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<document id="132D499FC4B1962E0B0741A4FE80E978" ID-DOI="10.4202/app.00220.2015" ID-ISSN="1732-2421" ID-Zenodo-Dep="10626349" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1707248875147" checkinUser="felipe" docAuthor="Klug, Christian, Baets, Kenneth De &amp; Korn, Dieter" docDate="2016" docId="03E087FCFFC8FFC518FCFC89FD8DFAF2" docLanguage="en" docName="ActaPalaeontolPol.61.1.1-14.pdf" docOrigin="Acta Palaeontologica Polonica 61 (1)" docSource="http://dx.doi.org/10.4202/app.00220.2015" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Entogonites bucheri Klug &amp; Baets &amp; Korn 2016, sp. nov." docType="treatment" docVersion="3" lastPageNumber="10" masterDocId="FFD9FF84FFC0FFCC186BFFE1FFD9FFAE" masterDocTitle="Exploring the limits of morphospace: Ontogeny and ecology of late Viséan ammonoids from the Tafilalt, Morocco" masterLastPageNumber="14" masterPageNumber="1" pageNumber="9" updateTime="1713401358581" updateUser="karina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CCC9C840E4C3DFA178C7D5E12B9B9412">Exploring the limits of morphospace: Ontogeny and ecology of late Viséan ammonoids from the Tafilalt, Morocco</mods:title>
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<mods:namePart id="7B428D1B350B80BE95A38987BB79A12D">Klug, Christian</mods:namePart>
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<mods:namePart id="300C25DE074FA3CC5235B61B7FB3327B">Baets, Kenneth De</mods:namePart>
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<mods:namePart id="9F59A272E04442D81CBB9373E935670B">Korn, Dieter</mods:namePart>
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<mods:title id="600447E452361EB635825A95A5EF3103">Acta Palaeontologica Polonica</mods:title>
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<mods:identifier id="3CAD868C33560ECE94FC5035327C3CC7" type="DOI">10.4202/app.00220.2015</mods:identifier>
<mods:identifier id="31FB2E42BDB013478797808A471E0F34" type="ISSN">1732-2421</mods:identifier>
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<treatment id="03E087FCFFC8FFC518FCFC89FD8DFAF2" LSID="urn:lsid:plazi:treatment:03E087FCFFC8FFC518FCFC89FD8DFAF2" httpUri="http://treatment.plazi.org/id/03E087FCFFC8FFC518FCFC89FD8DFAF2" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="C3536561FFC8FFC418FCFC89FDDDFC2A" box="[151,516,872,900]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BF636EAFFC8FFC418FCFC89FDDDFC2A" blockId="8.[151,516,872,900]" box="[151,516,872,900]" pageId="8" pageNumber="9">
<heading id="D0BE8186FFC8FFC418FCFC89FDDDFC2A" box="[151,516,872,900]" fontSize="12" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="4C494D69FFC8FFC418FCFC89FE43FC2A" ID-CoL="39YWP" authority="Klug &amp; Baets &amp; Korn, 2016" authorityName="Klug &amp; Baets &amp; Korn" authorityYear="2016" box="[151,410,872,900]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" order="Ammonoidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="bucheri" status="sp. nov.">
<emphasis id="B93DEAF8FFC8FFC418FCFC89FE43FC2A" box="[151,410,872,900]" italics="true" pageId="8" pageNumber="9">Entogonites bucheri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E5783FFC8FFC419C9FC89FDDDFC2A" box="[418,516,872,900]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3536561FFC8FFC518FCFC74FD8DFAF2" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="8BF636EAFFC8FFC418FCFC74FE99FC04" blockId="8.[151,320,917,938]" box="[151,320,917,938]" pageId="8" pageNumber="9">
<figureCitation id="13722A6FFFC8FFC418FCFC74FF33FC04" box="[151,234,917,938]" captionStart="Fig" captionStartId="3.[97,130,1529,1549]" captionTargetBox="[98,1481,159,1504]" captionTargetId="figure-147@3.[98,1481,159,1504]" captionTargetPageId="3" captionText="Fig. 2. Juvenile (neanic) specimens of Goniatites lazarus Korn, Klug, and Mapes, 2005, Entogonites saharensis Korn, Klug, and Mapes, 2005, and Entogonites bucheri sp. nov. All from the early late Viséan, 12 km SE of Dar Kaoua (Tafilalt, Morocco). A. PIMUZ 31512, two neanic G. lazarus specimens (white arrows) in ventral views (A1), note the associated E. saharensis and orthocones. Enlarged G. lazarus in lateral view (A2); the narrow umbilicus and the umbilical ridge (A3). B. PIMUZ 31512, a juvenile G. lazarus, a small juvenile and a subadult E. saharensis; G. lazarus in ventral (B1) and lateral (B2) views. C. PIMUZ 31520, detail of a fully grown Maxigoniatites saourensis (Pareyn, 1961) with G. lazarus, 3 juvenile E. saharensis, and a subadult E. saharensis and a hatchling of E. bucheri sp. nov. (arrowed); C2, detail of C1, showing G. lazarus and E. bucheri sp. nov. All specimens whitened with NH4Cl-sublimate. Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10626353" httpUri="https://zenodo.org/record/10626353/files/figure.png" pageId="8" pageNumber="9">Figs. 2C</figureCitation>
