
||
,
|
||
<date id="FFF21040FFCEFFFC314FFABFFE6C955D" box="[313,462,1342,1368]" pageId="6" pageNumber="527" value="2017-07-17">
|
||
<collectingDate id="EFB6E9A8FFCEFFFC314FFABFFE6C955D" box="[313,462,1342,1368]" pageId="6" pageNumber="527" value="2017-07-17">17 July 2017</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFCEFFFC30B1FAE2FD2195A5" blockId="6.[151,1437,1305,2017]" pageId="6" pageNumber="527">
|
||
<typeStatus id="54F78822FFCEFFFC30B1FAE2FE9A9578" box="[199,312,1379,1405]" pageId="6" pageNumber="527" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D4AFD09FFCEFFFC3133FAE3FE0D9579" box="[325,431,1378,1404]" pageId="6" pageNumber="527" type="male">18 males</specimenCount>
|
||
, CW 6.4–8.0 mm, CL
|
||
<quantity id="4CB49B65FFCEFFFC32CFFAE0FCE69578" box="[697,836,1377,1405]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="6.1" metricValueMin="5.1" pageId="6" pageNumber="527" unit="mm" value="5.6" valueMax="6.1" valueMin="5.1">5.1–6.1 mm</quantity>
|
||
,
|
||
<materialsCitation id="3B243CDDFFCEFFFC3324FAE2FDDD95A5" ID-GBIF-Occurrence="2806771305" collectingDate="2017-07-17" collectorName="S. Mitra" country="India" location="West Bengal" pageId="6" pageNumber="527" specimenCount="7" specimenCount-female="7" stateProvince="West Bengal" typeStatus="paratype">
|
||
<specimenCount id="9D4AFD09FFCEFFFC3324FAE2FC669579" box="[850,964,1378,1405]" pageId="6" pageNumber="527" type="female">7 females</specimenCount>
|
||
, CW
|
||
<quantity id="4CB49B65FFCEFFFC3471FAE3FB319578" box="[1031,1171,1378,1405]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="7.9" metricValueMin="6.2" pageId="6" pageNumber="527" unit="mm" value="7.05" valueMax="7.9" valueMin="6.2">6.2–7.9 mm</quantity>
|
||
, CL 5.0–
|
||
<quantity id="4CB49B65FFCEFFFC3489FAE0FAF59578" box="[1279,1367,1377,1405]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="6" pageNumber="527" unit="mm" value="5.9">5.9 mm</quantity>
|
||
(ZSI- C7126/2), collection data same as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356650BFFCEFFF230B1FA28FB409386" lastPageId="8" lastPageNumber="529" pageId="6" pageNumber="527" type="description">
|
||
<paragraph id="8BF33680FFCEFFFC30B1FA28FDB29754" blockId="6.[151,1437,1305,2017]" pageId="6" pageNumber="527">
|
||
<emphasis id="B938EA92FFCEFFFC30B1FA28FEFB95C1" bold="true" box="[199,345,1449,1476]" pageId="6" pageNumber="527">Description.</emphasis>
|
||
Carapace (
|
||
<figureCitation id="13772A05FFCEFFFC31AEFA2BFD9295C0" box="[472,560,1450,1477]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="6" pageNumber="527">Fig. 6A</figureCitation>
|
||
) subtrapezoidal, width 1.2–1.3 times length, anterolateral margins convergent towards anterior. Dorsal surface areolate, strongly sculptured, with several sulci/grooves; deep median groove running from front to posterior margin. Gastric region with four small but distinct lobules more or less centrally located, anterior to these and within median groove is a smaller, fainter 5
|
||
<superScript id="7C399BC8FFCEFFFC33F5F994FC329626" attach="left" box="[899,912,1557,1571]" fontSize="6" pageId="6" pageNumber="527">th</superScript>
|
||
lobule. Several grooves present. One oblique groove runs from mesogastric region towards external orbital angle; this groove connects with longer oblique groove at mesogastric region, which runs from there towards posterolateral angle; mesially, a third, shorter groove originates from cardiac region and also runs towards posterolateral angle. Lateral to these a near-vertical groove runs parallel to anterolateral margin of carapace, from just beneath anterolateral angle toward posterolateral margin, this groove does not bifurcate at its anterior end. Transverse groove present close to posterior margin, interrupted medially by areola of intestinal region; two parallel grooves run from this posterior groove to edge of gastric region, marking lateral edges of cardio-intestinal region. Side (branchial) wall of carapace with one longitudinal and two, much shorter, transverse grooves.
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFCEFFFD30B1F8DAFCAE90D2" blockId="6.[151,1437,1305,2017]" lastBlockId="7.[151,1437,153,467]" lastPageId="7" lastPageNumber="528" pageId="6" pageNumber="527">
|
||
Front relatively narrow (
|
||
<figureCitation id="13772A05FFCEFFFC31A1F8DBFD8C9770" box="[471,558,1882,1909]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="6" pageNumber="527">Fig. 6B</figureCitation>
|
||
), deflexed ventrally, appearing bilobate from dorsal view but margin entire and triangular in shape in frontal view. Supraorbital margins, granulate, continuing smoothly from base of front toward external orbital angle, which terminates in a small, acute, laterally directed tooth; this tooth separated from anterolateral margin of carapace by wide V-shaped indentation. Anterolateral margins subparallel, convergent anteriorly, slightly convex, much longer than posterolateral margins; posterolateral margins short, deeply concave; posterior margin slightly sinuous, central region slightly concave.
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFCFFFFD30B1FF60FBBA91D6" blockId="7.[151,1437,153,467]" pageId="7" pageNumber="528">
|
||
Eyes well developed (
|
||
<figureCitation id="13772A05FFCFFFFD31B3FF60FDBD90F9" box="[453,543,225,252]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="7" pageNumber="528">Fig. 6B</figureCitation>
|
||
), peduncles long, corneas bulging. Antennule with large basal article occupying most of antennular fossa, distal articles much smaller. Antenna small, tip of flagellum not reaching tip of eyes. Epistome well developed, central region broadly triangular. Immediately subjacent edges of pterygostome perpendicular to posterior border of epistome. Second maxilliped with penultimate article of endopod much expanded, Spoon tipped setae (
|
||
<figureCitation id="13772A05FFCFFFFD31F7FEF0FE7E918E" box="[385,476,369,395]" captionStart="FIGURE 7" captionStartId="9.[151,250,1809,1834]" captionTargetBox="[162,1423,219,1699]" captionTargetId="figure@9.[151,1436,211,1708]" captionTargetPageId="9" captionText="FIGURE 7. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal.A, pleon; D, spoon tipped seta of inner margin of merus of second maxilliped. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, pleon; E, spoon tipped setae of inner margin of merus of second maxilliped. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. C, pleon." figureDoi="http://doi.org/10.5281/zenodo.3936901" httpUri="https://zenodo.org/record/3936901/files/figure.png" pageId="7" pageNumber="528">Fig. 7D</figureCitation>
|
||
) on merus of second maxilliped six-tined at its distal end with bifurcate peduncles. Third maxillipeds large, leaving a slight median gap when closed together; merus larger than ischium, with inverted V-shaped sulcus; palp flattened; small, thin exopod concealed beneath endopod.
