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<document id="42721A195C76072F86722BB61AF709B2" ID-DOI="10.1111/zoj.12112" ID-ISSN="0024-4082" ID-Zenodo-Dep="10273730" ID-ZooBank="7BA11142-CBC1-4026-A578-EBAB6D2B6C0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630107647828" checkinUser="marcus" docAuthor="Mendoza Marroquín, Jorge I." docDate="2014" docId="03E087B45677FFC0FEED4FA7B3731603" docLanguage="en" docName="zoj12112.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 170 (4)" docSource="http://dx.doi.org/10.1111/zoj.12112" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Hemirrhagus papalotl Perez-Miles &amp; Locht 2003" docType="treatment" docVersion="3" lastPageNumber="664" masterDocId="FFD9FFCC566AFFDEFFD84973B208116D" masterDocTitle="Taxonomic revision of Hemirrhagus Simon, 1903 (Araneae: Theraphosidae, Theraphosinae), with description of five new species from Mexico" masterLastPageNumber="689" masterPageNumber="634" pageNumber="663" updateTime="1701826367839" updateUser="ExternalLinkService">
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<mods:title id="101440E9DDEDCC927DDE198CB809413E">Taxonomic revision of Hemirrhagus Simon, 1903 (Araneae: Theraphosidae, Theraphosinae), with description of five new species from Mexico</mods:title>
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<mods:namePart id="D3A93AC8850A6D1B6EBA74BB9E8D5728">Mendoza Marroquín, Jorge I.</mods:namePart>
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<mods:title id="CD1B5733E4E70DF59AC33F761E021831">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="5B2EE4B0A1B1C0765181830E26643D9B">2014</mods:date>
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<mods:title id="DE6BD36CC60784EE3B7D9F0125F6EDDD">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="EEFB9C08B2E0857E3ACC64CF892CA230">2014-03-28</mods:number>
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<mods:number id="5C6E2CA1E593839CBA3A4494E3C0D367">170</mods:number>
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<mods:identifier id="ED9F4DCD7B2B4441D11B993E393F8DC9" type="DOI">10.1111/zoj.12112</mods:identifier>
<mods:identifier id="C62FCCFD2A6B8E303A413FF1BEB02495" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03E087B45677FFC0FEED4FA7B3731603" LSID="urn:lsid:plazi:treatment:03E087B45677FFC0FEED4FA7B3731603" httpUri="http://treatment.plazi.org/id/03E087B45677FFC0FEED4FA7B3731603" lastPageId="30" lastPageNumber="664" pageId="29" pageNumber="663">
<subSubSection id="C35365295677FFC3FEED4FA7B0661664" pageId="29" pageNumber="663" type="nomenclature">
<paragraph id="8BF636A25677FFC3FEED4FA7B0591785" blockId="29.[218,684,1748,1834]" box="[309,593,1748,1769]" pageId="29" pageNumber="663">
<heading id="D0BE81CE5677FFC3FEED4FA7B0591785" box="[309,593,1748,1769]" centered="true" fontSize="9" level="2" pageId="29" pageNumber="663" reason="2">
<taxonomicName id="4C494D215677FFC3FEED4FA7B0591785" ID-CoL="6LQQS" authorityName="Perez-Miles &amp; Locht" authorityYear="2003" box="[309,593,1748,1769]" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="685" phylum="Arthropoda" rank="species" species="papalotl">
<emphasis id="B93DEAB05677FFC3FEED4FA7B0591785" box="[309,593,1748,1769]" italics="true" pageId="29" pageNumber="663">
<emphasis id="B93DEAB05677FFC3FEED4FA7B3D51785" box="[309,477,1748,1769]" italics="true" pageId="29" pageNumber="663">HEMIRRHAGUS</emphasis>
PAPALOTL
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF636A25677FFC3FEC04F87B0661664" blockId="29.[218,684,1748,1834]" box="[280,622,1780,1801]" pageId="29" pageNumber="663">
<heading id="D0BE81CE5677FFC3FEC04F87B0661664" box="[280,622,1780,1801]" centered="true" fontSize="8" level="2" pageId="29" pageNumber="663" reason="8">PÉREZ- MILES &amp; LOCHT, 2003</heading>
</paragraph>
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<subSubSection id="C35365295677FFC3FF024E67B0A41647" box="[218,684,1812,1834]" pageId="29" pageNumber="663" type="description">
<paragraph id="8BF636A25677FFC3FF024E67B0A41647" blockId="29.[218,684,1748,1834]" box="[218,684,1812,1834]" pageId="29" pageNumber="663">
(
<figureCitation id="13722A275677FFC3FF3A4E67B3531644" box="[226,347,1812,1833]" captionStart="Figure 1" captionStartId="3.[144,223,1334,1353]" captionTargetBox="[303,1262,196,1304]" captionTargetId="figure-144@3.[303,1263,196,1304]" captionTargetPageId="3" captionText="Figure 1. A–H, J–L, Hemirrhagus spp. habitus; I, Hemirrhagus papalotl egg-sac. A, Hemirrhagus chilango, male CNAN 4253, in life; B, C, Hemirrhagus eros, B, female, C, male; D, E, Hemirrhagus ocellatus, D, male CNAN 4457, in life, E, female; F, Hemirrhagus benzaa sp. nov., female; G–I, H. papalotl, G, male CNAN 4461, in life, H, female CNAN 4461, in life, I, female CNAN 4459 with fixed hammock egg-sac, in life; J, K, Hemirrhagus stygius, J, male CNAN 4462, in life, K, female CNAN 4462, in life; L, Hemirrhagus franckei sp. nov., holotype male CNAN-T0776, in life. Photos: A, C, D, E, G–K: J. Mendoza; B, F, L: E. Goyer." figureDoi="http://doi.org/10.5281/zenodo.10273732" httpUri="https://zenodo.org/record/10273732/files/figure.png" pageId="29" pageNumber="663">FIGS 1G–I</figureCitation>
,
<figureCitation id="13722A275677FFC3FEB24E67B3BF1644" box="[362,439,1812,1833]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="29" pageNumber="663">20A–J</figureCitation>
,
<figureCitation id="13722A275677FFC3FE1D4E67B0101644" box="[453,536,1812,1833]" captionStart="Figure 21" captionStartId="20.[164,243,733,752]" captionTargetBox="[387,1219,197,704]" captionTargetId="figure-142@20.[387,1219,197,704]" captionTargetPageId="20" captionText="Figure 21. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460, left palpal bulb. A, ventral view. B, dorsal view. C, retrolateral view. D, prolateral view. Abbreviations: SA, subapical keel; VG, ventral groove. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10273790" httpUri="https://zenodo.org/record/10273790/files/figure.png" pageId="29" pageNumber="663">21A–D</figureCitation>
