135 lines
16 KiB
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135 lines
16 KiB
XML
<document id="168FB1397F1C62B75BD6D16974E19A0B" ID-DOI="10.11646/zootaxa.4006.1.2" ID-GBIF-Dataset="182580d9-4ce0-4f8a-9dc4-c8d9e059b25b" ID-ISSN="1175-5326" ID-Zenodo-Dep="244069" ID-ZooBank="E8EEFA24-F07F-4729-8D4B-215E2BA2CDD3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461197205770" checkinUser="plazi" docAuthor="Zhou, Lan & Chen, Hong-Wei" docDate="2015" docId="03E087E3FFE46E68FF38406CCB4FF99A" docLanguage="en" docName="zt04006p058.pdf" docOrigin="Zootaxa 4006 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucophenga mutabilis" docType="treatment" docVersion="9" masterDocId="FFD9FF9BFFE06E6CFFAF420DCA31FFF7" masterDocTitle="The genus Leucophenga (Diptera, Drosophilidae), part V: the mutabilis species group from East Asia, with morphological and molecular evidence" updateTime="1698600967833" updateUser="plazi">
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<mods:titleInfo id="8538AE5DC137DAD502C32E56F1983949">
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<mods:title id="085942936F0B75863F0F6A7DF30A563B">The genus Leucophenga (Diptera, Drosophilidae), part V: the mutabilis species group from East Asia, with morphological and molecular evidence</mods:title>
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<mods:namePart id="E56853DA07C7FAF9AB23D55301945D0A">Zhou, Lan</mods:namePart>
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<mods:namePart id="20FEEC2EAA0849B19C9694D52EBFED77">Chen, Hong-Wei</mods:namePart>
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<mods:typeOfResource id="AE839226D352231146BD9E7CEB351664">text</mods:typeOfResource>
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<mods:title id="ED81C32FA0C65D1B48AFD311A9A98B07">Zootaxa</mods:title>
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<mods:part id="5DA67B89099A5C887E14D5CD0B426B1C">
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<mods:date id="8E2FEFC3BA8ADD1F4790C215687ED922">2015</mods:date>
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<mods:detail id="2B44AE6BBED02C228E39C757AA8B2FE0" type="volume">
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<mods:number id="F1FE313B2C6840185E3A01664EF2A789">4006</mods:number>
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<mods:number id="FD27977B141C011485FCF8F93192B8C8">1</mods:number>
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<mods:classification id="9C1A3B5390FB15CA2D0D8AB303B10AB5">journal volume</mods:classification>
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<mods:identifier id="FAD623DA9C7B1D2CD6D9710864154469" type="DOI">10.11646/zootaxa.4006.1.2</mods:identifier>
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<treatment id="03E087E3FFE46E68FF38406CCB4FF99A" ID-DOI="http://doi.org/10.5281/zenodo.5628999" ID-GBIF-Taxon="119629453" ID-Zenodo-Dep="5628999" LSID="urn:lsid:plazi:treatment:03E087E3FFE46E68FF38406CCB4FF99A" httpUri="http://treatment.plazi.org/id/03E087E3FFE46E68FF38406CCB4FF99A" pageId="4">
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<subSubSection id="C353657EFFE46E68FF38406CCBDFFD8C" box="[151,494,608,635]" pageId="4" type="nomenclature">
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<paragraph id="8BF636F5FFE46E68FF38406CCBDFFD8C" blockId="4.[151,494,608,635]" box="[151,494,608,635]" pageId="4">
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<heading id="D0BE8199FFE46E68FF38406CCBDFFD8C" bold="true" box="[151,494,608,635]" fontSize="11" level="1" pageId="4" reason="1">
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<emphasis id="B93DEAE7FFE46E68FF38406CCBDFFD8C" bold="true" box="[151,494,608,635]" pageId="4">
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The
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<taxonomicName id="4C494D76FFE46E68FF60406DCB0EFD8D" ID-CoL="8CJGL" box="[207,319,608,634]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="4" phylum="Arthropoda" rank="species" species="mutabilis">
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<emphasis id="B93DEAE7FFE46E68FF60406DCB0EFD8D" bold="true" box="[207,319,608,634]" italics="true" pageId="4">mutabilis</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C353657EFFE46E68FF3840A5CEA2FD49" box="[151,1171,680,702]" pageId="4" type="reference_group">
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<paragraph id="8BF636F5FFE46E68FF3840A5CEA2FD49" blockId="4.[151,1171,680,702]" box="[151,1171,680,702]" pageId="4">
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<treatmentCitationGroup id="AB5911DBFFE46E68FF3840A5CEA2FD49" box="[151,1171,680,702]" pageId="4">
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<taxonomicName id="4C494D76FFE46E68FF3840A5CBB2FD49" ID-CoL="8CJGL" box="[151,387,680,702]" class="Insecta" family="Drosophilidae" genus="Leucophenga" kingdom="Animalia" order="Diptera" pageId="4" phylum="Arthropoda" rank="species" species="mutabilis">