|
||
</paragraph>
|
||
<caption id="DF336608FFCFFFFD30E1F829FB1997E0" ID-DOI="http://doi.org/10.5281/zenodo.3936899" ID-Zenodo-Dep="3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="7" pageNumber="528" startId="7.[151,250,1960,1985]" targetBox="[165,1422,489,1935]" targetPageId="7">
|
||
<paragraph id="8BF33680FFCFFFFD30E1F829FB1997E0" blockId="7.[151,1436,1960,2021]" pageId="7" pageNumber="528">
|
||
<emphasis id="B938EA92FFCFFFFD30E1F829FEB197C4" bold="true" box="[151,275,1960,1985]" pageId="7" pageNumber="528">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C4C4D03FFCFFFFD316CF828FE7197C4" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[282,467,1961,1985]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFCFFFFD316CF828FE7197C4" box="[282,467,1961,1985]" italics="true" pageId="7" pageNumber="528">Dotilla fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFCFFFFD31AFF828FDB197C4" bold="true" box="[473,531,1961,1985]" pageId="7" pageNumber="528">
|
||
<taxonomicNameLabel id="A20B57E9FFCFFFFD31AFF828FDB197C4" box="[473,531,1961,1985]" pageId="7" pageNumber="528" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF33680FFC0FFF230B1FF18FA2D90FE" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
Thoracic sternum (
|
||
<figureCitation id="13772A05FFC0FFF231EBFF18FE5790B1" box="[413,501,153,180]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="8" pageNumber="529">Fig. 6D</figureCitation>
|
||
) without tympana; immediately beneath anterior margin of sternite 4 two large triangular teeth present, positioned on either side of sternite 4 and anterior to tip of sternoabdominal cavity. Sternal tubercle of sternopleonal locking mechanism located on anterior part of sternite 5 just next to suture with sternite 4.
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC0FFF230B1FE84FD6391D6" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
Chelipeds (P1) (
|
||
<figureCitation id="13772A05FFC0FFF231F5FE84FE7C9125" box="[387,478,261,288]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="8" pageNumber="529">Fig. 6A</figureCitation>
|
||
) symmetrical, homochelous. Merus with oval tympanum on dorsal surface. Carpus ovate, inflated, external surface finely granulate. Palm of chela (
|
||
<figureCitation id="13772A05FFC0FFF233F3FEA8FC7D9141" box="[901,991,297,324]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="8" pageNumber="529">Fig. 6C</figureCitation>
|
||
) subrectangular, inflated, superior and inferior margins convex, external surfaces finely granulate; a distinct ridge of small granules positioned on palm external surface next to inferior margin, continuing into and running along most of the length of fixed finger. Fingers relatively straight, distinctly longer than palm, cutting margins with small teeth but never with pronounced lobes; dactylus with granulate ridge on external surface.
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC0FFF230B1FE5CFF40923A" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
Ambulatory legs (
|
||
<figureCitation id="13772A05FFC0FFF231EEFE5CFE5691FD" box="[408,500,477,504]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="8" pageNumber="529">Fig. 6A</figureCitation>
|
||
) (P2–P5) relatively stout, moderately long. Meri with oval tympana on both flat surfaces. P5 dactylus slender, with row of long stiff setae on dorsal margin; approximately twice the length of propodus.
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC0FFF230B1FDC8FCC992CA" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
Male pleon (
|
||
<figureCitation id="13772A05FFC0FFF2312FFDC8FE1D9266" box="[345,447,585,611]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="8" pageNumber="529">Figs. 6D</figureCitation>
|
||
,
|
||
<figureCitation id="13772A05FFC0FFF231BBFDC8FE539266" box="[461,497,585,611]" captionStart="FIGURE 7" captionStartId="9.[151,250,1809,1834]" captionTargetBox="[162,1423,219,1699]" captionTargetId="figure@9.[151,1436,211,1708]" captionTargetPageId="9" captionText="FIGURE 7. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal.A, pleon; D, spoon tipped seta of inner margin of merus of second maxilliped. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, pleon; E, spoon tipped setae of inner margin of merus of second maxilliped. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. C, pleon." figureDoi="http://doi.org/10.5281/zenodo.3936901" httpUri="https://zenodo.org/record/3936901/files/figure.png" pageId="8" pageNumber="529">7A</figureCitation>
|
||
) with six freely articulated somites (pleomeres) and telson. Anterior margin of 4
|
||
<superScript id="7C399BC8FFC0FFF235E6FDC9FA3F9253" attach="right" box="[1424,1437,584,598]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
|
||
pleomere, slightly concave, overlapping 5
|
||
<superScript id="7C399BC8FFC0FFF2321DFDEDFDDA927F" attach="left" box="[619,632,620,634]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
|
||
pleomere with band of stiff setae on extending along the whole width of anterior margin. Telson with broadly rounded tip, slightly convex lateral margins. Male and female pleons similarly narrow and triangular, with the female pleon only slightly wider.
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC0FFF230B1FD58FEDE933E" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
G1 (
|
||
<figureCitation id="13772A05FFC0FFF2308FFD58FEF392F6" box="[249,337,729,755]" captionStart="FIGURE 8" captionStartId="10.[151,250,1806,1831]" captionTargetBox="[161,1428,388,1776]" captionTargetId="figure@10.[151,1436,381,1782]" captionTargetPageId="10" captionText="FIGURE 8. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2). A, left G1; E, left G2. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, left G1; F, left G2. Dotilla intermedia De Man, 1888 paralectotype, male (CW 4.2 mm, CL 3.5 mm) (ZSI-C8236/6), Sullivan I., Mergui Archipelago. C, left G1. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. D, left G1; G, left G2." figureDoi="http://doi.org/10.5281/zenodo.3936903" httpUri="https://zenodo.org/record/3936903/files/figure.png" pageId="8" pageNumber="529">Fig. 8A</figureCitation>
|
||
) slender, sinuous, somewhat S-shaped, tapering towards distal tip; distal tip mostly without setae, row of stiff setae arranged sparsely along lateral and mesial margins on distal two-thirds. G2 (
|
||
<figureCitation id="13772A05FFC0FFF234B3FD7CFAB99312" box="[1221,1307,765,791]" captionStart="FIGURE 8" captionStartId="10.[151,250,1806,1831]" captionTargetBox="[161,1428,388,1776]" captionTargetId="figure@10.[151,1436,381,1782]" captionTargetPageId="10" captionText="FIGURE 8. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2). A, left G1; E, left G2. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, left G1; F, left G2. Dotilla intermedia De Man, 1888 paralectotype, male (CW 4.2 mm, CL 3.5 mm) (ZSI-C8236/6), Sullivan I., Mergui Archipelago. C, left G1. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. D, left G1; G, left G2." figureDoi="http://doi.org/10.5281/zenodo.3936903" httpUri="https://zenodo.org/record/3936903/files/figure.png" pageId="8" pageNumber="529">Fig. 8E</figureCitation>
|
||
) stout with rounded apical lobe.