,
<figureCitation id="13722A275677FFC3FDFE4E67B0721644" box="[550,634,1812,1833]" captionStart="Figure 22" captionStartId="20.[164,243,1397,1416]" captionTargetBox="[323,1283,828,1367]" captionTargetId="figure-182@20.[323,1283,828,1367]" captionTargetPageId="20" captionText="Figure 22. A–C, Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, holotype female LAAH; D, female CNAN 4459. A, ocular tubercle, dorsal view. B, opisthosoma, dorsal view. C, spermathecae, ventral view. D, spermathecae, ventral view. Scale bars = 2 mm (B), 1 mm (D), 0.5 mm (A, C)." figureDoi="http://doi.org/10.5281/zenodo.10273792" httpUri="https://zenodo.org/record/10273792/files/figure.png" pageId="29" pageNumber="663">22A–D</figureCitation>
,
<figureCitation id="13722A275677FFC3FD504E67B0AD1647" box="[648,677,1812,1834]" captionStart="Figure 48" captionStartId="51.[144,223,1012,1031]" captionTargetBox="[151,1418,203,974]" captionTargetId="figure-352@51.[144,1425,196,982]" captionTargetPageId="51" captionText="Figure 48. Known distribution of Hemirrhagus stygius (Gertsch, 1971), Hemirrhagus elliotti (Gertsch, 1973), Hemirrhagus embolulatus sp. nov., Hemirrhagus eros Pérez-Miles &amp; Locht, 2003, Hemirrhagus chilango Pérez-Miles &amp; Locht, 2003, Hemirrhagus nahuanus (Gertsch, 1982), Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, Hemirrhagus grieta (Gertsch, 1982), Hemirrhagus coztic Pérez-Miles &amp; Locht, 2003, and Hemirrhagus perezmilesi García-Villafuerte &amp; Locht, 2010." figureDoi="http://doi.org/10.5281/zenodo.10273862" httpUri="https://zenodo.org/record/10273862/files/figure.png" pageId="29" pageNumber="663">48</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35365295677FFC3FF484E49B3801603" pageId="29" pageNumber="663" type="reference_group">
<paragraph id="8BF636A25677FFC3FF484E49B3801603" blockId="29.[144,759,1850,1903]" pageId="29" pageNumber="663">
<treatmentCitationGroup id="AB59118C5677FFC3FF484E49B3801603" pageId="29" pageNumber="663">
<treatmentCitation id="0AE810B35677FFC3FF484E49B2B41603" author="Perez-Miles F &amp; Locht A" page="372" pageId="29" pageNumber="663" year="2003">
<taxonomicName id="4C494D215677FFC3FF484E49B2B41603" ID-CoL="6LQQS" authority="Perez-Miles &amp; Locht, 2003: 372" authorityName="Perez-Miles &amp; Locht" authorityPageNumber="372" authorityYear="2003" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="663" phylum="Arthropoda" rank="species" species="papalotl">
<emphasis id="B93DEAB05677FFC3FF484E49B39E1622" box="[144,406,1850,1871]" italics="true" pageId="29" pageNumber="663">Hemirrhagus papalotl</emphasis>
<bibRefCitation id="EFD84B535677FFC3FE7E4E49B2B41603" author="Perez-Miles F &amp; Locht A" pageId="29" pageNumber="663" pagination="365 - 375" refId="ref35444" refString="Perez-Miles F, Locht A. 2003. Revision and cladistic analysis of the genus Hemirrhagus Simon, 1903 (Araneae, Theraphosidae, Theraphosinae). Bulletin of the British Arachnological Society 12: 365 - 375." type="journal article" year="2003">Pérez-Miles &amp; Locht, 2003: 372</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 35–38 (D
<specimenCount id="9D4FFD2B5677FFC3FEBC4E2BB3711603" box="[356,377,1880,1902]" pageId="29" pageNumber="663" type="female">♀</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35365295677FFC0FCFE49B5B3731603" lastPageId="30" lastPageNumber="664" pageId="29" pageNumber="663" type="materials_examined">
<paragraph id="8BF636A25677FFC3FCFE49B5B7881074" blockId="29.[806,1421,197,281]" pageId="29" pageNumber="663">
<emphasis id="B93DEAB05677FFC3FCFE49B5B1C711B6" box="[806,975,198,219]" italics="true" pageId="29" pageNumber="663">Type material:</emphasis>
<materialsCitation id="3B213CFF5677FFC3FC0449B5B7881074" collectingDate="1980-11-29" collectionCode="LAAH" collectorName="I. Cascalera. Examined." country="Mexico" location="Gruta de Aguacachil." municipality="Mpio. Taxco de Alarcon" pageId="29" pageNumber="663" specimenCount="2" specimenCount-female="2" stateProvince="Guerrero" typeStatus="holotype">
<typeStatus id="54F288005677FFC3FC0449B5B64A11B1" box="[988,1090,198,220]" pageId="29" pageNumber="663" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4FFD2B5677FFC3FB8849B6B66D11B6" box="[1104,1125,197,219]" pageId="29" pageNumber="663" type="female">♀</specimenCount>
and
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<specimenCount id="9D4FFD2B5677FFC3FAC449B6B73911B6" box="[1308,1329,197,219]" pageId="29" pageNumber="663" type="female">♀</specimenCount>
<collectionCode id="ED58AE675677FFC3FAE649B5B78111B6" box="[1342,1417,198,219]" pageId="29" pageNumber="663">LAAH</collectionCode>
,
<collectingCountry id="F35E76325677FFC3FCFE4996B19B1197" box="[806,915,229,250]" name="Mexico" pageId="29" pageNumber="663">MEXICO</collectingCountry>
:
<collectingRegion id="498DF8405677FFC3FC454996B6001197" box="[925,1032,229,250]" country="Mexico" name="Guerrero" pageId="29" pageNumber="663">Guerrero</collectingRegion>
,
<collectingMunicipality id="6B92ACD85677FFC3FBCB4997B7141197" box="[1043,1308,228,250]" pageId="29" pageNumber="663">Mpio. Taxco de Alarcón</collectingMunicipality>
,
<location id="8E9660795677FFC3FAFF4996B1A41074" LSID="urn:lsid:plazi:treatment:03E087B45677FFC0FEED4FA7B3731603:8E9660795677FFC3FAFF4996B1A41074" country="Mexico" municipality="Mpio. Taxco de Alarcon" name="Gruta de Aguacachil." pageId="29" pageNumber="663" stateProvince="Guerrero">Gruta de Aguacachil.</location>
<date id="FFF710625677FFC3FC6D4870B6231074" box="[949,1067,259,281]" pageId="29" pageNumber="663" value="1980-11-29">
<collectingDate id="EFB3E98A5677FFC3FC6D4870B6231074" box="[949,1067,259,281]" pageId="29" pageNumber="663" value="1980-11-29">29.xi.1980</collectingDate>
</date>
, col.