,
<figureCitation id="13722A6FFFC8FFC4189FFC74FECAFC04" box="[244,275,917,938]" captionStart="Fig" captionStartId="6.[151,184,1526,1546]" captionTargetBox="[153,1533,162,1499]" captionTargetId="figure-140@6.[153,1534,162,1499]" captionTargetPageId="6" captionText="Fig. 5. Ammonoids from the early late Viséan, 12 km SE of Dar Kaoua (Tafilalt, Morocco). A, C. Calygirtyoceras darkaouaense Korn, Klug, and Mapes, 1999. A. PIMUZ 31516 (A1), Goniatites lazarus with a juvenile specimen in oblique-ventral (A2), lateral (A3), and ventral (A4) views. C. PIMUZ 31513, fragmentary adult in lateral view (C1), cross section (C2). B. Entogonites bucheri sp. nov., PIMUZ 31509, holotype in lateral (B1) and ventral (B2) views. All specimens whitened with NH4Cl-sublimate." figureDoi="http://doi.org/10.5281/zenodo.10626359" httpUri="https://zenodo.org/record/10626359/files/figure.png" pageId="8" pageNumber="9">5A</figureCitation>
,
<figureCitation id="13722A6FFFC8FFC41976FC74FEE5FC04" box="[285,316,917,938]" captionStart="Fig" captionStartId="7.[98,132,1850,1870]" captionTargetBox="[100,1480,166,1820]" captionTargetId="figure-12@7.[100,1481,891,1820]" captionTargetPageId="7" captionText="Fig. 6. Mass occurrence of juvenile ammonoids from the early late Viséan, 12 km SE of Dar Kaoua (Tafilalt, Morocco). A. PIMUZ 31518 with 4 specimens of Nomismoceras sp., 3 Prolecanites sp., 4 Entogonites saharensis Korn, Klug, and Mapes, 2005, 1 Calygirtyoceras darkaouaense Korn, Klug, and Mapes, 1999, and 2 Bollandites sp. B. PIMUZ 31508, overview; many taxa are indicated in the figure. All specimens whitened with NH4Clsublimate." figureDoi="http://doi.org/10.5281/zenodo.10626361" httpUri="https://zenodo.org/record/10626361/files/figure.png" pageId="8" pageNumber="9">6C</figureCitation>
.
</paragraph>
<paragraph id="8BF636EAFFC8FFC418FCFC20FF34FBA1" blockId="8.[151,824,961,1039]" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418FCFC20FF27FC78" box="[151,254,961,982]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
: Honoring Hugo Bucher (
<collectingRegion id="498DF808FFC8FFC41A68FC20FD9FFC78" box="[515,582,961,982]" country="Switzerland" name="Zuerich" pageId="8" pageNumber="9">Zürich</collectingRegion>
) for his numerous, thorough and profound contributions to ammonoid paleobiology and stratigraphy.
</paragraph>
<caption id="DF366662FFC8FFC41B35FC58FC71FBDB" ID-DOI="http://doi.org/10.5281/zenodo.10626363" ID-Zenodo-Dep="10626363" httpUri="https://zenodo.org/record/10626363/files/figure.png" pageId="8" pageNumber="9" startId="8.[862,895,953,973]" targetBox="[865,1533,160,929]" targetPageId="8" targetType="figure">
<paragraph id="8BF636EAFFC8FFC41B35FC58FC71FBDB" blockId="8.[862,1535,953,1141]" pageId="8" pageNumber="9">
Fig. 7. Morphospace (UWI, umbilical width index; WER, whorl expansion rate; CWI, conch width index) with data of 3015 species of Palaeozoic ammonoids with additional points of the ammonoids from the early late Viséan of the Tafilalt (Morocco). Note that data points of juveniles (empty symbols) of
<taxonomicName id="4C494D69FFC8FFC41BBFFBC8FB34FB93" authorityName="Korn, Klug, and Mapes" authorityYear="1999" box="[980,1261,1065,1085]" class="Cephalopoda" family="Girtyoceratidae" genus="Calygirtyoceras" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="darkaouaense">
<emphasis id="B93DEAF8FFC8FFC41BBFFBC8FB34FB93" box="[980,1261,1065,1085]" italics="true" pageId="8" pageNumber="9">Calygirtyoceras darkaouaense</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D69FFC8FFC41D75FBC8FA11FB93" authorityName="Korn, Klug, and Mapes" authorityYear="2005" box="[1310,1480,1065,1085]" class="Cephalopoda" family="Goniatitidae" genus="Goniatites" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="lazarus">
<emphasis id="B93DEAF8FFC8FFC41D75FBC8FA11FB93" box="[1310,1480,1065,1085]" italics="true" pageId="8" pageNumber="9">Goniatites lazarus</emphasis>
</taxonomicName>
lie at the edge or outside of the cloud of 3015 species. Graph produced using JMP 11.