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<emphasis id="B93DEAE7FFE46E68FF3840A5CBB2FD49" box="[151,387,680,702]" italics="true" pageId="4">Leucophenga mutabilis</emphasis>
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</taxonomicName>
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species group:
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<treatmentCitation id="0AE810E4FFE46E68FD8D40A5C8FCFD49" author="Bachli" box="[546,717,680,702]" page="77" pageId="4" year="1971">
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<bibRefCitation id="EFD84B04FFE46E68FD8D40A5C896FD49" author="Bachli" box="[546,679,680,702]" pageId="4" refString="Bachli, G. (1971) Leucophenga und Paraleucophenga (Diptera: Brachycera) Fam. Drosophilidae. Exploration du Parc National de l' Upemba, Fascicule 71, 1 - 192 + pls. 1 - 38. [Bruxelles]" type="journal article" year="1971">Bächli, 1971</bibRefCitation>
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: 77
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</treatmentCitation>
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;
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<treatmentCitation id="0AE810E4FFE46E68FD7840A5C9BEFD49" author="Okada" box="[727,911,680,702]" page="803" pageId="4" year="1989">
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<bibRefCitation id="EFD84B04FFE46E68FD7840A5C96DFD49" author="Okada" box="[727,860,680,702]" pageId="4" refString="Okada, H. (1989) A revision of the Leucophenga mutabilis species-group (Diptera: Drosophilidae) of the Oriental and adjacent regions. Japan Journal of Entomology, 57, 803 - 813." type="journal article" year="1989">Okada, 1989</bibRefCitation>
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: 803
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</treatmentCitation>
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;
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<treatmentCitation id="0AE810E4FFE46E68FC3640A5CEA1FD49" author="Fartyal" box="[921,1168,680,702]" page="407" pageId="4" year="2005">
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<bibRefCitation id="EFD84B04FFE46E68FC3640A5CE6CFD49" author="Fartyal" box="[921,1117,680,702]" pageId="4" refString="Fartyal, R. S., Singh, B. K. & Toda, M. J. (2005) Review of the genus Leucophenga Mik (Diptera: Drosophilidae) in India, with descriptions of five new species from northern India. Entomological Science, 8, 405 - 417. http: // dx. doi. org / 10.1111 / j. 1479 - 8298.2005.00140. x" type="journal article" year="2005">
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Fartyal
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<emphasis id="B93DEAE7FFE46E68FC4840A4CE25FD49" box="[999,1044,680,702]" italics="true" pageId="4">et al</emphasis>
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., 2005
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</bibRefCitation>
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: 407
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C353657EFFE46E68FF3840E7CE76FCB9" pageId="4" type="diagnosis">
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<paragraph id="8BF636F5FFE46E68FF3840E7CE76FCB9" blockId="4.[151,1437,746,1645]" pageId="4">
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<emphasis id="B93DEAE7FFE46E68FF3840E7CB3BFCF4" bold="true" box="[151,266,746,771]" pageId="4">Diagnosis</emphasis>
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(modified from
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<bibRefCitation id="EFD84B04FFE46E68FE6640E7C865FCF4" author="Okada" box="[457,596,746,771]" pageId="4" refString="Okada, H. (1989) A revision of the Leucophenga mutabilis species-group (Diptera: Drosophilidae) of the Oriental and adjacent regions. Japan Journal of Entomology, 57, 803 - 813." type="journal article" year="1989">Okada 1989</bibRefCitation>
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). Palpus brown, sexually dimorphic (large, with 2–4 thin setae in female, slender, with a row of 5–7 setae in male) (
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<figureCitation id="13722A70FFE46E68FDD64103C8C8FCD1" box="[633,761,782,807]" captionStart="FIGURE 2. A, B" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[183,1407,213,1781]" captionTargetId="figure@5.[151,1436,193,1802]" captionTargetPageId="5" captionText="FIGURE 2. A, B. Head of Leucophenga angusta (A, male; B, female); C. wing of L. striatipennis (male); D. wing of L. taiwanensis (male); E, F. thorax of L. angusta; G, H. thorax of L. bellula; I, J. thorax of L. magnipalpis." httpUri="https://zenodo.org/record/244071/files/figure.png" pageId="4">Figs 2A, B</figureCitation>
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,
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<figureCitation id="13722A70FFE46E68FCA94103C925FCD0" box="[774,788,782,807]" captionStart="FIGURE 5" captionStartId="8.[151,250,1982,2004]" captionTargetBox="[166,1423,650,1945]" captionTargetId="figure@8.[151,1437,647,1961]" captionTargetPageId="8" captionText="FIGURE 5. Leucophenga angusta Okada, 1956, male: A. Head of male; B. epandrium, surstylus and cercus (lateral view); C. hypandrium, paramere, aedeagus and aedeagal apodeme (lateral view). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/244074/files/figure.png" pageId="4">5</figureCitation>
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A); proclinate orbital seta (Orb 1) close to ocellar triangle (
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<figureCitation id="13722A70FFE46E68FF304138CB2CFCBA" box="[159,285,821,846]" captionStart="FIGURE 2. A, B" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[183,1407,213,1781]" captionTargetId="figure@5.[151,1436,193,1802]" captionTargetPageId="5" captionText="FIGURE 2. A, B. Head of Leucophenga angusta (A, male; B, female); C. wing of L. striatipennis (male); D. wing of L. taiwanensis (male); E, F. thorax of L. angusta; G, H. thorax of L. bellula; I, J. thorax of L. magnipalpis." httpUri="https://zenodo.org/record/244071/files/figure.png" pageId="4">Figs 2A, B</figureCitation>
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,
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<figureCitation id="13722A70FFE46E68FE874138CB07FCB9" box="[296,310,821,846]" captionStart="FIGURE 5" captionStartId="8.[151,250,1982,2004]" captionTargetBox="[166,1423,650,1945]" captionTargetId="figure@8.[151,1437,647,1961]" captionTargetPageId="8" captionText="FIGURE 5. Leucophenga angusta Okada, 1956, male: A. Head of male; B. epandrium, surstylus and cercus (lateral view); C. hypandrium, paramere, aedeagus and aedeagal apodeme (lateral view). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/244074/files/figure.png" pageId="4">5</figureCitation>
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A); cercus with 10 long setae near posterior margin (
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<figureCitation id="13722A70FFE46E68FC2B4138C9FAFCB9" box="[900,971,821,846]" captionStart="FIGURE 5" captionStartId="8.[151,250,1982,2004]" captionTargetBox="[166,1423,650,1945]" captionTargetId="figure@8.[151,1437,647,1961]" captionTargetPageId="8" captionText="FIGURE 5. Leucophenga angusta Okada, 1956, male: A. Head of male; B. epandrium, surstylus and cercus (lateral view); C. hypandrium, paramere, aedeagus and aedeagal apodeme (lateral view). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/244074/files/figure.png" pageId="4">Figs 5</figureCitation>
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B, 6–11A).
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</paragraph>
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</subSubSection>
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<subSubSection id="C353657EFFE46E68FF684154CB4FF99A" pageId="4" type="description">
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<paragraph id="8BF636F5FFE46E68FF684154C8FBF9D2" blockId="4.[151,1437,746,1645]" pageId="4">
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<emphasis id="B93DEAE7FFE46E68FF684154CB68FC85" bold="true" box="[199,345,857,882]" pageId="4">Description.</emphasis>
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Eyes red to brownish red. Postocellar setae distinctly thick. Ocellar triangle brown to black, mostly with 2 pair of setae above ocellar setae. Frons narrower than 1/3 width of head, nearly aequilate, with a few minute setulae medially. All orbital setae large; proclinate orbital setae and anterior reclinate orbital setae close together (
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<figureCitation id="13722A70FFE46E68FEA941C8CB7DFC29" box="[262,332,965,990]" captionStart="FIGURE 5" captionStartId="8.[151,250,1982,2004]" captionTargetBox="[166,1423,650,1945]" captionTargetId="figure@8.[151,1437,647,1961]" captionTargetPageId="8" captionText="FIGURE 5. Leucophenga angusta Okada, 1956, male: A. Head of male; B. epandrium, surstylus and cercus (lateral view); C. hypandrium, paramere, aedeagus and aedeagal apodeme (lateral view). Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/244074/files/figure.png" pageId="4">Fig. 5</figureCitation>