<collectorName id="26BC53745677FFC3FBB24870B7881074" box="[1130,1408,259,281]" pageId="29" pageNumber="663">I. Cascalera. Examined.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BF636A25677FFC3FCFE4833B7411326" blockId="29.[806,1422,320,587]" pageId="29" pageNumber="663">
<emphasis id="B93DEAB05677FFC3FCFE4833B6941038" box="[806,1180,320,341]" italics="true" pageId="29" pageNumber="663">Additional material examined:</emphasis>
<materialsCitation id="3B213CFF5677FFC3FB774833B78110FE" collectingDate="1980-08-03" collectionCode="LAAH" collectorName="J. Palacios" country="Mexico" location="Gruta de Ahuacachil" municipality="Mpio. Taxco de Alarcon" pageId="29" pageNumber="663" specimenCount="1" specimenCount-immature="1" stateProvince="Guerrero">
<specimenCount id="9D4FFD2B5677FFC3FB774833B74A103B" box="[1199,1346,320,342]" pageId="29" pageNumber="663" type="immature">1 immature</specimenCount>
male
<collectionCode id="ED58AE675677FFC3FCFE482CB1791019" box="[806,881,351,372]" pageId="29" pageNumber="663">LAAH</collectionCode>
,
<collectingCountry id="F35E76325677FFC3FC58482CB1E51019" box="[896,1005,351,372]" name="Mexico" pageId="29" pageNumber="663">MEXICO</collectingCountry>
:
<collectingRegion id="498DF8405677FFC3FC23482CB66E1019" box="[1019,1126,351,372]" country="Mexico" name="Guerrero" pageId="29" pageNumber="663">Guerrero</collectingRegion>
,
<collectingMunicipality id="6B92ACD85677FFC3FBAC482CB7811018" box="[1140,1417,351,373]" pageId="29" pageNumber="663">Mpio. Taxco de Alarcón</collectingMunicipality>
,
<location id="8E9660795677FFC3FCFE480DB62B10FE" LSID="urn:lsid:plazi:treatment:03E087B45677FFC0FEED4FA7B3731603:8E9660795677FFC3FCFE480DB62B10FE" box="[806,1059,381,403]" country="Mexico" municipality="Mpio. Taxco de Alarcon" name="Gruta de Ahuacachil" pageId="29" pageNumber="663" stateProvince="Guerrero">Gruta de Ahuacachil</location>
,
<date id="FFF710625677FFC3FBEE480EB6B510FE" box="[1078,1213,381,403]" pageId="29" pageNumber="663" value="1980-08-03">
<collectingDate id="EFB3E98A5677FFC3FBEE480EB6B510FE" box="[1078,1213,381,403]" pageId="29" pageNumber="663" value="1980-08-03">03.viii.1980</collectingDate>
</date>
, col.
<collectorName id="26BC53745677FFC3FADD480DB78110FE" box="[1285,1417,381,403]" pageId="29" pageNumber="663">J. Palacios</collectorName>
</materialsCitation>
;
<materialsCitation id="3B213CFF5677FFC3FCFE48EFB75D10BC" collectingDate="2012-02-03" collectionCode="CNAN" country="Mexico" location="Gruta de Zacatecolotla" municipality="Mpio. Taxco de Alarcon" pageId="29" pageNumber="663" specimenCode="CNAN 4458" specimenCount="1" specimenCount-female="1" stateProvince="Guerrero">
<specimenCount id="9D4FFD2B5677FFC3FCFE48EFB15E10DF" box="[806,854,412,434]" pageId="29" pageNumber="663" type="female">1 ♀</specimenCount>
<specimenCode id="DBEF9ED95677FFC3FCBA48EFB1FC10DF" box="[866,1012,412,434]" collectionCode="CNAN" pageId="29" pageNumber="663">CNAN 4458</specimenCode>
,
<collectingCountry id="F35E76325677FFC3FBDE48EFB67B10DC" box="[1030,1139,412,433]" name="Mexico" pageId="29" pageNumber="663">MEXICO</collectingCountry>
:
<collectingRegion id="498DF8405677FFC3FB5B48EFB6E610DC" box="[1155,1262,412,433]" country="Mexico" name="Guerrero" pageId="29" pageNumber="663">Guerrero</collectingRegion>
,
<collectingMunicipality id="6B92ACD85677FFC3FB2748EFB1A110BC" pageId="29" pageNumber="663">Mpio. Taxco de Alarcón</collectingMunicipality>
,
<location id="8E9660795677FFC3FC6448C8B6C710BC" LSID="urn:lsid:plazi:treatment:03E087B45677FFC0FEED4FA7B3731603:8E9660795677FFC3FC6448C8B6C710BC" box="[956,1231,443,465]" country="Mexico" municipality="Mpio. Taxco de Alarcon" name="Gruta de Zacatecolotla" pageId="29" pageNumber="663" stateProvince="Guerrero">Gruta de Zacatecolotla</location>
,
<date id="FFF710625677FFC3FB3A48C8B75D10BC" box="[1250,1365,443,465]" pageId="29" pageNumber="663" value="2012-02-03">
<collectingDate id="EFB3E98A5677FFC3FB3A48C8B75D10BC" box="[1250,1365,443,465]" pageId="29" pageNumber="663" value="2012-02-03">03.ii.2012</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B213CFF5677FFC3FABF48C8B1BE1082" collectionCode="R" collectorName="Monjaraz" country="Mexico" pageId="29" pageNumber="663" specimenCount="1" stateProvince="Guerrero">
col.
<collectionCode id="ED58AE675677FFC3FCFE48A9B1331082" box="[806,827,474,495]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29" pageNumber="663">R</collectionCode>
.