</paragraph>
</caption>
<paragraph id="8BF636EAFFC8FFC418FCFBFAFE3CFBE3" blockId="8.[151,824,1051,1101]" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418FCFBFAFEC3FB9E" box="[151,282,1051,1072]" italics="true" pageId="8" pageNumber="9">Type material</emphasis>
:
<materialsCitation id="3B213CB7FFC8FFC4194CFBFAFD0FFB9E" box="[295,726,1051,1072]" collectionCode="PIMUZ" pageId="8" pageNumber="9" specimenCode="PIMUZ 31509" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F28848FFC8FFC4194CFBFAFE5AFB9E" box="[295,387,1051,1072]" pageId="8" pageNumber="9">Holotype</typeStatus>
, subadult specimen,
<specimenCode id="DBEF9E91FFC8FFC41A23FBFAFD0FFB9E" box="[584,726,1051,1072]" collectionCode="PIMUZ" country="Switzerland" httpUri="http://grbio.org/cool/25wz-z17r" name="Palaontologisches Institut und Museum der Universitat Zurich" pageId="8" pageNumber="9">PIMUZ 31509</specimenCode>
</materialsCitation>
<materialsCitation id="3B213CB7FFC8FFC41ABDFBFAFE3CFBE3" collectionCode="PIMUZ" pageId="8" pageNumber="9" specimenCode="PIMUZ 31508" specimenCount="1" typeStatus="paratype">
,
<typeStatus id="54F28848FFC8FFC41AB4FBFAFCEDFB9E" box="[735,820,1051,1072]" pageId="8" pageNumber="9">paratype</typeStatus>
, juvenile specimen,
<specimenCode id="DBEF9E91FFC8FFC4193AFBD9FE38FBE3" box="[337,481,1080,1101]" collectionCode="PIMUZ" country="Switzerland" httpUri="http://grbio.org/cool/25wz-z17r" name="Palaontologisches Institut und Museum der Universitat Zurich" pageId="8" pageNumber="9">PIMUZ 31508</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF636EAFFC8FFC418FCFBB8FE6AFB24" blockId="8.[151,825,1113,1162]" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418FCFBB8FECFFBC0" box="[151,278,1113,1134]" italics="true" pageId="8" pageNumber="9">Type locality</emphasis>
:
<quantity id="4CB19B0FFFC8FFC4194DFBB8FEBEFBC0" box="[294,359,1113,1134]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="km" value="12.0">12 km</quantity>
southeast of Dar Kaoua, southeast of Erfoud, eastern Anti–Atlas,
<collectingCountry id="F35E767AFFC8FFC4193EFB94FE76FB24" box="[341,431,1141,1162]" name="Morocco" pageId="8" pageNumber="9">Morocco</collectingCountry>
.
</paragraph>
<paragraph id="8BF636EAFFC8FFC418FCFB76FD11FB02" blockId="8.[151,712,1175,1196]" box="[151,712,1175,1196]" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418FCFB76FECAFB02" box="[151,275,1175,1196]" italics="true" pageId="8" pageNumber="9">Type horizon</emphasis>
: Early late Viséan (
<taxonomicName id="4C494D69FFC8FFC419BAFB76FD9DFB02" authorityName="Kittl" authorityYear="1904" box="[465,580,1175,1196]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="B93DEAF8FFC8FFC419BAFB76FD9DFB02" box="[465,580,1175,1196]" italics="true" pageId="8" pageNumber="9">Entogonites</emphasis>
</taxonomicName>
assemblage).
</paragraph>
<paragraph id="8BF636EAFFC8FFC418FCFB5EFDD4FAD4" blockId="8.[151,824,1215,1402]" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418FCFB5EFEDAFB76" box="[151,259,1215,1240]" italics="true" pageId="8" pageNumber="9">Material.</emphasis>
—
<materialsCitation id="3B213CB7FFC8FFC4194BFB5EFDD1FAD4" collectionCode="PIMUZ" pageId="8" pageNumber="9" specimenCode="PIMUZ 31509, PIMUZ 31512, PIMUZ 31520" specimenCount="5" specimenCount-juvenile="3" typeStatus="holotype">
6 individuals:
<specimenCount id="9D4FFD63FFC8FFC419A6FB5EFE02FB76" box="[461,475,1215,1240]" pageId="8" pageNumber="9">2</specimenCount>
specimens in
<specimenCode id="DBEF9E91FFC8FFC41AFBFB5EFCE1FB76" box="[656,824,1215,1240]" collectionCode="PIMUZ" country="Switzerland" httpUri="http://grbio.org/cool/25wz-z17r" name="Palaontologisches Institut und Museum der Universitat Zurich" pageId="8" pageNumber="9">PIMUZ 31509</specimenCode>
(the subadult holotype and a smaller individual);
<specimenCount id="9D4FFD63FFC8FFC41AA3FB01FD0FFB57" box="[712,726,1248,1273]" pageId="8" pageNumber="9">3</specimenCount>
juvenile to subadult specimens in
<specimenCode id="DBEF9E91FFC8FFC419A7FAE1FDA9FAB7" box="[460,624,1280,1305]" collectionCode="PIMUZ" country="Switzerland" httpUri="http://grbio.org/cool/25wz-z17r" name="Palaontologisches Institut und Museum der Universitat Zurich" pageId="8" pageNumber="9">PIMUZ 31512</specimenCode>
(associated with about 100 other juvenile ammonoid conchs); a very small juvenile conch in
<specimenCode id="DBEF9E91FFC8FFC4190FFAA0FDDEFAF4" box="[356,519,1345,1370]" collectionCode="PIMUZ" country="Switzerland" httpUri="http://grbio.org/cool/25wz-z17r" name="Palaontologisches Institut und Museum der Universitat Zurich" pageId="8" pageNumber="9">PIMUZ 31520</specimenCode>
(in the body chamber of a mature conch of
<taxonomicName id="4C494D69FFC8FFC4193EFA80FE24FAD4" authorityName="Korn, Klug &amp; Mapes" authorityYear="1999" box="[341,509,1377,1402]" class="Cephalopoda" family="Maxigoniatitidae" genus="Maxigoniatites" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="B93DEAF8FFC8FFC4193EFA80FE24FAD4" box="[341,509,1377,1402]" italics="true" pageId="8" pageNumber="9">Maxigoniatites</emphasis>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF636EAFFC8FFC418FCFA6DFD13F9E9" blockId="8.[151,824,1420,1607]" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418FCFA6DFE63FA0B" box="[151,442,1420,1445]" italics="true" pageId="8" pageNumber="9">
Diagnosis.—
<taxonomicName id="4C494D69FFC8FFC4195AFA6DFE63FA0B" authorityName="Kittl" authorityYear="1904" box="[305,442,1420,1445]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Entogonites</taxonomicName>
</emphasis>
with largely regularly coiled whorls. Only the third whorl displays faint tetragonal coiling. Expansion rate of juvenile and adult whorls exceeds 2.0. The uw/dm ratio is below 0.4. Ornamentation begins after the nepionic constriction with simple ribs, followed by bifurcating and intercalatory ribs (&gt;30 per whorl).