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC0FFF230B1FCC5FB409386" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
<emphasis id="B938EA92FFC0FFF230B1FCC5FEE3935A" bold="true" box="[199,321,836,863]" pageId="8" pageNumber="529">Variation:</emphasis>
|
||
Two large triangular teeth present on the anterior margin of sternite 4 are present in the larger adult males (CW at least 7.0 mm) but such structures are not observed in smaller males (CW <7.0 mm).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356650BFFC0FFF230B1FC0CFC0393CE" pageId="8" pageNumber="529" type="etymology">
|
||
<paragraph id="8BF33680FFC0FFF230B1FC0CFC0393CE" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
<emphasis id="B938EA92FFC0FFF230B1FC0CFEEC93A2" bold="true" box="[199,334,909,935]" pageId="8" pageNumber="529">Etymology.</emphasis>
|
||
The specific epithet,
|
||
<taxonomicName id="4C4C4D03FFC0FFF2324AFC0CFD1093A2" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[572,690,909,935]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="fraternalis">
|
||
<emphasis id="B938EA92FFC0FFF2324AFC0CFD1093A2" box="[572,690,909,935]" italics="true" pageId="8" pageNumber="529">fraternalis</emphasis>
|
||
</taxonomicName>
|
||
(Latin, adj., pertaining to a brother) alludes to the initial conflation of this species with its congener,
|
||
<taxonomicName id="4C4C4D03FFC0FFF2327DFC33FD0493CE" authorityName="De Man" authorityYear="1888" box="[523,678,945,971]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC0FFF2327DFC33FD0493CE" box="[523,678,945,971]" italics="true" pageId="8" pageNumber="529">D. intermedia</emphasis>
|
||
</taxonomicName>
|
||
. Used as an adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356650BFFC0FFF130B1FC54FBC792A9" lastPageId="11" lastPageNumber="532" pageId="8" pageNumber="529" type="discussion">
|
||
<paragraph id="8BF33680FFC0FFF230B1FC54FEFB950A" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
<emphasis id="B938EA92FFC0FFF230B1FC54FE9993EA" bold="true" box="[199,315,981,1007]" pageId="8" pageNumber="529">Remarks.</emphasis>
|
||
<taxonomicName id="4C4C4D03FFC0FFF23133FC54FDB793EA" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[325,533,981,1007]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC0FFF23133FC54FDB793EA" box="[325,533,981,1007]" italics="true" pageId="8" pageNumber="529">Dotilla fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC0FFF23269FC54FDC393EA" bold="true" box="[543,609,981,1007]" pageId="8" pageNumber="529">
|
||
<taxonomicNameLabel id="A20B57E9FFC0FFF23269FC54FDC393EA" box="[543,609,981,1007]" pageId="8" pageNumber="529" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
basically corresponds to the “low form” of
|
||
<taxonomicName id="4C4C4D03FFC0FFF23410FC57FF469416" authority="sensu Kemp (1919)" authorityName="sensu Kemp" authorityYear="1919" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC0FFF23410FC57FAEF93EA" box="[1126,1357,981,1007]" italics="true" pageId="8" pageNumber="529">D. intermedia sensu</emphasis>
|
||
<bibRefCitation id="EFDD4B71FFC0FFF2352EFC54FF469416" author="Kemp, S." pageId="8" pageNumber="529" pagination="305 - 348" refId="ref7192" refString="Kemp, S. (1919) Notes on Crustacea Decapoda in the Indian Museum. XII. Scopimerinae. Records of the Indian Museum, 16, 305 - 348. https: // doi. org / 10.5962 / bhl. part. 25926" type="journal article" year="1919">Kemp (1919)</bibRefCitation>
|
||
</taxonomicName>
|
||
, who provided the following distinguishing morphological characters: “In the ‘low’ male, the first abdominal sternum [= thoracic sternite 4] bears anteriorly a pair of large outstanding triangular teeth; these are in advance of the limit of the abdominal trough [= sternopleonal cavity] and are thus placed considerably further forward than the ridges in the ‘high’ male. The dactylus of the chela bears a low rounded lobe near the base of its prehensile edge (further back than in the ‘high’ male) and there is no lobe or large tooth on the fixed finger. The copulatory appendage [= G1] is more slender, strongly sinuous, and terminates in a fine point which is turned inwards and does not bear conspicuous setae.” (
|
||
<bibRefCitation id="EFDD4B71FFC0FFF231C9FB50FD2194EE" author="Kemp, S." box="[447,643,1233,1259]" pageId="8" pageNumber="529" pagination="305 - 348" refId="ref7192" refString="Kemp, S. (1919) Notes on Crustacea Decapoda in the Indian Museum. XII. Scopimerinae. Records of the Indian Museum, 16, 305 - 348. https: // doi. org / 10.5962 / bhl. part. 25926" type="journal article" year="1919">Kemp 1919: 331</bibRefCitation>
|
||
, figs. 10b, c, f).The present material agrees with this description (see
|
||
<figureCitation id="13772A05FFC0FFF230E1FB74FEB9950A" box="[151,283,1269,1295]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="8" pageNumber="529">Figs. 6C, D</figureCitation>
|
||
,
|
||
<figureCitation id="13772A05FFC0FFF23150FB74FEEE950A" box="[294,332,1269,1295]" captionStart="FIGURE 8" captionStartId="10.[151,250,1806,1831]" captionTargetBox="[161,1428,388,1776]" captionTargetId="figure@10.[151,1436,381,1782]" captionTargetPageId="10" captionText="FIGURE 8. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2). A, left G1; E, left G2. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, left G1; F, left G2. Dotilla intermedia De Man, 1888 paralectotype, male (CW 4.2 mm, CL 3.5 mm) (ZSI-C8236/6), Sullivan I., Mergui Archipelago. C, left G1. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. D, left G1; G, left G2." figureDoi="http://doi.org/10.5281/zenodo.3936903" httpUri="https://zenodo.org/record/3936903/files/figure.png" pageId="8" pageNumber="529">8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC0FFF230B1FA98FE23962A" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
Aside from these differences,
|
||
<taxonomicName id="4C4C4D03FFC0FFF23263FA9BFD0E9536" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[533,684,1305,1331]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC0FFF23263FA9BFD0E9536" box="[533,684,1305,1331]" italics="true" pageId="8" pageNumber="529">D. fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC0FFF232C5FA98FD539536" bold="true" box="[691,753,1305,1331]" pageId="8" pageNumber="529">
|
||
<taxonomicNameLabel id="A20B57E9FFC0FFF232C5FA98FD539536" box="[691,753,1305,1331]" pageId="8" pageNumber="529" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can further be distinguished from
|
||