<collectorName id="26BC53745677FFC3FC9F48AAB1BE1082" box="[839,950,473,495]" pageId="29" pageNumber="663">Monjaraz</collectorName>
</materialsCitation>
;
<materialsCitation id="3B213CFF5677FFC3FC1A48AAB63E1327" collectingDate="2012-04-21" collectionCode="CNAN" collectorName="J. Mendoza &amp; G. Contreras" country="Mexico" location="Gruta de Aguacachil" municipality="Mpio. Taxco de Alarcon" pageId="29" pageNumber="663" specimenCode="CNAN4459, CNAN 4460, CNAN 4461" specimenCount="16" specimenCount-female="11" specimenCount-male="5" stateProvince="Guerrero">
<specimenCount id="9D4FFD2B5677FFC3FC1A48AAB1F11082" box="[962,1017,473,495]" pageId="29" pageNumber="663" type="female">11 ♀</specimenCount>
and
<specimenCount id="9D4FFD2B5677FFC3FBEC48AAB66A1082" box="[1076,1122,473,495]" pageId="29" pageNumber="663" type="male">5 ♂</specimenCount>
<specimenCode id="DBEF9ED95677FFC3FBB248AAB6F81082" box="[1130,1264,473,495]" collectionCode="CNAN" pageId="29" pageNumber="663">CNAN4459</specimenCode>
,
<specimenCode id="DBEF9ED95677FFC3FB2448AAB7811082" box="[1276,1417,473,495]" collectionCode="CNAN" pageId="29" pageNumber="663">CNAN 4460</specimenCode>
,
<specimenCode id="DBEF9ED95677FFC3FCFE488BB1B31363" box="[806,955,504,526]" collectionCode="CNAN" pageId="29" pageNumber="663">CNAN 4461</specimenCode>
,
<collectingCountry id="F35E76325677FFC3FC08488BB6351360" box="[976,1085,504,525]" name="Mexico" pageId="29" pageNumber="663">MEXICO</collectingCountry>
:
<collectingRegion id="498DF8405677FFC3FB89488BB6B41360" box="[1105,1212,504,525]" country="Mexico" name="Guerrero" pageId="29" pageNumber="663">Guerrero</collectingRegion>
,
<collectingMunicipality id="6B92ACD85677FFC3FB08488BB1891341" pageId="29" pageNumber="663">Mpio. Taxco de Alarcón</collectingMunicipality>
,
<location id="8E9660795677FFC3FC404B65B6951341" LSID="urn:lsid:plazi:treatment:03E087B45677FFC0FEED4FA7B3731603:8E9660795677FFC3FC404B65B6951341" box="[920,1181,534,556]" country="Mexico" municipality="Mpio. Taxco de Alarcon" name="Gruta de Aguacachil" pageId="29" pageNumber="663" stateProvince="Guerrero">Gruta de Aguacachil</location>
,
<date id="FFF710625677FFC3FB6C4B65B7201341" box="[1204,1320,534,556]" pageId="29" pageNumber="663" value="2012-04-21">
<collectingDate id="EFB3E98A5677FFC3FB6C4B65B7201341" box="[1204,1320,534,556]" pageId="29" pageNumber="663" value="2012-04-21">21.iv.2012</collectingDate>
</date>
, col.
<collectorName id="26BC53745677FFC3FAA04B64B1871326" pageId="29" pageNumber="663">J. Mendoza</collectorName>
,
<collectorName id="26BC53745677FFC3FC454B46B63E1327" box="[925,1078,565,586]" pageId="29" pageNumber="663">G. Contreras</collectorName>
</materialsCitation>
,
<materialsCitation id="3B213CFF5677FFC3FB9C4B46B74E1326" box="[1092,1350,565,587]" collectionCode="R" collectorName="Monjaraz, D &amp; Ortiz" country="Mexico" pageId="29" pageNumber="663" specimenCount="1" stateProvince="Guerrero">
<collectionCode id="ED58AE675677FFC3FB9C4B46B6511327" box="[1092,1113,565,586]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29" pageNumber="663">R</collectionCode>
.
<collectorName id="26BC53745677FFC3FBBE4B46B6F11327" box="[1126,1273,565,587]" pageId="29" pageNumber="663">Monjaraz, D</collectorName>
.
<collectorName id="26BC53745677FFC3FADF4B46B74E1326" box="[1287,1350,565,587]" pageId="29" pageNumber="663">Ortíz</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF636A25677FFC3FCFE4B01B7851466" blockId="29.[806,1422,626,1291]" pageId="29" pageNumber="663">
<emphasis id="B93DEAB05677FFC3FCFE4B01B61813EA" box="[806,1040,626,647]" italics="true" pageId="29" pageNumber="663">Emended diagnosis:</emphasis>
The following character combination is diagnostic for
<taxonomicName id="4C494D215677FFC3FBE14BE2B6B113CB" authorityName="Perez-Miles &amp; Locht" authorityYear="2003" box="[1081,1209,657,678]" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="663" phylum="Arthropoda" rank="species" species="papalotl">
<emphasis id="B93DEAB05677FFC3FBE14BE2B6B113CB" box="[1081,1209,657,678]" italics="true" pageId="29" pageNumber="663">H. papalotl</emphasis>
</taxonomicName>
. Male palpal bulb with slender embolus, longer than tegulum,
<collectionCode id="ED58AE675677FFC3FAF74BDCB75813A9" box="[1327,1360,687,708]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="29" pageNumber="663">SA</collectionCode>
ends at embolus retrolateral face (
<figureCitation id="13722A275677FFC3FB5E4BBDB7061389" box="[1158,1294,718,740]" captionStart="Figure 21" captionStartId="20.[164,243,733,752]" captionTargetBox="[387,1219,197,704]" captionTargetId="figure-142@20.[387,1219,197,704]" captionTargetPageId="20" captionText="Figure 21. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460, left palpal bulb. A, ventral view. B, dorsal view. C, retrolateral view. D, prolateral view. Abbreviations: SA, subapical keel; VG, ventral groove. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10273790" httpUri="https://zenodo.org/record/10273790/files/figure.png" pageId="29" pageNumber="663">Fig. 21A, C</figureCitation>
); VG deep (
<figureCitation id="13722A275677FFC3FCF64B9EB19B126E" box="[814,915,749,771]" captionStart="Figure 21" captionStartId="20.[164,243,733,752]" captionTargetBox="[387,1219,197,704]" captionTargetId="figure-142@20.[387,1219,197,704]" captionTargetPageId="20" captionText="Figure 21. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460, left palpal bulb. A, ventral view. B, dorsal view. C, retrolateral view. D, prolateral view. Abbreviations: SA, subapical keel; VG, ventral groove. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10273790" httpUri="https://zenodo.org/record/10273790/files/figure.png" pageId="29" pageNumber="663">Fig. 21D</figureCitation>
). The posterior curvature of
<collectionCode id="ED58AE675677FFC3FB2D4B9EB71E126F" box="[1269,1302,749,770]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="29" pageNumber="663">SA</collectionCode>
occurs at the base of the embolus and forms an angle of 90° (
<figureCitation id="13722A275677FFC3FCF64A59B19A122D" box="[814,914,810,832]" captionStart="Figure 21" captionStartId="20.[164,243,733,752]" captionTargetBox="[387,1219,197,704]" captionTargetId="figure-142@20.[387,1219,197,704]" captionTargetPageId="20" captionText="Figure 21. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460, left palpal bulb. A, ventral view. B, dorsal view. C, retrolateral view. D, prolateral view. Abbreviations: SA, subapical keel; VG, ventral groove. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10273790" httpUri="https://zenodo.org/record/10273790/files/figure.png" pageId="29" pageNumber="663">Fig. 21C</figureCitation>