</paragraph>
<paragraph id="8BF636EAFFC8FFC41B35FB6FFC7FFA09" blockId="8.[862,1535,1166,1608]" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC41B35FB6FFC36FB09" box="[862,1007,1166,1191]" italics="true" pageId="8" pageNumber="9">Description.</emphasis>
—The small subevolute specimen PIMUZ 31520 measures about
<quantity id="4CB19B0FFFC8FFC41C05FB4FFB11FB69" box="[1134,1224,1198,1223]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="9" unit="mm" value="1.8">1.8 mm</quantity>
in diameter. It comprises the initial chamber and the initial 2.5 whorls (
<figureCitation id="13722A6FFFC8FFC41D0DFB2FFA66FB49" box="[1382,1471,1230,1255]" captionStart="Fig" captionStartId="3.[97,130,1529,1549]" captionTargetBox="[98,1481,159,1504]" captionTargetId="figure-147@3.[98,1481,159,1504]" captionTargetPageId="3" captionText="Fig. 2. Juvenile (neanic) specimens of Goniatites lazarus Korn, Klug, and Mapes, 2005, Entogonites saharensis Korn, Klug, and Mapes, 2005, and Entogonites bucheri sp. nov. All from the early late Viséan, 12 km SE of Dar Kaoua (Tafilalt, Morocco). A. PIMUZ 31512, two neanic G. lazarus specimens (white arrows) in ventral views (A1), note the associated E. saharensis and orthocones. Enlarged G. lazarus in lateral view (A2); the narrow umbilicus and the umbilical ridge (A3). B. PIMUZ 31512, a juvenile G. lazarus, a small juvenile and a subadult E. saharensis; G. lazarus in ventral (B1) and lateral (B2) views. C. PIMUZ 31520, detail of a fully grown Maxigoniatites saourensis (Pareyn, 1961) with G. lazarus, 3 juvenile E. saharensis, and a subadult E. saharensis and a hatchling of E. bucheri sp. nov. (arrowed); C2, detail of C1, showing G. lazarus and E. bucheri sp. nov. All specimens whitened with NH4Cl-sublimate. Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10626353" httpUri="https://zenodo.org/record/10626353/files/figure.png" pageId="8" pageNumber="9">Fig. 2C</figureCitation>
). The embryonic shell measures less than
<quantity id="4CB19B0FFFC8FFC41C98FB0FFAEDFAA9" box="[1267,1332,1262,1287]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="mm" value="1.0">1 mm</quantity>
and has a smooth surface. Only one side is exposed and thus, the thickness cannot be measured. At the presumed position of the nepionic constriction, the fine ornament commences. At
<quantity id="4CB19B0FFFC8FFC41DC0FAAFFA27FAC9" box="[1451,1534,1358,1383]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="9" unit="mm" value="1.8">1.8 mm</quantity>
diameter, there are 25 simple and rectiradiate ribs per half whorl.
</paragraph>
<paragraph id="8BF636EAFFC8FFC51BEFFA4EFEFEFEF5" blockId="8.[862,1535,1166,1608]" lastBlockId="9.[98,771,160,865]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
The ontogenetically slightly older specimen PIMUZ 31508 measures
<quantity id="4CB19B0FFFC8FFC41C71FA2EFB85FA46" box="[1050,1116,1487,1512]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="mm" value="5.0">5 mm</quantity>
in diameter. Its conch is subinvolute with a high whorl expansion rate (
<tableCitation id="C6CB0351FFC8FFC41C8DFA0EFAE7F9A6" box="[1254,1342,1519,1544]" captionStart="Table 4" captionStartId="8.[152,205,1632,1653]" captionTargetId="graphics-507@8.[151,1536,1730,1953]" captionTargetPageId="8" captionText="Table 4. Measurements (in mm) and ratios of Entogonites saharensis and E. bucheri sp. nov. Abbreviations: ah, apertural height; D, the distance between the generating curve and the coiling axis; dm, diameter; S, the shape of the generating curve; uw, umbilical width; W, whorl expansion rate; WER, whorl expansion rate; ww, whorl width (measured between ribs)." pageId="8" pageNumber="9">Table 4</tableCitation>
). As in the other specimen, the conch is only partially exposed and hence, conch thickness is unknown. The tetragonal coiling is only very faintly developed in early whorls and the last of the preserved (preadult) whorls is completely regularly coiled. At
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diameter, there are 20 primary ribs per half whorl. Most of these ribs bifurcate near the midflank, but there are a few trifurcating ribs as well (or bifurcating with an associated intercalatory rib).
</paragraph>
<caption id="DF366662FFC8FFC418F3F981FCA0F91E" pageId="8" pageNumber="9" startId="8.[152,205,1632,1653]" targetBox="[172,1519,1738,1950]" targetIsTable="true" targetPageId="8" targetType="table">
<paragraph id="8BF636EAFFC8FFC418F3F981FCA0F91E" blockId="8.[151,1535,1632,1712]" pageId="8" pageNumber="9">
Table 4. Measurements (in mm) and ratios of
<taxonomicName id="4C494D69FFC8FFC41A25F981FCF4F9DB" authorityName="Korn, Klug, and Mapes" authorityYear="2005" box="[590,813,1632,1653]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="saharensis">
<emphasis id="B93DEAF8FFC8FFC41A25F981FCF4F9DB" box="[590,813,1632,1653]" italics="true" pageId="8" pageNumber="9">Entogonites saharensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D69FFC8FFC41B30F981FC66F9DB" authority="Klug &amp; Baets &amp; Korn, 2016" authorityName="Klug &amp; Baets &amp; Korn" authorityYear="2016" box="[859,959,1632,1653]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="bucheri" status="sp. nov.">
<emphasis id="B93DEAF8FFC8FFC41B30F981FC66F9DB" box="[859,959,1632,1653]" italics="true" pageId="8" pageNumber="9">E. bucheri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E5783FFC8FFC41BAEF981FBC9F9DB" box="[965,1040,1632,1653]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations: ah, apertural height; D, the distance between the generating curve and the coiling axis; dm, diameter; S, the shape of the generating curve; uw, umbilical width; W, whorl expansion rate; WER, whorl expansion rate; ww, whorl width (measured between ribs).