<emphasis id="B938EA92FFC0FFF23406FA9BFAEE9536" box="[1136,1356,1305,1331]" italics="true" pageId="8" pageNumber="529">
|
||
<taxonomicName id="4C4C4D03FFC0FFF23406FA9BFAA99536" authorityName="sensu Kemp" authorityYear="1919" box="[1136,1291,1305,1331]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" sensu="stricto" species="intermedia">D. intermedia</taxonomicName>
|
||
<taxonomicNameLabel id="A20B57E9FFC0FFF23567FA9BFAEE9536" box="[1297,1356,1306,1331]" pageId="8" pageNumber="529" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by: (1) the abruptly perpendicular orientation of the pterygostome margin immediately subjacent to the posterior margin of the epistome (
|
||
<figureCitation id="13772A05FFC0FFF23120FAE0FE0F9579" box="[342,429,1377,1404]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="8" pageNumber="529">Fig. 6B</figureCitation>
|
||
) (
|
||
<emphasis id="B938EA92FFC0FFF231B1FAE3FDAD957E" box="[455,527,1378,1403]" italics="true" pageId="8" pageNumber="529">versus</emphasis>
|
||
gradually oblique in
|
||
<taxonomicName id="4C4C4D03FFC0FFF23374FAE3FC3C957E" authorityName="sensu Kemp" authorityYear="1919" box="[770,926,1377,1403]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC0FFF23374FAE3FC3C957E" box="[770,926,1377,1403]" italics="true" pageId="8" pageNumber="529">D. intermedia</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="13772A05FFC0FFF233D8FAE0FBB2957E" box="[942,1040,1377,1403]" captionStart="FIGURE 1" captionStartId="2.[151,250,1495,1520]" captionTargetBox="[154,1433,181,1470]" captionTargetId="figure@2.[154,1433,181,1470]" captionTargetPageId="2" captionText="FIGURE 1. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal.A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936889" httpUri="https://zenodo.org/record/3936889/files/figure.png" pageId="8" pageNumber="529">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="13772A05FFC0FFF2346BFAE0FBE2957E" box="[1053,1088,1377,1403]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1991]" captionTargetBox="[158,1430,520,1940]" captionTargetId="figure@3.[158,1430,520,1940]" captionTargetPageId="3" captionText="FIGURE 2. Dotilla intermedia De Man, 1888 paralectotype, male (CW 4.2 mm, CL 3.5 mm) (ZSI-C8236/6), Sullivan I., Mergui Archipelago.A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936891" httpUri="https://zenodo.org/record/3936891/files/figure.png" pageId="8" pageNumber="529">2B</figureCitation>
|
||
,
|
||
<figureCitation id="13772A05FFC0FFF2343BFAE0FBD2957E" box="[1101,1136,1377,1403]" captionStart="FIGURE 3" captionStartId="4.[151,248,1608,1633]" captionTargetBox="[161,1426,181,1581]" captionTargetId="figure@4.[161,1426,181,1581]" captionTargetPageId="4" captionText="FIGURE 3. Dotilla intermedia De Man, 1888 paralectotype, female (CW 4.3 mm, CL 3.6 mm) (ZSI-C8236/6), Sullivan I., Mergui Archipelago.A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936893" httpUri="https://zenodo.org/record/3936893/files/figure.png" pageId="8" pageNumber="529">3B</figureCitation>
|
||
); (2) the male telson has a relatively broader tip and slightly convex lateral margins (
|
||
<figureCitation id="13772A05FFC0FFF2335CFA04FC2D959A" box="[810,911,1413,1439]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="8" pageNumber="529">Figs. 6D</figureCitation>
|
||
,
|
||
<figureCitation id="13772A05FFC0FFF233EAFA04FC1D959A" box="[924,959,1413,1439]" captionStart="FIGURE 7" captionStartId="9.[151,250,1809,1834]" captionTargetBox="[162,1423,219,1699]" captionTargetId="figure@9.[151,1436,211,1708]" captionTargetPageId="9" captionText="FIGURE 7. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal.A, pleon; D, spoon tipped seta of inner margin of merus of second maxilliped. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, pleon; E, spoon tipped setae of inner margin of merus of second maxilliped. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. C, pleon." figureDoi="http://doi.org/10.5281/zenodo.3936901" httpUri="https://zenodo.org/record/3936901/files/figure.png" pageId="8" pageNumber="529">7A</figureCitation>
|
||
) (
|
||
<emphasis id="B938EA92FFC0FFF233AFFA07FB83959A" box="[985,1057,1414,1439]" italics="true" pageId="8" pageNumber="529">versus</emphasis>
|
||
tip of telson narrower and lateral margins slightly concave in
|
||
<taxonomicName id="4C4C4D03FFC0FFF231AEFA2BFDD795C6" authorityName="sensu Kemp" authorityYear="1919" box="[472,629,1449,1475]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC0FFF231AEFA2BFDD795C6" box="[472,629,1449,1475]" italics="true" pageId="8" pageNumber="529">D. intermedia</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="13772A05FFC0FFF232F0FA28FD4B95C6" box="[646,745,1449,1475]" captionStart="FIGURE 1" captionStartId="2.[151,250,1495,1520]" captionTargetBox="[154,1433,181,1470]" captionTargetId="figure@2.[154,1433,181,1470]" captionTargetPageId="2" captionText="FIGURE 1. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal.A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936889" httpUri="https://zenodo.org/record/3936889/files/figure.png" pageId="8" pageNumber="529">Figs. 1D</figureCitation>
|
||
,
|
||
<figureCitation id="13772A05FFC0FFF23281FA28FCBE95C6" box="[759,796,1449,1475]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1991]" captionTargetBox="[158,1430,520,1940]" captionTargetId="figure@3.[158,1430,520,1940]" captionTargetPageId="3" captionText="FIGURE 2. Dotilla intermedia De Man, 1888 paralectotype, male (CW 4.2 mm, CL 3.5 mm) (ZSI-C8236/6), Sullivan I., Mergui Archipelago.A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936891" httpUri="https://zenodo.org/record/3936891/files/figure.png" pageId="8" pageNumber="529">2D</figureCitation>
|
||
,
|
||
<figureCitation id="13772A05FFC0FFF2335FFA28FCEE95C6" box="[809,844,1449,1475]" captionStart="FIGURE 7" captionStartId="9.[151,250,1809,1834]" captionTargetBox="[162,1423,219,1699]" captionTargetId="figure@9.[151,1436,211,1708]" captionTargetPageId="9" captionText="FIGURE 7. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal.A, pleon; D, spoon tipped seta of inner margin of merus of second maxilliped. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, pleon; E, spoon tipped setae of inner margin of merus of second maxilliped. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. C, pleon." figureDoi="http://doi.org/10.5281/zenodo.3936901" httpUri="https://zenodo.org/record/3936901/files/figure.png" pageId="8" pageNumber="529">7B</figureCitation>
|
||
; cf.