). Embolus strongly curved retrolaterally on distal third (
<figureCitation id="13722A275677FFC3FC284A3BB6721233" box="[1008,1146,840,862]" captionStart="Figure 21" captionStartId="20.[164,243,733,752]" captionTargetBox="[387,1219,197,704]" captionTargetId="figure-142@20.[387,1219,197,704]" captionTargetPageId="20" captionText="Figure 21. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460, left palpal bulb. A, ventral view. B, dorsal view. C, retrolateral view. D, prolateral view. Abbreviations: SA, subapical keel; VG, ventral groove. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10273790" httpUri="https://zenodo.org/record/10273790/files/figure.png" pageId="29" pageNumber="663">Fig. 21A, B</figureCitation>
). Metatarsus I curved (
<figureCitation id="13722A275677FFC3FCF64A14B1821210" box="[814,906,871,893]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="29" pageNumber="663">Fig. 20I</figureCitation>
). Ocular tubercle slightly reduced and PME slightly reduced; periocular pigmentation only strongly marked on anterior eye row (
<figureCitation id="13722A275677FFC3FB034AD7B74C12D4" box="[1243,1348,932,954]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="29" pageNumber="663">Figs 20D</figureCitation>
,
<figureCitation id="13722A275677FFC3FA894AD7B78A12D7" box="[1361,1410,932,954]" captionStart="Figure 22" captionStartId="20.[164,243,1397,1416]" captionTargetBox="[323,1283,828,1367]" captionTargetId="figure-182@20.[323,1283,828,1367]" captionTargetPageId="20" captionText="Figure 22. A–C, Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, holotype female LAAH; D, female CNAN 4459. A, ocular tubercle, dorsal view. B, opisthosoma, dorsal view. C, spermathecae, ventral view. D, spermathecae, ventral view. Scale bars = 2 mm (B), 1 mm (D), 0.5 mm (A, C)." figureDoi="http://doi.org/10.5281/zenodo.10273792" httpUri="https://zenodo.org/record/10273792/files/figure.png" pageId="29" pageNumber="663">22A</figureCitation>
). Urticating setae arranged in one dorsomedian patch, brown in colour, with anterior and posterior margins notched medially, with rounded edges (butterflyshaped) (
<figureCitation id="13722A275677FFC3FC4E4D6DB1F61559" box="[918,1022,1054,1076]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="29" pageNumber="663">Figs 20C</figureCitation>
,
<figureCitation id="13722A275677FFC3FBCA4D6CB64A1558" box="[1042,1090,1055,1077]" captionStart="Figure 22" captionStartId="20.[164,243,1397,1416]" captionTargetBox="[323,1283,828,1367]" captionTargetId="figure-182@20.[323,1283,828,1367]" captionTargetPageId="20" captionText="Figure 22. A–C, Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, holotype female LAAH; D, female CNAN 4459. A, ocular tubercle, dorsal view. B, opisthosoma, dorsal view. C, spermathecae, ventral view. D, spermathecae, ventral view. Scale bars = 2 mm (B), 1 mm (D), 0.5 mm (A, C)." figureDoi="http://doi.org/10.5281/zenodo.10273792" httpUri="https://zenodo.org/record/10273792/files/figure.png" pageId="29" pageNumber="663">22B</figureCitation>
). Spermathecae receptacles are straight or nearly so; receptacles can vary from short and wide to large and thin (
<figureCitation id="13722A275677FFC3FB2A4D2FB789151C" box="[1266,1409,1116,1138]" captionStart="Figure 21" captionStartId="20.[164,243,733,752]" captionTargetBox="[387,1219,197,704]" captionTargetId="figure-142@20.[387,1219,197,704]" captionTargetPageId="20" captionText="Figure 21. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460, left palpal bulb. A, ventral view. B, dorsal view. C, retrolateral view. D, prolateral view. Abbreviations: SA, subapical keel; VG, ventral groove. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10273790" httpUri="https://zenodo.org/record/10273790/files/figure.png" pageId="29" pageNumber="663">Fig. 21C, D</figureCitation>
).
<taxonomicName id="4C494D215677FFC3FCFE4D09B62B15E2" authorityName="Perez-Miles &amp; Locht" authorityYear="2003" box="[806,1059,1146,1167]" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="663" phylum="Arthropoda" rank="species" species="papalotl">
<emphasis id="B93DEAB05677FFC3FCFE4D09B62B15E2" box="[806,1059,1146,1167]" italics="true" pageId="29" pageNumber="663">Hemirrhagus papalotl</emphasis>
</taxonomicName>
differs from most
<taxonomicName id="4C494D215677FFC3FB2D4D09B17E15C2" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="663" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93DEAB05677FFC3FB2D4D09B78515E2" box="[1269,1421,1146,1167]" italics="true" pageId="29" pageNumber="663">Hemirrhagus</emphasis>
species
</taxonomicName>
by the posterior curvature of
<collectionCode id="ED58AE675677FFC3FAD64DEAB73A15C3" box="[1294,1330,1177,1198]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="29" pageNumber="663">SA</collectionCode>
, which forms an angle of 90°. It differs from
<taxonomicName id="4C494D215677FFC3FB064DCBB75115A1" authorityName="Perez-Miles &amp; Locht" authorityYear="2003" box="[1246,1369,1207,1229]" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="663" phylum="Arthropoda" rank="species" species="gertschi">
<emphasis id="B93DEAB05677FFC3FB064DCBB75115A1" box="[1246,1369,1207,1229]" italics="true" pageId="29" pageNumber="663">H. gertschi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D215677FFC3FCFE4DA4B1911586" authority="Mendoza Marroquín, 2014" authorityName="Mendoza Marroquín" authorityYear="2014" box="[806,921,1238,1260]" class="Arachnida" family="Theraphosidae" genus="Hemirrhagus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="663" phylum="Arthropoda" rank="species" species="valdezi" status="sp. nov.">
<emphasis id="B93DEAB05677FFC3FCFE4DA4B1911586" box="[806,921,1238,1260]" italics="true" pageId="29" pageNumber="663">H. valdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57CB5677FFC3FC734DA5B6021586" box="[939,1034,1238,1259]" pageId="29" pageNumber="663" rank="species">sp. nov.</taxonomicNameLabel>
by the urticating setae being arranged in one dorsomedian, butterfly-shaped patch.