</paragraph>
</caption>
<paragraph id="8BF636EAFFC8FFC418AAF92BFB26F830" pageId="8" pageNumber="9">
<table id="F949C44AFFC8003318C7F92BFA36F830" box="[172,1519,1738,1950]" gridcols="13" gridrows="7" pageId="8" pageNumber="9">
<tr id="357934A8FFC8003318C7F92BFA36F971" box="[172,1519,1738,1759]" gridrow="0" pageId="8" pageNumber="9">
<th id="76A85DD4FFC8003318C7F92BFEC6F971" box="[172,287,1738,1759]" gridcol="0" gridrow="0" pageId="8" pageNumber="9">Species</th>
<th id="76A85DD4FFC80033193BF92BFDE4F971" box="[336,573,1738,1759]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">Specimen</th>
<th id="76A85DD4FFC800331A01F92BFD4DF971" box="[618,660,1738,1759]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">dm</th>
<th id="76A85DD4FFC800331ADEF92BFD06F971" box="[693,735,1738,1759]" gridcol="3" gridrow="0" pageId="8" pageNumber="9">ww</th>
<th id="76A85DD4FFC800331B6AF92BFCF2F971" box="[769,811,1738,1759]" gridcol="4" gridrow="0" pageId="8" pageNumber="9">uw</th>
<th id="76A85DD4FFC800331B27F92BFCAFF971" box="[844,886,1738,1759]" gridcol="5" gridrow="0" pageId="8" pageNumber="9">ah</th>
<th id="76A85DD4FFC800331BF7F92BFC08F971" box="[924,977,1738,1759]" gridcol="6" gridrow="0" pageId="8" pageNumber="9">WER</th>
<th id="76A85DD4FFC800331B91F92BFB9BF971" box="[1018,1090,1738,1759]" gridcol="7" gridrow="0" pageId="8" pageNumber="9">ww/dm</th>
<th id="76A85DD4FFC800331C02F92BFB7CF971" box="[1129,1189,1738,1759]" gridcol="8" gridrow="0" pageId="8" pageNumber="9">ah/dm</th>
<th id="76A85DD4FFC800331CA2F92BFAD5F971" box="[1225,1292,1738,1759]" gridcol="9" gridrow="0" pageId="8" pageNumber="9">uw/dm</th>
<th id="76A85DD4FFC800331D45F92BFA81F971" box="[1326,1368,1738,1759]" gridcol="10" gridrow="0" pageId="8" pageNumber="9">W</th>
<th id="76A85DD4FFC800331D11F92BFA7DF971" box="[1402,1444,1738,1759]" gridcol="11" gridrow="0" pageId="8" pageNumber="9">D</th>
<th id="76A85DD4FFC800331DAEF92BFA36F971" box="[1477,1519,1738,1759]" gridcol="12" gridrow="0" pageId="8" pageNumber="9">S</th>
</tr>
<tr id="357934A8FFC8003318C7F90BFA36F951" box="[172,1519,1770,1791]" gridrow="1" pageId="8" pageNumber="9" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-3="1" rowspan-7="1">
<td id="76A85DD4FFC80033193BF90BFDE4F951" box="[336,573,1770,1791]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">PIMUZ 31520</td>
<td id="76A85DD4FFC800331A01F90BFD4DF951" box="[618,660,1770,1791]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">7.67</td>
<td id="76A85DD4FFC800331B6AF90BFCF2F951" box="[769,811,1770,1791]" gridcol="4" gridrow="1" pageId="8" pageNumber="9">3.60</td>
<td id="76A85DD4FFC800331B27F90BFCAFF951" box="[844,886,1770,1791]" gridcol="5" gridrow="1" pageId="8" pageNumber="9">1.63</td>
<td id="76A85DD4FFC800331BF7F90BFC08F951" box="[924,977,1770,1791]" gridcol="6" gridrow="1" pageId="8" pageNumber="9">1.61</td>
<td id="76A85DD4FFC800331C02F90BFB7CF951" box="[1129,1189,1770,1791]" gridcol="8" gridrow="1" pageId="8" pageNumber="9">0.21</td>
<td id="76A85DD4FFC800331CA2F90BFAD5F951" box="[1225,1292,1770,1791]" gridcol="9" gridrow="1" pageId="8" pageNumber="9">0.47</td>
</tr>
<tr id="357934A8FFC8003318C7F8EBFA36F88F" box="[172,1519,1802,1825]" gridrow="2" pageId="8" pageNumber="9" rowspan-8="1">
<th id="76A85DD4FFC8003318C7F8EBFEC6F88F" box="[172,287,1802,1825]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418C7F8EDFEC6F88F" box="[172,287,1804,1825]" italics="true" pageId="8" pageNumber="9">Entogonites</emphasis>
</th>
<td id="76A85DD4FFC80033193BF8EBFDE4F88F" box="[336,573,1802,1825]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">MB.C.5301, holotype</td>
<td id="76A85DD4FFC800331A01F8EBFD4DF88F" box="[618,660,1802,1825]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">9.35</td>
<td id="76A85DD4FFC800331ADEF8EBFD06F88F" box="[693,735,1802,1825]" gridcol="3" gridrow="2" pageId="8" pageNumber="9">3.05</td>