|
||
<bibRefCitation id="EFDD4B71FFC0FFF23309FA28FBAA95C6" author="Kemp, S." box="[895,1032,1449,1475]" pageId="8" pageNumber="529" pagination="305 - 348" refId="ref7192" refString="Kemp, S. (1919) Notes on Crustacea Decapoda in the Indian Museum. XII. Scopimerinae. Records of the Indian Museum, 16, 305 - 348. https: // doi. org / 10.5962 / bhl. part. 25926" type="journal article" year="1919">Kemp 1919</bibRefCitation>
|
||
: fig. 10e); and (3) the spoon-tipped setae on the merus of the second maxilliped are six-tined at their distal end, and are with bifurcate peduncles (
|
||
<figureCitation id="13772A05FFC0FFF23519FA4CFF19960E" captionStart="FIGURE 7" captionStartId="9.[151,250,1809,1834]" captionTargetBox="[162,1423,219,1699]" captionTargetId="figure@9.[151,1436,211,1708]" captionTargetPageId="9" captionText="FIGURE 7. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal.A, pleon; D, spoon tipped seta of inner margin of merus of second maxilliped. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, pleon; E, spoon tipped setae of inner margin of merus of second maxilliped. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. C, pleon." figureDoi="http://doi.org/10.5281/zenodo.3936901" httpUri="https://zenodo.org/record/3936901/files/figure.png" pageId="8" pageNumber="529">Fig. 7D</figureCitation>
|
||
) (
|
||
<emphasis id="B938EA92FFC0FFF230A2FA73FEBE960E" box="[212,284,1522,1547]" italics="true" pageId="8" pageNumber="529">versus</emphasis>
|
||
spoon-tipped setae on merus of second maxilliped five-tined at distal end, with simple peduncle in
|
||
<taxonomicName id="4C4C4D03FFC0FFF235F7FA73FEB2962A" authorityName="sensu Kemp" authorityYear="1919" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC0FFF235F7FA73FEB2962A" italics="true" pageId="8" pageNumber="529">D. intermedia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="13772A05FFC0FFF23168F994FED7962A" box="[286,373,1557,1583]" captionStart="FIGURE 7" captionStartId="9.[151,250,1809,1834]" captionTargetBox="[162,1423,219,1699]" captionTargetId="figure@9.[151,1436,211,1708]" captionTargetPageId="9" captionText="FIGURE 7. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal.A, pleon; D, spoon tipped seta of inner margin of merus of second maxilliped. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, pleon; E, spoon tipped setae of inner margin of merus of second maxilliped. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. C, pleon." figureDoi="http://doi.org/10.5281/zenodo.3936901" httpUri="https://zenodo.org/record/3936901/files/figure.png" pageId="8" pageNumber="529">Fig. 7E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC0FFF230B1F9B8FAD29776" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
The statement by
|
||
<bibRefCitation id="EFDD4B71FFC0FFF231F9F9B8FD869656" author="Kemp, S." box="[399,548,1593,1619]" pageId="8" pageNumber="529" pagination="305 - 348" refId="ref7192" refString="Kemp, S. (1919) Notes on Crustacea Decapoda in the Indian Museum. XII. Scopimerinae. Records of the Indian Museum, 16, 305 - 348. https: // doi. org / 10.5962 / bhl. part. 25926" type="journal article" year="1919">Kemp (1919)</bibRefCitation>
|
||
that his “high” and “low” forms are conspecific is not supported by the present study. He used as basis for this pronouncement merely the similarity in the sculpturing of their dorsal carapace and their co-occurrence in the same general habitat. He considered the stark differences in the G1 morphology as an ultimately physiological difference between breeding (high form) and non-breeding (low form) males. There is, as discussed earlier in the remarks for both species, ample morphological evidence that suggests that the ‘low’ and ‘high’ forms are distinct species. Foremost among these are the differences in the armature of the male sternite 4, and the drastic differences in the morphology of the G1, of which the latter is rather suggestive of reproductive isolation, and which in turn is usually a reliable character for the discrimination of species (
|
||
<bibRefCitation id="EFDD4B71FFC0FFF23463F8B4FB07974A" author="Guinot, D." box="[1045,1189,1845,1871]" pageId="8" pageNumber="529" pagination="1 - 354" refId="ref7058" refString="Guinot, D. (1979) Morphologie et phylogenese des brachyoures. Memoires du Museum national d'Histoire naturelle, Series A, 112, 1 - 354." type="journal article" year="1979">Guinot 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDD4B71FFC0FFF234C6F8B4FAED974A" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[1200,1359,1845,1871]" pageId="8" pageNumber="529" pagination="1 - 286" refId="ref7269" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part 1. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
|
||
Ng
|
||
<emphasis id="B938EA92FFC0FFF234AEF8B7FAAD974A" box="[1240,1295,1845,1871]" italics="true" pageId="8" pageNumber="529">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDD4B71FFC0FFF2352FF8B4FEBF9776" author="Davie, P. J. F. & Guinot, D. & Ng, P. K. L." pageId="8" pageNumber="529" pagination="11 - 163" refId="ref6079" refString="Davie, P. J. F., Guinot, D. & Ng, P. K. L. (2015 a) Anatomy and functional morphology of Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. & Von Vaupel Klein, C. (Eds.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Crustacea. Vol. 9 C-I, Decapoda: Brachyura. Part 1. Brill, Leiden, pp. 11 - 163. https: // doi. org / 10.1163 / 9789004190832 _ 004" type="book chapter" year="2015">
|
||
Davie
|
||
<emphasis id="B938EA92FFC0FFF230E1F8DBFF6A9776" box="[151,200,1881,1907]" italics="true" pageId="8" pageNumber="529">et al</emphasis>
|
||
. 2015a
|
||
</bibRefCitation>
|
||
, b). Further studies on the genetics and behavior of these species should confirm their distinctness.
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC0FFF230B1F8FCFF5297DA" blockId="8.[151,1437,153,2015]" pageId="8" pageNumber="529">
|
||
<bibRefCitation id="EFDD4B71FFC0FFF230B1F8FCFE5B9792" author="Dev Roy, M. K. & Bhadra, S." box="[199,505,1916,1943]" pageId="8" pageNumber="529" pagination="357 - 535" refId="ref6627" refString="Dev Roy, M. K. & Bhadra, S. (2005) Marine and Estuarine crabs. In: Director ZSI, Kolkata (Ed.), Fauna of Andhra Pardesh, State Fauna Series. Part 5. Zoological Survey of India, Kolkata, pp. 357 - 535." type="book chapter" year="2005">Dev Roy & Bhadra (2005)</bibRefCitation>
|
||
reported “high” and “low” forms of
|
||
<taxonomicName id="4C4C4D03FFC0FFF233D5F8FCFBD69792" authorityName="sensu Kemp" authorityYear="1919" box="[931,1140,1917,1943]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC0FFF233D5F8FCFBD69792" box="[931,1140,1917,1943]" italics="true" pageId="8" pageNumber="529">Dotilla intermedia</emphasis>
|
||
</taxonomicName>
|
||
from the coast of
|
||
<collectingRegion id="4988F862FFC0FFF2353EF8FCFF5197BE" country="India" name="Andhra Pradesh" pageId="8" pageNumber="529">Andhra Pradesh</collectingRegion>
|
||
, they distinguished those two ‘forms’ well, but did not formally recognise the “low” form as a separate species.
|
||
</paragraph>
|
||
<caption id="DF336608FFC1FFF330E1F890FF7597BF" ID-DOI="http://doi.org/10.5281/zenodo.3936901" ID-Zenodo-Dep="3936901" httpUri="https://zenodo.org/record/3936901/files/figure.png" pageId="9" pageNumber="530" startId="9.[151,250,1809,1834]" targetBox="[162,1423,219,1699]" targetPageId="9">
|
||
<paragraph id="8BF33680FFC1FFF330E1F890FF7597BF" blockId="9.[151,1437,1809,1978]" pageId="9" pageNumber="530">
|
||
<emphasis id="B938EA92FFC1FFF330E1F890FEB0972F" bold="true" box="[151,274,1809,1834]" pageId="9" pageNumber="530">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C4C4D03FFC1FFF3316EF893FE73972F" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[280,465,1810,1834]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC1FFF3316EF893FE73972F" box="[280,465,1810,1834]" italics="true" pageId="9" pageNumber="530">Dotilla fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC1FFF331A1F893FDB2972F" bold="true" box="[471,528,1810,1834]" pageId="9" pageNumber="530">
|
||
<taxonomicNameLabel id="A20B57E9FFC1FFF331A1F893FDB2972F" box="[471,528,1810,1834]" pageId="9" pageNumber="530" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, pleon; D, spoon tipped seta of inner margin of merus of second maxilliped.