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A). Arista plumose, with long branches. Face usually brownish to brown; facial carina absent. Clypeus brownish medially, dark brown laterally. Vibrissa prominent; subvibrissal setae small. Gena and postgena narrow. Mesonotum yellowish to yellowish brown. Postpronotal lobe yellow, with 2–4 long setae and a few of shorter setae. Acrostichal setulae in ca. 12–16 irregular rows. Prescutellar setae large. Pleura mostly yellowish and with brown to black patches. Katepisternal with small setae medially, and 2 large ones anteriorly and posteriorly, respectively. Subscutellum somewhat swollen. Wing hyaline, sometimes infuscate, lacking patch. Basal medialcubital crossvein absent. Costal vein between R2+3 and R4+5 distally with 4–8 peg-like spinules on ventral surface; R2+3 slightly curved; R4+5 and M1 nearly parallel distally. Legs yellowish except for dark brown on knees of mid and hindlegs. Abdominal tergites yellow, with variable, black patterns. Male terminalia: Epandrium usually pubescent, with several setae around antero-ventral corner to posterior margin and developed apodeme along anterior margin. Surstylus usually broad and flat, nearly entirely pubescent, with several setae on outer and inner surface. Cercus separated from epandrium, pubescent and setigerous. Hypandrium (gonopod in
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<bibRefCitation id="EFD84B04FFE46E68FBBB4755CEECFA86" author="Bachli" box="[1044,1245,1368,1393]" pageId="4" refString="Bachli, G., Vilela, C. R., Andersson, S. & Saura, A. (2004) The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, 39, 1 - 362. [Brill, Leiden, New York]" type="journal article" year="2004">
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Bächli
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<emphasis id="B93DEAE7FFE46E68FBCB4754CEADFA87" box="[1124,1180,1368,1392]" italics="true" pageId="4">et al.</emphasis>
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2004
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</bibRefCitation>
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) anteriorly fused to aedeagal apodeme, laterally broad, usually with 1 paramedian seta distally. Gonopods (dorsal arch in
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<bibRefCitation id="EFD84B04FFE46E68FA984771CACAFA4E" author="Bachli" pageId="4" refString="Bachli, G., Vilela, C. R., Andersson, S. & Saura, A. (2004) The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, 39, 1 - 362. [Brill, Leiden, New York]" type="journal article" year="2004">
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Bächli
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||
<emphasis id="B93DEAE7FFE46E68FA274770CA85FA4F" italics="true" pageId="4">et al.</emphasis>
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||
2004
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||
</bibRefCitation>
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||
) fused with each other, forming slightly triangular plate, antero-ventrally with curved, median rod. Paramere (outer paraphysis in
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<bibRefCitation id="EFD84B04FFE46E68FE4347C9C884FA2A" author="Bachli" box="[492,693,1476,1501]" pageId="4" refString="Bachli, G., Vilela, C. R., Andersson, S. & Saura, A. (2004) The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, 39, 1 - 362. [Brill, Leiden, New York]" type="journal article" year="2004">
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Bächli
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<emphasis id="B93DEAE7FFE46E68FD9347C8C845FA2B" box="[572,628,1476,1500]" italics="true" pageId="4">et al.</emphasis>
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2004
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</bibRefCitation>
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) contiguous to arm of aedeagal apodeme basally, usually distorted basally, with a few sensilla distally. Aedeagal basal bridges (inner paraphysis in
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<bibRefCitation id="EFD84B04FFE46E68FB8847E5CEC5F9F6" author="Bachli" box="[1063,1268,1512,1537]" pageId="4" refString="Bachli, G., Vilela, C. R., Andersson, S. & Saura, A. (2004) The Drosophilidae (Diptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, 39, 1 - 362. [Brill, Leiden, New York]" type="journal article" year="2004">
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Bächli
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<emphasis id="B93DEAE7FFE46E68FBD747E4CE83F9F7" box="[1144,1202,1512,1536]" italics="true" pageId="4">et al.</emphasis>
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2004
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</bibRefCitation>
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) contiguous to median rod of gonopods, bifurcated basomedially.
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</paragraph>
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<paragraph id="8BF636F5FFE46E68FF68443CCB4FF99A" blockId="4.[151,1437,746,1645]" pageId="4">In the following species descriptions, only characters that depart from the above universal characters are provided for brevity.</paragraph>
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</subSubSection>
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||
</treatment>
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</document> |