<td id="76A85DD4FFC800331B6AF8EBFCF2F88F" box="[769,811,1802,1825]" gridcol="4" gridrow="2" pageId="8" pageNumber="9">4.55</td>
<td id="76A85DD4FFC800331B27F8EBFCAFF88F" box="[844,886,1802,1825]" gridcol="5" gridrow="2" pageId="8" pageNumber="9">2.05</td>
<td id="76A85DD4FFC800331BF7F8EBFC08F88F" box="[924,977,1802,1825]" gridcol="6" gridrow="2" pageId="8" pageNumber="9">1.64</td>
<td id="76A85DD4FFC800331B91F8EBFB9BF88F" box="[1018,1090,1802,1825]" gridcol="7" gridrow="2" pageId="8" pageNumber="9">0.33</td>
<td id="76A85DD4FFC800331CA2F8EBFAD5F88F" box="[1225,1292,1802,1825]" gridcol="9" gridrow="2" pageId="8" pageNumber="9">0.49</td>
<td id="76A85DD4FFC800331D45F8EBFA81F88F" box="[1326,1368,1802,1825]" gridcol="10" gridrow="2" pageId="8" pageNumber="9">1.67</td>
<td id="76A85DD4FFC800331D11F8EBFA7DF88F" box="[1402,1444,1802,1825]" gridcol="11" gridrow="2" pageId="8" pageNumber="9">0.49</td>
<td id="76A85DD4FFC800331DAEF8EBFA36F88F" box="[1477,1519,1802,1825]" gridcol="12" gridrow="2" pageId="8" pageNumber="9">1.13</td>
</tr>
<tr id="357934A8FFC8003318C7F8C9FA36F891" box="[172,1519,1832,1855]" gridrow="3" pageId="8" pageNumber="9" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76A85DD4FFC8003318C7F8C9FEC6F891" box="[172,287,1832,1855]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418C7F8C9FEC6F893" box="[172,287,1832,1853]" italics="true" pageId="8" pageNumber="9">saharaensis</emphasis>
</th>
<td id="76A85DD4FFC80033193BF8C9FDE4F891" box="[336,573,1832,1855]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">MB.C.5304, paratype</td>
<td id="76A85DD4FFC800331A01F8C9FD4DF891" box="[618,660,1832,1855]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">11.2</td>
<td id="76A85DD4FFC800331B6AF8C9FCF2F891" box="[769,811,1832,1855]" gridcol="4" gridrow="3" pageId="8" pageNumber="9">5.4</td>
<td id="76A85DD4FFC800331CA2F8C9FAD5F891" box="[1225,1292,1832,1855]" gridcol="9" gridrow="3" pageId="8" pageNumber="9">0.48</td>
</tr>
<tr id="357934A8FFC8003318C7F8A8FA36F8F0" box="[172,1519,1865,1886]" gridrow="4" pageId="8" pageNumber="9" rowspan-0="1" rowspan-5="1" rowspan-8="1">
<td id="76A85DD4FFC80033193BF8A8FDE4F8F0" box="[336,573,1865,1886]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">MB.C.5305, paratype</td>
<td id="76A85DD4FFC800331A01F8A8FD4DF8F0" box="[618,660,1865,1886]" gridcol="2" gridrow="4" pageId="8" pageNumber="9">2.2</td>
<td id="76A85DD4FFC800331ADEF8A8FD06F8F0" box="[693,735,1865,1886]" gridcol="3" gridrow="4" pageId="8" pageNumber="9">1.15</td>
<td id="76A85DD4FFC800331B6AF8A8FCF2F8F0" box="[769,811,1865,1886]" gridcol="4" gridrow="4" pageId="8" pageNumber="9">1.05</td>
<td id="76A85DD4FFC800331BF7F8A8FC08F8F0" box="[924,977,1865,1886]" gridcol="6" gridrow="4" pageId="8" pageNumber="9">1.58</td>
<td id="76A85DD4FFC800331B91F8A8FB9BF8F0" box="[1018,1090,1865,1886]" gridcol="7" gridrow="4" pageId="8" pageNumber="9">0.52</td>
<td id="76A85DD4FFC800331CA2F8A8FAD5F8F0" box="[1225,1292,1865,1886]" gridcol="9" gridrow="4" pageId="8" pageNumber="9">0.48</td>
<td id="76A85DD4FFC800331D45F8A8FA81F8F0" box="[1326,1368,1865,1886]" gridcol="10" gridrow="4" pageId="8" pageNumber="9">1.53</td>
<td id="76A85DD4FFC800331D11F8A8FA7DF8F0" box="[1402,1444,1865,1886]" gridcol="11" gridrow="4" pageId="8" pageNumber="9">0.43</td>
<td id="76A85DD4FFC800331DAEF8A8FA36F8F0" box="[1477,1519,1865,1886]" gridcol="12" gridrow="4" pageId="8" pageNumber="9">1.67</td>
</tr>
<tr id="357934A8FFC8003318C7F888FA36F82E" box="[172,1519,1897,1920]" gridrow="5" pageId="8" pageNumber="9" rowspan-10="1" rowspan-11="1" rowspan-12="1">
<th id="76A85DD4FFC8003318C7F888FEC6F82E" box="[172,287,1897,1920]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418C7F88AFEC6F82E" box="[172,287,1899,1920]" italics="true" pageId="8" pageNumber="9">Entogonites</emphasis>
</th>
<td id="76A85DD4FFC80033193BF888FDE4F82E" box="[336,573,1897,1920]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">PIMUZ 31509, holotype</td>