|
||
<taxonomicName id="4C4C4D03FFC1FFF33320F8B7FB0E974B" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" box="[854,1196,1846,1870]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC1FFF33320F8B7FBB0974B" box="[854,1042,1846,1870]" italics="true" pageId="9" pageNumber="530">Dotilla intermedia</emphasis>
|
||
<bibRefCitation id="EFDD4B71FFC1FFF3346FF8B7FB0E974B" author="De Man, J. G." box="[1049,1196,1846,1870]" pageId="9" pageNumber="530" pagination="1 - 312" refId="ref6453" refString="De Man, J. G. (1888) Report of the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, FRS, Superintendent of the Museum, Journal of the Linnaean Society, London, Zoology, 22, 1 - 312. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1888">De Man, 1888</bibRefCitation>
|
||
</taxonomicName>
|
||
male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, pleon; E, spoon tipped setae of inner margin of merus of second maxilliped.
|
||
<taxonomicName id="4C4C4D03FFC1FFF330E1F8FFFD909793" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[151,562,1917,1942]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="clepsydrodactylus">
|
||
<emphasis id="B938EA92FFC1FFF330E1F8FFFE3F9793" box="[151,413,1918,1942]" italics="true" pageId="9" pageNumber="530">Dotilla clepsydrodactylus</emphasis>
|
||
<bibRefCitation id="EFDD4B71FFC1FFF331D3F8FCFD909793" author="Alcock, A." box="[421,562,1917,1942]" pageId="9" pageNumber="530" pagination="279 - 456" refId="ref5901" refString="Alcock, A. (1900) Materials for carcinological fauna of India, No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, 69, 279 - 456. https: // doi. org / 10.5962 / bhl. title. 15344" type="journal article" year="1900">Alcock, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. C, pleon.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF33680FFC2FFF030B1FF18FA3E90FE" blockId="10.[151,1436,152,359]" pageId="10" pageNumber="531">
|
||
Earlier
|
||
<bibRefCitation id="EFDD4B71FFC2FFF0316EFF18FE1890B6" author="Bairagi, N." box="[280,442,152,179]" pageId="10" pageNumber="531" pagination="263 - 288" refId="ref6028" refString="Bairagi, N. (1995) Ocypodidae: Decapoda: Crustacea. In: Director ZSI, Kolkata (Ed.), Hugli Matla Estuary, West Bengal. Esturaine Ecosystem Series. Part 2. Zoological Survey Of India, Kolkata, pp. 263 - 288." type="book chapter" year="1995">Bairagi (1995)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="4C4C4D03FFC2FFF03250FF1BFD1D90B6" authorityName="De Man" authorityYear="1888" box="[550,703,153,179]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC2FFF03250FF1BFD1D90B6" box="[550,703,153,179]" italics="true" pageId="10" pageNumber="531">D. intermedia</emphasis>
|
||
</taxonomicName>
|
||
from Sagar Island,
|
||
<collectingRegion id="4988F862FFC2FFF033E2FF18FB8690B6" box="[916,1060,153,179]" country="India" name="West Bengal" pageId="10" pageNumber="531">West Bengal</collectingRegion>
|
||
.While he did not use the morphological distinctions of Kemp, he noted that all the specimens possess a pair of large triangular teeth positioned anteriorly on the thoracic sternum, which clearly indicate that the crabs are more properly identified as
|
||
<taxonomicName id="4C4C4D03FFC2FFF034B4FF63FAFB90FE" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[1218,1369,225,251]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC2FFF034B4FF63FAFB90FE" box="[1218,1369,225,251]" italics="true" pageId="10" pageNumber="531">D. fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B57E9FFC2FFF03529FF60FA3E90FE" box="[1375,1436,225,251]" pageId="10" pageNumber="531" rank="species">
|
||
<emphasis id="B938EA92FFC2FFF03529FF60FA3790FE" bold="true" box="[1375,1429,225,251]" pageId="10" pageNumber="531">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC2FFF030B1FE84FE3F9162" blockId="10.[151,1436,152,359]" pageId="10" pageNumber="531">
|
||
According to the observation of
|
||
<bibRefCitation id="EFDD4B71FFC2FFF0325BFE84FD66911A" author="Kemp, S." box="[557,708,261,287]" pageId="10" pageNumber="531" pagination="305 - 348" refId="ref7192" refString="Kemp, S. (1919) Notes on Crustacea Decapoda in the Indian Museum. XII. Scopimerinae. Records of the Indian Museum, 16, 305 - 348. https: // doi. org / 10.5962 / bhl. part. 25926" type="journal article" year="1919">Kemp (1919)</bibRefCitation>
|
||
, the ‘low’ forms should be small in size (carapace length of males rarely exceed
|
||
<quantity id="4CB49B65FFC2FFF03142FEA9FE2C9146" box="[308,398,296,323]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="531" unit="mm" value="4.5">4.5 mm</quantity>
|
||
), but current observations show that males of
|
||
<taxonomicName id="4C4C4D03FFC2FFF033E6FEABFB8A9146" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[912,1064,297,323]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC2FFF033E6FEABFB8A9146" box="[912,1064,297,323]" italics="true" pageId="10" pageNumber="531">D. fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC2FFF03459FEA8FBCC9146" bold="true" box="[1071,1134,297,323]" pageId="10" pageNumber="531">
|
||
<taxonomicNameLabel id="A20B57E9FFC2FFF03459FEA8FBCC9146" box="[1071,1134,297,323]" pageId="10" pageNumber="531" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be larger, reaching
|
||
<quantity id="4CB49B65FFC2FFF0350FFEA8FF619162" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="10" pageNumber="531" unit="mm" value="6.1">6.1 mm</quantity>
|
||
in carapace length.
|
||
</paragraph>
|
||
<caption id="DF336608FFC2FFF030E1F88FFE0497B2" ID-DOI="http://doi.org/10.5281/zenodo.3936903" ID-Zenodo-Dep="3936903" httpUri="https://zenodo.org/record/3936903/files/figure.png" pageId="10" pageNumber="531" startId="10.[151,250,1806,1831]" targetBox="[161,1428,388,1776]" targetPageId="10">
|
||
<paragraph id="8BF33680FFC2FFF030E1F88FFE0497B2" blockId="10.[151,1437,1806,1975]" pageId="10" pageNumber="531">
|
||
<emphasis id="B938EA92FFC2FFF030E1F88FFEB19722" bold="true" box="[151,275,1806,1831]" pageId="10" pageNumber="531">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C4C4D03FFC2FFF0316FF88EFE719722" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[281,467,1807,1831]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC2FFF0316FF88EFE719722" box="[281,467,1807,1831]" italics="true" pageId="10" pageNumber="531">Dotilla fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC2FFF031AFF88EFDB09722" bold="true" box="[473,530,1807,1831]" pageId="10" pageNumber="531">
|
||
<taxonomicNameLabel id="A20B57E9FFC2FFF031AFF88EFDB09722" box="[473,530,1807,1831]" pageId="10" pageNumber="531" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2). A, left G1; E, left G2.
|
||
<taxonomicName id="4C4C4D03FFC2FFF03522F88EFE3D974E" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC2FFF03522F88EFEA7974E" italics="true" pageId="10" pageNumber="531">Dotilla intermedia</emphasis>
|
||
<bibRefCitation id="EFDD4B71FFC2FFF0317AF8B5FE3D974E" author="De Man, J. G." box="[268,415,1843,1867]" pageId="10" pageNumber="531" pagination="1 - 312" refId="ref6453" refString="De Man, J. G. (1888) Report of the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, FRS, Superintendent of the Museum, Journal of the Linnaean Society, London, Zoology, 22, 1 - 312. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1888">De Man, 1888</bibRefCitation>
|
||
</taxonomicName>
|
||
male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, left G1; F, left G2.