</paragraph>
<paragraph id="8BF636A25677FFC3FCFE4C41B6AE14C9" blockId="29.[806,1422,1330,1902]" pageId="29" pageNumber="663">
<emphasis id="B93DEAB05677FFC3FCFE4C41B1A6142A" box="[806,942,1330,1351]" italics="true" pageId="29" pageNumber="663">Description:</emphasis>
Male
<collectionCode id="ED58AE675677FFC3FBD54C41B651142A" box="[1037,1113,1330,1351]" pageId="29" pageNumber="663">CNAN</collectionCode>
4461: body length (not including chelicerae and spinnerets) 23.46, carapace: 10.16 long, 8.96 wide. Caput not elevated; fovea recurved, width 1.875 (
<figureCitation id="13722A275677FFC3FBEE4CFDB69214C9" box="[1078,1178,1422,1444]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="29" pageNumber="663">Fig. 20A</figureCitation>
).
</paragraph>
<paragraph id="8BF636A25677FFC3FCE74CDEB7841603" blockId="29.[806,1422,1330,1902]" pageId="29" pageNumber="663">
Eyes: anterior eye row procurved, posterior eye row recurved. Periocular pigmentation only strongly marked on anterior eye row. PLE slightly reduced. Eye sizes and interocular distances: AME 0.275; ALE 0.375; PME 0.15; PLE 0.175; AME–AME 0.15; AME– ALE 0.075; PME–PME 0.75; PME–PLE 0.075; ALE– PLE 0.1. Ocular tubercle reduced, width 1.525; length 0.875; clypeus lacking (
<figureCitation id="13722A275677FFC3FBF44FF1B69917F5" box="[1068,1169,1666,1688]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="29" pageNumber="663">Fig. 20D</figureCitation>
). Labium: length 1.45; width 1.85; with 23 cuspules. Maxilla inner corner with approximately 130 cuspules (
<figureCitation id="13722A275677FFC3FB604FB3B71417BB" box="[1208,1308,1728,1750]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="29" pageNumber="663">Fig. 20E</figureCitation>
). Cheliceral promargin with ten teeth (proximal to distal: firstthird large, fourth-fifth medium, sixth-eighth large, ninth-tenth small) (
<figureCitation id="13722A275677FFC3FBC04E68B675165C" box="[1048,1149,1819,1841]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="29" pageNumber="663">Fig. 20G</figureCitation>
). Sternum length 4.5. Sigillae circular, third and fourth pairs hardly visible; fourth pair once its length from margin (
<figureCitation id="13722A275677FFC3FB254E2BB7881603" box="[1277,1408,1880,1902]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="29" pageNumber="663">Fig. 20B, F</figureCitation>
).
</paragraph>
<paragraph id="8BF636A25674FFC0FF6449B5B3AA10DF" blockId="30.[164,779,198,1566]" pageId="30" pageNumber="664">
Leg formula:
<collectionCode id="ED58AE675674FFC0FE8F49B5B37B11B6" box="[343,371,198,219]" pageId="30" pageNumber="664">IV</collectionCode>
, I,
<collectionCode id="ED58AE675674FFC0FE4F49B5B3A511B6" box="[407,429,198,219]" pageId="30" pageNumber="664">II</collectionCode>
,
<collectionCode id="ED58AE675674FFC0FE6249B5B3D311B6" box="[442,475,198,219]" pageId="30" pageNumber="664">III</collectionCode>
. Length of legs and palpal segments (femur, patella, tibia, metatarsus, tarsus, total): I: 12.20, 5.80, 11.29, 11.13, 8.07, 48.49.
<collectionCode id="ED58AE675674FFC0FD284870B10F1075" box="[752,775,259,280]" pageId="30" pageNumber="664">II</collectionCode>
: 11.87, 5.94, 10.74, 11.19, 7.72, 47.46.
<collectionCode id="ED58AE675674FFC0FD814851B072105A" box="[601,634,290,311]" pageId="30" pageNumber="664">III</collectionCode>
: 11.64, 4.96, 10.05, 12.47, 8.00, 47.12.
<collectionCode id="ED58AE675674FFC0FE1F4832B3ED103B" box="[455,485,321,342]" pageId="30" pageNumber="664">IV</collectionCode>
: 13.78, 5.25, 12.95, 17.39, 9.54, 58.91. Palp: 8.32, 4.25, 7.04, -, 3.47, 23.08. Spinnerets:
<collectionCode id="ED58AE675674FFC0FEE8480DB36210FE" box="[304,362,382,403]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="30" pageNumber="664">PMS</collectionCode>
, 1.2 long, 1.1 apart; PLS, 2.4 basal, 1.8 middle, 2.7 distal.
</paragraph>
<paragraph id="8BF636A25674FFC0FF6448C8B3EC1341" blockId="30.[164,779,198,1566]" pageId="30" pageNumber="664">
Scopulae: tarsi I–
<collectionCode id="ED58AE675674FFC0FE5348C8B3A010BD" box="[395,424,443,464]" pageId="30" pageNumber="664">IV</collectionCode>
densely scopulate,
<collectionCode id="ED58AE675674FFC0FD4848C8B0A610BD" box="[656,686,443,464]" pageId="30" pageNumber="664">III</collectionCode>
and
<collectionCode id="ED58AE675674FFC0FD3748C8B10310BD" box="[751,779,443,464]" pageId="30" pageNumber="664">IV</collectionCode>
divided by strong band of setae. Metatarsi I–
<collectionCode id="ED58AE675674FFC0FD2F48A9B1021082" box="[759,778,474,495]" pageId="30" pageNumber="664">II</collectionCode>
densely scopulate;
<collectionCode id="ED58AE675674FFC0FE52488BB3A01360" box="[394,424,504,525]" pageId="30" pageNumber="664">III</collectionCode>
scopulate on distal half,
<collectionCode id="ED58AE675674FFC0FD37488BB1031360" box="[751,779,504,525]" pageId="30" pageNumber="664">IV</collectionCode>
scopulate on distal quarter.
</paragraph>
<paragraph id="8BF636A25674FFC0FF644B46B362126E" blockId="30.[164,779,198,1566]" pageId="30" pageNumber="664">
Tibia I with two apophyses, which do not originate from a common base, Pap reduced with one large spinose seta on ventral face, the spinose seta is bent apically and exceeds the apex of Pap; Rap with one large spinose seta on dorsal face, the spinose seta exceeds the apex of Rap (
<figureCitation id="13722A275674FFC0FE0E4BBDB051138E" box="[470,601,718,740]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="30" pageNumber="664">Fig. 20H, J</figureCitation>
). Metatarsus I curved (
<figureCitation id="13722A275674FFC0FEDA4B9FB356126E" box="[258,350,748,771]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="30" pageNumber="664">Fig. 20I</figureCitation>
).