<td id="76A85DD4FFC800331A01F888FD4DF82E" box="[618,660,1897,1920]" gridcol="2" gridrow="5" pageId="8" pageNumber="9">8.93</td>
<td id="76A85DD4FFC800331ADEF888FD06F82E" box="[693,735,1897,1920]" gridcol="3" gridrow="5" pageId="8" pageNumber="9">3.1</td>
<td id="76A85DD4FFC800331B6AF888FCF2F82E" box="[769,811,1897,1920]" gridcol="4" gridrow="5" pageId="8" pageNumber="9">2.86</td>
<td id="76A85DD4FFC800331B27F888FCAFF82E" box="[844,886,1897,1920]" gridcol="5" gridrow="5" pageId="8" pageNumber="9">2.68</td>
<td id="76A85DD4FFC800331BF7F888FC08F82E" box="[924,977,1897,1920]" gridcol="6" gridrow="5" pageId="8" pageNumber="9">2.04</td>
<td id="76A85DD4FFC800331B91F888FB9BF82E" box="[1018,1090,1897,1920]" gridcol="7" gridrow="5" pageId="8" pageNumber="9">0.35</td>
<td id="76A85DD4FFC800331C02F888FB7CF82E" box="[1129,1189,1897,1920]" gridcol="8" gridrow="5" pageId="8" pageNumber="9">0.3</td>
<td id="76A85DD4FFC800331CA2F888FAD5F82E" box="[1225,1292,1897,1920]" gridcol="9" gridrow="5" pageId="8" pageNumber="9">0.32</td>
</tr>
<tr id="357934A8FFC8003318C7F866FA36F830" box="[172,1519,1927,1950]" gridrow="6" pageId="8" pageNumber="9" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-3="1" rowspan-7="1">
<th id="76A85DD4FFC8003318C7F866FEC6F830" box="[172,287,1927,1950]" gridcol="0" gridrow="6" pageId="8" pageNumber="9">
<emphasis id="B93DEAF8FFC8FFC418AAF866FED3F832" box="[193,266,1927,1948]" italics="true" pageId="8" pageNumber="9">bucheri</emphasis>
</th>
<td id="76A85DD4FFC80033193BF866FDE4F830" box="[336,573,1927,1950]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">PIMUZ 31512</td>
<td id="76A85DD4FFC800331A01F866FD4DF830" box="[618,660,1927,1950]" gridcol="2" gridrow="6" pageId="8" pageNumber="9">5.0</td>
<td id="76A85DD4FFC800331B6AF866FCF2F830" box="[769,811,1927,1950]" gridcol="4" gridrow="6" pageId="8" pageNumber="9">1.84</td>
<td id="76A85DD4FFC800331B27F866FCAFF830" box="[844,886,1927,1950]" gridcol="5" gridrow="6" pageId="8" pageNumber="9">2.37</td>
<td id="76A85DD4FFC800331BF7F866FC08F830" box="[924,977,1927,1950]" gridcol="6" gridrow="6" pageId="8" pageNumber="9">2.13</td>
<td id="76A85DD4FFC800331C02F866FB7CF830" box="[1129,1189,1927,1950]" gridcol="8" gridrow="6" pageId="8" pageNumber="9">0.47</td>
<td id="76A85DD4FFC800331CA2F866FAD5F830" box="[1225,1292,1927,1950]" gridcol="9" gridrow="6" pageId="8" pageNumber="9">0.37</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF636EAFFC9FFC518ECFE83FE69FCCF" blockId="9.[98,771,160,865]" pageId="9" pageNumber="10">
The
<typeStatus id="54F28848FFC9FFC518D7FE83FEC6FED5" box="[188,287,354,379]" pageId="9" pageNumber="10">holotype</typeStatus>
PIMUZ 31509 has a diameter of almost
<quantity id="4CB19B0FFFC9FFC51A9FFE83FF4BFE35" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="10" unit="mm" value="9.0">9 mm</quantity>
. At this size, the subevolute conch is thinly discoidal with a high whorl expansion rate. The initial chamber measures approximately
<quantity id="4CB19B0FFFC9FFC5193FFE22FE74FE72" box="[340,429,451,476]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="10" unit="mm" value="0.4">0.4 mm</quantity>
across. The first two whorls are regularly coiled followed by one vaguely tetragonally coiled whorl. The last two whorls are again normally coiled. The ornamentation consists of 34 primary ribs on the last half whorl. Most of these ribs bifurcate in the middle of the flank, some are simple but then often associated with intercalatory ribs. The rib course begins with a deep ventral sinus, which is nearly twice as wide as it is deep measured from the tips of the adjacent ventrolateral projections, which are rounded and asymmetric. The ribs are the most strongly developed in the ventrolateral projections. Most of the flank is covered by a broad and shallow sinus. None of the specimens displays the suture line.