|
||
<taxonomicName id="4C4C4D03FFC2FFF03522F8B2FE02976A" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC2FFF03522F8B2FEA7976A" italics="true" pageId="10" pageNumber="531">Dotilla intermedia</emphasis>
|
||
<bibRefCitation id="EFDD4B71FFC2FFF0317AF8D9FE02976A" author="De Man, J. G." box="[268,416,1879,1903]" pageId="10" pageNumber="531" pagination="1 - 312" refId="ref6453" refString="De Man, J. G. (1888) Report of the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, FRS, Superintendent of the Museum, Journal of the Linnaean Society, London, Zoology, 22, 1 - 312. https: // doi. org / 10.1111 / j. 1096 - 3642.1887. tb 00027. x" type="journal article" year="1888">De Man, 1888</bibRefCitation>
|
||
</taxonomicName>
|
||
paralectotype, male (CW 4.2 mm, CL 3.5 mm) (ZSI-C8236/6), Sullivan I., Mergui Archipelago. C, left G1.
|
||
<taxonomicName id="4C4C4D03FFC2FFF0309AF8FAFD209796" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[236,642,1914,1939]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="531" phylum="Arthropoda" rank="species" species="clepsydrodactylus">
|
||
<emphasis id="B938EA92FFC2FFF0309AF8FAFE529796" box="[236,496,1915,1939]" italics="true" pageId="10" pageNumber="531">Dotilla clepsydrodactylus</emphasis>
|
||
<bibRefCitation id="EFDD4B71FFC2FFF03181F8FBFD209796" author="Alcock, A." box="[503,642,1914,1939]" pageId="10" pageNumber="531" pagination="279 - 456" refId="ref5901" refString="Alcock, A. (1900) Materials for carcinological fauna of India, No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, 69, 279 - 456. https: // doi. org / 10.5962 / bhl. title. 15344" type="journal article" year="1900">Alcock, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. D, left G1; G, left G2.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF33680FFC3FFF130B1FF18FC3E921E" blockId="11.[151,1437,153,684]" pageId="11" pageNumber="532">
|
||
<taxonomicName id="4C4C4D03FFC3FFF130B1FF18FE3090B6" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[199,402,153,179]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC3FFF130B1FF18FE3090B6" box="[199,402,153,179]" italics="true" pageId="11" pageNumber="532">Dotilla fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC3FFF131EEFF18FE7790B6" bold="true" box="[408,469,153,179]" pageId="11" pageNumber="532">
|
||
<taxonomicNameLabel id="A20B57E9FFC3FFF131EEFF18FE7790B6" box="[408,469,153,179]" pageId="11" pageNumber="532" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also has some morphological similarities with
|
||
<taxonomicName id="4C4C4D03FFC3FFF133ABFF1BFAA190B6" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[989,1283,153,179]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="blanfordi">
|
||
<emphasis id="B938EA92FFC3FFF133ABFF1BFBC790B6" box="[989,1125,153,179]" italics="true" pageId="11" pageNumber="532">D. blanfordi</emphasis>
|
||
<bibRefCitation id="EFDD4B71FFC3FFF1341CFF18FAA190B6" author="Alcock, A." box="[1130,1283,153,179]" pageId="11" pageNumber="532" pagination="279 - 456" refId="ref5901" refString="Alcock, A. (1900) Materials for carcinological fauna of India, No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, 69, 279 - 456. https: // doi. org / 10.5962 / bhl. title. 15344" type="journal article" year="1900">Alcock, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
, as both those species have 4 lobules on the gastric region of the carapace, a mid-dorsal groove extending from the front to the posterior margin, and a transverse posterior groove present on the posterior region of the carapace, and there is no tooth below the orbital angle. The new species differs from
|
||
<taxonomicName id="4C4C4D03FFC3FFF1335EFE87FC12911A" authorityName="Alcock" authorityYear="1900" box="[808,944,261,287]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="blanfordi">
|
||
<emphasis id="B938EA92FFC3FFF1335EFE87FC12911A" box="[808,944,261,287]" italics="true" pageId="11" pageNumber="532">D. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
in many morphological characters: (1) there is a single oblique groove running along the side of the cardiac region towards posterolateral angle of the carapace in
|
||
<taxonomicName id="4C4C4D03FFC3FFF130C0FECFFEE39162" authorityName="Alcock" authorityYear="1900" box="[182,321,333,359]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="blanfordi">
|
||
<emphasis id="B938EA92FFC3FFF130C0FECFFEE39162" box="[182,321,333,359]" italics="true" pageId="11" pageNumber="532">D. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDD4B71FFC3FFF13127FECCFE4A9162" author="Alcock, A." box="[337,488,333,359]" pageId="11" pageNumber="532" pagination="279 - 456" refId="ref5901" refString="Alcock, A. (1900) Materials for carcinological fauna of India, No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, 69, 279 - 456. https: // doi. org / 10.5962 / bhl. title. 15344" type="journal article" year="1900">Alcock 1900</bibRefCitation>
|
||
: fig. 3;
|
||
<bibRefCitation id="EFDD4B71FFC3FFF13234FECCFD699162" author="Kemp, S." box="[578,715,333,359]" pageId="11" pageNumber="532" pagination="305 - 348" refId="ref7192" refString="Kemp, S. (1919) Notes on Crustacea Decapoda in the Indian Museum. XII. Scopimerinae. Records of the Indian Museum, 16, 305 - 348. https: // doi. org / 10.5962 / bhl. part. 25926" type="journal article" year="1919">Kemp 1919</bibRefCitation>
|
||
: fig. 9e) (
|
||
<emphasis id="B938EA92FFC3FFF1334AFECFFC269162" box="[828,900,334,359]" italics="true" pageId="11" pageNumber="532">versus</emphasis>
|
||
two oblique grooves running along the side of cardiac region towards posterolateral angle of the carapace in
|
||
<taxonomicName id="4C4C4D03FFC3FFF13325FEF3FC4E918E" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[851,1004,369,395]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC3FFF13325FEF3FC4E918E" box="[851,1004,369,395]" italics="true" pageId="11" pageNumber="532">D. fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC3FFF13382FEF0FB91918E" bold="true" box="[1012,1075,369,395]" pageId="11" pageNumber="532">
|
||
<taxonomicNameLabel id="A20B57E9FFC3FFF13382FEF0FB91918E" box="[1012,1075,369,395]" pageId="11" pageNumber="532" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<figureCitation id="13772A05FFC3FFF13435FEF0FB3F9189" box="[1091,1181,369,396]" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="11" pageNumber="532">Fig. 6A</figureCitation>
|
||
); (2) the lower surface of the palm is not carinate (J. Trivedi, pers. obs.) (
|
||
<emphasis id="B938EA92FFC3FFF132B4FE17FCA891AA" box="[706,778,406,431]" italics="true" pageId="11" pageNumber="532">versus</emphasis>
|
||
lower surface of palm carinate in
|
||