</paragraph>
<paragraph id="8BF636A25674FFC0FF644A78B3F1124C" blockId="30.[164,779,198,1566]" box="[188,505,779,801]" pageId="30" pageNumber="664">Stridulatory setae: lacking.</paragraph>
<paragraph id="8BF636A25674FFC0FF644A59B0D412D7" blockId="30.[164,779,198,1566]" pageId="30" pageNumber="664">
Legs and palpi LSC in: palp trochanter retrolateral face, femur prolateral and retrolateral faces; leg I coxa retrolateral face, trochanter prolateral face, femur prolateral face; leg
<collectionCode id="ED58AE675674FFC0FE374AF5B00B12F6" box="[495,515,902,923]" pageId="30" pageNumber="664">II</collectionCode>
coxa prolateral face, trochanter prolateral face, femur prolateral face.
</paragraph>
<paragraph id="8BF636A25674FFC0FF644AB0B342153E" blockId="30.[164,779,198,1566]" pageId="30" pageNumber="664">
Chaetotaxy (left side): femora I 2p, 3r;
<collectionCode id="ED58AE675674FFC0FDA64AB0B09A12B5" box="[638,658,963,984]" pageId="30" pageNumber="664">II</collectionCode>
4p, 3r;
<collectionCode id="ED58AE675674FFC0FD344AB0B10212B5" box="[748,778,963,984]" pageId="30" pageNumber="664">III</collectionCode>
4p, 4r;
<collectionCode id="ED58AE675674FFC0FF224A91B31E129A" box="[250,278,994,1015]" pageId="30" pageNumber="664">IV</collectionCode>
6p, 5r; palp 2p, 3r; patellae none; tibiae I 2p, 8v, 4r;
<collectionCode id="ED58AE675674FFC0FEC44D73B3381578" box="[284,304,1024,1045]" pageId="30" pageNumber="664">II</collectionCode>
3p, 9v, 4r;
<collectionCode id="ED58AE675674FFC0FE684D73B3C61578" box="[432,462,1024,1045]" pageId="30" pageNumber="664">III</collectionCode>
4p, 10v, 6r;
<collectionCode id="ED58AE675674FFC0FD844D73B0701578" box="[604,632,1024,1045]" pageId="30" pageNumber="664">IV</collectionCode>
7p, 15v, 11r; palp 3p, 9v; metatarsi I 3v;
<collectionCode id="ED58AE675674FFC0FE3A4D6CB3FE1559" box="[482,502,1055,1076]" pageId="30" pageNumber="664">II</collectionCode>
1p, 6v, 1r;
<collectionCode id="ED58AE675674FFC0FDAD4D6CB09B1559" box="[629,659,1055,1076]" pageId="30" pageNumber="664">III</collectionCode>
5p, 9v, 4r;
<collectionCode id="ED58AE675674FFC0FF7C4D4EB2C8153F" box="[164,192,1085,1106]" pageId="30" pageNumber="664">IV</collectionCode>
7p, 12v, 6r.
</paragraph>
<paragraph id="8BF636A25674FFC0FF644D2FB3351581" blockId="30.[164,779,198,1566]" pageId="30" pageNumber="664">
Palp: embolus slender, longer than tegulum.
<collectionCode id="ED58AE675674FFC0FF7C4D09B2CD15E2" box="[164,197,1146,1167]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="30" pageNumber="664">SA</collectionCode>
ends at embolus retrolateral face. The posterior curvature of
<collectionCode id="ED58AE675674FFC0FE524DEAB3A315C3" box="[394,427,1177,1198]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="30" pageNumber="664">SA</collectionCode>
occurs at the base of the embolus and forms an angle of 90°. VG deep (
<figureCitation id="13722A275674FFC0FF744DA5B3391581" box="[172,305,1238,1260]" captionStart="Figure 21" captionStartId="20.[164,243,733,752]" captionTargetBox="[387,1219,197,704]" captionTargetId="figure-142@20.[387,1219,197,704]" captionTargetPageId="20" captionText="Figure 21. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460, left palpal bulb. A, ventral view. B, dorsal view. C, retrolateral view. D, prolateral view. Abbreviations: SA, subapical keel; VG, ventral groove. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10273790" httpUri="https://zenodo.org/record/10273790/files/figure.png" pageId="30" pageNumber="664">Fig. 21A–D</figureCitation>
).
</paragraph>
<paragraph id="8BF636A25674FFC0FF644D86B37114E8" blockId="30.[164,779,198,1566]" pageId="30" pageNumber="664">
Urticating setae:
<typeStatus id="54F288005674FFC0FE484D86B3C91467" box="[400,449,1269,1290]" pageId="30" pageNumber="664">type</typeStatus>
<collectionCode id="ED58AE675674FFC0FE164D86B3E21467" box="[462,490,1269,1290]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="30" pageNumber="664">VI</collectionCode>
arranged in one dorsomedian patch, brown in colour, with anterior and posterior margins notched medially, with rounded edges (butterfly-shaped). The margins are clearly defined (
<figureCitation id="13722A275674FFC0FED14C1CB36514E8" box="[265,365,1391,1413]" captionStart="Figure 20" captionStartId="19.[144,223,1353,1372]" captionTargetBox="[303,1263,195,1323]" captionTargetId="figure-192@19.[303,1263,195,1323]" captionTargetPageId="19" captionText="Figure 20. Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, male CNAN 4460. A, carapace, dorsal view. B, prosoma, ventral view showing the retrolateral projection of the coxa (RP) on coxae of all legs (arrows show this feature only on coxa of third and fourth legs). C, opisthosoma, dorsal view. D, ocular tubercle, dorsal view. E, labial and maxillary cuspules. F, sternum. G, cheliceral teeth, ventral view. H, tibial apophyses, ventral view. I, metatarsus I, prolateral view. J, tibial apophyses, prolateral view. Scale bars = 2 mm (A–C, F, H, I), 1 mm (E, G, J), 0.5 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10273786" httpUri="https://zenodo.org/record/10273786/files/figure.png" pageId="30" pageNumber="664">Fig. 20C</figureCitation>
).
</paragraph>
<paragraph id="8BF636A25674FFC0FF644CFDB3071773" blockId="30.[164,779,198,1566]" pageId="30" pageNumber="664">
Colour pattern: in ethanol the colour is brown, with the carapace, legs, and palps darker. In live specimens the general colour is very dark brown, with femora and carapace darker than rest of the body (
<figureCitation id="13722A275674FFC0FF744F7BB3091773" box="[172,257,1544,1566]" captionStart="Figure 1" captionStartId="3.[144,223,1334,1353]" captionTargetBox="[303,1262,196,1304]" captionTargetId="figure-144@3.[303,1263,196,1304]" captionTargetPageId="3" captionText="Figure 1. A–H, J–L, Hemirrhagus spp. habitus; I, Hemirrhagus papalotl egg-sac. A, Hemirrhagus chilango, male CNAN 4253, in life; B, C, Hemirrhagus eros, B, female, C, male; D, E, Hemirrhagus ocellatus, D, male CNAN 4457, in life, E, female; F, Hemirrhagus benzaa sp. nov., female; G–I, H. papalotl, G, male CNAN 4461, in life, H, female CNAN 4461, in life, I, female CNAN 4459 with fixed hammock egg-sac, in life; J, K, Hemirrhagus stygius, J, male CNAN 4462, in life, K, female CNAN 4462, in life; L, Hemirrhagus franckei sp. nov., holotype male CNAN-T0776, in life. Photos: A, C, D, E, G–K: J. Mendoza; B, F, L: E. Goyer." figureDoi="http://doi.org/10.5281/zenodo.10273732" httpUri="https://zenodo.org/record/10273732/files/figure.png" pageId="30" pageNumber="664">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="8BF636A25674FFC0FF7C4F36B3731603" blockId="30.[164,779,1605,1902]" pageId="30" pageNumber="664">
<emphasis id="B93DEAB05674FFC0FF7C4F36B0251737" box="[164,557,1605,1626]" italics="true" pageId="30" pageNumber="664">Distribution and natural history:</emphasis>
Known only from Gruta de Aguacachil/Zacatecolotla in
<collectingRegion id="498DF8405674FFC0FD444F17B10F1714" box="[668,775,1636,1657]" country="Mexico" name="Guerrero" pageId="30" pageNumber="664">Guerrero</collectingRegion>
,
<collectingCountry id="F35E76325674FFC0FF7C4FF1B2FC17F5" box="[164,244,1666,1688]" name="Mexico" pageId="30" pageNumber="664">México</collectingCountry>
(
<figureCitation id="13722A275674FFC0FED24FF1B35417F5" box="[266,348,1666,1688]" captionStart="Figure 48" captionStartId="51.[144,223,1012,1031]" captionTargetBox="[151,1418,203,974]" captionTargetId="figure-352@51.[144,1425,196,982]" captionTargetPageId="51" captionText="Figure 48. Known distribution of Hemirrhagus stygius (Gertsch, 1971), Hemirrhagus elliotti (Gertsch, 1973), Hemirrhagus embolulatus sp. nov., Hemirrhagus eros Pérez-Miles &amp; Locht, 2003, Hemirrhagus chilango Pérez-Miles &amp; Locht, 2003, Hemirrhagus nahuanus (Gertsch, 1982), Hemirrhagus papalotl Pérez-Miles &amp; Locht, 2003, Hemirrhagus grieta (Gertsch, 1982), Hemirrhagus coztic Pérez-Miles &amp; Locht, 2003, and Hemirrhagus perezmilesi García-Villafuerte &amp; Locht, 2010." figureDoi="http://doi.org/10.5281/zenodo.10273862" httpUri="https://zenodo.org/record/10273862/files/figure.png" pageId="30" pageNumber="664">Fig. 48</figureCitation>
). The species lives only inside the cave; specimens were collected on walls at a depth of
<quantity id="4CB19B475674FFC0FF1F4FB3B2F717B8" box="[199,255,1728,1749]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="664" unit="m" value="20.0">20 m</quantity>
from the entrance (
<figureCitation id="13722A275674FFC0FE264FCCB05D17BB" box="[510,597,1727,1750]" captionStart="Figure 1" captionStartId="3.[144,223,1334,1353]" captionTargetBox="[303,1262,196,1304]" captionTargetId="figure-144@3.[303,1263,196,1304]" captionTargetPageId="3" captionText="Figure 1. A–H, J–L, Hemirrhagus spp. habitus; I, Hemirrhagus papalotl egg-sac. A, Hemirrhagus chilango, male CNAN 4253, in life; B, C, Hemirrhagus eros, B, female, C, male; D, E, Hemirrhagus ocellatus, D, male CNAN 4457, in life, E, female; F, Hemirrhagus benzaa sp. nov., female; G–I, H. papalotl, G, male CNAN 4461, in life, H, female CNAN 4461, in life, I, female CNAN 4459 with fixed hammock egg-sac, in life; J, K, Hemirrhagus stygius, J, male CNAN 4462, in life, K, female CNAN 4462, in life; L, Hemirrhagus franckei sp. nov., holotype male CNAN-T0776, in life. Photos: A, C, D, E, G–K: J. Mendoza; B, F, L: E. Goyer." figureDoi="http://doi.org/10.5281/zenodo.10273732" httpUri="https://zenodo.org/record/10273732/files/figure.png" pageId="30" pageNumber="664">Fig. 1H</figureCitation>
). Females laid fixed hammock egg-sacs in the cave wall; these were fixed between two protuberances on the wall (
<figureCitation id="13722A275674FFC0FD694F8EB0F3167E" box="[689,763,1789,1811]" captionStart="Figure 1" captionStartId="3.[144,223,1334,1353]" captionTargetBox="[303,1262,196,1304]" captionTargetId="figure-144@3.[303,1263,196,1304]" captionTargetPageId="3" captionText="Figure 1. A–H, J–L, Hemirrhagus spp. habitus; I, Hemirrhagus papalotl egg-sac. A, Hemirrhagus chilango, male CNAN 4253, in life; B, C, Hemirrhagus eros, B, female, C, male; D, E, Hemirrhagus ocellatus, D, male CNAN 4457, in life, E, female; F, Hemirrhagus benzaa sp. nov., female; G–I, H. papalotl, G, male CNAN 4461, in life, H, female CNAN 4461, in life, I, female CNAN 4459 with fixed hammock egg-sac, in life; J, K, Hemirrhagus stygius, J, male CNAN 4462, in life, K, female CNAN 4462, in life; L, Hemirrhagus franckei sp. nov., holotype male CNAN-T0776, in life. Photos: A, C, D, E, G–K: J. Mendoza; B, F, L: E. Goyer." figureDoi="http://doi.org/10.5281/zenodo.10273732" httpUri="https://zenodo.org/record/10273732/files/figure.png" pageId="30" pageNumber="664">Fig. 1I</figureCitation>
). The females with egg-sacs were found close together, each protecting their egg sac. One of the egg-sacs contained
<specimenCount id="9D4FFD2B5674FFC0FEC44E2BB37F1603" box="[284,375,1880,1902]" pageId="30" pageNumber="664" type="egg">63 eggs</specimenCount>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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