</paragraph>
<paragraph id="8BF636EAFFC9FFC51809FC92FDA0FABF" blockId="9.[98,771,883,1297]" pageId="9" pageNumber="10">
<emphasis id="B93DEAF8FFC9FFC51809FC92FF15FC22" box="[98,204,883,908]" italics="true" pageId="9" pageNumber="10">Remarks.</emphasis>
—Although the suture is not visible, the combination of coiling, ornamentation and stratigraphic origin confirms the genus assignment. The new species differs from all other species in the extremely reduced tetragonal coiling of the inner whorls. Additionally, it has a clearly higher whorl expansion rate (values partially from the online database GONIAT (http://www.goniat.org, accessed
<date id="FFF7102AFFC9FFC51AE1FBD4FF18FBC1" pageId="9" pageNumber="10" value="2015-09-07">
September 7
<superScript id="7C3C9BA2FFC9FFC51804FBB3FFA6FBCD" attach="left" box="[111,127,1106,1123]" fontSize="7" pageId="9" pageNumber="10">th</superScript>
2015
</date>
) and
<bibRefCitation id="EFD84B1BFFC9FFC51963FBB7FE10FBC1" author="Korn, D. &amp; Jovanovic, D. &amp; Novak, M. &amp; Sudar, M. N." box="[264,457,1110,1135]" pageId="9" pageNumber="10" pagination="355 - 364" refId="ref9221" refString="Korn, D., Jovanovic, D., Novak, M., and Sudar, M. N. 2010. Early late Visean ammonoid faunas from the Jadar Block (NW Serbia). Geologica Carpathica 61: 355 - 364." type="journal article" year="2010">Korn et al. 2010</bibRefCitation>
) than all other species (
<taxonomicName id="4C494D69FFC9FFC51A81FBB7FF63FB21" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="bucheri">
<emphasis id="B93DEAF8FFC9FFC51A81FBB7FF63FB21" italics="true" pageId="9" pageNumber="10">E. bucheri</emphasis>
</taxonomicName>
, 2.0–2.15;
<taxonomicName id="4C494D69FFC9FFC51928FB97FE0DFB21" authorityName="Kittl" authorityYear="1904" box="[323,468,1142,1167]" class="Cephalopoda" family="Entogonitidae" genus="Tetragonites" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="grimmeri">
<emphasis id="B93DEAF8FFC9FFC51928FB97FE0DFB21" box="[323,468,1142,1167]" italics="true" pageId="9" pageNumber="10">E. grimmeri</emphasis>
</taxonomicName>
, 1.5–1.7;
<taxonomicName id="4C494D69FFC9FFC51A26FB97FD22FB21" box="[589,763,1142,1167]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="saharaensis">
<emphasis id="B93DEAF8FFC9FFC51A26FB97FD22FB21" box="[589,763,1142,1167]" italics="true" pageId="9" pageNumber="10">E. saharaensis</emphasis>
</taxonomicName>
, 1.5–1.65;
<taxonomicName id="4C494D69FFC9FFC518B9FB76FEB6FB1E" baseAuthorityName="Kullmann" baseAuthorityYear="1962" box="[210,367,1175,1200]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="tetragonus">
<emphasis id="B93DEAF8FFC9FFC518B9FB76FEB6FB1E" box="[210,367,1175,1200]" italics="true" pageId="9" pageNumber="10">E. tetragonus</emphasis>
</taxonomicName>
, 1.8–1.9) and, as can be expected from the covariation of these characters, the umbilicus is narrower (
<taxonomicName id="4C494D69FFC9FFC518B2FB39FE8BFB5F" box="[217,338,1240,1265]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="bucheri">
<emphasis id="B93DEAF8FFC9FFC518B2FB39FE8BFB5F" box="[217,338,1240,1265]" italics="true" pageId="9" pageNumber="10">E. bucheri</emphasis>
</taxonomicName>
, &lt;0.4;
<taxonomicName id="4C494D69FFC9FFC519CFFB39FDFBFB5F" authorityName="Gordon" authorityYear="1957" box="[420,546,1240,1265]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="borealis">
<emphasis id="B93DEAF8FFC9FFC519CFFB39FDFBFB5F" box="[420,546,1240,1265]" italics="true" pageId="9" pageNumber="10">E. borealis</emphasis>
</taxonomicName>
, 0.41;
<taxonomicName id="4C494D69FFC9FFC51A04FB39FD22FB5F" authorityName="Kittl" authorityYear="1904" box="[623,763,1240,1265]" class="Cephalopoda" family="Entogonitidae" genus="Tetragonites" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="grimmeri">
<emphasis id="B93DEAF8FFC9FFC51A04FB39FD22FB5F" box="[623,763,1240,1265]" italics="true" pageId="9" pageNumber="10">E. grimmeri</emphasis>
</taxonomicName>
, 0.45;
<taxonomicName id="4C494D69FFC9FFC518F5FB19FE9CFABF" box="[158,325,1272,1297]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="saharaensis">
<emphasis id="B93DEAF8FFC9FFC518F5FB19FE9CFABF" box="[158,325,1272,1297]" italics="true" pageId="9" pageNumber="10">E. saharaensis</emphasis>
</taxonomicName>
,&gt;0.48;
<taxonomicName id="4C494D69FFC9FFC519CBFB19FDE0FABF" baseAuthorityName="Kullmann" baseAuthorityYear="1962" box="[416,569,1272,1297]" class="Cephalopoda" family="Entogonitidae" genus="Entogonites" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="tetragonus">
<emphasis id="B93DEAF8FFC9FFC519CBFB19FDE0FABF" box="[416,569,1272,1297]" italics="true" pageId="9" pageNumber="10">E. tetragonus</emphasis>
</taxonomicName>
, 0.5).
</paragraph>
<paragraph id="8BF636EAFFC9FFC51809FAC2FD8DFAF2" blockId="9.[98,770,1315,1372]" pageId="9" pageNumber="10">
<emphasis id="B93DEAF8FFC9FFC51809FAC2FDD9FA92" box="[98,512,1315,1340]" italics="true" pageId="9" pageNumber="10">Stratigraphic and geographic range.</emphasis>
—Early late Viséan, so far only known from the Tafilalt (
<collectingCountry id="F35E767AFFC9FFC519B5FAA2FD91FAF2" box="[478,584,1347,1372]" name="Morocco" pageId="9" pageNumber="10">Morocco</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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