<taxonomicName id="4C4C4D03FFC3FFF134F3FE17FABE91AA" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[1157,1308,405,431]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC3FFF134F3FE17FABE91AA" box="[1157,1308,405,431]" italics="true" pageId="11" pageNumber="532">D. fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC3FFF13555FE14FAC391AA" bold="true" box="[1315,1377,405,431]" pageId="11" pageNumber="532">
|
||
<taxonomicNameLabel id="A20B57E9FFC3FFF13555FE14FAC391AA" box="[1315,1377,405,431]" pageId="11" pageNumber="532" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<figureCitation id="13772A05FFC3FFF13506FE14FF1991D1" captionStart="FIGURE 6" captionStartId="7.[151,250,1960,1985]" captionTargetBox="[165,1422,489,1935]" captionTargetId="figure@7.[165,1422,489,1935]" captionTargetPageId="7" captionText="FIGURE 6. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal. A, habitus, dorsal view; B, frontal view; C, right chela, outer view; D, thoracic sternum and pleon, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3936899" httpUri="https://zenodo.org/record/3936899/files/figure.png" pageId="11" pageNumber="532">Fig. 6C</figureCitation>
|
||
); and (3) the spoon-tipped setae on the merus of the second maxilliped are four-tined at their distal end, and are with a simple peduncle (
|
||
<bibRefCitation id="EFDD4B71FFC3FFF131DDFE5CFD9391F2" author="Vogel, F." box="[427,561,477,503]" pageId="11" pageNumber="532" pagination="225 - 234" refId="ref7876" refString="Vogel, F. (1984) Comparative and functional morphology of the spoon-tipped setae on the second maxillipeds in Dotilla Stimpson, 1858 (Decapoda: Brachyura: Ocypodidae) Crustaceana, 47, 225 - 234. https: // doi. org / 10.1163 / 156854084 X 00487" type="journal article" year="1984">Vogel 1984</bibRefCitation>
|
||
: fig. 4) (
|
||
<emphasis id="B938EA92FFC3FFF132E5FE5FFD7991F2" box="[659,731,478,503]" italics="true" pageId="11" pageNumber="532">versus</emphasis>
|
||
spoon-tipped setae on merus of second maxilliped six-tined at distal end, with bifurcate peduncles in
|
||
<taxonomicName id="4C4C4D03FFC3FFF1323EFD83FD42921E" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[584,736,513,539]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC3FFF1323EFD83FD42921E" box="[584,736,513,539]" italics="true" pageId="11" pageNumber="532">D. fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC3FFF1329EFD80FC84921E" bold="true" box="[744,806,513,539]" pageId="11" pageNumber="532">
|
||
<taxonomicNameLabel id="A20B57E9FFC3FFF1329EFD80FC84921E" box="[744,806,513,539]" pageId="11" pageNumber="532" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<figureCitation id="13772A05FFC3FFF13340FD80FC2D921E" box="[822,911,513,539]" captionStart="FIGURE 7" captionStartId="9.[151,250,1809,1834]" captionTargetBox="[162,1423,219,1699]" captionTargetId="figure@9.[151,1436,211,1708]" captionTargetPageId="9" captionText="FIGURE 7. Dotilla fraternalis n. sp., holotype, male (CW 7.9 mm, CL 5.9 mm) (ZSI-C7125/2), Digha, West Bengal.A, pleon; D, spoon tipped seta of inner margin of merus of second maxilliped. Dotilla intermedia De Man, 1888 male (CW 9.6 mm, CL 7.9 mm) (ZSI-C7963/2), Digha, West Bengal. B, pleon; E, spoon tipped setae of inner margin of merus of second maxilliped. Dotilla clepsydrodactylus Alcock, 1900, lectotype, male (CW 6.5 mm, CL 5.0 mm) (ZSI 438–45/7), False Point, Odisha. C, pleon." figureDoi="http://doi.org/10.5281/zenodo.3936901" httpUri="https://zenodo.org/record/3936901/files/figure.png" pageId="11" pageNumber="532">Fig. 7D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF33680FFC3FFF130B1FDA4FBC792A9" blockId="11.[151,1437,153,684]" pageId="11" pageNumber="532">
|
||
<taxonomicName id="4C4C4D03FFC3FFF130B1FDA4FE37923A" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[199,405,549,575]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC3FFF130B1FDA4FE37923A" box="[199,405,549,575]" italics="true" pageId="11" pageNumber="532">Dotilla fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938EA92FFC3FFF131E8FDA4FE7C923A" bold="true" box="[414,478,549,575]" pageId="11" pageNumber="532">
|
||
<taxonomicNameLabel id="A20B57E9FFC3FFF131E8FDA4FE7C923A" box="[414,478,549,575]" pageId="11" pageNumber="532" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is presently known from Digha Beach,
|
||
<collectingRegion id="4988F862FFC3FFF133DEFDA4FB95923A" box="[936,1079,549,575]" country="India" name="West Bengal" pageId="11" pageNumber="532">West Bengal</collectingRegion>
|
||
(
|
||
<typeStatus id="54F78822FFC3FFF1343EFDA4FBD8923A" box="[1096,1146,549,575]" pageId="11" pageNumber="532">type</typeStatus>
|
||
locality), and Chandipur,
|
||
<collectingRegion id="4988F862FFC3FFF130E1FDC8FF489266" box="[151,234,585,611]" country="India" name="Odisha" pageId="11" pageNumber="532">Odisha</collectingRegion>
|
||
. It occurs sympatrically with
|
||
<taxonomicName id="4C4C4D03FFC3FFF13236FDCBFD7F9266" authorityName="De Man" authorityYear="1888" box="[576,733,585,611]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC3FFF13236FDCBFD7F9266" box="[576,733,585,611]" italics="true" pageId="11" pageNumber="532">D. intermedia</emphasis>
|
||
</taxonomicName>
|
||
on the littoral zone at Digha (
|
||
<collectingRegion id="4988F862FFC3FFF1344DFDC8FB6D9266" box="[1083,1231,585,611]" country="India" name="West Bengal" pageId="11" pageNumber="532">West Bengal</collectingRegion>
|
||
,
|
||
<collectingCountry id="F35B7610FFC3FFF134ABFDC8FAB99261" box="[1245,1307,585,612]" name="India" pageId="11" pageNumber="532">India</collectingCountry>
|
||
). However, while
|
||
<taxonomicName id="4C4C4D03FFC3FFF130ABFDEFFEDA9282" authorityName="De Man" authorityYear="1888" box="[221,376,621,647]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="intermedia">
|
||
<emphasis id="B938EA92FFC3FFF130ABFDEFFEDA9282" box="[221,376,621,647]" italics="true" pageId="11" pageNumber="532">D. intermedia</emphasis>
|
||
</taxonomicName>
|
||
prefers the upper to middle littoral zone with a sandier substrate,
|
||
<taxonomicName id="4C4C4D03FFC3FFF1342BFDEFFB549282" authority="Mitra & Trivedi & Mendoza, 2020" authorityName="Mitra & Trivedi & Mendoza" authorityYear="2020" box="[1117,1270,621,647]" class="Malacostraca" family="Dotillidae" genus="Dotilla" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="532" phylum="Arthropoda" rank="species" species="fraternalis" status="sp. nov.">
|
||
<emphasis id="B938EA92FFC3FFF1342BFDEFFB549282" box="[1117,1270,621,647]" italics="true" pageId="11" pageNumber="532">D. fraternalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B57E9FFC3FFF1348BFDECFA9F9282" box="[1277,1341,621,647]" pageId="11" pageNumber="532" rank="species">
|
||
<emphasis id="B938EA92FFC3FFF1348BFDECFA979282" bold="true" box="[1277,1333,621,647]" pageId="11" pageNumber="532">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
tends to occupy the upper littoral zone, where the substrate is mostly silty (S. Mitra, pers. obs.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |